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Basidiomycota

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compatible nuclei following meiosis migrate into each basidiospore, which is then dispersed as a pre-existing dikaryon. Often such species form only two spores per basidium, but that too varies. Following meiosis, mitotic divisions can occur in the basidium. Multiple numbers of basidiospores can result, including odd numbers via degeneration of nuclei, or pairing up of nuclei, or lack of migration of nuclei. For example, the chanterelle genus
356:) outside of these, among the Basidiomycota. As now classified, the subphyla join and also cut across various obsolete taxonomic groups (see below) previously commonly used to describe Basidiomycota. According to a 2008 estimate, Basidiomycota comprise three subphyla (including six unassigned classes) 16 classes, 52 orders, 177 families, 1,589 genera, and 31,515 species. Wijayawardene et al. 2020 produced an update that recognized 19 classes ( 914: 1146:. can have one-, two-, three- or four-spored basidia under some circumstances. Occasionally, monokaryons of some taxa can form morphologically fully formed basidiomes and anatomically correct basidia and ballistic basidiospores in the absence of dikaryon formation, diploid nuclei, and meiosis. A rare few number of taxa have extended diploid lifecycles, but can be common species. Examples exist in the mushroom genera 1042: 31: 1054:
multiple compatibility factors. Plasmogamy between compatible individuals leads to delayed karyogamy leading to establishment of a dikaryon. The dikaryon is long lasting but ultimately gives rise to either fruitbodies with basidia or directly to basidia without fruitbodies. The paired dikaryon in the basidium fuse (i.e. karyogamy takes place). The diploid basidium begins the cycle again.
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which are tapered spine-like projections on basidia, and are typically curved, like the horns of a bull. In some Basidiomycota the spores are not ballistic, and the sterigmata may be straight, reduced to stubs, or absent. The basidiospores of these non-ballistosporic basidia may either bud off, or be
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Padamsee, M.; Kumar, T. K.; Riley, R.; Binder, M.; Boyd, A.; Calvo, A. M.; Furukawa, K.; Hesse, C.; Hohmann, S.; James, T. Y.; LaButti, K.; Lapidus, A.; Lindquist, E.; Lucas, S.; Miller, K.; Shantappa, S.; Grigoriev, I. V.; Hibbett, D. S.; McLaughlin, D. J.; Spatafora, J. W.; Aime, M. C. (Mar 2012).
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and resultant monokaryons, have nuclei that are compatible with 50% (if bipolar) or 25% (if tetrapolar) of their sister basidiospores (and their resultant monokaryons) because the mating genes must differ for them to be compatible. However, there are sometimes more than two possible alleles for a
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In summary, meiosis takes place in a diploid basidium. Each one of the four haploid nuclei migrates into its own basidiospore. The basidiospores are ballistically discharged and start new haploid mycelia called monokaryons. There are no males or females, rather there are compatible thalli with
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Many variations occur: some variations are self-compatible and spontaneously form dikaryons without a separate compatible thallus being involved. These fungi are said to be homothallic, versus the normal heterothallic species with mating types. Others are secondarily homothallic, in that two
1180:, four nuclei following meiosis remain in the basidium, but continually divide mitotically, each nucleus migrating into synchronously forming nonballistic basidiospores that are then pushed upwards by another set forming below them, resulting in four parallel chains of dry "basidiospores". 1102:
are examples of pathogenic basidiomycota. Such pathogens must be able to overcome the oxidative defenses of their respective hosts in order to produce a successful infection. The ability to undergo meiosis may provide a survival benefit for these fungi by promoting successful infection. A
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There are several genera classified in the Basidiomycota that are 1) poorly known, 2) have not been subjected to DNA analysis, or 3) if analysed phylogenetically do not group with as yet named or identified families, and have not been assigned to a specific family (i.e., they are
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Basidiomycota. In plant parasitic taxa, the saprotrophic phase is normally the yeast while the infectious stage is hyphal. However, there are examples of animal and human parasites where the species are dimorphic but it is the yeast-like state that is infectious. The genus
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because meiosis progresses synchronously in about 10 million cells within the mushroom cap, and the meiotic prophase stage is prolonged. Burns et al. studied the expression of genes involved in the 15-hour meiotic process, and found that the pattern of gene expression of
1292:), and haploid yeast-like conidia/basidiospores sometimes called sporidia, bud off laterally from each cell. In various smuts, the yeast phase may proliferate, or they may fuse, or they may infect plant tissue and become hyphal. In other smuts, such as 1239:
in dry chains in inverted cup-shaped bodies embedded in host tissue. These aeciospores then infect the second host, known as the primary or asexual host (in macrocyclic rusts). On the primary host a repeating spore stage is formed, numbered "II", the
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such as rain carry the spermatia from spermagonium to spermagonium, cross inoculating the mating types. Neither thallus is male or female. Once crossed, the dikaryons are established and a second spore stage is formed, numbered "I" and called
1202:) at their greatest complexity, produce five different types of spores on two different host plants in two unrelated host families. Such rusts are heteroecious (requiring two hosts) and macrocyclic (producing all five spores types). Wheat 1594:
Hibbett, David S.; Binder, Manfred; Bischoff, Joseph F.; Blackwell, Meredith; Cannon, Paul F.; Eriksson, Ove E.; Huhndorf, Sabine; James, Timothy; Kirk, Paul M. (May 2007). "A higher-level phylogenetic classification of the Fungi".
1280:. The teliospores are initially dikaryotic but become diploid via karyogamy. Meiosis takes place at the time of germination. A promycelium is formed that consists of a short hypha (equated to a basidium). In some smuts such as 1214:, which are miniature, flask-shaped, hollow, submicroscopic bodies embedded in the host tissue (such as a leaf). This stage, numbered "0", produces single-celled spores that ooze out in a sweet liquid and that act as nonmotile 1088:. These similarities in the patterns of expression led to the conclusion that the core expression program of meiosis has been conserved in these fungi for over half a billion years of evolution since these species diverged. 1021:. In most species, the basidiospores disperse and each can start a new haploid mycelium, continuing the lifecycle. Basidia are microscopic but they are often produced on or in multicelled large fructifications called 319:
reproducers. Basidiomycota that reproduce asexually (discussed below) can typically be recognized as members of this division by gross similarity to others, by the formation of a distinctive anatomical feature (the
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which are by then dikaryotic. Dikaryotic conidia may then form. Eventually the host is infected by infectious hyphae. Teliospores form in host tissue. Many variations on these general themes occur.
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Burns, C.; Stajich, J. E.; Rechtsteiner, A.; Casselton, L.; Hanlon, S. E.; Wilke, S. K.; Savytskyy, O. P.; Gathman, A. C.; Lilly, W. W.; Lieb, J. D.; Zolan, M. E.; Pukkila, P. J. (September 2010).
1345:, which are the asexual stages. Some Basidiomycota are only known as anamorphs. Many are yeasts, collectively called basidiomycetous yeasts to differentiate them from ascomycetous yeasts in the 1716: 1248:. Urediospores are dikaryotic and can infect the same host that produced them. They repeatedly infect this host over the growing season. At the end of the season, a fourth spore type, the 1276:
is the thick-walled, often darkly pigmented, ornate, teliospore that serves to survive harsh conditions such as overwintering and also serves to help disperse the fungus as dry
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follows shortly with the production of 4 haploid nuclei that migrate into 4 external, usually apical basidiospores. Variations occur, however. Typically the basidiospores are
564:, which includes mirror yeasts), and a mixture of odd, infrequently seen, or seldom recognized fungi, often parasitic on plants. The eight classes in the Pucciniomycotina are 519:. This sub-phyla also includes the "classic" mushrooms, polypores, corals, chanterelles, crusts, puffballs and stinkhorns. The three classes in the Agaricomycotina are the 997:
nuclear divisions. Variations are frequent and multiple. In a typical Basidiomycota lifecycle the long lasting dikaryons periodically (seasonally or occasionally) produce
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Li, Liande; Wright, Sara J.; Krystofova, Svetlana; Park, Gyungsoon; Borkovich, Katherine A. (2007). "Heterotrimeric G Protein Signaling in Filamentous Fungi".
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characteristic central feature of meiosis is recombination between homologous chromosomes. This process is associated with repair of DNA damage, particularly
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after meiosis, with each of the 4 cells bearing one basidiospore each. The basidiospores disperse and start the infection process on host 1 again.
1162:. Occasionally, basidiospores are not formed and parts of the "basidia" act as the dispersal agents, e.g. the peculiar mycoparasitic jelly fungus, 5629: 4717: 2178: 5775: 1662:
Wijayawardene, Nalin; Hyde, Kevin; Al-Ani, Laith Khalil Tawfeeq; Somayeh, Dolatabadi; Stadler, Marc; Haelewaters, Danny; et al. (2020).
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the nuclei migrate into the promycelium that becomes septate (i.e., divided into cellular compartments separated by cell walls called
2131:"Light microscopy of basidia, basidiospores, and nuclei in spores and hyphae of Filobasidiella neoformans (Cryptococcus neoformans)" 1987: 1449:
Moore, R. T. (1980). "Taxonomic proposals for the classification of marine yeasts and other yeast-like fungi including the smuts".
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Other variations occur: some as standard lifecycles (that themselves have variations within variations) within specific orders.
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to undergo meiosis may contribute to their virulence by repairing the oxidative DNA damage caused by their host's release of
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given locus, and in such species, depending on the specifics, over 90% of monokaryons could be compatible with each other.
3540: 3530: 3513: 2500: 2490: 2473: 2366: 1501: 1298:, the elongated haploid basidiospores form apically, often in compatible pairs that fuse centrally resulting in H-shaped 1755:"Genome and transcriptome sequencing of the halophilic fungus Wallemia ichthyophaga: haloadaptations present and absent" 57: 1932:; Heitman, J. (April 2005). "Sexual reproduction between partners of the same mating type in Cryptococcus neoformans". 4710: 1813: 921:
Unlike animals and plants which have readily recognizable male and female counterparts, Basidiomycota (except for the
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recognized three subphyla (Pucciniomycotina, Ustilaginomycotina, Agaricomycotina) and two other class level taxa (
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rusts complete their life-cycles on one host instead of two, and microcyclic rusts cut out one or more stages.
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The maintenance of the dikaryotic status in dikaryons in many Basidiomycota is facilitated by the formation of
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Scheme of a typical basidiocarp, the dipoid reproductive structure of a basidiomycete, showing fruiting body,
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that physically appear to help coordinate and re-establish pairs of compatible nuclei following synchronous
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Rodriguez-Carres, Marianela; Findley, Keisha; Sun, Sheng; Dietrich, Fred S.; Heitman, Joseph (2010-03-10).
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Michod, R. E.; Bernstein, H.; Nedelcu, A. M. (May 2008). "Adaptive value of sex in microbial pathogens".
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and then the compatible nuclei migrate into each other's mycelia and pair up with the resident nuclei.
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is not yet placed in a subdivision, but recent genomic evidence suggests that it is a sister group of
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The dimorphic Basidiomycota with yeast stages and the pleiomorphic rusts are examples of fungi with
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species can have two-, four-, six-, or eight-spored basidia, and the cultivated button mushroom,
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Zajc, J.; Liu, Y.; Dai, W.; Yang, Z.; Hu, J.; Gostinčar, C.; Gunde-Cimerman, N. (Sep 13, 2013).
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Prillinger, Hansjörg; Lopandic, Ksenija; Schweigkofler, Wolfgang; et al. (February 2002).
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forms basidia on hyphae but the main infectious stage is more commonly known by the
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Traditionally, the Basidiomycota were divided into two classes, now obsolete:
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Smuts with both a yeast phase and an infectious hyphal state are examples of
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is an example. By convention, the stages and spore states are numbered by
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or the entire "basidium" acts as a "spore", e.g. in some false puffballs (
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is a basidiomycete mushroom. It is particularly suited to the study of
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Nonetheless these former concepts continue to be used as two types of
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is delayed, so that the compatible nuclei remain in pairs, called a
499:(an obsolete morphological based class of Basidiomycota that formed 5582: 2345: 5608: 5197: 5069: 4631: 4271: 1232: 1040: 912: 904: 896: 308: 292: 288: 219: 5235: 4727: 3888: 5489: 4699: 3409: 2362: 1256:. In the Pucciniales, the basidia are cylindrical and become 3- 5569: 554:
include the rust fungi, the insect parasitic/symbiotic genus
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released via dissolution or disintegration of the basidia.
194: 188: 179: 173: 167: 152: 442:(alternatively called holobasidiomycetes), including true 226:. More specifically, Basidiomycota includes these groups: 977:) mating system. This results in the fact that following 161: 608:
are most (but not all) of the former smut fungi and the
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Kirk, P. M.; Cannon, P. F.; Stalpers, J. A. (2008).
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often has six-spored basidia, while some corticioid
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was similar to two other fungal species, the yeasts
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with respect to familial placement). These include:
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A 2007 classification, adopted by a coalition of 67
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The classes of the Ustilaginomycotina are the 287:Basidiomycota are filamentous fungi composed of 1870: 1868: 1494: 1492: 2065: 2063: 4711: 3421: 2374: 1272:The characteristic part of the life-cycle of 1033:, etc. Ballistic basidiospores are formed on 8: 495:include what had previously been called the 2253:"LUXGENE.COM: The glow-in-the-dark website" 901:Sexual reproduction cycle of basidiomycetes 315:. However, some Basidiomycota are obligate 170: 5486: 5030: 4968: 4740: 4736: 4718: 4704: 4696: 4581: 4411: 4402: 4390: 4297: 4261: 4231: 4178: 4169: 4156: 4042: 3971: 3925: 3912: 3895: 3882: 3855: 3835: 3831: 3760: 3718: 3675: 3671: 3642: 3580: 3576: 3565: 3561: 3428: 3414: 3406: 3238: 3093: 2993: 2964: 2766: 2652: 2640: 2609: 2551: 2524: 2381: 2367: 2359: 29: 20: 2154: 2105: 2095: 1904: 1894: 1782: 1772: 1689: 1679: 1610: 716:Ammirati, A.D.Parker & Matheny (2007) 1355:with its apple-seed-shaped and coloured 917:Cell cycle of a Dikaryotic basidiomycete 686:Bernicchia, Gorjón & Nakasone (2011) 1664:"Outline of Fungi and fungus-like taxa" 1441: 1017:, hence they are sometimes also called 766:Tkalčec, Mešić & Chun Y.Deng (2015) 560:, a former group of smut fungi (in the 4686:denotes groups with over 1,000 species 2202:: CS1 maint: archived copy as title ( 2195: 1519:10.1146/annurev.micro.61.080706.093432 7: 5671:05d71028-e72e-49e8-ad74-0211ea111628 961:male nor female. They have either a 1706:"Lecture 7 : Ustilaginomycets" 646:W.P.Wu, B.Sutton & Gange (1997) 432:) with multiple orders and genera. 307:(usually four). These specialized 14: 1988:Infection, Genetics and Evolution 1174:). In the human pathogenic genus 616:, the Entorrhizomycetes, and the 471:groupings, the "mushrooms" (e.g. 1860:Kirk, Cannon & Stalpers 2008 1848:Kirk, Cannon & Stalpers 2008 1741:Kirk, Cannon & Stalpers 2008 1650:Kirk, Cannon & Stalpers 2008 328:components, and definitively by 145: 56: 2355:at the Tree of Life Web Project 1722:from the original on 2020-07-30 836:Hjortstam & Ryvarden (2008) 696:Hjortstam & Ryvarden (2008) 4165:      4161:      3921:      3917:      1426:List of Basidiomycota families 1029:, variously called mushrooms, 937:being composed of filamentous 477:) and the non-mushrooms (e.g. 284:, the human pathogenic yeast. 1: 5776:Taxa named by Royall T. Moore 2147:10.1128/jb.128.1.445-455.1976 1502:Annual Review of Microbiology 756:Dhingra & Priyanka (2011) 2129:Erke, K. H. (October 1976). 2097:10.1371/journal.pone.0009620 2001:10.1016/j.meegid.2008.01.002 1896:10.1371/journal.pgen.1001135 1621:10.1016/j.mycres.2007.03.004 1365:with its sclerotium and its 806:Marvanová & Bärl. (2000) 676:Marvanová & Bärl. (1998) 656:Marvanová & Bärl. (2000) 303:that normally bear external 210:, constitute the subkingdom 1814:Fungal Genetics and Biology 214:(often referred to as the " 5799: 786:K.Ando & Tubaki (1984) 515:), as well as most of the 4967: 4739: 4735: 4730:phyla/divisions by domain 4662: 4414: 4389: 4234: 4181: 4155: 4045: 3974: 3911: 3898: 3881: 3834: 3830: 3763: 3721: 3674: 3645: 3579: 3564: 3560: 3444: 3392: 2554: 2404: 1827:10.1016/j.fgb.2012.01.007 1681:10.5943/mycosphere/11/1/8 1566:10.1007/s10453-011-9234-y 1482:Dictionary.com Unabridged 1085:Schizosaccharomyces pombe 117: 112: 53:Scientific classification 51: 37: 28: 23: 3891:     3887:     1774:10.1186/1471-2164-14-617 1368:Tilachlidiopsis racemosa 1235:, which form dikaryotic 1123:Variations in lifecycles 1079:Saccharomyces cerevisiae 981:, the resulting haploid 909:Basidiomycota life cycle 206:that, together with the 5766:Fungi by classification 4940:Thermodesulfobacteriota 3281:Cryptomycocolacomycetes 3074:Sporopachydermiomycetes 2889:Calcarisporiellomycetes 2332:Dictionary of the Fungi 2135:Journal of Bacteriology 1463:10.1515/bot-1980-230605 1362:Dendrocollybia racemosa 1329:Cryptococcus neoformans 1244:in dry pustules called 1117:reactive oxygen species 1093:Cryptococcus neoformans 578:Cryptomycocolacomycetes 378:Cryptomycocolacomycetes 222:. Members are known as 5781:Taxa described in 1980 2931:Palaeovesiarbusmycetes 2881:Calcarisporiellomycota 2848:Ramicandelaberomycetes 2665:Neocallimastigomycetes 2334:(10th ed.). CABI. 1218:, and also protruding 1198:, previously known as 1050: 918: 910: 902: 332:molecular analysis of 218:") within the kingdom 202:) is one of two large 5692:Paleobiology Database 5168:Neocallimastigomycota 3398:(polyphyletic group). 3026:Schizosaccharomycetes 2688:Monoblepharidomycetes 2657:Neocallimastigomycota 1713:Cpb-us-e1.wpmucdn.com 1044: 916: 908: 900: 474:Schizophyllum commune 113:Subdivisions/Classes 45:Kunstformen der Natur 4833:Coprothermobacterota 3314:Peribolosporomycetes 3261:Agaricostilbomycetes 3147:Collemopsidiomycetes 3021:Pneumocystidomycetes 2820:Entomophthoromycetes 2683:Hyaloraphidiomycetes 2564:Paramicrosporidiales 2230:on 27 September 2007 1598:Mycological Research 1405:Decapitatus flavidus 1105:double-strand breaks 566:Agaricostilbomycetes 449:Heterobasidiomycetes 362:Agaricostilbomycetes 38:Basidiomycetes from 4563:Gunflint microbiota 4324:Klebsormidiophyceae 4108:Hyphochytriomycetes 3291:Spiculogloeomycetes 3266:Cystobasidiomycetes 3079:Trigonopsidomycetes 3049:Alloascoideomycetes 3006:Archaeorhizomycetes 2941:Archaeosporomycetes 2807:Entomophthoromycota 2746:Blastocladiomycetes 2741:Physodermatomycetes 2707:Caulochytriomycetes 2547:Microsporidiomycota 2088:2010PLoSO...5.9620R 2028:Chemical Immunology 1956:10.1038/nature03448 1948:2005Natur.434.1017L 1335:Cryptococcus gattii 1226:and probably other 1064:Coprinopsis cinerea 582:Cystobasidiomycetes 414:Spiculogloeomycetes 382:Cystobasidiomycetes 293:basidiomycota-yeast 5163:Blastocladiomycota 4334:Coleochaetophyceae 3714:    3710:    3638:    3634:    3367:Bartheletiomycetes 3319:Monilielliomycetes 3306:Ustilaginomycotina 3286:Atractiellomycetes 3271:Microbotryomycetes 3251:Tritirachiomycetes 3200:Laboulbeniomycetes 2936:Paraglomeromycetes 2871:Mortierellomycetes 2843:Dimargaritomycetes 2797:Basidiobolomycetes 2733:Blastocladiomycota 2675:Monoblepharomycota 2351:2020-11-26 at the 1862:, pp. 717–718 1283:Mycosarcoma maydis 1107:. The ability of 1099:Mycosarcoma maydis 1051: 1025:or basidiomes, or 933:which are usually 919: 911: 903: 893:Typical life cycle 826:Ghob.-Nejh. (2010) 662:Anthoseptobasidium 606:Ustilaginomycotina 600:Ustilaginomycotina 586:Microbotryomycetes 570:Atractiellomycetes 562:Microbotryomycetes 480:Mycosarcoma maydis 440:Homobasidiomycetes 422:Tritirachiomycetes 406:Monilielliomycetes 398:Microbotryomycetes 370:Bartheletiomycetes 366:Atractiellomycetes 131:Ustilaginomycotina 5743: 5742: 5679:Open Tree of Life 5492:Taxon identifiers 5483: 5482: 5479: 5478: 5457: 5456: 5020: 5019: 4963: 4962: 4955:Verrucomicrobiota 4693: 4692: 4658: 4657: 4594:Spironematellidae 4575:Meteora sporadica 4530: 4529: 4526: 4525: 4459: 4458: 4385: 4384: 4381: 4380: 4377: 4376: 4373: 4372: 4369: 4368: 4365: 4364: 4361: 4360: 4357: 4356: 4285:Prasinodermophyta 4221: 4220: 4151: 4150: 4147: 4146: 4143: 4142: 4139: 4138: 4135: 4134: 4032: 4031: 3868:Centroplasthelida 3826: 3825: 3822: 3821: 3818: 3817: 3814: 3813: 3810: 3809: 3806: 3805: 3802: 3801: 3704: 3703: 3628: 3627: 3403: 3402: 3388: 3387: 3384: 3383: 3380: 3379: 3334:Exobasidiomycetes 3329:Ustilaginomycetes 3324:Malasseziomycetes 3276:Classiculomycetes 3222: 3221: 3218: 3217: 3167:Xylobotryomycetes 3127:Candelariomycetes 2977:Entorrhizomycetes 2954: 2953: 2863:Mortierellomycota 2789:Basidiobolomycota 2754: 2753: 2718:Sanchytriomycetes 2630: 2629: 2601: 2600: 2597: 2596: 2040:10.1159/000058868 2034:(9413): 207–295. 1942:(7036): 1017–21. 1399:Mycena citricolor 1352:Collybia tuberosa 1143:Agaricus bisporus 991:clamp connections 887: 877: 867: 857: 856:W.Y.Zhuang (2008) 847: 837: 827: 817: 807: 797: 787: 777: 767: 757: 747: 737: 727: 726:P. Roberts (2006) 717: 707: 697: 687: 677: 667: 657: 647: 618:Ustilaginomycetes 614:Exobasidiomycetes 574:Classiculomycetes 426:Ustilaginomycetes 394:Malasseziomycetes 390:Exobasidiomycetes 374:Classiculomycetes 354:Entorrhizomycetes 138: 137: 108: 16:Division of fungi 5788: 5736: 5735: 5723: 5722: 5710: 5709: 5700: 5699: 5687: 5686: 5674: 5673: 5664: 5663: 5651: 5650: 5638: 5637: 5625: 5624: 5612: 5611: 5599: 5598: 5586: 5585: 5573: 5572: 5560: 5559: 5547: 5546: 5534: 5533: 5532: 5519: 5518: 5517: 5487: 5231:Anthocerotophyta 5110:Hemimastigophora 5031: 5002:Nitrososphaerota 4969: 4945:Thermomicrobiota 4885:Kiritimatiellota 4880:Ignavibacteriota 4845:Deferribacterota 4808:Campylobacterota 4793:Bdellovibrionota 4741: 4737: 4720: 4713: 4706: 4697: 4640:Collodictyonidae 4586:Hemimastigophora 4582: 4541: 4412: 4403: 4399: 4391: 4340:Zygnematophyceae 4298: 4278: 4262: 4232: 4179: 4170: 4166: 4157: 4043: 3972: 3926: 3922: 3913: 3896: 3892: 3883: 3856: 3843: 3836: 3832: 3786:Choanoflagellata 3761: 3719: 3715: 3676: 3672: 3668: 3643: 3639: 3581: 3577: 3573: 3566: 3562: 3473: 3453: 3430: 3423: 3416: 3407: 3396:fungi imperfecti 3243:Pucciniomycotina 3239: 3190:Geoglossomycetes 3132:Coniocybomycetes 3094: 3054:Dipodascomycetes 3041:Saccharomycotina 2998:Taphrinomycotina 2994: 2969:Entorrhizomycota 2965: 2853:Kickxellomycetes 2815:Neozygitomycetes 2767: 2723:Chytridiomycetes 2653: 2641: 2610: 2552: 2525: 2433: 2413: 2396:classification, 2383: 2376: 2369: 2360: 2335: 2318: 2317: 2315: 2314: 2305:. Archived from 2299: 2293: 2292: 2290: 2289: 2280:. Archived from 2274: 2268: 2267: 2265: 2264: 2255:. Archived from 2249: 2240: 2239: 2237: 2235: 2229: 2223:. Archived from 2222: 2214: 2208: 2207: 2201: 2193: 2191: 2190: 2181:. Archived from 2175: 2169: 2168: 2158: 2126: 2120: 2119: 2109: 2099: 2067: 2058: 2057: 2055: 2054: 2019: 2013: 2012: 1982: 1976: 1975: 1925: 1919: 1918: 1908: 1898: 1872: 1863: 1857: 1851: 1845: 1839: 1838: 1803: 1797: 1796: 1786: 1776: 1750: 1744: 1738: 1732: 1731: 1729: 1727: 1721: 1710: 1702: 1696: 1695: 1693: 1683: 1659: 1653: 1652:, pp. 78–79 1647: 1641: 1640: 1614: 1591: 1585: 1584: 1582: 1580: 1545: 1539: 1538: 1496: 1487: 1486: 1473: 1467: 1466: 1446: 1421:Forest pathology 1220:receptive hyphae 886:P.Roberts (1994) 885: 875: 865: 855: 845: 835: 825: 815: 805: 795: 785: 775: 765: 755: 745: 735: 732:Dacryomycetopsis 725: 715: 706:P.Roberts (2005) 705: 695: 685: 675: 665: 655: 645: 552:Pucciniomycotina 546:Pucciniomycotina 503:layers on their 451:, including the 322:clamp connection 201: 200: 197: 196: 193: 190: 187: 184: 181: 178: 175: 172: 169: 166: 163: 160: 157: 154: 151: 126:Pucciniomycotina 103: 61: 60: 33: 21: 5798: 5797: 5791: 5790: 5789: 5787: 5786: 5785: 5746: 5745: 5744: 5739: 5731: 5726: 5718: 5713: 5705: 5703: 5695: 5690: 5682: 5677: 5669: 5667: 5659: 5654: 5646: 5641: 5633: 5628: 5620: 5615: 5607: 5602: 5594: 5589: 5581: 5576: 5568: 5563: 5555: 5550: 5542: 5537: 5528: 5527: 5522: 5513: 5512: 5507: 5494: 5484: 5475: 5453: 5404:Gnathostomulida 5309:Xenacoelomorpha 5275: 5271:Flowering plant 5226:Marchantiophyta 5192: 5158:Chytridiomycota 5144: 5016: 4959: 4915:Planctomycetota 4875:Gemmatimonadota 4860:Elusimicrobiota 4753:Acidobacteriota 4731: 4724: 4694: 4689: 4654: 4626: 4617:Ancyromonadidae 4603: 4580: 4535: 4522: 4513:Malawimonadidae 4499: 4455: 4430: 4394: 4353: 4296: 4269: 4268: 4260: 4217: 4192: 4160: 4131: 4084: 4056: 4028: 3991: 3961: 3916: 3907: 3886: 3877: 3854: 3839: 3798: 3772: 3750: 3709: 3700: 3664: 3659: 3633: 3624: 3569: 3556: 3555: 3471: 3449: 3440: 3434: 3404: 3399: 3376: 3372:Tremellomycetes 3362:Wallemiomycetes 3344:Agaricomycotina 3338: 3300: 3296:Pucciniomycetes 3230: 3214: 3210:Sordariomycetes 3171: 3162:Lecanoromycetes 3152:Dothideomycetes 3142:Arthoniomycetes 3113: 3083: 3069:Saccharomycetes 3035: 3031:Taphrinomycetes 3011:Neolectomycetes 2989: 2981: 2950: 2916: 2907:Endogonomycetes 2893: 2875: 2857: 2830:Kickxellomycota 2824: 2801: 2783: 2762: 2750: 2727: 2698:Chytridiomycota 2692: 2669: 2648: 2626: 2614:Aphelidiomycota 2606:Aphelidiomyceta 2593: 2584:Metchnikovellea 2579:Chytridiopsidea 2567: 2558:Morellosporales 2541: 2516: 2515: 2431: 2409: 2400: 2387: 2353:Wayback Machine 2342: 2329: 2326: 2321: 2312: 2310: 2301: 2300: 2296: 2287: 2285: 2276: 2275: 2271: 2262: 2260: 2251: 2250: 2243: 2233: 2231: 2227: 2220: 2216: 2215: 2211: 2194: 2188: 2186: 2179:"Archived copy" 2177: 2176: 2172: 2128: 2127: 2123: 2069: 2068: 2061: 2052: 2050: 2021: 2020: 2016: 1984: 1983: 1979: 1927: 1926: 1922: 1889:(9): e1001135. 1874: 1873: 1866: 1858: 1854: 1846: 1842: 1805: 1804: 1800: 1752: 1751: 1747: 1739: 1735: 1725: 1723: 1719: 1708: 1704: 1703: 1699: 1661: 1660: 1656: 1648: 1644: 1612:10.1.1.626.9582 1593: 1592: 1588: 1578: 1576: 1547: 1546: 1542: 1498: 1497: 1490: 1477:"Basidiomycota" 1475: 1474: 1470: 1451:Botanica Marina 1448: 1447: 1443: 1439: 1431:Mating in fungi 1417: 1411:called gemmae. 1295:Tilletia caries 1270: 1189: 1165:Tetragoniomyces 1160:Physalacriaceae 1125: 1060: 895: 890: 822:Punctulariopsis 746:Vellinga (2011) 626: 624:Genera included 602: 594:Pucciniomycetes 548: 540:Agaricomycotina 536:Wallemiomycetes 529:Tremellomycetes 493:Agaricomycotina 489: 487:Agaricomycotina 430:Wallemiomycetes 418:Tremellomycetes 410:Pucciniomycetes 350:Wallemiomycetes 342: 148: 144: 121:Agaricomycotina 102: 55: 17: 12: 11: 5: 5796: 5795: 5792: 5784: 5783: 5778: 5773: 5768: 5763: 5758: 5748: 5747: 5741: 5740: 5738: 5737: 5724: 5711: 5701: 5688: 5675: 5665: 5652: 5639: 5626: 5613: 5600: 5587: 5574: 5561: 5548: 5535: 5520: 5504: 5502: 5496: 5495: 5490: 5481: 5480: 5477: 5476: 5469: 5467: 5464:Incertae sedis 5459: 5458: 5455: 5454: 5452: 5451: 5446: 5441: 5436: 5431: 5426: 5421: 5416: 5411: 5409:Micrognathozoa 5406: 5401: 5399:Acanthocephala 5396: 5391: 5386: 5381: 5376: 5371: 5366: 5361: 5356: 5351: 5346: 5341: 5336: 5331: 5326: 5321: 5316: 5311: 5306: 5301: 5296: 5291: 5285: 5283: 5277: 5276: 5274: 5273: 5268: 5263: 5258: 5253: 5248: 5243: 5241:Lycopodiophyta 5238: 5233: 5228: 5223: 5218: 5213: 5208: 5202: 5200: 5194: 5193: 5191: 5190: 5185: 5180: 5175: 5170: 5165: 5160: 5154: 5152: 5146: 5145: 5143: 5142: 5137: 5132: 5127: 5122: 5117: 5112: 5107: 5102: 5097: 5092: 5087: 5082: 5077: 5072: 5067: 5062: 5057: 5055:Ancyromonadida 5052: 5047: 5041: 5039: 5028: 5022: 5021: 5018: 5017: 5015: 5014: 5009: 5007:Thermoproteota 5004: 4999: 4992: 4985: 4977: 4975: 4965: 4964: 4961: 4960: 4958: 4957: 4952: 4947: 4942: 4937: 4932: 4927: 4922: 4920:Pseudomonadota 4917: 4912: 4907: 4902: 4897: 4895:Mycoplasmatota 4892: 4890:Lentisphaerota 4887: 4882: 4877: 4872: 4870:Fusobacteriota 4867: 4865:Fibrobacterota 4862: 4857: 4852: 4847: 4842: 4835: 4830: 4825: 4820: 4815: 4810: 4805: 4800: 4795: 4790: 4785: 4780: 4775: 4770: 4768:Armatimonadota 4765: 4760: 4758:Actinomycetota 4755: 4749: 4747: 4733: 4732: 4725: 4723: 4722: 4715: 4708: 4700: 4691: 4690: 4688: 4687: 4681: 4674: 4670:excluded from 4663: 4660: 4659: 4656: 4655: 4653: 4652: 4647: 4642: 4636: 4634: 4628: 4627: 4625: 4624: 4622:Planomonadidae 4619: 4613: 4611: 4609:Ancyromonadida 4605: 4604: 4602: 4601: 4599:Paramastigidae 4596: 4590: 4588: 4579: 4578: 4571: 4565: 4559: 4551: 4544: 4542: 4538:Incertae sedis 4532: 4531: 4528: 4527: 4524: 4523: 4521: 4520: 4515: 4509: 4507: 4501: 4500: 4498: 4497: 4492: 4487: 4482: 4477: 4469: 4467: 4461: 4460: 4457: 4456: 4454: 4453: 4446: 4440: 4438: 4432: 4431: 4429: 4428: 4421: 4415: 4409: 4400: 4387: 4386: 4383: 4382: 4379: 4378: 4375: 4374: 4371: 4370: 4367: 4366: 4363: 4362: 4359: 4358: 4355: 4354: 4352: 4351: 4343: 4336: 4331: 4326: 4321: 4314: 4306: 4304: 4295: 4294: 4287: 4281: 4279: 4259: 4258: 4251: 4246: 4241: 4235: 4229: 4227:Archaeplastida 4223: 4222: 4219: 4218: 4216: 4215: 4210: 4202: 4200: 4194: 4193: 4191: 4190: 4182: 4176: 4167: 4153: 4152: 4149: 4148: 4145: 4144: 4141: 4140: 4137: 4136: 4133: 4132: 4130: 4129: 4124: 4117: 4110: 4105: 4100: 4094: 4092: 4086: 4085: 4083: 4082: 4077: 4072: 4066: 4064: 4058: 4057: 4055: 4054: 4046: 4040: 4034: 4033: 4030: 4029: 4027: 4026: 4021: 4018:Dinoflagellata 4014: 4012:Chrompodellida 4009: 4001: 3999: 3993: 3992: 3990: 3989: 3982: 3975: 3969: 3963: 3962: 3960: 3959: 3952: 3945: 3940: 3934: 3932: 3923: 3909: 3908: 3906: 3905: 3899: 3893: 3879: 3878: 3876: 3875: 3870: 3864: 3862: 3853: 3852: 3846: 3844: 3841:Diaphoretickes 3828: 3827: 3824: 3823: 3820: 3819: 3816: 3815: 3812: 3811: 3808: 3807: 3804: 3803: 3800: 3799: 3797: 3796: 3788: 3782: 3780: 3774: 3773: 3771: 3770: 3764: 3758: 3752: 3751: 3749: 3748: 3743: 3738: 3730: 3722: 3716: 3706: 3705: 3702: 3701: 3699: 3698: 3690: 3684: 3682: 3669: 3661: 3660: 3658: 3657: 3652: 3646: 3640: 3630: 3629: 3626: 3625: 3623: 3622: 3617: 3612: 3607: 3600: 3595: 3589: 3587: 3574: 3558: 3557: 3554: 3553: 3552: 3551: 3550: 3549: 3546:Mesomycetozoea 3543: 3538: 3528: 3518: 3517: 3516: 3511: 3506: 3501: 3496: 3495: 3494: 3482:Diaphoretickes 3479: 3474: 3469: 3464: 3459: 3454: 3446: 3445: 3442: 3441: 3439:classification 3435: 3433: 3432: 3425: 3418: 3410: 3401: 3400: 3393: 3390: 3389: 3386: 3385: 3382: 3381: 3378: 3377: 3375: 3374: 3369: 3364: 3359: 3357:Agaricomycetes 3354: 3352:Dacrymycetales 3348: 3346: 3340: 3339: 3337: 3336: 3331: 3326: 3321: 3316: 3310: 3308: 3302: 3301: 3299: 3298: 3293: 3288: 3283: 3278: 3273: 3268: 3263: 3258: 3253: 3247: 3245: 3236: 3224: 3223: 3220: 3219: 3216: 3215: 3213: 3212: 3207: 3202: 3197: 3192: 3187: 3181: 3179: 3177:Sordariomyceta 3173: 3172: 3170: 3169: 3164: 3159: 3157:Eurotiomycetes 3154: 3149: 3144: 3139: 3137:Lichinomycetes 3134: 3129: 3123: 3121: 3119:Dothideomyceta 3115: 3114: 3112: 3111: 3106: 3104:Orbiliomycetes 3100: 3098: 3091: 3089:Pezizomycotina 3085: 3084: 3082: 3081: 3076: 3071: 3066: 3061: 3056: 3051: 3045: 3043: 3037: 3036: 3034: 3033: 3028: 3023: 3018: 3013: 3008: 3002: 3000: 2991: 2983: 2982: 2980: 2979: 2973: 2971: 2962: 2956: 2955: 2952: 2951: 2949: 2948: 2946:Glomeromycetes 2943: 2938: 2933: 2926: 2924: 2918: 2917: 2915: 2914: 2909: 2903: 2901: 2895: 2894: 2892: 2891: 2885: 2883: 2877: 2876: 2874: 2873: 2867: 2865: 2859: 2858: 2856: 2855: 2850: 2845: 2840: 2838:Zoopagomycetes 2834: 2832: 2826: 2825: 2823: 2822: 2817: 2811: 2809: 2803: 2802: 2800: 2799: 2793: 2791: 2785: 2784: 2782: 2781: 2779:Olpidiomycetes 2775: 2773: 2764: 2763:(paraphyletic) 2756: 2755: 2752: 2751: 2749: 2748: 2743: 2737: 2735: 2729: 2728: 2726: 2725: 2720: 2715: 2709: 2702: 2700: 2694: 2693: 2691: 2690: 2685: 2679: 2677: 2671: 2670: 2668: 2667: 2661: 2659: 2650: 2649:(paraphyletic) 2638: 2632: 2631: 2628: 2627: 2625: 2624: 2618: 2616: 2607: 2603: 2602: 2599: 2598: 2595: 2594: 2592: 2591: 2586: 2581: 2575: 2573: 2569: 2568: 2566: 2565: 2562: 2561:Nucleophagales 2559: 2555: 2549: 2543: 2542: 2540: 2539: 2537:Rozellomycetes 2533: 2531: 2522: 2518: 2517: 2514: 2513: 2512: 2511: 2510: 2509: 2506:Mesomycetozoea 2503: 2498: 2488: 2478: 2477: 2476: 2471: 2466: 2461: 2456: 2455: 2454: 2442:Diaphoretickes 2439: 2434: 2429: 2424: 2419: 2414: 2406: 2405: 2402: 2401: 2388: 2386: 2385: 2378: 2371: 2363: 2357: 2356: 2341: 2340:External links 2338: 2337: 2336: 2325: 2322: 2320: 2319: 2294: 2269: 2241: 2209: 2170: 2141:(1): 445–455. 2121: 2059: 2014: 1977: 1920: 1864: 1852: 1840: 1798: 1745: 1733: 1697: 1654: 1642: 1605:(5): 509–547. 1586: 1540: 1511:Annual Reviews 1488: 1485:(Online). n.d. 1468: 1440: 1438: 1435: 1434: 1433: 1428: 1423: 1416: 1413: 1313:Filobasidiella 1269: 1266: 1208:Roman numerals 1188: 1185: 1158:, both in the 1124: 1121: 1059: 1056: 1019:ballistospores 976: 972: 968: 964: 960: 894: 891: 889: 888: 878: 868: 858: 848: 838: 828: 818: 808: 798: 788: 778: 768: 758: 748: 738: 728: 718: 708: 698: 692:Brevicellopsis 688: 678: 668: 658: 648: 637: 632:incertae sedis 625: 622: 601: 598: 547: 544: 521:Agaricomycetes 509:Gasteromycetes 488: 485: 465: 464: 446: 358:Agaricomycetes 341: 340:Classification 338: 224:basidiomycetes 136: 135: 134: 133: 128: 123: 115: 114: 110: 109: 97: 93: 92: 87: 83: 82: 77: 73: 72: 67: 63: 62: 49: 48: 35: 34: 26: 25: 24:Basidiomycota 15: 13: 10: 9: 6: 4: 3: 2: 5794: 5793: 5782: 5779: 5777: 5774: 5772: 5769: 5767: 5764: 5762: 5759: 5757: 5756:Basidiomycota 5754: 5753: 5751: 5734: 5729: 5725: 5721: 5716: 5712: 5708: 5707:basidiomycota 5702: 5698: 5693: 5689: 5685: 5680: 5676: 5672: 5666: 5662: 5657: 5653: 5649: 5644: 5640: 5636: 5631: 5627: 5623: 5618: 5614: 5610: 5605: 5601: 5597: 5592: 5588: 5584: 5579: 5575: 5571: 5566: 5562: 5558: 5553: 5549: 5545: 5540: 5536: 5531: 5530:Basidiomycota 5525: 5521: 5516: 5510: 5506: 5505: 5503: 5501: 5500:Basidiomycota 5497: 5493: 5488: 5474: 5473: 5468: 5466: 5465: 5460: 5450: 5447: 5445: 5442: 5440: 5437: 5435: 5432: 5430: 5427: 5425: 5422: 5420: 5417: 5415: 5412: 5410: 5407: 5405: 5402: 5400: 5397: 5395: 5392: 5390: 5387: 5385: 5382: 5380: 5377: 5375: 5372: 5370: 5367: 5365: 5362: 5360: 5357: 5355: 5352: 5350: 5347: 5345: 5342: 5340: 5337: 5335: 5332: 5330: 5327: 5325: 5322: 5320: 5317: 5315: 5312: 5310: 5307: 5305: 5302: 5300: 5297: 5295: 5292: 5290: 5287: 5286: 5284: 5282: 5278: 5272: 5269: 5267: 5264: 5262: 5259: 5257: 5254: 5252: 5249: 5247: 5244: 5242: 5239: 5237: 5234: 5232: 5229: 5227: 5224: 5222: 5219: 5217: 5214: 5212: 5209: 5207: 5204: 5203: 5201: 5199: 5195: 5189: 5188:Basidiomycota 5186: 5184: 5181: 5179: 5176: 5174: 5173:Glomeromycota 5171: 5169: 5166: 5164: 5161: 5159: 5156: 5155: 5153: 5151: 5147: 5141: 5138: 5136: 5135:Stramenopiles 5133: 5131: 5128: 5126: 5123: 5121: 5118: 5116: 5113: 5111: 5108: 5106: 5103: 5101: 5098: 5096: 5093: 5091: 5088: 5086: 5083: 5081: 5078: 5076: 5073: 5071: 5068: 5066: 5063: 5061: 5060:Apusomonadida 5058: 5056: 5053: 5051: 5048: 5046: 5043: 5042: 5040: 5037: 5032: 5029: 5027: 5023: 5013: 5010: 5008: 5005: 5003: 5000: 4997: 4996:Nanoarchaeota 4993: 4990: 4986: 4983: 4982:Euryarchaeota 4979: 4978: 4976: 4974: 4970: 4966: 4956: 4953: 4951: 4948: 4946: 4943: 4941: 4938: 4936: 4933: 4931: 4930:Spirochaetota 4928: 4926: 4925:Rhodothermota 4923: 4921: 4918: 4916: 4913: 4911: 4908: 4906: 4903: 4901: 4898: 4896: 4893: 4891: 4888: 4886: 4883: 4881: 4878: 4876: 4873: 4871: 4868: 4866: 4863: 4861: 4858: 4856: 4855:Dictyoglomota 4853: 4851: 4848: 4846: 4843: 4840: 4839:Cyanobacteria 4836: 4834: 4831: 4829: 4828:Chrysiogenota 4826: 4824: 4823:Chloroflexota 4821: 4819: 4816: 4814: 4811: 4809: 4806: 4804: 4803:Calditrichota 4801: 4799: 4798:Caldisericota 4796: 4794: 4791: 4789: 4786: 4784: 4781: 4779: 4776: 4774: 4773:Atribacterota 4771: 4769: 4766: 4764: 4761: 4759: 4756: 4754: 4751: 4750: 4748: 4746: 4742: 4738: 4734: 4729: 4721: 4716: 4714: 4709: 4707: 4702: 4701: 4698: 4685: 4682: 4679: 4675: 4673: 4669: 4666:¹traditional 4665: 4664: 4661: 4651: 4648: 4646: 4645:Mantamonadida 4643: 4641: 4638: 4637: 4635: 4633: 4629: 4623: 4620: 4618: 4615: 4614: 4612: 4610: 4606: 4600: 4597: 4595: 4592: 4591: 4589: 4587: 4583: 4577: 4576: 4572: 4570: 4566: 4564: 4560: 4558: 4557: 4552: 4550: 4546: 4545: 4543: 4540: 4539: 4533: 4519: 4516: 4514: 4511: 4510: 4508: 4506: 4502: 4496: 4493: 4491: 4488: 4486: 4483: 4481: 4478: 4476: 4475: 4471: 4470: 4468: 4466: 4462: 4452: 4451: 4447: 4445: 4444:Heterolobosea 4442: 4441: 4439: 4437: 4436:Discicristata 4433: 4427: 4426: 4422: 4420: 4417: 4416: 4413: 4410: 4408: 4404: 4401: 4397: 4392: 4388: 4349: 4348: 4344: 4342: 4341: 4337: 4335: 4332: 4330: 4327: 4325: 4322: 4320: 4319: 4315: 4313: 4312: 4308: 4307: 4305: 4303: 4299: 4293: 4292: 4288: 4286: 4283: 4282: 4280: 4276: 4275: 4267: 4266:Viridiplantae 4263: 4257: 4256: 4252: 4250: 4247: 4245: 4242: 4240: 4237: 4236: 4233: 4230: 4228: 4224: 4214: 4211: 4209: 4208: 4204: 4203: 4201: 4199: 4195: 4189: 4188: 4187:Microheliella 4184: 4183: 4180: 4177: 4175: 4171: 4168: 4164: 4158: 4154: 4128: 4127:Actinophryida 4125: 4123: 4122: 4118: 4116: 4115: 4111: 4109: 4106: 4104: 4101: 4099: 4096: 4095: 4093: 4091: 4087: 4081: 4078: 4076: 4073: 4071: 4068: 4067: 4065: 4063: 4059: 4053: 4052: 4048: 4047: 4044: 4041: 4039: 4038:Stramenopiles 4035: 4025: 4022: 4020: 4019: 4015: 4013: 4010: 4008: 4007: 4003: 4002: 4000: 3998: 3994: 3988: 3987: 3983: 3980: 3977: 3976: 3973: 3970: 3968: 3964: 3958: 3957: 3953: 3951: 3950: 3946: 3944: 3941: 3939: 3936: 3935: 3933: 3931: 3927: 3924: 3920: 3914: 3910: 3904: 3901: 3900: 3897: 3894: 3890: 3884: 3880: 3874: 3871: 3869: 3866: 3865: 3863: 3861: 3857: 3851: 3848: 3847: 3845: 3842: 3837: 3833: 3829: 3794: 3793: 3789: 3787: 3784: 3783: 3781: 3779: 3775: 3769: 3766: 3765: 3762: 3759: 3757: 3753: 3747: 3744: 3742: 3741:Ichthyosporea 3739: 3737: 3736: 3731: 3729: 3728: 3724: 3723: 3720: 3717: 3713: 3707: 3696: 3695: 3691: 3689: 3688:Rotosphaerida 3686: 3685: 3683: 3681: 3677: 3673: 3670: 3667: 3662: 3656: 3653: 3651: 3650:Apusomonadida 3648: 3647: 3644: 3641: 3637: 3631: 3621: 3618: 3616: 3613: 3611: 3608: 3606: 3605: 3601: 3599: 3596: 3594: 3591: 3590: 3588: 3586: 3582: 3578: 3575: 3572: 3567: 3563: 3559: 3547: 3544: 3542: 3539: 3537: 3534: 3533: 3532: 3529: 3527: 3524: 3523: 3522: 3519: 3515: 3512: 3510: 3509:Stramenopiles 3507: 3505: 3502: 3500: 3497: 3493: 3490: 3489: 3488: 3485: 3484: 3483: 3480: 3478: 3475: 3472:(major groups 3470: 3468: 3465: 3463: 3460: 3458: 3455: 3452: 3448: 3447: 3443: 3438: 3431: 3426: 3424: 3419: 3417: 3412: 3411: 3408: 3397: 3391: 3373: 3370: 3368: 3365: 3363: 3360: 3358: 3355: 3353: 3350: 3349: 3347: 3345: 3341: 3335: 3332: 3330: 3327: 3325: 3322: 3320: 3317: 3315: 3312: 3311: 3309: 3307: 3303: 3297: 3294: 3292: 3289: 3287: 3284: 3282: 3279: 3277: 3274: 3272: 3269: 3267: 3264: 3262: 3259: 3257: 3254: 3252: 3249: 3248: 3246: 3244: 3240: 3237: 3234: 3229: 3228:Basidiomycota 3225: 3211: 3208: 3206: 3203: 3201: 3198: 3196: 3195:Leotiomycetes 3193: 3191: 3188: 3186: 3185:Xylonomycetes 3183: 3182: 3180: 3178: 3174: 3168: 3165: 3163: 3160: 3158: 3155: 3153: 3150: 3148: 3145: 3143: 3140: 3138: 3135: 3133: 3130: 3128: 3125: 3124: 3122: 3120: 3116: 3110: 3109:Pezizomycetes 3107: 3105: 3102: 3101: 3099: 3095: 3092: 3090: 3086: 3080: 3077: 3075: 3072: 3070: 3067: 3065: 3064:Pichiomycetes 3062: 3060: 3057: 3055: 3052: 3050: 3047: 3046: 3044: 3042: 3038: 3032: 3029: 3027: 3024: 3022: 3019: 3017: 3016:Novakomycetes 3014: 3012: 3009: 3007: 3004: 3003: 3001: 2999: 2995: 2992: 2988: 2984: 2978: 2975: 2974: 2972: 2970: 2966: 2963: 2961: 2957: 2947: 2944: 2942: 2939: 2937: 2934: 2932: 2928: 2927: 2925: 2923: 2922:Glomeromycota 2919: 2913: 2912:Mucoromycetes 2910: 2908: 2905: 2904: 2902: 2900: 2896: 2890: 2887: 2886: 2884: 2882: 2878: 2872: 2869: 2868: 2866: 2864: 2860: 2854: 2851: 2849: 2846: 2844: 2841: 2839: 2836: 2835: 2833: 2831: 2827: 2821: 2818: 2816: 2813: 2812: 2810: 2808: 2804: 2798: 2795: 2794: 2792: 2790: 2786: 2780: 2777: 2776: 2774: 2772: 2771:Olpidiomycota 2768: 2765: 2761: 2757: 2747: 2744: 2742: 2739: 2738: 2736: 2734: 2730: 2724: 2721: 2719: 2716: 2714: 2710: 2708: 2704: 2703: 2701: 2699: 2695: 2689: 2686: 2684: 2681: 2680: 2678: 2676: 2672: 2666: 2663: 2662: 2660: 2658: 2654: 2651: 2646: 2642: 2639: 2637: 2633: 2623: 2620: 2619: 2617: 2615: 2611: 2608: 2604: 2590: 2587: 2585: 2582: 2580: 2577: 2576: 2574: 2572:Microsporidia 2570: 2563: 2560: 2557: 2556: 2553: 2550: 2548: 2544: 2538: 2535: 2534: 2532: 2530: 2529:Rozellomycota 2526: 2523: 2521:Rozellomyceta 2519: 2507: 2504: 2502: 2499: 2497: 2494: 2493: 2492: 2489: 2487: 2484: 2483: 2482: 2479: 2475: 2472: 2470: 2469:Stramenopiles 2467: 2465: 2462: 2460: 2457: 2453: 2450: 2449: 2448: 2445: 2444: 2443: 2440: 2438: 2435: 2432:(major groups 2430: 2428: 2425: 2423: 2420: 2418: 2415: 2412: 2408: 2407: 2403: 2399: 2398:fungal orders 2395: 2391: 2384: 2379: 2377: 2372: 2370: 2365: 2364: 2361: 2354: 2350: 2347: 2346:Basidiomycota 2344: 2343: 2339: 2333: 2328: 2327: 2323: 2309:on 2006-12-21 2308: 2304: 2298: 2295: 2284:on 2006-12-21 2283: 2279: 2273: 2270: 2259:on 2007-09-28 2258: 2254: 2248: 2246: 2242: 2226: 2219: 2213: 2210: 2205: 2199: 2185:on 2007-10-25 2184: 2180: 2174: 2171: 2166: 2162: 2157: 2152: 2148: 2144: 2140: 2136: 2132: 2125: 2122: 2117: 2113: 2108: 2103: 2098: 2093: 2089: 2085: 2081: 2077: 2073: 2066: 2064: 2060: 2049: 2045: 2041: 2037: 2033: 2029: 2025: 2018: 2015: 2010: 2006: 2002: 1998: 1995:(3): 267–85. 1994: 1990: 1989: 1981: 1978: 1973: 1969: 1965: 1961: 1957: 1953: 1949: 1945: 1941: 1937: 1936: 1931: 1924: 1921: 1916: 1912: 1907: 1902: 1897: 1892: 1888: 1884: 1883: 1882:PLOS Genetics 1878: 1871: 1869: 1865: 1861: 1856: 1853: 1850:, p. 581 1849: 1844: 1841: 1836: 1832: 1828: 1824: 1821:(3): 217–26. 1820: 1816: 1815: 1810: 1802: 1799: 1794: 1790: 1785: 1780: 1775: 1770: 1766: 1762: 1761: 1756: 1749: 1746: 1742: 1737: 1734: 1718: 1714: 1707: 1701: 1698: 1692: 1687: 1682: 1677: 1674:: 1060–1456. 1673: 1669: 1665: 1658: 1655: 1651: 1646: 1643: 1638: 1634: 1630: 1626: 1622: 1618: 1613: 1608: 1604: 1600: 1599: 1590: 1587: 1575: 1571: 1567: 1563: 1559: 1555: 1551: 1544: 1541: 1536: 1532: 1528: 1524: 1520: 1516: 1512: 1508: 1504: 1503: 1495: 1493: 1489: 1484: 1483: 1478: 1472: 1469: 1464: 1460: 1456: 1452: 1445: 1442: 1436: 1432: 1429: 1427: 1424: 1422: 1419: 1418: 1414: 1412: 1410: 1407: 1406: 1401: 1400: 1395: 1392: 1391: 1386: 1385: 1384:Hohenbuehelia 1380: 1376: 1375: 1370: 1369: 1364: 1363: 1358: 1354: 1353: 1348: 1344: 1339: 1337: 1336: 1331: 1330: 1325: 1324: 1319: 1315: 1314: 1308: 1303: 1301: 1297: 1296: 1291: 1290: 1285: 1284: 1279: 1275: 1267: 1265: 1263: 1259: 1255: 1251: 1247: 1243: 1238: 1234: 1229: 1225: 1221: 1217: 1213: 1209: 1205: 1201: 1197: 1193: 1186: 1184: 1181: 1179: 1178: 1173: 1172: 1167: 1166: 1161: 1157: 1156: 1151: 1150: 1145: 1144: 1139: 1138: 1133: 1132: 1122: 1120: 1118: 1114: 1110: 1109:C. neoformans 1106: 1101: 1100: 1095: 1094: 1089: 1087: 1086: 1081: 1080: 1075: 1070: 1066: 1065: 1057: 1055: 1048: 1043: 1039: 1036: 1032: 1028: 1024: 1020: 1016: 1012: 1008: 1004: 1000: 996: 992: 987: 984: 983:basidiospores 980: 974: 970: 966: 962: 958: 956: 952: 948: 944: 940: 936: 932: 928: 924: 915: 907: 899: 892: 884: 883: 879: 874: 873: 869: 864: 863: 859: 854: 853: 849: 844: 843: 839: 834: 833: 829: 824: 823: 819: 816:Corner (1966) 814: 813: 809: 804: 803: 799: 796:Wakef. (1934) 794: 793: 789: 784: 783: 779: 776:Oberw. (1990) 774: 773: 769: 764: 763: 759: 754: 753: 749: 744: 743: 739: 734: 733: 729: 724: 723: 719: 714: 713: 709: 704: 703: 699: 694: 693: 689: 684: 683: 679: 674: 673: 669: 664: 663: 659: 654: 653: 652:Anguillomyces 649: 644: 643: 639: 638: 636: 634: 633: 623: 621: 619: 615: 611: 610:Exobasidiales 607: 599: 597: 595: 591: 587: 583: 579: 575: 571: 567: 563: 559: 558: 557:Septobasidium 553: 545: 543: 541: 537: 532: 530: 526: 522: 518: 514: 510: 506: 502: 498: 497:Hymenomycetes 494: 486: 484: 482: 481: 476: 475: 470: 462: 458: 454: 450: 447: 445: 441: 438: 437: 436: 433: 431: 427: 423: 419: 415: 411: 407: 403: 399: 395: 391: 387: 383: 379: 375: 371: 367: 363: 359: 355: 351: 347: 339: 337: 335: 331: 327: 323: 318: 314: 313:basidiospores 310: 306: 302: 298: 294: 290: 285: 283: 282: 277: 276:mirror yeasts 273: 269: 265: 261: 257: 253: 249: 245: 241: 240:bracket fungi 237: 233: 229: 225: 221: 217: 213: 209: 205: 199: 142: 141:Basidiomycota 132: 129: 127: 124: 122: 119: 118: 116: 111: 106: 101: 100:Basidiomycota 98: 95: 94: 91: 88: 85: 84: 81: 78: 75: 74: 71: 68: 65: 64: 59: 54: 50: 47: 46: 41: 40:Ernst Haeckel 36: 32: 27: 22: 19: 5761:Fungus phyla 5499: 5470: 5462: 5384:Orthonectida 5379:Gastrotricha 5354:Nematomorpha 5329:Chaetognatha 5319:Hemichordate 5246:Pteridophyta 5187: 5115:Malawimonada 4989:Korarchaeota 4950:Thermotogota 4935:Synergistota 4910:Nitrospirota 4905:Nitrospinota 4850:Deinococcota 4783:Bacteroidota 4683: 4678:paraphyletic 4573: 4554: 4536: 4505:Malawimonada 4485:Anaeromonada 4480:Anaeramoebae 4472: 4448: 4425:Tsukubamonas 4423: 4345: 4338: 4329:Charophyceae 4316: 4309: 4302:Streptophyta 4289: 4273: 4253: 4249:Rhodelphidia 4207:Palpitomonas 4205: 4185: 4174:Pancryptista 4119: 4112: 4049: 4016: 4004: 3984: 3954: 3949:Foraminifera 3947: 3790: 3733: 3727:Tunicaraptor 3725: 3692: 3666:Opisthokonta 3602: 3531:Opisthokonta 3256:Mixiomycetes 3227: 3205:Sareomycetes 2899:Mucoromycota 2713:Coenomycetes 2491:Opisthokonta 2331: 2311:. Retrieved 2307:the original 2297: 2286:. Retrieved 2282:the original 2272: 2261:. Retrieved 2257:the original 2232:. Retrieved 2225:the original 2212: 2187:. Retrieved 2183:the original 2173: 2138: 2134: 2124: 2082:(3): e9620. 2079: 2075: 2051:. Retrieved 2031: 2027: 2017: 1992: 1986: 1980: 1939: 1933: 1923: 1886: 1880: 1855: 1843: 1818: 1812: 1801: 1764: 1760:BMC Genomics 1758: 1748: 1743:, p. 13 1736: 1724:. Retrieved 1712: 1700: 1671: 1667: 1657: 1645: 1602: 1596: 1589: 1577:. Retrieved 1560:(2): 83–97. 1557: 1554:Aerobiologia 1553: 1543: 1506: 1500: 1480: 1471: 1454: 1450: 1444: 1403: 1397: 1390:Nematoctonus 1388: 1382: 1372: 1366: 1360: 1350: 1340: 1333: 1327: 1323:Cryptococcus 1321: 1311: 1304: 1293: 1287: 1281: 1271: 1242:urediospores 1190: 1182: 1177:Cryptococcus 1175: 1169: 1163: 1153: 1147: 1141: 1135: 1129: 1126: 1112: 1108: 1097: 1091: 1090: 1083: 1077: 1073: 1062: 1061: 1052: 1049:and basidia. 1023:basidiocarps 1005:) to form a 988: 967:unifactorial 920: 880: 876:Corda (1837) 870: 860: 850: 846:Vánky (2008) 840: 832:Radulodontia 830: 820: 812:Paraphelaria 810: 800: 790: 780: 772:Kryptastrina 770: 760: 752:Hallenbergia 750: 740: 730: 720: 710: 700: 690: 680: 670: 660: 650: 640: 630: 627: 603: 590:Mixiomycetes 555: 549: 533: 525:Dacrymycetes 490: 478: 472: 469:growth habit 466: 434: 402:Mixiomycetes 386:Dacrymycetes 343: 334:DNA sequence 330:phylogenetic 291:(except for 286: 281:Cryptococcus 279: 256:chanterelles 223: 216:higher fungi 215: 140: 139: 99: 86:Subkingdom: 43: 18: 5617:iNaturalist 5524:Wikispecies 5439:Brachiopoda 5414:Cycliophora 5359:Onychophora 5334:Kinorhyncha 5256:Ginkgophyta 5251:Cycadophyta 5216:Chlorophyta 5206:Glaucophyta 4900:Myxococcota 4818:Chlorobiota 4813:Chlamydiota 4569:Chitinozoan 4495:Parabasalia 4347:Embryophyta 4311:Chlorokybus 4291:Chlorophyta 4239:Glaucophyta 4213:Cryptophyta 4103:Pirsoniales 4070:Bicosoecida 4051:Platysulcus 4024:Perkinsozoa 4006:Apicomplexa 3979:Colponemida 3746:Pluriformea 3615:Archamoebea 3604:Eumycetozoa 3059:Lipomycetes 2990:(sac fungi) 2589:Microsporea 2390:Opisthokont 1930:Hull, C. M. 1691:10481/61998 1513:: 423–452. 1387:with their 1379:rhizomorphs 1377:with their 1320:yeast name 1254:promycelium 1237:aeciospores 1196:Pucciniales 1171:Scleroderma 1131:Craterellus 1027:fruitbodies 975:bifactorial 955:monokaryons 927:Pucciniales 866:Rick (1958) 802:Nodulospora 782:Microstella 762:Hymenoporus 736:Rick (1958) 712:Cleistocybe 702:Celatogloea 666:Rick (1943) 642:Anastomyces 517:jelly fungi 513:fruitbodies 505:fruitbodies 346:mycologists 311:are called 260:earth stars 248:jelly fungi 5750:Categories 5472:Parakaryon 5434:Entoprocta 5369:Arthropoda 5364:Tardigrada 5344:Priapulida 5339:Loricifera 5324:Echinoderm 5294:Ctenophora 5266:Gnetophyta 5221:Charophyta 5211:Rhodophyta 5183:Ascomycota 5178:Zygomycota 5120:Metamonada 5095:Euglenozoa 5080:Ciliophora 4788:Balneolota 4763:Aquificota 4650:Rigifilida 4549:Acritarchs 4474:Barthelona 4465:Metamonada 4450:Euglenozoa 4318:Mesostigma 4274:sensu lato 4255:Rhodophyta 4114:Ochrophyta 4075:Placidozoa 3986:Ciliophora 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Index


Ernst Haeckel
Kunstformen der Natur
Scientific classification
Edit this classification
Eukaryota
Fungi
Dikarya
Basidiomycota
R.T.Moore
Agaricomycotina
Pucciniomycotina
Ustilaginomycotina
/bəˌsɪdi.mˈktə/
divisions
Ascomycota
Dikarya
Fungi
agarics
puffballs
stinkhorns
bracket fungi
polypores
jelly fungi
boletes
chanterelles
earth stars
smuts
bunts
rusts

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