113:
763:
1514:
145:
672:
839:
2722:
2739:
112:
502:. However, it has been proven that this assimilation is mistaken, as it is mainly based on superficial resemblances, leaving the latter awaiting a more in-depth study to determine its relationships with other representatives of the group. Moreover, studies published post-2018 prove that
600:-like features than other gorgonopsians. It quickly became apparent that the species is no more related to these same cynodonts than the other gorgonopsians, and in 1940, only two years after the publication of his article, one of Hartmann-Weinberg's employees, the reputed
741:
Other specimens have been attributed to this species, but no concrete affiliation has been worked out to date. For example, the well-preserved skull cataloged SAM-PK-K10034, mentioned briefly in a study concerning the nocturnal vision of non-mammalian
1765:
Gebauer, Eva V. I. (2014), "Re-assessment of the
Taxonomic Position of the Specimen GPIT/RE/7113 (Sauroctonus parringtoni comb. Nov., Gorgonopsia)", in Kammerer, Christian F.; Angielczyk, Kenneth D.; Fröbisch, Jörg (eds.),
728:, a claim that she would confirm again in 2014. However, this affiliation is formally rejected in 2018, as new classifications based on cranial anatomical traits show that derived gorgonopsians are separated into two
1420:
is known from the locality of Sjomin Ravine, located in the republic of
Tatarstan, in the northeast of European Russia. This fossil site, discovered in 1938 and dating from the end of the Middle Permian
1401:. A study published in 2020 shows that several prehistoric predators with this comparable morphology would have had various possible prey-killing techniques. Among the gorgonopsians,
767:
562:
121:
907:
in an uncertain position among the gorgonopsians, with some seeing it as close to the
African genera, while others consider it to belong to a lineage in its own right.
657:
where the first known skull was discovered. Among these fossils remains figures PIN 156/6, a specimen containing a crushed skull, some anterior part of the postcranial
2900:
1739:
2816:
2910:
2890:
2880:
2875:
2245:
1639:
2905:
2133:
2039:
1982:
2084:
1783:
483:
and postcranial elements, currently all recorded from
Tatarstan. These elements show that the animal was a mid-sized gorgonopsian.
1397:
One of the most recognizable features among gorgonopsians is the presence of long, saber-like canines on the upper and lower
2885:
788:
is 22.5 cm long, indicating that it should be a medium-sized gorgonopsian. The skull is narrow posteriorly with small
724:. It is from 2007 that the paleontologist Eva V. I. Gebauer reclassifies it as an African representative of the genus
144:
2895:
2082:
Christian F. Kammerer; Bruce S. Rubidge (2022). "The earliest gorgonopsians from the Karoo Basin of South Africa".
1631:
858:
2238:
711:
389:
2721:
1409:, indicating that it would not have been as specialized in hunting as the other representatives of the group.
895:
within the
Gorgonopsidae in 2003, without assigning it to any subtaxon. In his 2007 thesis, Gebauer considers
1741:
Phylogeny and
Evolution of the Gorgonopsia with a Special Reference to the Skull and Skeleton of GPIT/RE/7113
1720:] (in Russian). Vol. 143. Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR. pp. 1–226.
865:
within the family
Gorgonopsidae, and in the subfamily Cynariopinae, alongside various African genera such as
2060:
586:
2759:
1871:
Bendel, Eva-Maria; Kammerer, Christian F.; Kardjilov, Nikolay; Fernandez, Vincent; Fröbisch, Jörg (2018).
707:
2422:
2127:
Stephan
Lautenschlager; Borja Figueirido; Daniel D. Cashmore; Eva-Maria Bendel; Thomas L. Stubbs (2020).
930:, in particular for certain shared cranial characteristics, more precisely for close contact between the
2842:
1957:
F. von Huene (1950). "Die
Theriodontier des ostafrikanischen Ruhuhu-Gebietes in der Tübinger Sammlung".
900:
762:
910:
It is from 2018 that paleontologists
Christian Kammerer and Vladimir Masyutin definitively reclassify
2870:
2742:
2231:
2093:
690:, mainly represented by cranial material. In 1950, a relatively complete skeleton, discovered in the
1519:
699:
375:
296:
1748:
2327:
2209:
2109:
888:
307:
139:
2847:
2829:
2170:
2152:
2035:
2009:
1910:
1850:
1779:
1689:
1635:
1617:
793:
2834:
2797:
2201:
2160:
2142:
2101:
1999:
1991:
1900:
1890:
1840:
1830:
1771:
1621:
823:
671:
838:
789:
691:
428:
1770:, Vertebrate Paleobiology and Paleoanthropology, Springer Netherlands, pp. 185–207,
2097:
2034:. Encyclopedia of Paleoherpetology. Vol. 17 B/I. Stuttgart: Gustav Fischer Verlag.
1935:
2698:
2591:
2475:
2165:
2128:
2004:
1977:
1905:
1872:
1845:
1810:
1559:
1494:
1074:
931:
920:
873:
827:
734:
654:
619:
514:
472:
424:
251:
2864:
2584:
2553:
2468:
2213:
2189:
2129:"Morphological convergence obscures functional diversity in sabre-toothed carnivores"
2113:
2105:
1528:
1467:
1463:
1143:
1091:
716:
638:
610:
601:
468:
463:
383:
345:
339:
63:
1978:"Nocturnality in synapsids predates the origin of mammals by over 100 million years"
2705:
2676:
2659:
2632:
2511:
2450:
2401:
1472:
1299:
994:
802:
797:
577:
570:
546:
449:
444:
417:
328:
260:
649:, "murderer" or "killer"), to literally give "lizard killer", in reference to its
1895:
1775:
1658:
A. P. Bystrow (1955). "A gorgonopsian from the Upper Permian beds of the Volga".
2791:
2683:
2669:
2571:
2560:
2532:
2489:
2415:
2408:
2387:
2365:
2339:
2254:
1535:
1489:
1476:
1458:
1315:
1249:
1160:
1026:
956:
926:
675:
The specimen GPIT/RE/7113, previously assigned to a second species of the genus
557:. One of the fossils described in his paper is an incomplete but well-preserved
508:
494:, which would make this genus the only known gorgonopsian to have lived in both
231:
218:
38:
2190:"A National Geopark in the Republic of Tatarstan (Russia): a Feasibility Study"
2625:
2618:
2609:
2598:
2296:
2205:
1509:
1498:
1485:
1453:
1434:
1422:
631:
420:
202:
83:
48:
32:
2782:
2156:
2058:
Mikhail F. Ivakhnenko (2003). "Eotherapsids from the East European placket".
1693:
857:
has changed many times over the decades. In 1974, the Russian paleontologist
2690:
2546:
2539:
2525:
2443:
2436:
2312:
2287:
1627:
1564:
1481:
1437:
1225:
1201:
1177:
970:
942:
879:
867:
771:
650:
627:
550:
506:
belongs to a grouping of Russian gorgonopsians, placed alongside the genera
436:
369:
189:
156:
125:
88:
2174:
2147:
2013:
1995:
1914:
1854:
1444:, represented by isolated bones and fragmentary skeletons, as well as rare
2808:
1405:
is found to have had a reduced jaw opening, unlike in the closely related
2776:
2639:
2278:
1881:
1688:: Éditions du Centre national de la recherche scientifique. p. 416.
1441:
850:
743:
695:
658:
593:
487:
176:
78:
73:
58:
53:
43:
2032:
Theriodontia I: Phthinosuchia, Biarmosuchia, Eotitanosuchia, Gorgonopsia
938:. This classification will be immediately followed by the next studies.
2821:
2518:
1835:
1711:
1266:
819:
566:
440:
410:
93:
68:
710:. In 1970, the skeleton was reidentified by the French paleontologist
471:, before being definitively classified in a separate genus erected by
461:
was designated as such until 1940, when it was assigned to the genus
2272:
811:
698:, cataloged GPIT/RE/7113, was described by the German paleontologist
597:
554:
542:
541:
paleontologist Alexandra Paulinovna Hartmann-Weinberg described some
538:
499:
495:
432:
166:
2753:
1620:; Mikhail A. Shishkin; David M. Unwin; Evgenii N. Kurochkin (2000).
1589:
A. P. Hartmann-Weinberg (1938). "Gorgonopsians as time indicators".
592:
is incorrectly named due to its supposed morphological closeness to
565:
156/5. Hartmann-Weinberg considers this specimen to belong to a new
630:
of the taxon a proposal that will be quickly recognized later. The
1685:
1445:
935:
837:
815:
774:
761:
729:
670:
605:
573:
558:
480:
458:
413:
128:
903:
Gorgonopsidae. These repetitive classification changes have left
686:
are only recorded in Russian territory and belong to the species
1430:
1426:
2757:
2265:
2227:
2223:
1398:
749:
1747:(PhD). Eberhard-Karls University of Tübingen. Archived from
914:
in a group of Russian gorgonopsians, alongside the genera
732:
of Russian and African origin, leaving GPIT/RE/7113 as an
622:
proposed a separate genus for the species, which he named
2030:
Denise Sigogneau-Russell (1989). Peter Wellnhofer (ed.).
2188:
M. S. Vdovets; V. V. Silantiev; V. V. Mozzherin (2010).
1452:, the main tetrapods identified in the area include the
883:. In a book published in 1989, Sigogneau-Russell leaves
2025:
2023:
949:
within Gorgonopsia follows Kammerer and Rubidge, 2022:
1682:
Révision systématique des gorgonopsiens sud-africains
796:
is elongated, the cranial arches are narrow, and the
596:, which have a large number of progressive and more
267:
2766:
2657:
2608:
2570:
2499:
2460:
2385:
2338:
2311:
887:as a gorgonopsian whose placement is undetermined.
2053:
2051:
1819:) from the Permian Kotelnich locality of Russia"
1612:
1610:
1608:
1606:
1604:
826:. There are also 4 to 6 postcanine teeth on the
2077:
2075:
1705:
1703:
626:, and designates the specimen PIN 156/5 as the
618:. In 1955, after the analysis of the material,
475:(in 1955). The most complete, known fossils of
1804:
1802:
1800:
1798:
1796:
1794:
522:, due to some shared cranial characteristics.
2239:
1959:Neues Jahrbuch für Geologie und Paläontologie
1760:
1758:
1733:
1731:
1729:
1727:
1653:
1651:
439:, were initially believed to belong to a new
8:
1930:
1928:
1926:
1924:
1866:
1864:
1768:Early Evolutionary History of the Synapsida
1623:The age of dinosaurs in Russia and Mongolia
2754:
2576:
2503:
2393:
2382:
2317:
2308:
2262:
2246:
2232:
2224:
490:have been attributed to a second species,
111:
20:
2164:
2146:
2003:
1904:
1894:
1844:
1834:
1581:
1548:
364:
323:
1809:Kammerer, C. F.; Masyutin, V. (2018).
1675:
1673:
608:as a new species of the Russian genus
1976:K. D. Angielczyk; L. Schmitz (2014).
1873:"Cranial anatomy of the gorgonopsian
7:
486:Nearly complete remains listed from
1291:
1241:
1217:
1193:
1135:
1066:
1042:
1018:
1010:
986:
962:
952:
702:as a representative of the species
653:. All known specimens are from the
2134:Proceedings of the Royal Society B
1983:Proceedings of the Royal Society B
14:
2901:Lopingian genus first appearances
2085:Journal of African Earth Sciences
1680:Denise Sigogneau-Russell (1970).
818:, the bones poorly sculpted, and
2738:
2737:
2720:
2106:10.1016/j.jafrearsci.2022.104631
1512:
423:who lived during the end of the
143:
1433:with separated layers of fine
822:only slightly larger than the
1:
2911:Fossil taxa described in 1955
2891:Lopingian synapsids of Europe
2881:Lopingian synapsids of Africa
661:and numerous isolated bones.
2876:Prehistoric therapsid genera
1896:10.1371/journal.pone.0207367
1776:10.1007/978-94-007-6841-3_12
1710:Leonid P. Tatarinov (1974).
1440:, which contains fossils of
1425:), is mainly represented by
814:tuberosities that have many
2906:Lopingian genus extinctions
1877:based on CT-reconstruction"
16:Extinct genus of therapsids
2927:
1811:"Gorgonopsian therapsids (
1738:Gebauer, E. V. I. (2007).
1632:Cambridge University Press
859:Leonid Petrovich Tatarinov
800:is moderately high. As in
706:, being named in honor of
645:, "lizard") and κτόνος / (
549:having been discovered in
2733:
2718:
2579:
2506:
2396:
2381:
2362:
2320:
2307:
2261:
2206:10.1007/s12371-010-0010-0
1313:
1296:
1289:
1263:
1246:
1239:
1222:
1215:
1198:
1191:
1174:
1157:
1140:
1133:
1088:
1071:
1064:
1047:
1040:
1023:
1016:
1008:
991:
984:
967:
960:
738:within the latter group.
722:Aelurognathus parringtoni
704:Scymnognathus parringtoni
682:All confirmed remains of
313:
306:
287:
282:
266:
259:
140:Scientific classification
138:
119:
110:
23:
1591:Problems of Paleontology
945:showing the position of
712:Denise Sigogneau-Russell
616:Inostrancevia progressus
409:("lizard killer") is an
292:Sauroctonus? parringtoni
2061:Paleontological Journal
1718:Theriodonts of the USSR
1558:is now considered as a
766:Diagram reconstructing
583:Arctognathus progressus
334:Hartmann-Weinberg, 1938
277:Hartmann-Weinberg, 1938
2148:10.1098/rspb.2020.1818
1996:10.1098/rspb.2014.1642
846:
781:
708:Francis Rex Parrington
679:
272:Sauroctonus progressus
2843:Paleobiology Database
1660:Voprosy Paleontologii
1131:"African clade"
1014:"Russian clade"
841:
765:
674:
553:, in the present-day
1448:remains. Apart from
842:Life restoration of
746:, is referred to as
581:, designating it as
2886:Fossils of Tanzania
2098:2022JAfES.19404631K
1520:Paleontology portal
700:Friedrich von Huene
1875:Cynariops robustus
1836:10.7717/peerj.4954
889:Mikhail Ivakhnenko
847:
784:The skull size of
782:
680:
2896:Fossils of Russia
2858:
2857:
2830:Open Tree of Life
2760:Taxon identifiers
2751:
2750:
2729:
2728:
2716:
2715:
2653:
2652:
2649:
2648:
2377:
2376:
2373:
2372:
2358:
2357:
1641:978-0-521-55476-3
1618:Michael J. Benton
1389:
1388:
1380:
1379:
1371:
1370:
1362:
1361:
1353:
1352:
1344:
1343:
1335:
1334:
1326:
1325:
1278:
1277:
1121:
1120:
1112:
1111:
1103:
1102:
891:still classifies
714:as a specimen of
614:, renaming it as
604:reclassified the
402:
401:
393:
390:Sigogneau-Russell
379:
349:
335:
300:
255:
2918:
2851:
2850:
2838:
2837:
2825:
2824:
2812:
2811:
2802:
2801:
2800:
2787:
2786:
2785:
2755:
2741:
2740:
2724:
2577:
2504:
2394:
2383:
2318:
2309:
2302:
2301:
2263:
2248:
2241:
2234:
2225:
2218:
2217:
2185:
2179:
2178:
2168:
2150:
2124:
2118:
2117:
2079:
2070:
2069:
2055:
2046:
2045:
2027:
2018:
2017:
2007:
1973:
1967:
1966:
1954:
1948:
1947:
1932:
1919:
1918:
1908:
1898:
1889:(11): e0207367.
1868:
1859:
1858:
1848:
1838:
1806:
1789:
1788:
1762:
1753:
1752:
1746:
1735:
1722:
1721:
1707:
1698:
1697:
1677:
1668:
1667:
1655:
1646:
1645:
1614:
1599:
1598:
1586:
1569:
1553:
1522:
1517:
1516:
1515:
1484:, including the
1292:
1242:
1218:
1194:
1136:
1067:
1043:
1019:
1011:
987:
963:
953:
824:postcanine teeth
720:, being renamed
651:carnivorous diet
587:specific epithet
526:Research history
435:, discovered in
388:
374:
344:
333:
295:
269:
250:
243:
230:
217:
148:
147:
115:
105:
35:
29:Temporal range:
21:
2926:
2925:
2921:
2920:
2919:
2917:
2916:
2915:
2861:
2860:
2859:
2854:
2846:
2841:
2833:
2828:
2820:
2815:
2807:
2805:
2796:
2795:
2790:
2781:
2780:
2775:
2762:
2752:
2747:
2725:
2712:
2645:
2604:
2566:
2495:
2456:
2390:
2369:
2354:
2334:
2303:
2268:
2267:
2257:
2252:
2222:
2221:
2187:
2186:
2182:
2126:
2125:
2121:
2081:
2080:
2073:
2057:
2056:
2049:
2042:
2029:
2028:
2021:
1975:
1974:
1970:
1956:
1955:
1951:
1934:
1933:
1922:
1870:
1869:
1862:
1808:
1807:
1792:
1786:
1764:
1763:
1756:
1744:
1737:
1736:
1725:
1713:Териодонты СССР
1709:
1708:
1701:
1679:
1678:
1671:
1657:
1656:
1649:
1642:
1634:. p. 696.
1616:
1615:
1602:
1588:
1587:
1583:
1578:
1573:
1572:
1554:
1550:
1545:
1518:
1513:
1511:
1508:
1415:
1395:
1390:
1381:
1372:
1363:
1354:
1345:
1336:
1327:
1279:
1122:
1113:
1104:
836:
828:maxillary bones
760:
692:Usili Formation
669:
666:S.? parringtoni
637:comes from the
535:
528:
429:European Russia
427:in what is now
398:
363:
362:
360:S?. parringtoni
354:
322:
321:
278:
275:
249:
241:
228:
215:
142:
106:
104:
103:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
31:
30:
27:
17:
12:
11:
5:
2924:
2922:
2914:
2913:
2908:
2903:
2898:
2893:
2888:
2883:
2878:
2873:
2863:
2862:
2856:
2855:
2853:
2852:
2839:
2826:
2813:
2803:
2788:
2772:
2770:
2764:
2763:
2758:
2749:
2748:
2746:
2745:
2734:
2731:
2730:
2727:
2726:
2719:
2717:
2714:
2713:
2711:
2710:
2702:
2699:Paragalerhinus
2695:
2687:
2680:
2673:
2665:
2663:
2655:
2654:
2651:
2650:
2647:
2646:
2644:
2643:
2636:
2629:
2622:
2614:
2612:
2606:
2605:
2603:
2602:
2595:
2592:Ruhuhucerberus
2588:
2580:
2574:
2568:
2567:
2565:
2564:
2557:
2550:
2543:
2536:
2529:
2522:
2515:
2507:
2501:
2497:
2496:
2494:
2493:
2486:
2479:
2476:Pravoslavlevia
2472:
2464:
2462:
2458:
2457:
2455:
2454:
2447:
2440:
2433:
2419:
2412:
2405:
2397:
2391:
2386:
2379:
2378:
2375:
2374:
2371:
2370:
2363:
2360:
2359:
2356:
2355:
2353:
2352:
2351:
2350:
2344:
2342:
2336:
2335:
2333:
2332:
2331:
2330:
2321:
2315:
2305:
2304:
2300:
2299:
2290:
2281:
2275:
2266:
2259:
2258:
2253:
2251:
2250:
2243:
2236:
2228:
2220:
2219:
2180:
2141:(1935): 1–10.
2119:
2071:
2068:(S4): 339–465.
2047:
2041:978-3437304873
2040:
2019:
1968:
1949:
1920:
1860:
1815:gen. nov. and
1790:
1784:
1754:
1751:on 2012-07-22.
1723:
1699:
1669:
1647:
1640:
1600:
1580:
1579:
1577:
1574:
1571:
1570:
1560:junior synonym
1547:
1546:
1544:
1541:
1540:
1539:
1532:
1524:
1523:
1507:
1504:
1495:archosauriform
1414:
1411:
1394:
1391:
1387:
1386:
1383:
1382:
1378:
1377:
1374:
1373:
1369:
1368:
1365:
1364:
1360:
1359:
1356:
1355:
1351:
1350:
1347:
1346:
1342:
1341:
1338:
1337:
1333:
1332:
1329:
1328:
1324:
1323:
1320:
1319:
1312:
1309:
1308:
1305:
1304:
1295:
1290:
1288:
1285:
1284:
1281:
1280:
1276:
1275:
1272:
1271:
1262:
1259:
1258:
1255:
1254:
1245:
1240:
1238:
1235:
1234:
1231:
1230:
1221:
1216:
1214:
1211:
1210:
1207:
1206:
1197:
1192:
1190:
1187:
1186:
1183:
1182:
1173:
1170:
1169:
1166:
1165:
1156:
1153:
1152:
1149:
1148:
1139:
1134:
1132:
1128:
1127:
1124:
1123:
1119:
1118:
1115:
1114:
1110:
1109:
1106:
1105:
1101:
1100:
1097:
1096:
1087:
1084:
1083:
1080:
1079:
1075:Pravoslavlevia
1070:
1065:
1063:
1060:
1059:
1056:
1055:
1046:
1041:
1039:
1036:
1035:
1032:
1031:
1022:
1017:
1015:
1009:
1007:
1004:
1003:
1000:
999:
990:
985:
983:
980:
979:
976:
975:
966:
961:
959:
951:
941:The following
921:Pravoslavlevia
874:Scylacognathus
835:
834:Classification
832:
810:also has high
794:temporal fossa
759:
756:
735:incertae sedis
668:
663:
620:Alexey Bystrow
534:
529:
527:
524:
515:Pravoslavlevia
492:S. parringtoni
473:Alexey Bystrow
457:in 1938). The
425:Middle Permian
400:
399:
397:
396:
395:
394:
380:
357:
356:
355:
353:
352:
351:
350:
336:
316:
315:
314:
311:
310:
304:
303:
302:
301:
285:
284:
283:Other species
280:
279:
276:
264:
263:
257:
256:
239:
235:
234:
226:
222:
221:
213:
206:
205:
200:
193:
192:
187:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
136:
135:
117:
116:
108:
107:
99:
98:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
36:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2923:
2912:
2909:
2907:
2904:
2902:
2899:
2897:
2894:
2892:
2889:
2887:
2884:
2882:
2879:
2877:
2874:
2872:
2869:
2868:
2866:
2849:
2844:
2840:
2836:
2831:
2827:
2823:
2818:
2814:
2810:
2804:
2799:
2793:
2789:
2784:
2778:
2774:
2773:
2771:
2769:
2765:
2761:
2756:
2744:
2736:
2735:
2732:
2723:
2708:
2707:
2703:
2701:
2700:
2696:
2693:
2692:
2688:
2686:
2685:
2681:
2679:
2678:
2674:
2672:
2671:
2667:
2666:
2664:
2662:
2661:
2656:
2642:
2641:
2637:
2635:
2634:
2630:
2628:
2627:
2623:
2621:
2620:
2616:
2615:
2613:
2611:
2607:
2601:
2600:
2596:
2594:
2593:
2589:
2587:
2586:
2585:Aelurognathus
2582:
2581:
2578:
2575:
2573:
2569:
2563:
2562:
2558:
2556:
2555:
2551:
2549:
2548:
2544:
2542:
2541:
2537:
2535:
2534:
2530:
2528:
2527:
2523:
2521:
2520:
2516:
2514:
2513:
2509:
2508:
2505:
2502:
2500:African clade
2498:
2492:
2491:
2487:
2485:
2484:
2480:
2478:
2477:
2473:
2471:
2470:
2469:Inostrancevia
2466:
2465:
2463:
2461:Russian clade
2459:
2453:
2452:
2448:
2446:
2445:
2441:
2439:
2438:
2434:
2432:
2431:
2430:
2426:
2420:
2418:
2417:
2413:
2411:
2410:
2406:
2404:
2403:
2399:
2398:
2395:
2392:
2389:
2384:
2380:
2368:
2367:
2361:
2348:
2347:
2346:
2345:
2343:
2341:
2337:
2329:
2325:
2324:
2323:
2322:
2319:
2316:
2314:
2310:
2306:
2298:
2294:
2291:
2289:
2285:
2282:
2280:
2276:
2274:
2270:
2269:
2264:
2260:
2256:
2249:
2244:
2242:
2237:
2235:
2230:
2229:
2226:
2215:
2211:
2207:
2203:
2199:
2195:
2191:
2184:
2181:
2176:
2172:
2167:
2162:
2158:
2154:
2149:
2144:
2140:
2136:
2135:
2130:
2123:
2120:
2115:
2111:
2107:
2103:
2099:
2095:
2091:
2087:
2086:
2078:
2076:
2072:
2067:
2063:
2062:
2054:
2052:
2048:
2043:
2037:
2033:
2026:
2024:
2020:
2015:
2011:
2006:
2001:
1997:
1993:
1989:
1985:
1984:
1979:
1972:
1969:
1964:
1961:(in German).
1960:
1953:
1950:
1945:
1941:
1939:
1931:
1929:
1927:
1925:
1921:
1916:
1912:
1907:
1902:
1897:
1892:
1888:
1884:
1883:
1878:
1876:
1867:
1865:
1861:
1856:
1852:
1847:
1842:
1837:
1832:
1828:
1824:
1820:
1818:
1814:
1805:
1803:
1801:
1799:
1797:
1795:
1791:
1787:
1785:9789400768413
1781:
1777:
1773:
1769:
1761:
1759:
1755:
1750:
1743:
1742:
1734:
1732:
1730:
1728:
1724:
1719:
1715:
1714:
1706:
1704:
1700:
1695:
1691:
1687:
1684:(in French).
1683:
1676:
1674:
1670:
1665:
1661:
1654:
1652:
1648:
1643:
1637:
1633:
1629:
1625:
1624:
1619:
1613:
1611:
1609:
1607:
1605:
1601:
1596:
1592:
1585:
1582:
1575:
1567:
1566:
1561:
1557:
1556:Scymnognathus
1552:
1549:
1542:
1538:
1537:
1533:
1531:
1530:
1529:Inostrancevia
1526:
1525:
1521:
1510:
1505:
1503:
1501:
1500:
1496:
1492:
1491:
1487:
1483:
1479:
1478:
1474:
1470:
1469:
1468:Chroniosuchus
1465:
1464:reptiliomorph
1461:
1460:
1455:
1451:
1450:S. progressus
1447:
1443:
1439:
1436:
1432:
1428:
1424:
1419:
1418:S. progressus
1412:
1410:
1408:
1407:Inostrancevia
1404:
1400:
1392:
1385:
1384:
1376:
1375:
1367:
1366:
1358:
1357:
1349:
1348:
1340:
1339:
1331:
1330:
1322:
1321:
1318:
1317:
1311:
1310:
1307:
1306:
1303:
1302:
1301:
1294:
1293:
1287:
1286:
1283:
1282:
1274:
1273:
1270:
1269:
1268:
1261:
1260:
1257:
1256:
1253:
1252:
1251:
1244:
1243:
1237:
1236:
1233:
1232:
1229:
1228:
1227:
1220:
1219:
1213:
1212:
1209:
1208:
1205:
1204:
1203:
1196:
1195:
1189:
1188:
1185:
1184:
1181:
1180:
1179:
1172:
1171:
1168:
1167:
1164:
1163:
1162:
1155:
1154:
1151:
1150:
1147:
1146:
1145:
1138:
1137:
1130:
1129:
1126:
1125:
1117:
1116:
1108:
1107:
1099:
1098:
1095:
1094:
1093:
1092:Inostrancevia
1086:
1085:
1082:
1081:
1078:
1077:
1076:
1069:
1068:
1062:
1061:
1058:
1057:
1054:
1053:
1052:
1045:
1044:
1038:
1037:
1034:
1033:
1030:
1029:
1028:
1021:
1020:
1013:
1012:
1006:
1005:
1002:
1001:
998:
997:
996:
989:
988:
982:
981:
978:
977:
974:
973:
972:
965:
964:
958:
955:
954:
950:
948:
944:
939:
937:
933:
929:
928:
923:
922:
917:
916:Inostrancevia
913:
908:
906:
902:
898:
894:
890:
886:
882:
881:
876:
875:
870:
869:
864:
860:
856:
852:
845:
844:S. progressus
840:
833:
831:
829:
825:
821:
817:
813:
809:
805:
804:
799:
795:
791:
787:
780:
779:S. progressus
776:
773:
769:
764:
757:
755:
753:
751:
745:
739:
737:
736:
731:
727:
723:
719:
718:
717:Aelurognathus
713:
709:
705:
701:
697:
693:
689:
688:S. progressus
685:
678:
673:
667:
664:
662:
660:
656:
655:type locality
652:
648:
644:
640:
639:Ancient Greek
636:
633:
629:
625:
621:
617:
613:
612:
611:Inostrancevia
607:
603:
602:Ivan Yefremov
599:
595:
591:
588:
584:
580:
579:
575:
572:
571:South African
568:
564:
560:
556:
552:
548:
547:gorgonopsians
544:
540:
537:In 1938, the
533:
532:S. progressus
530:
525:
523:
521:
520:Inostrancevia
517:
516:
511:
510:
505:
504:S. progressus
501:
497:
493:
489:
484:
482:
478:
477:S. progressus
474:
470:
469:Ivan Yefremov
466:
465:
464:Inostrancevia
460:
456:
455:A. progressus
452:
451:
446:
445:South African
442:
438:
434:
430:
426:
422:
419:
415:
412:
408:
407:
391:
387:
385:
384:Aelurognathus
381:
377:
373:
371:
370:Scymnognathus
367:
366:
365:
361:
347:
343:
341:
340:Inostrancevia
337:
332:
330:
326:
325:
324:
320:
319:S. progressus
312:
309:
305:
298:
294:
293:
289:
288:
286:
281:
274:
273:
265:
262:
258:
253:
248:
247:
240:
237:
236:
233:
232:Gorgonopsidae
227:
224:
223:
220:
214:
211:
208:
207:
204:
201:
198:
195:
194:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
146:
141:
137:
134:
133:S. progressus
130:
127:
123:
118:
114:
109:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
34:
26:
22:
19:
2767:
2706:Wangwusaurus
2704:
2697:
2689:
2682:
2677:Broomisaurus
2675:
2668:
2660:Nomina dubia
2658:
2638:
2633:Leontosaurus
2631:
2624:
2617:
2597:
2590:
2583:
2559:
2552:
2545:
2538:
2531:
2524:
2517:
2512:Arctognathus
2510:
2488:
2482:
2481:
2474:
2467:
2451:Viatkogorgon
2449:
2442:
2435:
2428:
2424:
2421:
2414:
2407:
2402:Aelurosaurus
2400:
2364:
2292:
2283:
2200:(1): 25–37.
2197:
2193:
2183:
2138:
2132:
2122:
2089:
2083:
2065:
2059:
2031:
1987:
1981:
1971:
1962:
1958:
1952:
1943:
1937:
1886:
1880:
1874:
1826:
1822:
1817:Viatkogorgon
1816:
1812:
1767:
1749:the original
1740:
1717:
1712:
1681:
1663:
1659:
1622:
1594:
1590:
1584:
1563:
1555:
1551:
1534:
1527:
1497:
1488:
1475:
1473:temnospondyl
1466:
1457:
1449:
1417:
1416:
1413:Paleoecology
1406:
1402:
1396:
1393:Paleobiology
1314:
1300:Arctognathus
1298:
1297:
1265:
1264:
1248:
1247:
1224:
1223:
1200:
1199:
1176:
1175:
1159:
1158:
1142:
1141:
1090:
1089:
1073:
1072:
1050:
1049:
1048:
1025:
1024:
995:Viatkogorgon
993:
992:
969:
968:
946:
940:
925:
919:
915:
911:
909:
904:
896:
892:
884:
878:
872:
866:
862:
854:
853:position of
848:
843:
807:
803:Viatkogorgon
801:
798:dentary bone
785:
783:
778:
747:
740:
733:
725:
721:
715:
703:
687:
683:
681:
676:
665:
646:
642:
634:
623:
615:
609:
589:
582:
578:Arctognathus
576:
561:, cataloged
536:
531:
519:
513:
507:
503:
491:
485:
476:
462:
454:
450:Arctognathus
448:
431:. The first
418:gorgonopsian
405:
404:
403:
382:
368:
359:
358:Synonyms of
338:
329:Arctognathus
327:
318:
317:Synonyms of
291:
290:
271:
270:
261:Type species
245:
244:
209:
196:
183:
132:
120:Overview of
24:
18:
2871:Gorgonopsia
2798:Sauroctonus
2792:Wikispecies
2768:Sauroctonus
2684:Cerdorhinus
2670:Arctosuchus
2572:Rubidgeinae
2561:Smilesaurus
2533:Eriphostoma
2490:Suchogorgon
2483:Sauroctonus
2416:Cyonosaurus
2409:Aloposaurus
2388:Gorgonopsia
2366:Gorgonopsia
2340:Gorgonopsia
2255:Gorgonopsia
2194:Geoheritage
1938:Sauroctonus
1536:Suchogorgon
1490:Scutosaurus
1480:, and even
1477:Dvinosaurus
1459:Idelesaurus
1403:Sauroctonus
1316:Rubidgeinae
1250:Smilesaurus
1161:Eriphostoma
1051:Sauroctonus
1027:Suchogorgon
957:Gorgonopsia
947:Sauroctonus
927:Suchogorgon
912:Sauroctonus
905:Sauroctonus
897:Sauroctonus
893:Sauroctonus
885:Sauroctonus
863:Sauroctonus
861:classified
855:Sauroctonus
808:Sauroctonus
786:Sauroctonus
770:156/5, the
758:Description
752:parringtoni
726:Sauroctonus
684:Sauroctonus
677:Sauroctonus
635:Sauroctonus
624:Sauroctonus
545:remains of
509:Suchogorgon
406:Sauroctonus
386:parringtoni
372:parringtoni
246:Sauroctonus
219:Gorgonopsia
124:156/5, the
25:Sauroctonus
2865:Categories
2626:Dinogorgon
2619:Clelandina
2610:Rubidgeini
2599:Sycosaurus
2349:see below↓
2297:Therapsida
2092:: 104631.
1576:References
1499:Eorasaurus
1486:pareiasaur
1482:sauropsids
1456:therapsid
1454:dicynodont
1435:polymictic
1423:Capitanian
632:genus name
590:progressus
421:therapsids
342:progressus
331:progressus
203:Therapsida
33:Capitanian
2691:Leogorgon
2547:Lycaenops
2540:Gorgonops
2526:Cynariops
2444:Scylacops
2437:Nochnitsa
2328:Synapsida
2313:Synapsida
2288:Synapsida
2271:Kingdom:
2214:128952446
2157:1471-2954
2114:249977414
1965:: 47–136.
1944:Paleofile
1829:: e4954.
1813:Nochnitsa
1694:462558622
1628:Cambridge
1597:: 47–123.
1565:Gorgonops
1442:tetrapods
1438:sandstone
1226:Lycaenops
1202:Cynariops
1178:Gorgonops
971:Nochnitsa
943:cladogram
932:pterygoid
880:Scylacops
868:Cynariops
851:taxonomic
772:lectotype
744:synapsids
628:lectotype
594:cynodonts
551:Tatarstan
437:Tatarstan
376:von Huene
297:von Huene
190:Synapsida
163:Kingdom:
157:Eukaryota
126:lectotype
2806:BioLib:
2777:Wikidata
2743:Category
2640:Rubidgea
2279:Chordata
2277:Phylum:
2273:Animalia
2175:32993469
2014:25186003
1990:(1793).
1915:30485338
1882:PLOS ONE
1855:29900078
1506:See also
1493:and the
934:and the
899:to be a
820:incisors
696:Tanzania
659:skeleton
641:σαῦρος (
488:Tanzania
479:include
346:Yefremov
308:Synonyms
225:Family:
177:Chordata
173:Phylum:
167:Animalia
153:Domain:
2835:4946274
2822:4975315
2783:Q132968
2554:Phorcys
2519:Arctops
2166:7542828
2094:Bibcode
2005:4173690
1906:6261584
1846:5995105
1666:: 7–18.
1267:Arctops
1144:Phorcys
812:palatal
569:of the
567:species
481:cranial
453:(named
443:of the
441:species
433:fossils
411:extinct
252:Bystrow
238:Genus:
100:↓
2809:650849
2429:nasuta
2425:Dixeya
2212:
2173:
2163:
2155:
2112:
2038:
2012:
2002:
1913:
1903:
1853:
1843:
1782:
1692:
1638:
1471:, the
1462:, the
792:. The
790:orbits
647:któnos
643:saûros
598:mammal
585:. The
555:Russia
543:fossil
539:Soviet
500:Russia
496:Africa
447:genus
392:, 1970
378:, 1950
348:, 1940
299:, 1950
254:, 1955
2848:39129
2293:Clade
2284:Clade
2210:S2CID
2110:S2CID
1823:PeerJ
1745:(PDF)
1716:[
1686:Paris
1543:Notes
1446:flora
1431:silts
1427:clays
936:vomer
901:basal
816:teeth
775:skull
730:clade
606:taxon
574:genus
559:skull
459:taxon
414:genus
210:Clade
197:Clade
184:Clade
129:skull
2817:GBIF
2326:see
2171:PMID
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