1110:
3180:
that in Atz' lobster the pigmentation of this organ can be controlled independently of that integument. It would seem that the retention of essential pigment by the dominant white has been naturally selected and that presumably its whole phenotype has been selected. Arctic white races and species no doubt have this type of allele. True recessive albinos lack tyrosinase (Onslow, 1917; Foster, 1952; Barnicott, 1957; Fitzpatrick et al., 1958) or have the enzyme but no free tyrosine (Verne, 1926, p. 535; Fox and Vevers, 1960, p. 37) but in the dominant white the enzyme is present in the skin, under inhibition.
2051:
1131:. Mutations causing changes in carotenoid-based colour pigments are rare; melanine mutations occur much more frequently. Two types of melanin, eumelanin and phaeomelanin, are present in birds. In the skin and eyes, only eumelanin is present. In some bird species, the colour is completely caused by eumelanin, however, both types of melanin are found in most species. In birds, albinism has been defined as "a total lack of both melanins in feathers, eyes and skin as a result of an inherited absence of tyrosinase", however, this ignores the effects of other pigments and structural colours.
2072:
479:
805:
1217:
66:
2031:
905:) along the coastline of the southern Western Ghats (Kerala and Kanyakumari coast, India). Albinism was observed in jungle cats from the Amaravila area of Trivandrum district in the Kerala State. Albinism in jackals was observed from the Polooni area in Malappuram district and Chaliyam area of Calicut district (Kerala). As albinism is observed in those areas where the density of these mammals is comparatively low, it is concluded that continuous inbreeding could be the reason for expression of albinism.
751:
1230:
960:
495:
whilst taking most taxa into account, ignore others. So, "a person or animal with very pale skin, white hair or fur, and pink eyes caused by a medical condition that they were born with" and "a person or animal with white skin and hair and pink eyes" do not include feathers, scales or cuticles of birds, fish and invertebrates, nor do they include plants. Some definitions are too broad to be of much use, e.g. "an animal or plant with a marked deficiency in pigmentation".
1687:
582:. Mammals have a gene that codes for the presence of tyrosinase in cells – called the TYR gene. If this gene is altered or damaged, melanin cannot be reliably produced and the mammal becomes an albino. Besides the TYR gene, several other genes can cause albinism. This is because other hormones and proteins are involved in melanin production, the presence of which is genetically determined. In mice, a total of 100 genes are known to affect albinism.
763:
2350:
2233:
1319:) differ in their characteristics. Normal individuals of this species are superior to albinos in body weight and total length. Albinos crossed with other albinos require 11 days longer to spawn and produce smaller egg masses. These masses contain eggs of lighter weight with poorer hatchability than crosses of normal fish. The albino fish have lower survival rates than normal fish but dress-out percentages are nearly equal.
41:
555:
89:
824:) is unusual for mammals as it is rich in cones. Central cell densities are less than 5% lower in albino squirrels than in pigmented individuals. This relatively minor disruption to vision is thought to assist in the survivability of albino squirrels in the wild. This is supported by observations that the behaviour of albinos in the wild, e.g. leaping from branch to branch, is similar to pigmented squirrels.
694:
however, it has been stated that in studies where animals had many places to hide, predators captured albino and normally coloured animals at the same rate. Furthermore, in certain species, albino animals may be excluded from families or other groups, or rejected as mates. On the other hand, there is also evidence for positive selection for albinism in some animals, as well as cultural selection favoring
648:
193:
1205:
1792:
1713:". The alligator is partially blind because of lack of pigment in its eyes. The albino alligator hatched from the egg in 1995 in Florida, and was brought to the academy in 2008. This alligator would not have survived in the wild because its whiteness would have made it too easy a prey object. The only known albino alligators are in captivity. While extremely rare, white-coloured
2245:
2176:
1255:. The gross eye morphology, feeding and swimming behaviours between wild-type and albino zebrafish were indistinguishable, except under dim or bright light or low contrast. In mammals, albinism is occasionally associated with hearing impairments. However, when tested, there was no differences in responses between wild-type and albinistic European wels catfish (
413:. The structures themselves are colourless. Light typically passes through multiple layers and is reflected more than once. The multiple reflections compound one another and intensify the colours. Structural colour differs according to the observer's position whereas pigments appear the same regardless of the angle-of-view. Animals that show iridescence include
660:). In nearly all mammals, the overwhelming majority of photoreceptors are rods rather than cones, which detect color. Albinism specifically affects the rod cells, but the number and distribution of the cones is unaffected. In contrast, the retinas of birds are cone rich, meaning that the vision of albino birds is affected less than albino mammals.
148:. Albinism is a well-recognized phenomenon in molluscs, both in the shell and in the soft parts. It has been claimed by some that albinism can occur for a number of reasons aside from inheritance, including genetic mutations, diet, living conditions, age, disease, or injury. However, this is contrary to definitions where the condition is inherited.
1095:
3179:
gene of the palomino horse is dominant and there is a dominant white in many mammals so that dominance/recessivity is not necessarily positively correlated with excess/deficiency. the dominant white is not so deficient in pigment as the true albino and in particular as the normal eye-pigmentation, so
2134:
that is caught in the wild or cultivated for food. Normal
Japanese sea cucumbers start to develop pigmentation when they are about 1 cm long. The upperside becomes a dull, yellowish -brown to maroon and the underside a light brown. The body walls of adult, albino Japanese sea cucumbers contain
693:
in some environments, and are therefore less able to conceal themselves from their predators or prey. However, in other environments albino animals are less likely to be killed, and selection may favor them. The lower survival rate of animals with albinism in certain environments has been documented,
498:
Other definitions of albinism encompass most of the major animal taxa, but ignore the several other pigments that non-mammalian animals have and also structural colouration. For example, "Absence of the pigment melanin in the eyes, skin, hair, scales, or feathers." refers only to the pigment melanin.
494:
Definitions of albinism vary and are inconsistent. While they are clear and precise for humans and other mammals, this is because the majority of mammals have only one pigment, melanin. Many animals have pigments other than melanin, and some also have structural colours. Some definitions of albinism,
1805:
The incidence of albinism in frogs, salamanders, and newts is relatively higher than other taxa. It has been estimated that one in four hundred of these animals is albino. When albino tadpoles hatch, they are almost transparent. This may help camouflage them initially, however, after two weeks, when
1760:
are the most likely type of turtle to express albinism. The shells have an almost yellow colouration and they have pink eyes. For turtles, a pure white colour is nearly impossible, even with albinism. Albino turtles can have a longer lifespan than many other albino animals; their hard shells help to
718:
fish in the laboratory, i.e. with no predators, sufficient food supply, controlled temperatures, etc., found that albinos had considerably reduced viability; from 800 albino embryos, only 29 survived to full adulthood. Early studies on fish led some researchers to describe albinism as a "semi-lethal
534:
One definition states that "albinism, (from the Latin albus, meaning "white"), hereditary condition characterized by the absence of pigment in the eyes, skin, hair, scales, or feathers", however, this does not encompass invertebrates, nor does it include plants. Furthermore, it could be interpreted
3619:
In the Hopi culture, albinos were considered special and given special status. Because extensive exposure to sunlight could be damaging or deadly, Hopi male albinos did no agricultural work. Albinism was considered a positive trait rather than a negative physical condition, allowing albinos to have
3174:
Some dominant chromogenic mutations are known and in fact there are more than predicted by the
Haldane-Fisher theory of evolution of wild type dominance (Searle, 1968, p. 250)"..."Of 32 dominant mutations known in the mouse as many as nine are chromogenic (Hadorn, 1961, p. 132). This is a very high
601:
Albinism occurs throughout the animal kingdom. The condition is most commonly seen in birds, reptiles and amphibians, but more rarely seen in mammals and other taxa. It is often difficult to explain occasional occurrences, especially when only one documented incidence has occurred, such as only one
1122:
The most important pigments that determine plumage colouration in birds are melanines and carotenoids. The latter are ingested in food and transformed into colour pigments by enzymes. Aberrations in this pigmentation are mostly caused by food deficiencies and usually do not have a genetic basis.
626:
Melanin functions in the normal development of various parts of the eye, including the iris, retina, eye muscles, and optic nerve. The absence of melanin results in abnormal development of eyes and leads to problems with focusing, and depth perception. The eyes of albino animals appear red because
1744:
In snakes, partial absence of pigment is more common than absolute albinism. For snakes that are usually patterned in colours, they appear as a faint blue, peach or yellowish. In these cases, there has been a genetic mutation in the melanin and pigment delivery. The appearance comes from the
3588:
Indeed, two
Mendelian pigment phenotypes (albinism and brown) are both widespread, highly penetrant, and mediated through loss-of-function mutations in the genes Oca2 and Mc1r, respectively . Recent studies have provided evidence of selection for albinism as a means of increasing catecholamine
545:
A clear definition appears to be – "Congenital absence of any pigmentation or colouration in a person, animal, or plant, resulting in white hair and pink eyes in mammals." Whilst this does not state specifically that non-mammalian albino animals (or plants) are white, this can be inferred from
135:
Varied use and interpretation of the terms mean that written reports of albinistic animals can be difficult to verify. Albinism can reduce the survivability of an animal; for example, it has been suggested that albino alligators have an average survival span of only 24 hours due to the lack of
510:
The term "partial albino" is sometimes used in the literature. However, it has been stated that "A common misnomer is 'partial albino' – this is not possible since albinism affects the whole plumage of a bird, not just part" and the definition of albinism precludes the possibility of "partial
1787:
As with reptiles, many amphibians labeled as albino are, in fact, not completely lacking in all colour pigments. They are actually amelanistic, not albino. Amphibians have six types of chromatophore in their skin, i.e. melanophores, xanthophores, erythrophores, leucophores, cyanophores and
795:
Many individual albino mammals are in captivity and were caught while young. However, it is worth noting that the survival prospects of these creatures into adulthood outside of captivity are uncertain. It is doubtful whether these individuals would have survived to become adults without the
2019:
has been recognized to be a hereditary phenomenon at least since 1900. Albinism in molluscs can exist to a variable degree. Sometimes an individual snail has a normally pigmented body, but the shell is completely without the normal pigmentation because of a defect in the cells of the
970:
The costs of albinism for marine mammals may include reduced heat absorption in colder waters, poor camouflage from predators, increased sensitivity to sunlight, and impaired visual communication. Despite the costs, some individuals do reach adult age and breeding status.
2050:
3643:
There is no apparent reason why selection for albinism should have been greater at Morton Plains thanelse where in north-western
Victoria. It is likely that its high occurrence there is a founder effect from the domestic-type feral population established during the
2071:
2030:
1779:, however, breeders state that albinos are generally recognized by their lack of black pigment and having red eyes is not a requirement to be considered an albino. These three strains are called "tremper albinos", "rainwater albinos" and "bell albinos".
589:
horses, genes coding for coat whiteness are dominant, and this is also true for several arctic mammals who possess dominant white colors which are pseudo-albinistic. However, these mammals differ from truly recessive albinos in that they still produce
4322:
Protas, Meredith E; Hersey, Candace; Kochanek, Dawn; Zhou, Yi; Wilkens, Horst; Jeffery, William R; Zon, Leonard I; Borowsky, Richard; Tabin, Clifford J (2006). "Genetic analysis of cavefish reveals molecular convergence in the evolution of albinism".
680:
from the sun while allowing beneficial frequencies to enter the body. This means some animals may die from UV radiation due to a lack of protection. Albino humans must use an excessive amount of sunscreen, even if the sun is hidden behind the clouds.
106:
2218:
is the direct opposite of albinism. An unusually high level of melanin pigmentation (and sometimes absence of other types of pigment in species that have more than one) results in an appearance darker than non-melanistic specimens from the same
502:
Because of the various uses of different terms applied to colouration, some authors have indicated that the colour of the eyes is the defining characteristic of albinism, e.g. "This leads to a good diagnostic feature with which to distinguish
1177:
However, it has been argued that the definition of albinism precludes the possibility of "partial albinism" in which a mostly white bird shows some form of melanin pigmentation. "It is simply impossible, just like being 'partially pregnant'.
916:) found that albinism in this species conferred an advantage for the males compared to the wild-type; albino males had higher mount frequencies than wild-type males. In addition, the albinos had greater differential fertilizing capacity.
609:
Some species, such as white peacocks, swans and geese, are not believed to be true albinos, as they do not have red eyes, rather, their colouration is suggested to be the expression of a white fur or feather gene, not a lack of melanin.
2277:, usually on the base of the (normal) redwood tree from which it first grew. Only about sixty examples of albino redwoods are known. Additionally, an even smaller number of "chimeric" redwood trees have both normal and white needles.
1729:
with some colour tinges remaining on their bodies although looking at first like other albino creatures, thus creating the misconception that the reptiles are albino themselves when they are not. Four such alligators are kept at the
1273:) do not differ in hearing sensitivity from the normally pigmented and eyed surface-dwelling populations. Fish lack melanin in the inner ear, meaning that hearing in fishes is less likely to be affected by albinism than in mammals.
2145:
is the main carotenoid in marine crustaceans (and fish). It has been shown that adding astaxanthin to the feed can improve the skin and muscle colour of marine organisms and thereby increase their commercial and ornamental value.
179:, which can reduce survivability. Some plant variations may have white flowers or other parts. However, these plants are not totally devoid of chlorophyll. Terms associated with this phenomenon are "hypochromia" and "albiflora".
2260:, which can reduce survivability. Some plant variations may have white flowers or other parts. However, these plants are not totally devoid of chlorophyll. Terms associated with this phenomenon are "hypochromia" and "albiflora".
974:
Albino dolphins were first sighted in the Gulf of Mexico in 1962. Since 1994, three further individuals have been seen. These tend to be pink in colour due to blood vessels showing through the blubber and unpigmented skin.
655:
In all albino mammals studied, the centre of the retina is under-developed and there is a deficit of rod cells, which detect light levels; the central ganglion cell density is approximately 25% below normal (except for the
791:
strains, laboratory rabbits and ferrets. Some researchers have argued that albino animals are not always the best choice for scientific studies due to the consequences of albinism (e.g. hearing and visual impairments).
1698:, not albino. Reptiles often possess at least two pigments. Among the most common are xanthin (yellow) and erythrin (red). An amelanistic reptile therefore, may still have pale yellow, orange, or red pigmentation.
705:
The novelty of albino animals has occasionally led to their protection by groups such as the Albino
Squirrel Preservation Society. They have also been protected in studies on their ecology, sociology and behaviour.
1745:
inability for full colours to be present, such as black, red, brown and others. The eyes of an albino snake are typically red or pink. Albino snakes can remain in the sunlight for several hours with minimal harm.
635:
can be seen through the iris, which has no pigment to obscure this. Some albino animals may have pale-blue eyes due to other colour generating processes. Albino vertebrates exposed to intense light typically lose
3029:
136:
protection from UV radiation and their lack of camouflage to avoid predators. It is a common misconception that all albino animals have characteristic pink or red eyes (resulting from the lack of pigment in the
4895:
Hoste, B.; Simpson, S. J.; Tanaka, S.; De Loof, A.; Breuer, M. (2002). "A comparison of phase-related shifts in behavior and morphometrics of an albino strain, deficient in -corazonin, and a normally colored
2626:
Gunnarsson, Ulrika; Hellström, Anders R.; Tixier-Boichard, Michele; Minvielle, Francis; Bed'hom, Bertrand; Ito, Shin'ichi; Jensen, Per; Rattink, Annemieke; Vereijken, Addie; Andersson, Leif (February 2007).
511:
albinism" in which a mostly white bird shows some form of melanin pigmentation. "It is simply impossible, just like being 'partially pregnant' ". Conditions that are commonly termed "partial albino" include
570:(producing light reddish tans and blondes). The dark and light melanins have their influence either alone or in conjunction, making either plain or multi-coloured coats. Sometimes, in a condition called
5192:
4719:
4081:
507:
and albino individuals – the colour of the eye." However, there are multiple forms of albinism – currently seven types recognised for humans – most of which do not result in red or pink pupils.
1243:
have three types of chromatophores—iridophores, melanophores, and xanthophores—which produce silver, black, and yellow pigmentation respectively. Zebrafish that lack iridophores are known as
4367:
1164:
Total albinism – a simultaneous complete absence of melanin from the eyes, skin, and feathers. This is the rarest form. Only 7% of 1,847 cases of avian albinism examined was this type.
1281:
There are several reports of total albinism in both freshwater and marine fish, however, frequently captured albino fish are only reported in aquarium magazines and local newspapers.
1127:, which owe their distinct pink colour to the presence of red carotenoids in their natural food. When these carotenoids are in short supply, these birds appear white after the next
546:"...absence of any pigmentation or colouration..." Due to the varied use and interpretation of the term "albino", written reports of albinistic organisms can often not be verified.
4597:
538:
The lack of clarity about the term is furthered when the name of an animal includes the term "albino" although the animals (clearly) do not have the condition. For example, the
3472:
Consider albino animals of the arctic. Hungry predators may have unwittingly selected for the albino color variant by eating more of the easily visible dark-colored variants.
252:. What is perceived as a plant or animal's "colour" is the wavelengths of light that are not absorbed by the pigment, but instead are reflected. Biological pigments include
4938:
3890:
3044:
3407:"Reduced retinal deficits in an albino mammal with a cone rich retina: A study of the ganglion cell layer at the area centralis of pigmented and albino grey squirrels"
162:, resulting in reduced pigmentation. Type I oculocutaneous albinism (OCA1a) is the form most commonly recognised as 'albino' as this results in a complete absence of
2208:
differs from albinism in that the melanin is, at least, partially absent but the eyes retain their usual colour. Some leucistic animals are white or pale because of
3357:
Allison, W. T.; Hallows, T. E.; Johnson, T.; Hawryshyn, C.W.; Allen, D. M. (2006). "Photic history modifies susceptibility to retinal damage in albino trout".
2139:
is thinner in the albinos and contains fewer melanocytes. Albino individuals are similar to normal individuals in growth rate, digestion rate and fertility.
166:
in the skin, hair/fur/feathers, and pink pupils, however this has led many to assume that all albinos are pure white with pink pupils, which is not the case.
140:
allowing the blood vessels of the retina to be visible); this is not the case for some forms of albinism. Familiar albino animals include in-bred strains of
128:
in an animal or plant resulting in white hair, feathers, scales and skin and reddish pink or blue eyes. Individuals with the condition are referred to as
5204:
3102:
4738:
1109:
5278:
2007:(an amphibian) are also used widely in the laboratory as their transparent skin allows observation of the underlying tissues during limb regeneration.
2501:
562:
Melanin is an organic pigment that produces most of the colour seen in mammals. Depending on how it is created, melanin comes in two colour ranges,
4255:
3175:
percentage of the 40 coal-colour genes known among the mouse and implies that dominant mutations are rather characteristic of colour genes. The
1186:
As with other animals, it has been stated that for fish to be properly described as "albino", they must have a white body and pink or red eyes.
2161:) occasionally appear in laboratory colonies, however, these still contain green and yellow pigments. This albinism does not affect mortality.
978:
A report published in 2008 stated that in marine mammals, "anomalously white" individuals have been reported for 21 cetacean species and seven
249:
4054:
1741:. In Australia, a crocodile believed to be "part-albino" and nicknamed by people in the area as "Michael Jackson", attacked and killed a man.
417:
seashells, fish, and peacocks. These are just a few examples of animals with this quality, but it is most pronounced in the butterfly family.
5351:
4975:
4398:
4008:
3801:
3707:
3612:
3465:
3250:
3167:
3137:
2037:
1666:
There are several reports of albino lampreys and it has been estimated that this occurs at a frequency of one in 100,000 normal individuals.
2135:
only 0.24% melanin compared to 3.12% in normal adults. The difference in melanin content becomes visually apparent at 60 days of age. The
531:
that have been identified in humans (and confirmed in some other animals) and the use of the term "partial albino" is therefore misleading.
1932:) has been attributed to one of three mutations that created a dysfunctional tyrosinase. Two of those mutations involve an insertion of a
5112:
4592:
5463:
4671:
818:
It has been claimed that "Squirrels are the only known albino mammal to survive successfully in the wild". The retina of the squirrel (
346:
for colouration. The term chromatophore can also refer to coloured, membrane-associated vesicles found in some forms of photosynthetic
4024:
2449:
1330:
is a species that has evolved specialized characteristics in a series of independent caves. One of these is albinism linked to the
232:
2993:
4575:
1303:. Albinism has been reported in hagfish, lampreys, sharks, rays and numerous teleost fishes, e.g. catfishes, grunts or cyprinids.
1138:
and pink or red eyes. Albinism is only seen in about 1 of every 1,800 birds. The two most common species of albino birds are the
727:
Pigmentation disorders such as albinism are occasionally associated with hearing impairments in mice, rats, guinea pigs and cats.
4935:
3909:
2180:
4095:
3331:
3487:. The Louise Lindsey Merrick Texas Environment Series, No. 6, pp. 28–31. Texas A&M University Press, College Station (1983)
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3860:
2154:"Depigmented" arthropods have been found, usually in cave populations. "Albino" individuals of normally red citrus red mites (
5440:
2273:
are rare examples of an albino tree with white needles; despite its lack of chlorophyll it may grow to substantial size as a
487:
214:
55:
3633:
4295:
Bondari, K. (1984). "Comparative performance of albino and normally pigmented channel catfish in tanks, cages, and ponds".
3791:
3240:
2036:
An exhibit showing albino and normal specimens of nine local species of marine molluscs, both gastropods and bivalves (The
5431:
5387:
3257:
RSA was also used to identify markers associated with the dominant albino phenotype in a
Japanese strain of rainbow trout.
3001:
2369:
1906:
1702:
1031:, Louisiana; Carolina Snowball, a popular albino bottlenose dolphin displayed at the Miami Seaquarium in the early 1960s;
424:(reflective/iridescent): sometimes called "guanophores", reflect light using plates of crystalline chemochromes made from
2240:
seedling. The two seedlings are of the same age, but the albino stopped developing once the seed's reserve was exhausted.
358:
during embryonic development. Mature chromatophores are grouped into subclasses based on their colour under white light:
1924:, some of which have mutations that show no tyrosinase or DOPA oxidase activity in albinos. The albino phenotype of the
4619:
1753:
or diamondback snakes, are the most commonly affected by albinism often appearing to be a pinkish or yellowish colour.
154:(OCA) is a clearly defined set of seven types of genetic mutations which reduce or completely prevent the synthesis of
5489:
2116:
is a mutation causing a congenital lack of normal pigment; it is a similar phenomenon to albinism in other organisms.
1346:, several species of naturally occurring albino rays and sharks have been recorded. Furthermore, an albino individual
478:
5416:
3074:
1475:
1032:
848:
827:
A 2012 survey of the literature reported that in India, there were several records of albino mammals including the
4958:
Zhao, H.; Chen, M.; Yang, H. (2015). "Albinism". In Yang, Hongsheng; Hamel, Jean-Francois; Mercier, Annie (eds.).
203:
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5253:
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1925:
1525:
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677:
606:. In mammals, albinism occurs once in every 10,000 births, but in birds, the rate is once in every 1,764 births.
210:
2101:. The Okinawa strain of this species is deficient in -corazonin and is albino. One of the typical features of
1694:
Many reptiles labeled as albino are, in fact, not completely lacking in all colour pigments. They are actually
3455:
2686:"The Mouse p (pink-eyed dilution) and Human P Genes, Oculocutaneous Albinism Type 2 (OCA2), and Melanosomal pH"
2320:
804:
3999:
Fertl, D.; Rosel, P. E. (2008). "Albinism". In Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.).
3532:"A pleiotropic interaction between vision loss and hypermelanism in Astyanax mexicanus cave x surface hybrids"
2778:
1772:), approximately 12 cm (4.7 in) long and roughly 1 year old, had been found in Victoria, Australia.
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5286:
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1826:
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832:
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528:
151:
112:
95:
20:
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Most forms of albinism follow a recessive pattern of inheritance. However, this is not always the case. In
2575:
2126:
1957:
844:
2517:
144:(rats, mice and rabbits), but populations of naturally occurring albino animals exist in the wild, e.g.,
5170:
3276:
2738:"Albinism in plants: A major bottleneck in wide hybridization, androgenesis and doubled haploid culture"
2079:
2061:
1898:
1806:
their hindlegs begin to emerge, they become milky white. A survey in 2001 found hundreds of tiny albino
1147:
657:
5308:
4270:
1761:
prevent predation and other environmental challenges. Vision and sensory organs are slightly affected.
2169:
In some animals, albinism-like conditions may affect other pigments or pigment-production mechanisms:
1768:
was reported and photographed. In 2007, it was reported that an albino stumpy-tail lizard (possibly a
1229:
959:
750:
5337:
4821:
2513:
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pigments and incomplete differentiation of chloroplast membranes. Albinism in plants interferes with
1965:
1910:
1834:
820:
520:
175:
pigments and incomplete differentiation of chloroplast membranes. Albinism in plants interferes with
997:
Whales and dolphins also may appear white if extensively scarred, or covered with a fungus, such as
2390:
2327:
2319:
petitioned for an election to name their albino squirrel as the university's secondary mascot. The
2312:
2105:
is that they are gregarious locusts. However, the albino strain shows more solitarious behaviour.
1937:
1850:
1807:
1620:
1139:
808:
571:
406:
245:
65:
2857:
Taylor, J. D. (1969). "The effects of intermedin on the ultrastructure of amphibian iridophores".
2601:
2542:
618:
Melanin has several functions in most mammals and other animals; these are disrupted by albinism.
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5004:
4545:
4390:
4348:
4162:
3842:
3579:
3436:
3382:
3221:
2757:
2455:
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2199:
1810:, but when the researcher returned a few months later, not a single albino adult could be found.
1769:
1710:
1706:
1600:
1415:
1405:
1327:
1269:
1234:
1167:
Incomplete albinism – when melanin is not simultaneously absent from the eyes, skin and feathers.
1160:
In one study, albinism in birds has been categorised according to the extent of pigment absence.
1024:
788:
695:
637:
594:, and have normal eye pigmentation. In Japan, research has identified a dominant albinism in the
145:
141:
51:
47:
27:
2024:. Shells of certain mollusc species can be translucent when they lack the normal pigmentation.
3699:
2892:
5484:
5411:
5198:
5109:
4971:
4917:
4879:
4843:
4802:
4794:
4537:
4489:
4340:
4154:
4004:
3834:
3797:
3769:
3703:
3608:
3571:
3553:
3461:
3428:
3374:
3246:
3213:
3163:
3157:
3133:
3127:
2874:
2839:
2805:
2715:
2707:
2666:
2648:
2445:
2173:"Whiteface", a condition that affects some parrot species, is caused by a lack of psittacins.
1981:
1686:
1630:
1495:
1395:
1385:
1296:
840:
698:
in some human societies, which results in the increase of genes associated with albinism. The
402:
269:
4862:
4764:
1334:
gene, a known pigmentation gene. This trait has evolved independently in at least two caves.
5360:
5235:
5035:
4909:
4833:
4784:
4776:
4527:
4382:
4332:
4304:
4234:
4197:
4144:
3974:
3826:
3759:
3751:
3665:
3561:
3543:
3510:
3418:
3366:
3205:
3069:
2910:
2866:
2829:
2821:
2749:
2697:
2656:
2640:
2437:
2041:
2021:
1757:
1675:
1263:
940:
512:
275:
2065:,. Note that in the albino both the body and the shell are lacking the normal pigmentation.
762:
5116:
4942:
3669:
2968:
2349:
2281:
1818:
1570:
1347:
1312:
1059:
1020:
414:
257:
5226:
Kerr, E. A. (1963). "Inheritance of crinkle, variegation, and albinism in sweet cherry".
5055:
4765:"Genetic and metabolic bases of two "albino" phenotypes in the leopard frog,Rana pipiens"
3030:"Not every white bird is an albino: Sense and nonsense about colour aberrations in birds"
2569:
2441:
2232:
1546:
A study published in 2006 reported albinistic individuals of the following ray species:
1284:
The incidence of albinism can be artificially increased in fish by exposing the eggs to
248:
are substances produced by living organisms that have a colour resulting from selective
4967:
3764:
3739:
3692:
3566:
3531:
2940:
2834:
2809:
2661:
2628:
2405:
2400:
2355:
2270:
2257:
2248:
The foliage of an Albino redwood. Note the distinctive white coloration of the needles.
1580:
1355:
1343:
1300:
1257:
1143:
1051:
1028:
1016:
879:
871:
776:
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554:
447:
351:
253:
176:
137:
4913:
4694:
4645:
4149:
4132:
3423:
3406:
2702:
2685:
1788:
iridophores. An amelanistic amphibian therefore, may still have various pigmentation.
278:. Animals may have both biological pigments and structural colours, for example, some
40:
5478:
5123:
online archive; sourced
December 2006, actual authoring/publication date unspecified.
4308:
4256:"Albino zebra shark from the Indian ocean with comments on albinism in elasmobranchs"
4032:
3936:"Albinism in jungle cat and jackal along the coastline of the southern Western Ghats"
3830:
3755:
3583:
3515:
3498:
3194:"Genetic mapping of the dominant albino locus in rainbow trout (Oncorhynchus mykiss)"
2963:
2870:
2459:
2374:
2209:
1989:
1776:
1640:
1485:
1435:
1365:
1036:
999:
863:
595:
311:
4549:
4394:
4352:
4239:
4218:
4202:
4181:
4166:
4106:
3959:"Effects of albinism on copulatory behavior and sperm competition in prairie voles (
3846:
3817:
Creel, D. (June 1980). "Inappropriate use of albino animals as models in research".
3440:
3386:
2761:
1944:
of the TYR protein that is defective. The other mutation involves the deletion of a
527:. These conditions result from fundamentally different causes to the seven types of
88:
26:
This article is about the biological term. For the medical condition in humans, see
5445:
5421:
5088:
4822:"Spontaneous tyrosinase mutations identified in albinos of three wild frog species"
3225:
2737:
2285:
2202:
of melanin, resulting in reduction or absence of red and yellow pteridine pigments.
1945:
1941:
1842:
1726:
1560:
1455:
1285:
964:
909:
669:
628:
535:
that "...absence of pigment..." does not include an absence of structural colours.
371:
355:
2629:"Mutations in SLC45A2 Cause Plumage Color Variation in Chicken and Japanese Quail"
775:
Intentionally bred albinistic strains of some animal species are commonly used as
3635:
Founder
Effects in Some Victorian Wild Rabbit Oryctolaguscuniculus L. Populations
3602:
2825:
574:, they make multi-coloured individual hairs. The production of melanin occurs in
5309:"Albino And Leucistic White Animals Are Extremely Rare And Incredibly Beautiful"
4449:
Sandoval-Castillo, J.; Mariano-Melendez, E.; Villavicencio-Garayzar, C. (2006).
4082:"Albino Animals from Snowflake the white gorilla to White Diamond the alligator"
3589:
levels, which promote adaptive feeding and sleep behaviors in
Astyanax cavefish.
2644:
2253:
2192:
2142:
2084:
1866:
1858:
1765:
1750:
1590:
1535:
1515:
1505:
1151:
1069:
1043:
890:
647:
575:
567:
539:
463:
451:
410:
380:
362:
279:
192:
172:
159:
5165:
4562:
1791:
350:. Chromatophores are largely responsible for generating skin and eye colour in
5450:
5383:
5377:
4532:
4511:
4366:
Reum, J. C. P.; Paulsen, C. E.; Pietsch, T. W.; Parker-Stetter, S. L. (2008).
3548:
3370:
2810:"Mechanisms and behavioural functions of structural coloration in cephalopods"
2753:
2395:
2379:
2345:
2331:
2292:
2264:
2195:
2156:
2131:
2113:
2057:
1949:
1890:
1814:
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1734:
1718:
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983:
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673:
591:
579:
516:
459:
430:
421:
389:
366:
343:
331:
327:
4991:
Locket, N. A. (1986). "Albinism and eye structure in an
Australian scorpion,
4798:
3557:
2711:
2652:
5397:
5040:
5023:
3193:
2476:
2384:
2300:
2244:
2220:
2188:
2136:
1894:
1886:
1799:
1795:
1731:
1695:
1465:
1323:
1240:
1170:
Imperfect albinism – when melanin is reduced in the eyes, skin and feathers.
1064:
924:
920:
852:
641:
563:
504:
455:
375:
315:
155:
5056:"The Parblue Puzzle: Part 4 – Common Parblue Varieties: The Cockatiel [
4921:
4847:
4780:
4763:
Smith-Gill, Sandra J.; Richards, Christina M.; Nace, George W. (May 1972).
4344:
4158:
3773:
3575:
3378:
3217:
2935:
2843:
2719:
2670:
1173:
Partial albinism – when albinism is localized to certain areas of the body.
5119:
by Dr. Julie Feinstein of the American Museum of Natural History (NY), in
4820:
Miura, Ikuo; Tagami, Masataka; Fujitani, Takeshi; Ogata, Mitsuaki (2017).
4806:
4541:
3838:
3432:
3209:
3144:
The inheritance pattern for most forms of albinism is autosomal recessive.
2878:
2175:
1361:
Albinistic individuals of the following shark species have been reported:
542:
has this name because it is ash-grey whereas other gaur are almost black.
105:
4838:
4789:
4028:
2296:
2274:
2215:
1993:
1134:
An albino bird has a white beak, white plumage, non-coloured skin, white
1124:
979:
932:
586:
524:
467:
347:
299:
4620:"Albino crocodile that killed a fisherman shot dead near Darwin – video"
4131:
Ren, J. Q.; McCarthy, W. R.; Zhang, H.; Adolphc, A. R.; Lia, L. (2002).
3935:
5426:
5008:
3979:
3958:
2782:
2411:
2335:
2237:
2205:
2087:. Without its normal pigment, the shell of this species is translucent.
2016:
2004:
1997:
1985:
1977:
1973:
1933:
1917:
1738:
1722:
1550:
1292:
1154:
1039:
1012:
991:
944:
928:
898:
859:
483:
425:
384:
323:
217: in this section. Unsourced material may be challenged and removed.
163:
125:
4133:"Behavioral visual responses of wild-type and hypopigmented zebrafish"
811:, a blue-eyed western lowland gorilla caught in the wild with Type IV
702:
may explain why some animal populations become selected for albinism.
1953:
1914:
948:
836:
780:
715:
632:
434:
392:(blue): limited taxonomic range but found in some fish and amphibians
335:
5239:
4881:
Monograph of the Land & Freshwater Mollusca of the British Isles
4864:
Monograph of the Land & Freshwater Mollusca of the British Isles
3497:
Dobosz, ByS; Kohlmann, K.; Goryczko, K.; Kuzminski, H. (July 2008).
2252:
In plants, albinism is characterised by partial or complete loss of
310:, scales and cuticles, contain pigments in specialized cells called
171:
In plants, albinism is characterised by partial or complete loss of
5329:
5193:"A Creepy Monster of the Forest: The Albino, Vampiric Redwood Tree"
4336:
4223:(Siluriformes: Loricariidae) from an Atlantic Forest coastal basin"
1094:
5435:
5391:
4450:
2323:
in Kentucky also has a documented population of albino squirrels.
2243:
2231:
2174:
1790:
1228:
1128:
1047:
958:
936:
828:
803:
646:
603:
553:
477:
2736:
Kumari, M.; Clarke, H. J.; Small, I.; Siddique, K. H. M. (2009).
1709:, as of 2015, has on display an albino American alligator named "
4512:"A seminoma and a leiomyosarcoma in an albino African lungfish (
4061:
2097:
The neurohormone -corazonin induces darkening of the cuticle of
1331:
1135:
339:
319:
307:
291:
274:
Animals can appear coloured due to two mechanisms, pigments and
5333:
4451:"New records of albinism in two elasmobranchs: The tiger shark
2432:
Oetting, William S; Adams, David (2018). "Albinism: Genetics".
3620:
more children and thus increasing the frequency of the allele.
2263:
Plants that are pale simply from being in the dark are termed
779:
in biomedical research and also as pets. Examples include the
303:
295:
186:
3656:
Menzel, R. W. (1958). "Further notes on the albino catfish".
3280:
1721:
do exist in other places. However, most of these animals are
1251:, and those which lack both melanophores and iridophores are
982:
species but there were no known reports of anomalously white
437:(often guanine) to produce a reflective, shiny, white colour.
314:. These cells are found in a wide range of animals including
4182:"First record of partial albinism in two catfish species of
2311:
The Albino Squirrel Preservation Society was founded at the
2303:
in plants, even several seasons or years after applicating.
433:(reflective white): found in some fish, utilize crystalline
383:(black/brown): contain black and brown pigments such as the
2191:
is a condition common in reptiles and amphibians, in which
5064:
The Genetics of Colour in the Budgerigar and Other Parrots
2408:, unusually yellow pigmentation and lack of yellow pigment
1288:(e.g. arsenic, cadmium, copper, mercury, selenium, zinc).
743:
Examples of pet mammals artificially selected for albinism
5135:"It's no snow job – handful of redwoods are rare albinos"
4186:(Siluriformes: Ariidae) in an estuary of Southern Brazil"
3242:
Genome Mapping and Genomics in Fishes and Aquatic Animals
2596:
2594:
2592:
4387:
10.1898/1051-1733(2008)89[60:FROAAC]2.0.CO;2
3457:
Myths Of Gender: Biological Theories About Women And Men
374:(red): contain reddish pigments such as carotenoids and
2547:
National Organization for Albinism and Hypopigmentation
4217:
de Brito, Marcelo F. G.; Caramaschi, Érica P. (2005).
3499:"Growth and vitality in yellow forms of rainbow trout"
4003:(2nd ed.). Amsterdam: Elsevier. pp. 22–24.
3245:. Springer Science & Business Media. p. 29.
454:
along with several red and yellow pigments including
3641:. Field Naturalists' Club of Victoria. p. 100.
2326:
Albino animals are often kept as pets, for example,
1913:
tyrosinase gene which still has some tyrosinase and
862:
have been reported in several occasions including a
664:
Reduced protection from sunlight in albino creatures
5085:
LivingUnderworld.com:A Web Project About Amphibians
1197:
Examples of fish artificially selected for albinism
651:
A true albino squirrel. Note the reddish-pink eyes.
4591:
3691:
3132:. Lippincott Williams & Wilkins. p. 641.
2315:at Austin in 2001. Members of the society at the
2056:Normal (left) and albinistic (right) forms of the
1299:(flatfish), however, it is rarely reported in the
1291:In the wild, albinism is reasonably common in the
558:A hedgehog with albinism due to a genetic mutation
365:(yellow): contain yellow pigments in the forms of
3785:
3783:
2988:
2986:
2471:
2469:
3891:"A case of total albinism in the Bonnet Macaque
3889:Mahabal, Anil; Rane, P. D.; Pati, S. K. (2012).
3300:
3298:
2212:(pigment cell) defects, and do not lack melanin.
4576:"Albino alligator Claude to mark 15th Birthday"
3934:Sanil, R.; Shameer, T. T.; Easa, P. S. (2014).
3126:Schalock, Peter C.; Hsu, Jeffrey T. S. (2012).
2606:NORD (National Organization for Rare Disorders)
2280:Albinism has frequently occurred in progeny of
1115:Blue winged kookaburra with normal pigmentation
796:protection and care they receive in captivity.
5028:Annals of the Entomological Society of America
4368:"First record of an albino Chimaeriforme fish"
3239:Kocher, Thomas D.; Kole, Chittaranjan (2008).
2537:
2535:
446:The primary function of pigments in plants is
5345:
5024:"A recessive albinism in the citrus red mite"
4718:Modesti, A.; Aguzzi, S.; Manenti, R. (2011).
4713:
4711:
1678:has been reported on at least two occasions.
523:, or other depigmentation conditions such as
19:"Albino" redirects here. For other uses, see
8:
5279:"10 Most Incredible Albino Animals on Earth"
4695:"Guide to Leopard Gecko Morphs and Genetics"
4190:Brazilian Archives of Biology and Technology
2773:
2771:
342:, in contrast, have a class of cells called
4953:
4951:
4884:. Vol. 1. Taylor Brothers. p. 90.
4867:. Vol. 1. Taylor Brothers. p. 92.
4593:"White alligator is one of rarest in world"
4444:
4442:
4055:"Albino dolphin in northern Gulf of Mexico"
3460:(Rev. ed.). Basic Books. p. 172.
3400:
3398:
3396:
1756:Albino tortoises and turtles are uncommon;
1326:have populations that are albinistic. The
71:American alligator with normal pigmentation
5352:
5338:
5330:
5159:
5157:
4025:"Rare Pink Dolphin Seen in Louisiana Lake"
3733:
3731:
3729:
3096:
3094:
3092:
2543:"Information Bulletin – What is Albinism?"
1996:(Arg), respectively. These changes in the
1775:There are three known "albino" strains of
689:Animals with albinism may lack protective
578:in a complex process involving the enzyme
5166:"Albino redwoods hold scientific mystery"
5039:
4837:
4788:
4531:
4238:
4201:
4148:
3994:
3992:
3990:
3978:
3763:
3685:
3683:
3681:
3679:
3565:
3547:
3514:
3422:
3271:
3269:
3267:
3265:
2833:
2808:; Marshall, N. J.; Hanlon, R. T. (2009).
2701:
2660:
2083:showing the red oxygen-transport pigment
233:Learn how and when to remove this message
4510:Hubbard, G. B.; Fletcher, K. C. (1985).
3796:. Minneapolis: Lerner Publishing Group.
3023:
3021:
3019:
2731:
2729:
1685:
1267:). Similarly, Mexican blind cave fish (
401:Animals can also appear coloured due to
3884:
3882:
3867:. Texas Parks & Wildlife Department
3279:. Wonders of the Nature. Archived from
2462:– via John Wiley & Sons, Ltd.
2424:
2026:
1725:given that they have a general loss of
622:Abnormal eye development and appearance
566:(producing dark browns and blacks) and
4668:"Rare albino lizard found in backyard"
3819:Pharmacology Biochemistry and Behavior
2814:Journal of the Royal Society Interface
1905:) has been attributed to a failure in
1749:and snakes of larger types, such as a
5077:"Amphibian Biology & Physiology:
4600:from the original on January 12, 2022
4574:Guthrie, Julian (September 14, 2010)
4427:. ReefQuest Centre for Shark Research
3957:Dewsbury, D. A.; Ward, S. E. (1985).
3670:10.1093/oxfordjournals.jhered.a106828
3129:Lippincott's Primary Care Dermatology
2859:General and Comparative Endocrinology
2038:Bailey-Matthews National Shell Museum
1261:) and South American bronze catfish (
81:Examples of albino laboratory mammals
16:Disorder causing lack of pigmentation
7:
5164:Krieger, L. M. (November 28, 2010).
3698:. London: Chapman and Hall. p.
3604:Transmission and Population Genetics
2284:, Bing and Hedelfingen varieties of
1817:, albinism has been recorded in the
215:adding citations to reliable sources
5464:Pre-modern conceptions of whiteness
4672:Australian Broadcasting Corporation
3967:Bulletin of the Psychonomic Society
3405:Esteve, J. V.; Jeffery, G. (1998).
2442:10.1002/9780470015902.a0006081.pub3
2000:causes a dysfunctional tyrosinase.
1920:activity. This is in contrast with
1019:living off the coast of Australia;
5203:. December 8, 2010. Archived from
5133:Stienstra, T. (October 11, 2007).
4964:: History, Biology and Aquaculture
2742:Critical Reviews in Plant Sciences
1247:, those that lack melanophores as
14:
4722:Lissotriton vulgaris meridionalis
4490:"Rare Albino Whale Shark Spotted"
4180:Leal, M. E.; et al. (2013).
4096:"Albinism in the Great Gray Owl (
3192:Nakamura, K.; Ozaki (June 2001).
2703:10.1034/j.1600-0749.2001.140203.x
1889:have focused on mutations in the
1875:Lissotriton vulgaris meridionalis
354:animals and are generated in the
4720:"A case of complete albinism in
4608:– via www.telegraph.co.uk.
4565:. California Academy of Sciences
3756:10.1111/j.1469-7998.2010.00762.x
3738:Lechner, W.; Ladich, F. (1987).
3516:10.1046/j.1439-0426.2000.00147.x
2348:
2181:Audubon Aquarium of the Americas
2070:
2049:
2029:
1215:
1203:
1108:
1093:
761:
749:
290:Many animal body-parts, such as
191:
104:
87:
64:
39:
5054:Hesford, Clive (January 1998).
4769:Journal of Experimental Zoology
4263:Japanese Journal of Ichthyology
4240:10.1590/S1679-62252005000100009
4203:10.1590/S1516-89132013000200008
4001:Encyclopaedia of Marine Mammals
3530:Gross, Joshua (June 30, 2016).
3198:Molecular Genetics and Genomics
1921:
1885:Genetic studies of albinism in
1146:. Famous albino birds include "
668:Melanin protects the skin from
450:, which uses the green pigment
202:needs additional citations for
4031:. July 3, 2007. Archived from
3503:Journal of Applied Ichthyology
3454:Fausto-Sterling, Anne (2008).
3159:The Significance of Zoochromes
2915:Cambridge Dictionaries On-line
1311:Albino and normally pigmented
1011:Famous albino mammals include
602:albino gorilla and one albino
56:California Academy of Sciences
1:
5110:"Feather Colors: What We See"
4995:(Scorpiones, Scorpionidae)".
4914:10.1016/s0022-1910(02)00106-3
4878:Taylor, John William (1900).
4861:Taylor, John William (1900).
4150:10.1016/S0042-6989(01)00284-X
3424:10.1016/s0042-6989(97)00229-0
3305:Nasr, S. R. (July 15, 2008).
3002:British Trust for Ornithology
2684:Brilliant, Murray H. (2001).
2568:McCardle, Hayley (May 2012).
2179:A leucistic alligator at the
1871:Lissotriton vulgaris vulgaris
1865:) and two sub-species of the
1703:California Academy of Science
1690:"Albino" California kingsnake
124:is the congenital absence of
4902:Journal of Insect Physiology
4520:Journal of Wildlife Diseases
4309:10.1016/0044-8486(84)90295-3
4269:(2): 120–122. Archived from
3831:10.1016/0091-3057(80)90461-X
3601:Pierce, Benjamin A. (2008).
2871:10.1016/0016-6480(69)90157-9
2826:10.1098/rsif.2008.0366.focus
2571:Albinism in wild vertebrates
2387:, unusually red pigmentation
4826:Genes & Genetic Systems
4455:and the giant electric ray
4219:"An albino armored catfish
3690:Purdom, C. F., ed. (1993).
3632:Shepherd, Rosamond (1996).
2645:10.1534/genetics.106.063107
2124:The Japanese sea cucumber (
1940:, resulting in a truncated
919:Albinism can also occur in
5506:
5417:Color terminology for race
5228:Canadian Journal of Botany
4941:February 26, 2015, at the
3740:"How do albino fish hear?"
3694:Genetics and Fish Breeding
2375:Albino and white squirrels
1907:post-translational control
1476:Scalloped hammerhead shark
849:five-striped palm squirrel
267:
25:
18:
5459:
5407:
5367:
4737:: 395–396. Archived from
4533:10.7589/0090-3558-21.1.72
4488:Haith, Sara-Lise (2008).
3607:. Macmillan. p. A1.
3549:10.1186/s12862-016-0716-y
3371:10.1017/s0952523806231031
3162:. Springer. p. 325.
2941:Dictionary.com Unabridged
2754:10.1080/07352680903133252
2602:"Oculocutaneous Albinism"
2317:University of North Texas
2165:Related pigment disorders
1972:), a substitution from a
1930:Pelophylax nigromaculatus
1526:Whitespotted bamboo shark
1376:Broadnose sevengill shark
889:Albinism was observed in
678:electromagnetic radiation
3908:(12): 22. Archived from
3536:BMC Evolutionary Biology
2321:University of Louisville
2198:is affected rather than
2183:, New Orleans, Louisiana
2077:Albino freshwater snail
1460:Carcharhinus amboinensis
1430:Carcharhinus amboinensis
1123:Well-known examples are
1027:living in and around in
5140:San Francisco Chronicle
5121:Birder's World Magazine
5115:March 28, 2009, at the
4580:San Francisco Chronicle
4375:Northwestern Naturalist
4227:Neotropical Ichthyology
4094:Alaja, P.; Mikkola, H.
3156:Needham, A. E. (2012).
3075:Encyclopædia Britannica
3043:: 79–89. Archived from
2994:"Leucisam and albinism"
1827:gold-striped salamander
1611:Giant oceanic manta ray
1530:Chiloscyllium plagiosum
1446:Narrownose smooth-hound
1056:Jamestown, North Dakota
833:lesser mouse-tailed bat
813:oculocutaneous albinism
672:radiation in sunlight.
550:Mechanism and frequency
529:oculocutaneous albinism
152:Oculocutaneous albinism
113:oculocutaneous albinism
96:oculocutaneous albinism
21:Albino (disambiguation)
5022:Wilkes, J. T. (1963).
4997:Journal of Arachnology
4962:Apostichopus japonicus
4781:10.1002/jez.1401800203
2576:Texas State University
2291:Some herbicides (e.g.
2249:
2241:
2184:
2127:Apostichopus japonicus
1863:Ichthyosaura alpestris
1802:
1691:
1555:Myliobatis californica
1390:Carcharodon carcharias
1380:Notorynchus cepedianus
1237:
1222:Albino peacock cichlid
1058:, and inspiration for
967:
908:A study on albinistic
845:northern palm squirrel
815:
652:
559:
513:neural crest disorders
491:
183:Biological colouration
5258:Utah State University
5171:San Jose Mercury News
5058:Nymphicus hollandicus
5041:10.1093/aesa/56.6.792
4744:on September 24, 2015
4596:. February 16, 2009.
3790:Halls, K. M. (2004).
3332:"Albinism in animals"
3210:10.1007/s004380100464
3103:"Color on, color off"
3028:de Grouw, H. (2006).
2690:Pigment Cell Research
2247:
2235:
2178:
2080:Biomphalaria glabrata
2062:Pseudofusulus varians
1831:Chioglossa lusitanica
1823:Salamandra salamandra
1794:
1737:in the U.S. state of
1689:
1420:Orectolobus japonicus
1400:Mustelus californicus
1232:
1190:Artificially selected
1007:Famous albino mammals
962:
807:
736:Artificially selected
658:eastern gray squirrel
650:
557:
481:
5207:on September 6, 2019
4970:. pp. 211–226.
4936:Fruit Fly Phenotypes
4934:Service, Elizabeth.
4839:10.1266/ggs.16-00061
4425:"Albinism in sharks"
4221:Schizolecis guntheri
4035:on February 24, 2010
3961:Microtus ochrogaster
3895:(Geoffroy) from Goa"
3483:Ilo Hiler, Albinos.
3307:"How albinism works"
2897:MacMillan Dictionary
2820:(Suppl 2): 149–163.
2514:The Georgia Aquarium
2370:Albinism in chickens
2299:) can cause partial
1988:(Gly) changes to an
1970:Fejervarya kawamurai
1835:Italian crested newt
1808:plains leopard frogs
1540:Stegostoma fasciatum
1440:Triakis semifasciata
1410:Hemitriakis japanica
1050:for White Cloud), a
914:Microtus ochrogaster
821:Sciurus carolinensis
521:Waardenburg syndrome
490:) with her offspring
211:improve this article
5289:on January 20, 2015
5075:Miller, Jessica J.
4993:Urodacus yaschenkoi
4276:on February 2, 2015
4254:Nakaya, K. (1973).
3915:on February 8, 2015
3902:Zoos' Print Journal
3658:Journal of Heredity
3359:Visual Neuroscience
3336:Science Illustrated
3283:on January 28, 2015
3004:. December 13, 2011
2785:on February 2, 2015
2520:on January 20, 2015
2510:GeorgiaAquarium.org
2481:The Free Dictionary
2328:African clawed frog
2313:University of Texas
1938:frameshift mutation
1851:Iberian ribbed newt
1847:Triturus marmoratus
1764:In 2012, an albino
1621:Ocellate spot skate
1358:has been reported.
1317:Ictalurus punctatus
1277:Naturally occurring
1140:common house martin
800:Naturally occurring
407:coherent scattering
246:Biological pigments
5490:Animal coat colors
5254:"Herbicide Injury"
4898:Locusta migratoria
4514:Protopterus dolloi
3980:10.3758/bf03329782
3744:Journal of Zoology
2549:. January 27, 2018
2506:Albino Alligators"
2365:Albinism in humans
2250:
2242:
2185:
2103:Locusta migratoria
2099:Locusta migratoria
1803:
1770:shingleback lizard
1707:Steinhart Aquarium
1692:
1635:Dasyatis americana
1601:Giant electric ray
1595:Rhinoptera bonasus
1575:Dasyatis pastinaca
1500:Nebrius ferigineus
1416:Japanese wobbegong
1406:Japanese topeshark
1370:Cetorhinus maximus
1352:Hydrolagus colliei
1328:Mexican cave tetra
1270:Astyanax mexicanus
1238:
1025:bottlenose dolphin
968:
816:
789:Sprague Dawley rat
653:
627:the colour of the
560:
492:
397:Structural colours
282:with white wings.
276:structural colours
264:Animal colouration
146:Mexican cave tetra
142:laboratory animals
52:American alligator
28:Albinism in humans
5472:
5471:
5412:Fitzpatrick scale
5234:(10): 1395–1404.
4977:978-0-12-799953-1
4960:The Sea Cucumber
4731:Herpetology Notes
4693:Sage, P. (2006).
4626:. August 20, 2014
4457:Narcine entemedor
4453:Galeocerdo cuvier
4112:on August 2, 2018
4100:) and Other Owls"
4010:978-0-12-373553-9
3803:978-1-58196-019-8
3721:albinism in fish.
3709:978-0-412-33040-7
3614:978-1-4292-1118-5
3467:978-0-7867-2390-4
3311:HowStuffWorks.com
3252:978-3-540-73837-4
3169:978-3-642-80766-4
3139:978-1-4511-4877-0
1998:polypeptide chain
1982:missense mutation
1962:Glandirana rugosa
1893:gene. The albino
1855:Pleurodeles waltl
1839:Triturus carnifex
1758:Sulcata tortoises
1631:Southern stingray
1605:Narcine entemedor
1510:Galeocerdo cuvier
1496:Tawny nurse shark
1490:Squalus acanthias
1450:Mustelus schmitti
1396:Grey smooth-hound
1386:Great white shark
1354:) from the order
1338:In Chondrichthyes
1307:In Actinopterygii
1297:Pleuronectiformes
1295:, especially the
841:common palm civet
723:Hearing disorders
710:Reduced viability
442:Plant colouration
403:structural colour
270:Animal coloration
250:colour absorption
243:
242:
235:
94:Mice with Type I
5497:
5398:Dark/Black/Ebony
5378:Light/White/Fair
5361:Human skin color
5354:
5347:
5340:
5331:
5324:
5323:
5321:
5319:
5305:
5299:
5298:
5296:
5294:
5285:. Archived from
5275:
5269:
5268:
5266:
5264:
5250:
5244:
5243:
5223:
5217:
5216:
5214:
5212:
5199:DiscoverMagazine
5189:
5183:
5182:
5180:
5178:
5161:
5152:
5151:
5149:
5147:
5130:
5124:
5107:
5101:
5100:
5098:
5096:
5087:. Archived from
5072:
5066:
5052:
5046:
5045:
5043:
5019:
5013:
5012:
4988:
4982:
4981:
4955:
4946:
4932:
4926:
4925:
4892:
4886:
4885:
4875:
4869:
4868:
4858:
4852:
4851:
4841:
4817:
4811:
4810:
4792:
4760:
4754:
4753:
4751:
4749:
4743:
4728:
4715:
4706:
4705:
4703:
4701:
4690:
4684:
4683:
4681:
4679:
4664:
4658:
4657:
4655:
4653:
4642:
4636:
4635:
4633:
4631:
4616:
4610:
4609:
4607:
4605:
4595:
4588:
4582:
4572:
4566:
4560:
4554:
4553:
4535:
4507:
4501:
4500:
4498:
4496:
4485:
4479:
4478:
4476:
4474:
4446:
4437:
4436:
4434:
4432:
4420:
4414:
4413:
4411:
4409:
4404:on March 4, 2016
4403:
4397:. Archived from
4372:
4363:
4357:
4356:
4319:
4313:
4312:
4292:
4286:
4285:
4283:
4281:
4275:
4260:
4251:
4245:
4244:
4242:
4214:
4208:
4207:
4205:
4177:
4171:
4170:
4152:
4128:
4122:
4121:
4119:
4117:
4111:
4105:. Archived from
4104:
4091:
4085:
4079:
4073:
4072:
4070:
4068:
4059:
4051:
4045:
4044:
4042:
4040:
4029:Fox News Channel
4021:
4015:
4014:
3996:
3985:
3984:
3982:
3954:
3948:
3947:
3931:
3925:
3924:
3922:
3920:
3914:
3899:
3886:
3877:
3876:
3874:
3872:
3865:Young Naturalist
3857:
3851:
3850:
3814:
3808:
3807:
3787:
3778:
3777:
3767:
3735:
3724:
3723:
3718:
3716:
3697:
3687:
3674:
3673:
3653:
3647:
3646:
3640:
3629:
3623:
3622:
3598:
3592:
3591:
3569:
3551:
3527:
3521:
3520:
3518:
3494:
3488:
3485:Young Naturalist
3481:
3475:
3474:
3451:
3445:
3444:
3426:
3402:
3391:
3390:
3354:
3348:
3347:
3345:
3343:
3328:
3322:
3321:
3319:
3317:
3302:
3293:
3292:
3290:
3288:
3277:"Albino animals"
3273:
3260:
3259:
3236:
3230:
3229:
3189:
3183:
3182:
3153:
3147:
3146:
3123:
3117:
3116:
3114:
3112:
3107:
3098:
3087:
3086:
3084:
3082:
3066:
3060:
3059:
3057:
3055:
3050:on July 13, 2017
3049:
3034:
3025:
3014:
3013:
3011:
3009:
2990:
2981:
2980:
2978:
2976:
2960:
2954:
2953:
2951:
2949:
2932:
2926:
2925:
2923:
2921:
2907:
2901:
2900:
2889:
2883:
2882:
2854:
2848:
2847:
2837:
2804:Mäthger, L. M.;
2801:
2795:
2794:
2792:
2790:
2781:. Archived from
2775:
2766:
2765:
2733:
2724:
2723:
2705:
2681:
2675:
2674:
2664:
2623:
2617:
2616:
2614:
2612:
2598:
2587:
2586:
2584:
2582:
2565:
2559:
2558:
2556:
2554:
2539:
2530:
2529:
2527:
2525:
2516:. Archived from
2498:
2492:
2491:
2489:
2487:
2473:
2464:
2463:
2429:
2358:
2353:
2352:
2307:In human culture
2074:
2053:
2042:Sanibel, Florida
2033:
2011:In invertebrates
1952:that code for a
1676:African lungfish
1674:Albinism in the
1670:In Sarcopterygii
1625:Okamejei kenojei
1264:Corydoras aeneus
1219:
1207:
1112:
1097:
765:
753:
405:, the result of
238:
231:
227:
224:
218:
195:
187:
111:Rat with Type I
108:
91:
68:
43:
5505:
5504:
5500:
5499:
5498:
5496:
5495:
5494:
5475:
5474:
5473:
5468:
5455:
5403:
5363:
5358:
5328:
5327:
5317:
5315:
5307:
5306:
5302:
5292:
5290:
5277:
5276:
5272:
5262:
5260:
5252:
5251:
5247:
5240:10.1139/b63-122
5225:
5224:
5220:
5210:
5208:
5191:
5190:
5186:
5176:
5174:
5163:
5162:
5155:
5145:
5143:
5132:
5131:
5127:
5117:Wayback Machine
5108:
5104:
5094:
5092:
5091:on May 27, 2007
5074:
5073:
5069:
5053:
5049:
5021:
5020:
5016:
4990:
4989:
4985:
4978:
4957:
4956:
4949:
4943:Wayback Machine
4933:
4929:
4894:
4893:
4889:
4877:
4876:
4872:
4860:
4859:
4855:
4819:
4818:
4814:
4762:
4761:
4757:
4747:
4745:
4741:
4726:
4717:
4716:
4709:
4699:
4697:
4692:
4691:
4687:
4677:
4675:
4666:
4665:
4661:
4651:
4649:
4644:
4643:
4639:
4629:
4627:
4618:
4617:
4613:
4603:
4601:
4590:
4589:
4585:
4573:
4569:
4561:
4557:
4509:
4508:
4504:
4494:
4492:
4487:
4486:
4482:
4472:
4470:
4448:
4447:
4440:
4430:
4428:
4422:
4421:
4417:
4407:
4405:
4401:
4370:
4365:
4364:
4360:
4325:Nature Genetics
4321:
4320:
4316:
4294:
4293:
4289:
4279:
4277:
4273:
4258:
4253:
4252:
4248:
4216:
4215:
4211:
4179:
4178:
4174:
4137:Vision Research
4130:
4129:
4125:
4115:
4113:
4109:
4102:
4093:
4092:
4088:
4084:. The Telegraph
4080:
4076:
4066:
4064:
4057:
4053:
4052:
4048:
4038:
4036:
4023:
4022:
4018:
4011:
3998:
3997:
3988:
3956:
3955:
3951:
3933:
3932:
3928:
3918:
3916:
3912:
3897:
3888:
3887:
3880:
3870:
3868:
3859:
3858:
3854:
3816:
3815:
3811:
3804:
3789:
3788:
3781:
3737:
3736:
3727:
3714:
3712:
3710:
3689:
3688:
3677:
3655:
3654:
3650:
3638:
3631:
3630:
3626:
3615:
3600:
3599:
3595:
3529:
3528:
3524:
3496:
3495:
3491:
3482:
3478:
3468:
3453:
3452:
3448:
3411:Vision Research
3404:
3403:
3394:
3356:
3355:
3351:
3341:
3339:
3330:
3329:
3325:
3315:
3313:
3304:
3303:
3296:
3286:
3284:
3275:
3274:
3263:
3253:
3238:
3237:
3233:
3191:
3190:
3186:
3170:
3155:
3154:
3150:
3140:
3125:
3124:
3120:
3110:
3108:
3105:
3101:Binkley, S. K.
3100:
3099:
3090:
3080:
3078:
3068:
3067:
3063:
3053:
3051:
3047:
3032:
3027:
3026:
3017:
3007:
3005:
2992:
2991:
2984:
2974:
2972:
2969:Merriam-Webster
2962:
2961:
2957:
2947:
2945:
2934:
2933:
2929:
2919:
2917:
2909:
2908:
2904:
2891:
2890:
2886:
2856:
2855:
2851:
2803:
2802:
2798:
2788:
2786:
2777:
2776:
2769:
2735:
2734:
2727:
2683:
2682:
2678:
2625:
2624:
2620:
2610:
2608:
2600:
2599:
2590:
2580:
2578:
2567:
2566:
2562:
2552:
2550:
2541:
2540:
2533:
2523:
2521:
2500:
2499:
2495:
2485:
2483:
2475:
2474:
2467:
2452:
2431:
2430:
2426:
2421:
2416:
2354:
2347:
2344:
2309:
2282:Black Tartarian
2271:Albino redwoods
2230:
2167:
2152:
2122:
2095:
2088:
2075:
2066:
2054:
2045:
2034:
2013:
1883:
1819:fire salamander
1785:
1684:
1672:
1664:
1615:Manta birostris
1585:Torpedo torpedo
1571:Common stingray
1520:Rhincodon typus
1348:spotted ratfish
1340:
1313:channel catfish
1309:
1279:
1227:
1226:
1225:
1224:
1223:
1220:
1212:
1211:
1208:
1199:
1198:
1192:
1184:
1120:
1119:
1118:
1117:
1116:
1113:
1105:
1104:
1098:
1089:
1088:
1082:
1060:Herman Melville
1009:
957:
802:
777:model organisms
773:
772:
771:
770:
769:
766:
758:
757:
754:
745:
744:
738:
733:
725:
712:
687:
666:
629:red blood cells
624:
616:
552:
476:
444:
415:mother of pearl
399:
288:
272:
266:
258:flower pigments
239:
228:
222:
219:
208:
196:
185:
119:
118:
117:
116:
115:
109:
100:
99:
98:
92:
83:
82:
76:
75:
74:
73:
72:
69:
60:
59:
58:
44:
31:
24:
17:
12:
11:
5:
5503:
5501:
5493:
5492:
5487:
5477:
5476:
5470:
5469:
5467:
5466:
5460:
5457:
5456:
5454:
5453:
5448:
5443:
5438:
5429:
5424:
5419:
5414:
5408:
5405:
5404:
5402:
5401:
5395:
5381:
5375:
5368:
5365:
5364:
5359:
5357:
5356:
5349:
5342:
5334:
5326:
5325:
5300:
5270:
5245:
5218:
5184:
5153:
5125:
5102:
5067:
5047:
5034:(6): 792–795.
5014:
5003:(1): 101–115.
4983:
4976:
4968:Academic Press
4947:
4927:
4908:(8): 791–801.
4887:
4870:
4853:
4832:(4): 189–196.
4812:
4775:(2): 157–167.
4755:
4707:
4685:
4659:
4646:"Albino anole"
4637:
4611:
4583:
4567:
4555:
4502:
4480:
4438:
4423:Martin, R. A.
4415:
4358:
4337:10.1038/ng1700
4331:(1): 107–111.
4314:
4303:(4): 293–301.
4287:
4246:
4233:(1): 123–125.
4209:
4196:(2): 237–240.
4172:
4143:(3): 293–299.
4123:
4098:Strix nebulosa
4086:
4074:
4046:
4016:
4009:
3986:
3949:
3926:
3893:Macaca radiata
3878:
3852:
3825:(6): 969–967.
3809:
3802:
3793:Albino Animals
3779:
3750:(3): 186–192.
3725:
3708:
3675:
3664:(6): 284–293.
3648:
3624:
3613:
3593:
3522:
3509:(3): 117–120.
3489:
3476:
3466:
3446:
3417:(6): 937–940.
3392:
3349:
3323:
3294:
3261:
3251:
3231:
3204:(4): 687–693.
3184:
3168:
3148:
3138:
3118:
3088:
3061:
3015:
2982:
2955:
2927:
2902:
2884:
2865:(3): 405–416.
2849:
2796:
2767:
2748:(6): 393–409.
2725:
2676:
2639:(2): 867–877.
2618:
2588:
2574:(MSc thesis).
2560:
2531:
2493:
2465:
2450:
2423:
2422:
2420:
2417:
2415:
2414:
2409:
2406:Xanthochromism
2403:
2401:Mongolian spot
2398:
2393:
2388:
2382:
2377:
2372:
2367:
2361:
2360:
2359:
2356:Animals portal
2343:
2340:
2308:
2305:
2258:photosynthesis
2229:
2226:
2225:
2224:
2213:
2203:
2186:
2166:
2163:
2151:
2148:
2121:
2120:In echinoderms
2118:
2094:
2091:
2090:
2089:
2076:
2069:
2067:
2055:
2048:
2046:
2035:
2028:
2012:
2009:
1980:(A) creates a
1956:(Lys). In the
1882:
1879:
1784:
1781:
1683:
1680:
1671:
1668:
1663:
1662:In Hyperoartia
1660:
1659:
1658:
1648:
1638:
1628:
1618:
1608:
1598:
1588:
1581:Common torpedo
1578:
1568:
1558:
1544:
1543:
1533:
1523:
1513:
1503:
1493:
1483:
1480:Sphyrna lewini
1473:
1463:
1453:
1443:
1433:
1423:
1413:
1403:
1393:
1383:
1373:
1356:Chimaeriformes
1344:Chondrichthyes
1339:
1336:
1308:
1305:
1278:
1275:
1258:Silurus glanis
1249:albino mutants
1221:
1214:
1213:
1210:Albino catfish
1209:
1202:
1201:
1200:
1196:
1195:
1194:
1193:
1191:
1188:
1183:
1180:
1175:
1174:
1171:
1168:
1165:
1144:American robin
1114:
1107:
1106:
1099:
1092:
1091:
1090:
1086:
1085:
1084:
1083:
1081:
1078:
1029:Calcasieu Lake
1017:humpback whale
1008:
1005:
988:Enhydra lutris
956:
955:Marine mammals
953:
880:bonnet macaque
872:rhesus macaque
801:
798:
767:
760:
759:
755:
748:
747:
746:
742:
741:
740:
739:
737:
734:
732:
729:
724:
721:
711:
708:
700:founder effect
686:
685:Survival value
683:
676:block harmful
665:
662:
638:photoreceptors
623:
620:
615:
612:
551:
548:
475:
472:
448:photosynthesis
443:
440:
439:
438:
428:
398:
395:
394:
393:
387:
378:
369:
352:poikilothermic
312:chromatophores
287:
284:
268:Main article:
265:
262:
254:plant pigments
241:
240:
199:
197:
190:
184:
181:
177:photosynthesis
110:
103:
102:
101:
93:
86:
85:
84:
80:
79:
78:
77:
70:
63:
62:
61:
45:
38:
37:
36:
35:
34:
15:
13:
10:
9:
6:
4:
3:
2:
5502:
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5488:
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5480:
5465:
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5409:
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5288:
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5280:
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4800:
4796:
4791:
4790:2027.42/38071
4786:
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4778:
4774:
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4759:
4756:
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4725:
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4673:
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4604:September 19,
4599:
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3333:
3327:
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3312:
3308:
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3258:
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3215:
3211:
3207:
3203:
3199:
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3178:
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3160:
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3145:
3141:
3135:
3131:
3130:
3122:
3119:
3104:
3097:
3095:
3093:
3089:
3077:
3076:
3071:
3065:
3062:
3046:
3042:
3038:
3037:Dutch Birding
3031:
3024:
3022:
3020:
3016:
3003:
2999:
2995:
2989:
2987:
2983:
2971:
2970:
2965:
2959:
2956:
2944:(Online). n.d
2943:
2942:
2937:
2931:
2928:
2916:
2912:
2906:
2903:
2898:
2894:
2888:
2885:
2880:
2876:
2872:
2868:
2864:
2860:
2853:
2850:
2845:
2841:
2836:
2831:
2827:
2823:
2819:
2815:
2811:
2807:
2806:Denton, E. J.
2800:
2797:
2784:
2780:
2774:
2772:
2768:
2763:
2759:
2755:
2751:
2747:
2743:
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2726:
2721:
2717:
2713:
2709:
2704:
2699:
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2677:
2672:
2668:
2663:
2658:
2654:
2650:
2646:
2642:
2638:
2634:
2630:
2622:
2619:
2607:
2603:
2597:
2595:
2593:
2589:
2577:
2573:
2572:
2564:
2561:
2548:
2544:
2538:
2536:
2532:
2519:
2515:
2511:
2507:
2505:
2497:
2494:
2482:
2478:
2472:
2470:
2466:
2461:
2457:
2453:
2451:9780470016176
2447:
2443:
2439:
2435:
2428:
2425:
2418:
2413:
2410:
2407:
2404:
2402:
2399:
2397:
2394:
2392:
2389:
2386:
2383:
2381:
2378:
2376:
2373:
2371:
2368:
2366:
2363:
2362:
2357:
2351:
2346:
2341:
2339:
2337:
2333:
2329:
2324:
2322:
2318:
2314:
2306:
2304:
2302:
2298:
2294:
2289:
2287:
2283:
2278:
2276:
2272:
2268:
2266:
2261:
2259:
2255:
2246:
2239:
2234:
2227:
2222:
2217:
2214:
2211:
2210:chromatophore
2207:
2204:
2201:
2197:
2194:
2190:
2187:
2182:
2177:
2172:
2171:
2170:
2164:
2162:
2160:
2158:
2149:
2147:
2144:
2140:
2138:
2133:
2129:
2128:
2119:
2117:
2115:
2111:
2106:
2104:
2100:
2092:
2086:
2082:
2081:
2073:
2068:
2064:
2063:
2059:
2052:
2047:
2043:
2039:
2032:
2027:
2025:
2023:
2018:
2010:
2008:
2006:
2001:
1999:
1995:
1992:(Asp) and an
1991:
1990:aspartic acid
1987:
1984:, in which a
1983:
1979:
1975:
1971:
1967:
1964:) and in the
1963:
1959:
1958:wrinkled frog
1955:
1951:
1947:
1943:
1939:
1935:
1931:
1927:
1923:
1919:
1916:
1912:
1908:
1904:
1900:
1896:
1892:
1888:
1880:
1878:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1848:
1844:
1840:
1836:
1832:
1828:
1824:
1820:
1816:
1815:Salamandridae
1811:
1809:
1801:
1797:
1793:
1789:
1783:In amphibians
1782:
1780:
1778:
1777:leopard gecko
1773:
1771:
1767:
1762:
1759:
1754:
1752:
1748:
1742:
1740:
1736:
1733:
1728:
1724:
1720:
1716:
1712:
1708:
1704:
1699:
1697:
1688:
1681:
1679:
1677:
1669:
1667:
1661:
1656:
1652:
1649:
1646:
1642:
1641:Thornback ray
1639:
1636:
1632:
1629:
1626:
1622:
1619:
1616:
1612:
1609:
1606:
1602:
1599:
1596:
1592:
1589:
1586:
1582:
1579:
1576:
1572:
1569:
1566:
1562:
1559:
1556:
1552:
1549:
1548:
1547:
1541:
1537:
1534:
1531:
1527:
1524:
1521:
1517:
1514:
1511:
1507:
1504:
1501:
1497:
1494:
1491:
1487:
1486:Spiny dogfish
1484:
1481:
1477:
1474:
1471:
1467:
1464:
1461:
1457:
1454:
1451:
1447:
1444:
1441:
1437:
1436:Leopard shark
1434:
1431:
1427:
1424:
1421:
1417:
1414:
1411:
1407:
1404:
1402:) – two cases
1401:
1397:
1394:
1391:
1387:
1384:
1381:
1377:
1374:
1372:) – two cases
1371:
1367:
1366:Basking shark
1364:
1363:
1362:
1359:
1357:
1353:
1349:
1345:
1342:In the class
1337:
1335:
1333:
1329:
1325:
1320:
1318:
1314:
1306:
1304:
1302:
1301:elasmobranchs
1298:
1294:
1289:
1287:
1282:
1276:
1274:
1272:
1271:
1266:
1265:
1260:
1259:
1254:
1250:
1246:
1242:
1236:
1235:giant gourami
1231:
1218:
1206:
1189:
1187:
1181:
1179:
1172:
1169:
1166:
1163:
1162:
1161:
1158:
1156:
1153:
1149:
1145:
1141:
1137:
1132:
1130:
1126:
1111:
1103:
1096:
1079:
1077:
1075:
1071:
1067:
1066:
1061:
1057:
1053:
1049:
1045:
1041:
1038:
1037:Barcelona Zoo
1034:
1030:
1026:
1022:
1018:
1014:
1006:
1004:
1002:
1001:
1000:Lacazia loboi
995:
993:
989:
985:
981:
976:
972:
966:
961:
954:
952:
950:
946:
942:
938:
934:
930:
926:
922:
917:
915:
911:
910:prairie voles
906:
904:
900:
896:
892:
887:
885:
881:
877:
873:
869:
865:
864:toque macaque
861:
856:
854:
850:
846:
842:
838:
834:
830:
825:
823:
822:
814:
810:
806:
799:
797:
793:
790:
786:
782:
778:
768:Albino rabbit
764:
756:Albino ferret
752:
735:
730:
728:
722:
720:
717:
709:
707:
703:
701:
697:
696:albino people
692:
684:
682:
679:
675:
671:
663:
661:
659:
649:
645:
643:
639:
634:
630:
621:
619:
613:
611:
607:
605:
599:
597:
596:rainbow trout
593:
588:
583:
581:
577:
573:
569:
565:
556:
549:
547:
543:
541:
536:
532:
530:
526:
522:
518:
514:
508:
506:
500:
496:
489:
485:
480:
473:
471:
469:
465:
461:
457:
453:
449:
441:
436:
432:
429:
427:
423:
420:
419:
418:
416:
412:
409:perceived as
408:
404:
396:
391:
388:
386:
382:
379:
377:
373:
372:erythrophores
370:
368:
364:
361:
360:
359:
357:
353:
349:
345:
341:
337:
333:
329:
325:
321:
317:
313:
309:
305:
301:
297:
293:
285:
283:
281:
277:
271:
263:
261:
259:
255:
251:
247:
237:
234:
226:
216:
212:
206:
205:
200:This section
198:
194:
189:
188:
182:
180:
178:
174:
169:
168:
165:
161:
157:
153:
149:
147:
143:
139:
133:
131:
127:
123:
114:
107:
97:
90:
67:
57:
53:
50:", an albino
49:
42:
33:
29:
22:
5318:November 18,
5316:. Retrieved
5313:AnimalsGuide
5312:
5303:
5291:. Retrieved
5287:the original
5282:
5273:
5263:November 24,
5261:. Retrieved
5257:
5248:
5231:
5227:
5221:
5211:November 23,
5209:. Retrieved
5205:the original
5196:
5187:
5177:November 23,
5175:. Retrieved
5169:
5144:. Retrieved
5138:
5128:
5120:
5105:
5093:. Retrieved
5089:the original
5084:
5078:
5070:
5063:
5057:
5050:
5031:
5027:
5017:
5000:
4996:
4992:
4986:
4963:
4959:
4930:
4905:
4901:
4897:
4890:
4880:
4873:
4863:
4856:
4829:
4825:
4815:
4772:
4768:
4758:
4746:. Retrieved
4739:the original
4734:
4730:
4721:
4698:. Retrieved
4688:
4676:. Retrieved
4662:
4650:. Retrieved
4640:
4628:. Retrieved
4624:The Guardian
4623:
4614:
4602:. Retrieved
4586:
4579:
4570:
4558:
4526:(1): 72–74.
4523:
4519:
4513:
4505:
4495:November 19,
4493:. Retrieved
4483:
4471:. Retrieved
4469:(2): 191–192
4466:
4462:
4456:
4452:
4429:. Retrieved
4418:
4406:. Retrieved
4399:the original
4378:
4374:
4361:
4328:
4324:
4317:
4300:
4296:
4290:
4278:. Retrieved
4271:the original
4266:
4262:
4249:
4230:
4226:
4220:
4212:
4193:
4189:
4183:
4175:
4140:
4136:
4126:
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2286:sweet cherry
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2150:In arachnids
2141:
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2112:mutation in
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2060:
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2014:
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1961:
1929:
1909:in a single
1903:Rana pipiens
1902:
1899:leopard frog
1884:
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1854:
1846:
1843:marbled newt
1838:
1830:
1822:
1813:In European
1812:
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1798:with albino
1786:
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1727:pigmentation
1700:
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1645:Raja clavata
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1561:Common skate
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464:anthocyanins
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400:
381:melanophores
363:xanthophores
356:neural crest
289:
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229:
220:
209:Please help
204:verification
201:
170:
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129:
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5394:(III–IV–V.)
5384:Beige/Olive
5380:(I–II–III.)
5293:January 20,
5146:December 6,
5095:January 18,
4748:January 27,
4473:February 2,
4431:February 2,
4408:February 2,
4297:Aquaculture
4280:January 26,
4116:January 24,
4067:January 30,
3919:January 29,
3715:February 2,
3342:January 24,
3316:January 23,
3287:January 24,
3111:January 22,
3081:January 27,
3054:January 24,
3008:January 31,
2975:January 31,
2920:January 31,
2789:January 28,
2779:"Albiflora"
2581:January 24,
2524:January 20,
2486:January 31,
2332:guinea pigs
2254:chlorophyll
2193:xanthophore
2143:Astaxanthin
2114:fruit flies
2085:haemoglobin
1950:nucleotides
1867:smooth newt
1859:alpine newt
1747:Corn snakes
1696:amelanistic
1682:In reptiles
1655:Raja naevus
1591:Cownose ray
1536:Zebra shark
1516:Whale shark
1506:Tiger shark
1470:Lamna nasus
1245:roy mutants
1152:Bristol Zoo
1087:Kookaburras
1070:sperm whale
1044:Mahpiya Ska
895:Felis chaus
891:jungle cats
809:"Snowflake"
719:mutation".
714:Studies on
674:Melanosomes
576:melanocytes
568:pheomelanin
540:albino gaur
460:carotenoids
452:chlorophyll
431:leucophores
422:iridophores
411:iridescence
390:cyanophores
367:carotenoids
344:melanocytes
332:cephalopods
328:crustaceans
280:butterflies
173:chlorophyll
160:pheomelanin
5479:Categories
4945:. unc.edu.
4630:August 11,
3542:(1): 145.
3070:"Albinism"
2964:"Albinism"
2477:"Albinism"
2419:References
2396:Leukoderma
2380:Dyschromia
2293:glyphosate
2236:An albino
2196:metabolism
2157:Panonychus
2132:echinoderm
2093:In insects
2058:land snail
1976:(G) to an
1891:tyrosinase
1887:amphibians
1735:theme park
1719:alligators
1715:crocodiles
1651:Cuckoo ray
1565:Raja batis
1512:) – embryo
1426:Java shark
1322:Some wild
1102:kookaburra
1074:Mocha Dick
984:sea otters
963:An albino
927:, such as
925:monotremes
921:marsupials
884:M. radiata
876:M. mulatta
783:mouse and
731:In mammals
691:camouflage
592:tyrosinase
580:tyrosinase
517:piebaldism
474:Definition
456:porphyrins
316:amphibians
223:April 2023
4900:strain".
4799:0022-104X
4563:The Swamp
4381:: 60–62.
3871:March 21,
3861:"Albinos"
3584:255789720
3558:1471-2148
2948:August 7,
2712:1600-0749
2653:0016-6731
2460:239781411
2385:Erythrism
2301:chlorosis
2297:triazines
2265:etiolated
2228:In plants
2221:gene pool
2200:synthesis
2189:Axanthism
2137:epidermis
1966:rice frog
1926:pond frog
1911:recessive
1895:phenotype
1800:phenotype
1796:Wood Frog
1732:Gatorland
1723:leucistic
1705:, in the
1466:Porbeagle
1324:cave fish
1241:Zebrafish
1125:flamingos
1072:known as
1065:Moby-Dick
1062:'s novel
1033:Snowflake
992:sirenians
945:wallabies
933:kangaroos
868:M. sinica
853:wild boar
642:apoptosis
564:eumelanin
505:leucistic
468:betalains
376:pteridine
156:eumelanin
5485:Albinism
5113:Archived
4939:Archived
4922:12770057
4848:28674275
4700:March 8,
4678:March 8,
4652:March 8,
4598:Archived
4550:41540530
4395:86529521
4353:21421177
4345:16341223
4184:Genidens
4167:15140090
4159:11809482
3946:: 23–25.
3940:Cat News
3847:27084719
3774:21552308
3576:27363593
3441:20900399
3387:26337580
3379:16597348
3218:11459189
3177:dilation
2936:"albino"
2911:"Albino"
2893:"Albino"
2844:19091688
2762:85298264
2720:11310796
2671:17151254
2633:Genetics
2611:June 23,
2553:June 23,
2342:See also
2336:peacocks
2275:parasite
2216:Melanism
2130:) is an
2017:molluscs
1994:arginine
1948:, three
1881:Genetics
1293:teleosts
1148:Snowdrop
1142:and the
1080:In birds
980:pinniped
929:echidnas
860:macaques
587:palomino
525:vitiligo
515:such as
385:melanins
348:bacteria
324:reptiles
300:feathers
286:Pigments
122:Albinism
5427:Caboclo
5400:(V–VI.)
5283:Scribol
5079:Caudata
5009:3705557
4807:4623610
4542:3981753
3839:7403210
3765:3083522
3567:4929771
3433:9624442
3226:7868741
2998:BTO.org
2879:5769930
2835:2706477
2662:1800597
2436:: 1–8.
2412:Leucism
2238:Rebutia
2206:Leucism
2005:axolotl
2003:Albino
1986:glycine
1978:adenine
1974:guanine
1942:isoform
1936:(T), a
1934:thymine
1922:mammals
1918:oxidase
1897:of the
1739:Florida
1551:Bat ray
1233:Albino
1182:In fish
1155:penguin
1100:Albino
1052:buffalo
1040:gorilla
1013:Migaloo
949:wombats
941:possums
899:jackals
878:), and
858:Albino
640:due to
631:in the
484:wallaby
482:Albino
435:purines
426:guanine
336:Mammals
164:melanin
130:albinos
126:melanin
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5441:Bronze
5372:Albino
5197:Blogs.
5060:]"
5007:
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4648:. 2012
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1954:lysine
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937:koalas
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785:Wistar
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716:medaka
633:retina
572:agouti
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5451:Olive
5446:Black
5436:Pardo
5432:Brown
5422:White
5392:Pardo
5388:Brown
5005:JSTOR
4742:(PDF)
4727:(PDF)
4546:S2CID
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4391:S2CID
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4349:S2CID
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