114:
137:
4370:
4942:
4389:
4935:
683:
4949:
4957:
4382:
1908:
new taxon could potentially be placed outside
Baurusuchidae because it is not a descendant of the most recent common ancestor of baurusuchids, but would still be a baurusuchian because it is more closely related to baurusuchids than it is to other crocodyliforms. For now, however, Baurusuchidae and Baurusuchia are almost identical in scope, with Baurusuchia also including
1834:. They adopted the name Baurusuchia in a phylogenetic sense to distinguish baurusuchids from related crocodyliforms. Baurusuchia was first erected as an infraorder in 1968, but in the 2011 analysis it was found to be in an identical position to Baurusuchidae in the final tree. The only difference between Baurusuchidae and Baurusuchia is that the former is a
4376:
2757:
living crocodilians (the closest living relatives of baurusuchids), eggs undergo extrinsic degradation to allow hatchlings to easily break through their shells. The fossils indicate that baurusuchid hatchlings probably broke through thin egg shells rather than shells that had been degraded over their incubation period.
1907:
In contrast to the node-based
Baurusuchidae, the stem-based Baurusuchia does not include a common ancestor and all its descendants, but rather all forms more closely related to a specific baurusuchid than a non-baurusuchid. As a stem-based taxon, Baurusuchia is more inclusive than Baurusuchidae; a
2756:
was named on the basis of these remains. The eggs are about twice as long as they are wide and have blunt ends. At about a quarter of a millimeter in thickness, the shells are relatively thin. Some eggs may have already hatched by the time they were buried, but none show extensive degradation. In
3647:
Ismar De Souza
Carvalho; Vicente De Paula Antunes Teixeira; Mara Lúcia Da Fonseca Ferraz; Luiz Carlos Borges Ribeiro; Agustín Guillermo Martinelli; Francisco Macedo Neto; Joseph J. W. Sertich; Gabriel Cardoso Cunha; Isabella Cardoso Cunha & Patrícia Fonseca Ferraz (2011).
1450:. Peirosauridae, a family of small terrestrial crocodyliforms, had often been placed in or near Neosuchia in previous studies. The assignment of sebecids to Sebecia placed the family closer to Neosuchia than Notosuchia. In this study, baurusuchids were split up, with
2774:
Piacentini
Pinheiro, André E.; Pereira, Paulo Victor Luiz Gomes da Costa; de Souza, Rafael G.; Brum, Arthur S.; Lopes, Ricardo T.; Machado, Alessandra S.; Bergqvist, Lílian P.; Simbras, Felipe M. (2018-08-01). Piras, Paolo (ed.).
805:. Several features were used to unite these groups: a deep snout, a ziphodont dentition, a curved tooth row, an enlarged canine-like dentary tooth that fits into a deep notch in the upper jaw, and a groove on the lower jaw.
2364:. In addition to this, the study conducted a phylogenetic analysis to resolve the affinites of the new taxon and provide a reference phylogeny for the newly defined clades. The cladogram of this analysis is shown below.
843:
were later assigned to
Baurusuchidae, and phylogenetic analyses encompassing these taxa continued to find Baurusuchidae to be closely related to Sebecidae. Both families were allied with notosuchians in the larger group
1804:
deep within
Notosuchia, just as the larger group Sebecosuchia once was, while the remaining sebecosuchian genera have been placed more distantly in Metasuchia. A new baurusuchid called
787:
have deep snouts and ziphodont dentitions (teeth that are serrated and laterally compressed). Other forms were later found that had a close appearance to these two genera, among them
3298:
Gasparini, Z.B.; Chiappe, L.M.; Fernandez, M. (1991). "A new
Senonian peirosaurid (Crocodylomorpha) from Argentina and a sinopsis of the South American Cretaceous crocodilians".
3014:
Wilson, J.A.; Malkane, M.S.; Gingerich, P.D. (2001). "New crocodyliform (Reptilia, Mesoeucrocodylia) from the Upper
Cretaceous Pab Formation of Vitakri, Balochistan (Pakistan)".
5078:
812:
analyses within the past decades have supported a close relationship between the two families. Baurusuchids and sebecosuchids are both early members of the
831:). Sebecosuchians, which include both baurusuchids and sebecosuchids, were found to be closely related to notosuchians in several studies. The new genera
1423:
Later studies noted many features that distinguished baurusuchids from sebecosuchids. Sebecosuchids were often considered to be more closely related to
5132:
5065:
3596:
Ruiz, Juan V.; Bronzati, Mario; Ferreira, Gabriel S.; Martins, Kawan C.; Queiroz, Marcos V.; Langer, Max C.; Montefeltro, Felipe C. (2021-02-16).
3557:
2855:"3D skull modelling and description of a new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous (Bauru Basin) of Brazil"
3745:
5137:
4984:
1915:
based on their reference phylogenies. Other analyses however, have recovered additional taxa within
Baurusuchia outside of Baurusuchidae. (
1464:) placed as sebecians. Therefore, the family Baurusuchidae was paraphyletic. Below is a modified cladogram from Larsson and Sues (2007):
1435:
formed a clade with notosuchians to the exclusion of other ziphosuchians. This placement has been upheld by recent analyses, which place
5142:
2979:
Martinelli, A.G.; Pais, D.F. (2008). "A new baurusuchid crocodyliform (Archosauria) from the Late
Cretaceous of Patagonia (Argentina)".
848:, composed of ziphodont crocodyliforms. More recently, sebecosuchians - including baurusuchids - have been placed within Notosuchia as
5147:
1810:
was named in 2011, and a comprehensive phylogenetic analysis of baurusuchids was conducted along with its description. Montefeltro
5157:
3241:"A new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous of Brazil and the phylogeny of Baurusuchidae"
1894:
1129:
grouping, or a grouping that includes some descendants of a common ancestor but not all. Sebecids formed an assemblage of
1868:; these genera are all of its known descendants. Baurusuchia is formally defined as the most inclusive clade containing
136:
3165:
2529:
1865:
1101:
4941:
732:
4369:
3738:
3416:
2577:
4388:
3489:
3393:
2642:
2409:
4934:
3511:
Clark, J.M. (1997). "Patterns of evolution in Mesozoic Crocodyliformes". In Fraser, N.C.; Sues, H.-D. (eds.).
4948:
3602:(Notosuchia, Sphagesauridae) from the Late Cretaceous of Brazil and the evolutionary history of Sphagesauria"
3476:
Turner, A.H.; Calvo, J.O. (2005). "A new sebecosuchian crocodyliform from the Late Cretaceous of Patagonia".
1133:
sebecosuchians, while baurusuchids remained a valid grouping of derived sebecosuchians. Below is a modified
3688:"Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil"
5005:
2553:
1100:
was established to include baurusuchids and sebecids. Phylogenetically, Baurusuchoidea was defined as the
4956:
1431:, while baurusuchids were thought to be a more distantly related clade. In a 1999 phylogenetic analysis,
5152:
5104:
3830:
2484:
2385:
1963:
1848:
1130:
660:
547:
3575:
113:
5052:
4381:
3731:
3699:
3686:
Oliveira, C.E.M.; Santucci, R.M.; Andrade, M.B.; Fulfaro, V.J.; Basílo, J.A.F.; Benton, M.J. (2011).
3435:
3350:
3307:
3252:
3184:
2988:
2926:
2792:
2735:
2625:
2601:
2467:
2443:
1900:
1888:
1870:
1859:
1852:
762:
750:
724:
558:
551:
304:
284:
3654:
gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil"
3081:
gen. et sp. nov., a new Late Cretaceous baurusuchid (Crocodyliformes) from the Bauru Basin, Brazil"
2912:"Tetrapods from the Upper Cretaceous (Turonian-Maastrichtian) Bauru Group of Brazil: a reappraisal"
1882:
1876:
571:
5043:
4684:
3629:
3493:
3397:
2429:
2361:
2105:
1935:
1112:
and all of its descendants while Baurusuchidae was defined as the most recent common ancestor of
579:
564:
460:
131:
5109:
4375:
5091:
4244:
3974:
3621:
3451:
3280:
3200:
2892:
2874:
2828:
2810:
5096:
5083:
3333:
Ortega, F.; Buscaloni, A.D; Gasaparini, Z. (1996). "Reinterpretation and new denomination of
4828:
3790:
3707:
3668:
3613:
3565:
3535:
3485:
3443:
3389:
3358:
3315:
3270:
3260:
3192:
3099:
3055:
2996:
2934:
2882:
2866:
2818:
2800:
1839:
1835:
514:
510:
4887:
4862:
4700:
4673:
4618:
4593:
4586:
4540:
4493:
4467:
4279:
4251:
3990:
2175:
2120:
2084:
2055:
1806:
1226:
1118:
795:
789:
688:
641:
635:
629:
599:
518:
480:
448:
397:
387:
345:
230:
217:
3380:(Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina)".
3703:
3649:
3439:
3354:
3311:
3256:
3188:
3095:
3092:
2992:
2930:
2796:
4903:
4853:
4815:
4761:
4739:
4625:
4563:
4510:
4237:
4161:
4140:
4038:
3955:
3275:
3240:
3121:
Rio de Janeiro, Departamento Nacional da Produção Mineral, Notas preliminares e estudos
2887:
2823:
2776:
1097:
747:
366:
323:
3447:
3362:
3196:
2911:
5126:
4896:
4869:
4835:
4796:
4781:
4754:
4747:
4693:
4659:
4652:
4643:
4570:
4482:
4452:
4445:
4314:
4230:
4223:
4207:
4133:
4105:
3712:
3687:
3633:
3597:
3526:
Pol, D. (1999). "Basal Mesoeucrocodylian relationships: new clues to old conflicts".
3060:
3039:
2854:
2752:
2515:
2357:
2351:
2334:
2160:
2031:
1939:
1923:
1719:
1547:
1447:
758:
705:
623:
575:
522:
470:
428:
418:
408:
334:
71:
3497:
3401:
682:
4876:
4772:
4726:
4517:
4460:
4286:
4258:
4216:
4175:
4168:
4152:
4125:
4118:
4080:
4017:
3815:
3539:
3319:
1514:
1211:
1175:
1126:
902:
849:
809:
766:
723:, was assigned to the family in May, 2011. It is known from the Turonian-Santonian
262:
204:
3617:
2938:
3265:
2805:
5037:
4789:
4707:
4666:
4578:
4550:
4437:
4307:
4293:
4196:
4094:
4045:
4031:
4024:
4010:
3983:
3941:
3910:
3892:
3879:
3842:
3754:
2245:
2228:
2204:
2189:
2137:
1694:
1578:
1428:
1337:
1308:
1291:
1250:
1023:
956:
926:
845:
839:
833:
828:
801:
647:
621:, five other South American crocodyliforms have been assigned to Baurusuchidae:
602:
490:
438:
355:
123:
46:
3000:
704:
is the youngest genus that has been assigned to Baurusuchidae, and is from the
675:
are known only from fragmentary remains. Both genera are from the Santonian of
4842:
4804:
4524:
4503:
4475:
4419:
4406:
4337:
4300:
4272:
4265:
4189:
4073:
4066:
4059:
3963:
3799:
3784:
3672:
3142:, representative of a peculiar suborder of fossil Crocodilia from Patagonia".
3104:
3076:
2781:
Roxo, 1936: Historical approach, systematic, and description by new materials"
2007:
1992:
1917:
1497:
1483:
1471:
1198:
1145:
878:
863:
820:
816:
246:
91:
56:
5028:
3625:
3455:
3204:
2878:
2814:
4632:
4182:
4052:
4001:
3948:
2747:
1910:
1662:
1635:
1607:
1460:
1424:
1354:
1159:
1134:
824:
770:
676:
614:
538:
526:
376:
191:
148:
96:
40:
28:
3570:
3284:
3208:
2960:
from the red sandstones of the Territory of Neuquén (Argentine Republic)".
2896:
2832:
2356:
and proposed formal definitions for the clades Baurusuchia, Baurusuchidae,
2853:
Darlim, Gustavo; Montefeltro, Felipe C.; Langer, Max C. (September 2021).
712:. It was named in 2001 but has since been reassigned as a basal member of
5022:
3778:
3376:
Ortega, F. Z.; Buscalioni, A. D.; Calvo, J. O. (2000). "A new species of
709:
610:
534:
507:
168:
86:
81:
66:
61:
51:
3421:, a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil"
5070:
4087:
3927:
1677:
1620:
1443:
1267:
986:
775:
713:
656:
178:
101:
76:
3459:
2870:
1814:(2011) found Baurusuchidae to be a monophyletic group with the genera
3772:
3016:
Contributions from the Museum of Paleontology, University of Michigan
2910:
Candeiro, C.R.A.; Martinelli, A.G.; Avilla, L.S.; Rich, T.H. (2006).
2739:
1970:
may also be associated with a juvenile individual. The material that
744:
606:
530:
158:
32:
4999:
3119:
Price, L.I. (1945). "A new reptile from the Cretaceous of Brazil".
3044:(Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco"
1454:
placed as a more basal metasuchian and the remaining baurusuchids (
3170:, a new Crocodylomorpha from the Bauru Basin (Cretaceous), Brazil"
1843:
813:
681:
542:
521:. It is a group of terrestrial hypercarnivorous crocodilians from
2734:
In 2011, fossilized eggs were described from the Late Cretaceous
5057:
852:
members of the clade. Below is a modified cladogram from Ortega
5003:
4973:
4404:
3877:
3765:
3727:
3723:
1962:
is not a member of either of these subfamilies, but the most
827:(larger, often semiaquatic crocodyliforms, including living
594:
was the first, being the namesake of the family. Remains of
3490:
10.1671/0272-4634(2005)025[0087:ANSCFT]2.0.CO;2
3394:
10.1671/0272-4634(2000)020[0057:ANSOAC]2.0.CO;2
663:, although it was only recently assigned to Baurusuchidae.
3239:
Montefeltro, F.C.; Larsson, H.C.E.; Langer, M.C. (2011).
3515:. Cambridge: Cambridge University Press. pp. 84–97.
3513:
In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods
2952:
Woodward, A.S. (1896). ". On two Mesozoic crocodilians,
2742:
that may have been laid by a baurusuchid, most probably
1974:
is based on has been suggested to be a juvenile form of
1934:(2011) also divided Baurusuchidae into two subfamilies,
727:
of the Bauru Basin of Brazil. Soon after, the new genus
1966:
baurusuchid. Many of the unique features that separate
3415:
Carvalho, I.S.; Ribeiro, L.C.B.; Avilla, L.S. (2004).
3040:"Cranial osteology and phylogenetic relationships of
3164:
Carvalho, I.S.; Campos, A.C.A.; Nobre, P.H. (2005).
590:
Several genera have been assigned to Baurusuchidae.
486:
476:
466:
456:
444:
434:
424:
414:
404:
393:
383:
372:
362:
351:
341:
330:
319:
5012:
4886:
4852:
4814:
4771:
4725:
4683:
4642:
4604:
4549:
4538:
4492:
4417:
4336:
4206:
4151:
4104:
4000:
3973:
3925:
3890:
3841:
3814:
3144:Bulletin of the American Museum of Natural History
1842:. Baurusuchidae is defined as the least inclusive
655:was the first known baurusuchid, named in 1896 by
537:. Baurusuchidae has been, in accordance with the
1125:In a 2005 analysis, Sebecidae was found to be a
856:(2000) placing baurusuchids within Notosuchia:
3739:
2777:"Reassessment of the enigmatic crocodyliform
769:, which united Baurusuchidae with the family
8:
2962:Anales del Museo de la Plata (Paleontología)
541:, officially defined as the least inclusive
3558:"Cretaceous crocodyliforms from the Sahara"
1446:was erected. Sebecia included sebecids and
731:was named from the Campanian–Maastrichtian
5000:
4970:
4820:
4731:
4610:
4555:
4546:
4429:
4425:
4414:
4401:
4333:
4110:
3933:
3902:
3898:
3887:
3874:
3820:
3811:
3762:
3746:
3732:
3724:
112:
20:
3711:
3569:
3556:Sereno, P. C.; Larsson, H. C. E. (2009).
3274:
3264:
3103:
3059:
3048:Zoological Journal of the Linnean Society
2886:
2822:
2804:
700:, was assigned to Baurusuchidae in 2001.
3234:
3232:
3230:
3228:
3159:
3157:
2342:, and was not included in the analysis.
1864:As in all node-based clades, there is a
823:(mainly terrestrial crocodyliforms) and
3551:
3549:
3038:Larsson, H. C. E.; Sues, H.-D. (2007).
3033:
3031:
3029:
2766:
3337:(Middle Eocene; Issel, France) as cf.
1981:Below is a cladogram from Montefeltro
1978:, and the two taxa may be synonymous.
743:The family Baurusuchidae was named by
2338:, was named just a few months before
563:Baurusuchids have been placed in the
7:
3075:Carvalho, I.S.; et al. (2011).
2848:
2846:
2844:
2842:
2349:(2021) described a new baurusuchid,
3606:Journal of Systematic Palaeontology
2617:
2593:
2569:
2545:
2521:
2459:
2435:
2425:
2401:
2377:
2367:
2220:
2196:
2167:
2112:
2100:
2076:
2047:
2023:
1999:
1988:
1669:
1657:
1627:
1600:
1570:
1563:
1539:
1489:
1478:
1467:
1329:
1283:
1242:
1218:
1191:
1152:
1140:
1002:
978:
948:
918:
894:
870:
859:
3528:Journal of Vertebrate Paleontology
3478:Journal of Vertebrate Paleontology
3382:Journal of Vertebrate Paleontology
3300:Journal of Vertebrate Paleontology
14:
3419:Uberabasuchus terrificus sp. nov.
4955:
4947:
4940:
4933:
4387:
4380:
4374:
4368:
3713:10.1111/j.1475-4983.2010.01028.x
3341:(Crocodylomorpha, Metasuchia)".
3061:10.1111/j.1096-3642.2007.00271.x
1800:More recent studies have nested
135:
5133:Late Cretaceous crocodylomorphs
1427:, a group that includes modern
819:, which includes the subgroups
3540:10.1080/02724634.1999.10011202
3320:10.1080/02724634.1991.10011401
2332:A sixth genus of baurusuchid,
1942:. Pissarrachampsinae includes
1137:from Turner and Calvo (2005):
733:Vale do Rio do Peixe Formation
1:
3618:10.1080/14772019.2021.1888815
3448:10.1016/S1342-937X(05)71079-0
3363:10.1016/S0016-6995(96)80037-4
3197:10.1016/S1342-937X(05)70259-8
2958:Cynodontosuchus (genus novum)
2939:10.1016/j.cretres.2006.05.002
1950:while Baurusuchinae includes
1895:Montealtosuchus arrudacamposi
3266:10.1371/journal.pone.0021916
2806:10.1371/journal.pone.0199984
1442:In 2007, a new clade called
1122:and all of its descendants.
5138:Terrestrial crocodylomorphs
4985:Terrestrial crocodylomorphs
2530:Aphaurosuchus escharafacies
1866:most recent common ancestor
1102:most recent common ancestor
735:, also in the Bauru Basin.
5174:
5143:Turonian first appearances
3335:Atacisaurus crassiproratus
3001:10.1016/j.crpv.2008.05.002
773:(represented by the genus
16:Extinct family of reptiles
5148:Maastrichtian extinctions
4980:
4969:
4931:
4823:
4734:
4613:
4558:
4432:
4428:
4413:
4400:
4366:
4332:
4113:
3936:
3905:
3901:
3886:
3873:
3823:
3810:
3761:
3673:10.11646/zootaxa.2871.1.2
3105:10.11646/zootaxa.2871.1.2
2779:"Goniopholis" paulistanus
2639:
2622:
2615:
2598:
2591:
2578:Stratiotosuchus maxhechti
2574:
2567:
2550:
2543:
2526:
2519:
2481:
2464:
2457:
2440:
2433:
2423:
2406:
2399:
2382:
2375:
2242:
2225:
2218:
2201:
2194:
2172:
2165:
2134:
2117:
2110:
2098:
2081:
2074:
2052:
2045:
2028:
2021:
2004:
1997:
1717:
1691:
1674:
1667:
1655:
1632:
1625:
1605:
1598:
1575:
1568:
1561:
1544:
1537:
1511:
1494:
1487:
1476:
1351:
1334:
1327:
1305:
1288:
1281:
1264:
1247:
1240:
1223:
1216:
1196:
1189:
1172:
1157:
1150:
1096:In 2004, the superfamily
1020:
1007:
1000:
983:
976:
953:
946:
923:
916:
899:
892:
875:
868:
598:have been found from the
316:
311:
132:Scientific classification
130:
120:
111:
23:
3168:Baurusuchus salgadoensis
2954:Notosuchus (genus novum)
2643:Baurusuchus salgadoensis
2410:Gondwanasuchus scabrosus
124:Baurusuchus salgadoensis
3138:Colbert, E.H. (1946). "
667:was described in 2008.
617:in age. In addition to
5158:Pseudosuchian families
3571:10.3897/zookeys.28.325
3534:(3, Supplement): 69A.
2981:Comptes Rendus Palevol
2753:Bauruoolithus fragilis
2554:Aplestosuchus sordidus
693:
517:that lived during the
5105:Paleobiology Database
2485:Wargosuchus australis
2386:Cynodontosuchus rothi
1849:Cynodontosuchus rothi
685:
661:Arthur Smith Woodward
609:in deposits that are
548:Cynodontosuchus rothi
3652:Campinasuchus dinizi
3079:Campinasuchus dinizi
3042:Hamadasuchus rebouli
2736:Adamantina Formation
2626:Baurusuchus pachecoi
2602:Baurusuchus albertoi
2468:Pissarrachampsa sera
2444:Campinasuchus dinizi
1901:Crocodylus niloticus
1889:Araripesuchus gomesi
1871:Baurusuchus pachecoi
1860:Baurusuchus pachecoi
1853:Pissarrachampsa sera
1838:and the latter is a
763:Edwin Harris Colbert
751:Llewellyn Ivor Price
725:Adamantina Formation
574:have been proposed:
559:Baurusuchus pachecoi
552:Pissarrachampsa sera
3704:2011Palgy..54..309O
3440:2004GondR...7..975C
3355:1996Geobi..29..353O
3312:1991JVPal..11..316G
3257:2011PLoSO...621916M
3189:2005GondR...8...11C
2993:2008CRPal...7..371M
2931:2006CrRes..27..923C
2919:Cretaceous Research
2797:2018PLoSO..1399984P
1883:Sphagesaurus huenei
1877:Sebecus icaeorhinus
1439:within Notosuchia.
1009:Itaborai crocodile
753:in 1945 to include
4975:Related categories
4685:Pissarrachampsinae
3598:"A new species of
2859:Journal of Anatomy
2430:Pissarrachampsinae
2362:Pissarrachampsinae
2106:Pissarrachampsinae
1936:Pissarrachampsinae
1787:
1083:
765:erected the group
694:
580:Pissarrachampsinae
515:mesoeucrocodylians
461:Pissarrachampsinae
5120:
5119:
5092:Open Tree of Life
5006:Taxon identifiers
4997:
4996:
4993:
4992:
4965:
4964:
4929:
4928:
4925:
4924:
4921:
4920:
4917:
4916:
4913:
4912:
4721:
4720:
4717:
4716:
4534:
4533:
4396:
4395:
4364:
4363:
4360:
4359:
4356:
4355:
4328:
4327:
4324:
4323:
4245:Colhuehuapisuchus
3975:Mahajangasuchidae
3921:
3920:
3869:
3868:
3865:
3864:
3861:
3860:
3428:Gondwana Research
3177:Gondwana Research
2871:10.1111/joa.13442
2726:
2725:
2717:
2716:
2708:
2707:
2699:
2698:
2690:
2689:
2681:
2680:
2672:
2671:
2663:
2662:
2654:
2653:
2505:
2504:
2496:
2495:
2329:
2328:
2320:
2319:
2311:
2310:
2302:
2301:
2293:
2292:
2284:
2283:
2275:
2274:
2266:
2265:
2257:
2256:
2149:
2148:
1797:
1796:
1784:
1783:
1775:
1774:
1766:
1765:
1757:
1756:
1748:
1747:
1739:
1738:
1730:
1729:
1706:
1705:
1526:
1525:
1420:
1419:
1411:
1410:
1402:
1401:
1393:
1392:
1384:
1383:
1375:
1374:
1366:
1365:
1093:
1092:
1080:
1079:
1071:
1070:
1062:
1061:
1053:
1052:
1044:
1043:
1035:
1034:
501:
500:
307:
287:
5165:
5113:
5112:
5100:
5099:
5087:
5086:
5074:
5073:
5061:
5060:
5048:
5047:
5046:
5033:
5032:
5031:
5001:
4971:
4960:
4959:
4952:
4951:
4945:
4944:
4937:
4829:Adamantinasuchus
4821:
4732:
4611:
4556:
4547:
4430:
4426:
4415:
4402:
4391:
4385:
4384:
4378:
4372:
4334:
4111:
3934:
3903:
3899:
3888:
3875:
3821:
3812:
3805:
3804:
3791:Mesoeucrocodylia
3763:
3748:
3741:
3734:
3725:
3718:
3717:
3715:
3683:
3677:
3676:
3658:
3644:
3638:
3637:
3593:
3587:
3586:
3584:
3583:
3574:. Archived from
3573:
3553:
3544:
3543:
3523:
3517:
3516:
3508:
3502:
3501:
3473:
3467:
3466:
3464:
3458:. Archived from
3425:
3412:
3406:
3405:
3373:
3367:
3366:
3330:
3324:
3323:
3295:
3289:
3288:
3278:
3268:
3236:
3223:
3222:
3220:
3219:
3213:
3207:. Archived from
3174:
3161:
3152:
3151:
3135:
3129:
3128:
3116:
3110:
3109:
3107:
3072:
3066:
3065:
3063:
3035:
3024:
3023:
3011:
3005:
3004:
2976:
2970:
2969:
2949:
2943:
2942:
2916:
2907:
2901:
2900:
2890:
2850:
2837:
2836:
2826:
2808:
2771:
2618:
2594:
2570:
2546:
2522:
2460:
2436:
2426:
2402:
2378:
2368:
2221:
2197:
2168:
2113:
2101:
2077:
2048:
2024:
2000:
1989:
1840:stem-based taxon
1836:node-based taxon
1788:
1670:
1658:
1642:
1628:
1601:
1585:
1571:
1564:
1540:
1490:
1479:
1468:
1330:
1284:
1243:
1219:
1192:
1153:
1141:
1084:
1003:
979:
963:
949:
933:
919:
895:
871:
860:
696:A fourth genus,
488:
478:
468:
458:
446:
436:
426:
416:
406:
395:
385:
374:
364:
353:
343:
332:
321:
302:
297:
282:
277:
261:
245:
140:
139:
116:
106:
43:
27:Temporal range:
21:
5173:
5172:
5168:
5167:
5166:
5164:
5163:
5162:
5123:
5122:
5121:
5116:
5108:
5103:
5095:
5090:
5082:
5077:
5069:
5064:
5056:
5051:
5042:
5041:
5036:
5027:
5026:
5021:
5008:
4998:
4989:
4976:
4961:
4954:
4946:
4939:
4909:
4888:Caipirasuchinae
4882:
4863:Armadillosuchus
4848:
4810:
4767:
4713:
4701:Pissarrachampsa
4679:
4674:Stratiotosuchus
4638:
4619:Cynodontosuchus
4600:
4594:Razanandrongobe
4587:Pehuenchesuchus
4543:
4541:Xenodontosuchia
4530:
4494:Candidodontidae
4488:
4468:Razanandrongobe
4422:
4409:
4392:
4379:
4352:
4320:
4280:Montealtosuchus
4252:Gasparinisuchus
4202:
4147:
4100:
3996:
3991:Mahajangasuchus
3969:
3930:
3917:
3895:
3882:
3857:
3837:
3806:
3768:
3767:
3757:
3752:
3722:
3721:
3685:
3684:
3680:
3656:
3646:
3645:
3641:
3595:
3594:
3590:
3581:
3579:
3555:
3554:
3547:
3525:
3524:
3520:
3510:
3509:
3505:
3475:
3474:
3470:
3462:
3434:(4): 975–1002.
3423:
3414:
3413:
3409:
3375:
3374:
3370:
3332:
3331:
3327:
3297:
3296:
3292:
3238:
3237:
3226:
3217:
3215:
3211:
3172:
3163:
3162:
3155:
3137:
3136:
3132:
3118:
3117:
3113:
3091:(2871): 19–42.
3074:
3073:
3069:
3037:
3036:
3027:
3013:
3012:
3008:
2978:
2977:
2973:
2951:
2950:
2946:
2914:
2909:
2908:
2904:
2852:
2851:
2840:
2791:(8): e0199984.
2773:
2772:
2768:
2763:
2732:
2727:
2718:
2709:
2700:
2691:
2682:
2673:
2664:
2655:
2506:
2497:
2340:Pissarrachampsa
2330:
2321:
2312:
2303:
2294:
2285:
2276:
2267:
2258:
2229:B. salgadoensis
2176:Stratiotosuchus
2150:
2121:Pissarrachampsa
2085:Cynodontosuchus
2056:Armadillosuchus
1972:Cynodontosuchus
1968:Cynodontosuchus
1960:Cynodontosuchus
1952:Stratiotosuchus
1944:Pissarrachampsa
1828:Stratiotosuchus
1824:Pissarrachampsa
1820:Cynodontosuchus
1807:Pissarrachampsa
1798:
1786:
1785:
1776:
1767:
1758:
1749:
1740:
1731:
1707:
1636:
1579:
1527:
1421:
1412:
1403:
1394:
1385:
1376:
1367:
1227:Pehuenchesuchus
1119:Stratiotosuchus
1094:
1082:
1081:
1072:
1063:
1054:
1045:
1036:
957:
927:
796:Stratiotosuchus
790:Cynodontosuchus
761:paleontologist
741:
729:Pissarrachampsa
698:Stratiotosuchus
689:Pissarrachampsa
669:Cynodontosuchus
659:paleontologist
653:Cynodontosuchus
642:Stratiotosuchus
636:Pissarrachampsa
630:Cynodontosuchus
600:Late Cretaceous
588:
533:) and possibly
519:Late Cretaceous
481:Pissarrachampsa
449:Stratiotosuchus
398:Razanandrongobe
388:Pehuenchesuchus
346:Cynodontosuchus
301:
295:
281:
275:
259:
243:
231:Crocodyliformes
218:Crocodylomorpha
134:
107:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
38:
37:
36:
25:
17:
12:
11:
5:
5171:
5169:
5161:
5160:
5155:
5150:
5145:
5140:
5135:
5125:
5124:
5118:
5117:
5115:
5114:
5101:
5088:
5075:
5062:
5049:
5034:
5018:
5016:
5010:
5009:
5004:
4995:
4994:
4991:
4990:
4988:
4987:
4981:
4978:
4977:
4974:
4967:
4966:
4963:
4962:
4932:
4930:
4927:
4926:
4923:
4922:
4919:
4918:
4915:
4914:
4911:
4910:
4908:
4907:
4904:Morrinhosuchus
4900:
4892:
4890:
4884:
4883:
4881:
4880:
4873:
4866:
4858:
4856:
4854:Sphagesaurinae
4850:
4849:
4847:
4846:
4839:
4832:
4824:
4818:
4816:Sphagesauridae
4812:
4811:
4809:
4808:
4801:
4793:
4786:
4777:
4775:
4769:
4768:
4766:
4765:
4762:Eptalofosuchus
4758:
4751:
4744:
4740:Brasileosaurus
4735:
4729:
4723:
4722:
4719:
4718:
4715:
4714:
4712:
4711:
4704:
4697:
4689:
4687:
4681:
4680:
4678:
4677:
4670:
4663:
4656:
4648:
4646:
4640:
4639:
4637:
4636:
4629:
4626:Gondwanasuchus
4622:
4614:
4608:
4602:
4601:
4599:
4598:
4590:
4583:
4575:
4567:
4564:Chimaerasuchus
4559:
4553:
4544:
4539:
4536:
4535:
4532:
4531:
4529:
4528:
4521:
4514:
4511:Lavocatchampsa
4507:
4499:
4497:
4490:
4489:
4487:
4486:
4479:
4472:
4464:
4457:
4449:
4442:
4433:
4423:
4418:
4411:
4410:
4405:
4398:
4397:
4394:
4393:
4367:
4365:
4362:
4361:
4358:
4357:
4354:
4353:
4351:
4350:
4349:
4348:
4342:
4340:
4330:
4329:
4326:
4325:
4322:
4321:
4319:
4318:
4311:
4304:
4297:
4290:
4283:
4276:
4269:
4262:
4255:
4248:
4241:
4238:Bayomesasuchus
4234:
4227:
4220:
4212:
4210:
4204:
4203:
4201:
4200:
4193:
4186:
4179:
4172:
4165:
4162:Barreirosuchus
4157:
4155:
4149:
4148:
4146:
4145:
4141:Stolokrosuchus
4137:
4130:
4122:
4114:
4108:
4102:
4101:
4099:
4098:
4091:
4084:
4077:
4070:
4063:
4056:
4049:
4042:
4039:Dentaneosuchus
4035:
4028:
4021:
4014:
4006:
4004:
3998:
3997:
3995:
3994:
3987:
3979:
3977:
3971:
3970:
3968:
3967:
3960:
3956:Trematochampsa
3952:
3945:
3937:
3931:
3926:
3923:
3922:
3919:
3918:
3916:
3915:
3906:
3896:
3891:
3884:
3883:
3878:
3871:
3870:
3867:
3866:
3863:
3862:
3859:
3858:
3856:
3855:
3854:
3853:
3847:
3845:
3839:
3838:
3836:
3835:
3834:
3833:
3824:
3818:
3808:
3807:
3803:
3802:
3793:
3787:
3781:
3775:
3766:
3759:
3758:
3753:
3751:
3750:
3743:
3736:
3728:
3720:
3719:
3698:(2): 309–321.
3678:
3639:
3612:(4): 265–287.
3588:
3545:
3518:
3503:
3468:
3465:on 2011-07-06.
3407:
3368:
3349:(3): 353–364.
3325:
3306:(3): 316–333.
3290:
3224:
3153:
3130:
3111:
3067:
3054:(4): 533–567.
3025:
3022:(12): 321–336.
3006:
2987:(6): 371–381.
2971:
2944:
2925:(6): 923–946.
2902:
2865:(3): 622–662.
2838:
2765:
2764:
2762:
2759:
2731:
2728:
2724:
2723:
2720:
2719:
2715:
2714:
2711:
2710:
2706:
2705:
2702:
2701:
2697:
2696:
2693:
2692:
2688:
2687:
2684:
2683:
2679:
2678:
2675:
2674:
2670:
2669:
2666:
2665:
2661:
2660:
2657:
2656:
2652:
2651:
2648:
2647:
2638:
2635:
2634:
2631:
2630:
2621:
2616:
2614:
2611:
2610:
2607:
2606:
2597:
2592:
2590:
2587:
2586:
2583:
2582:
2573:
2568:
2566:
2563:
2562:
2559:
2558:
2549:
2544:
2542:
2539:
2538:
2535:
2534:
2525:
2520:
2518:
2512:
2511:
2508:
2507:
2503:
2502:
2499:
2498:
2494:
2493:
2490:
2489:
2480:
2477:
2476:
2473:
2472:
2463:
2458:
2456:
2453:
2452:
2449:
2448:
2439:
2434:
2432:
2424:
2422:
2419:
2418:
2415:
2414:
2405:
2400:
2398:
2395:
2394:
2391:
2390:
2381:
2376:
2374:
2366:
2327:
2326:
2323:
2322:
2318:
2317:
2314:
2313:
2309:
2308:
2305:
2304:
2300:
2299:
2296:
2295:
2291:
2290:
2287:
2286:
2282:
2281:
2278:
2277:
2273:
2272:
2269:
2268:
2264:
2263:
2260:
2259:
2255:
2254:
2251:
2250:
2241:
2238:
2237:
2234:
2233:
2224:
2219:
2217:
2214:
2213:
2210:
2209:
2200:
2195:
2193:
2185:
2184:
2181:
2180:
2171:
2166:
2164:
2156:
2155:
2152:
2151:
2147:
2146:
2143:
2142:
2133:
2130:
2129:
2126:
2125:
2116:
2111:
2109:
2099:
2097:
2094:
2093:
2090:
2089:
2080:
2075:
2073:
2065:
2064:
2061:
2060:
2051:
2046:
2044:
2041:
2040:
2037:
2036:
2027:
2022:
2020:
2017:
2016:
2013:
2012:
2003:
1998:
1996:
1987:
1795:
1794:
1791:
1790:
1782:
1781:
1778:
1777:
1773:
1772:
1769:
1768:
1764:
1763:
1760:
1759:
1755:
1754:
1751:
1750:
1746:
1745:
1742:
1741:
1737:
1736:
1733:
1732:
1728:
1727:
1724:
1723:
1716:
1713:
1712:
1709:
1708:
1704:
1703:
1700:
1699:
1690:
1687:
1686:
1683:
1682:
1673:
1668:
1666:
1656:
1654:
1651:
1650:
1647:
1646:
1631:
1626:
1624:
1616:
1615:
1612:
1611:
1604:
1599:
1597:
1594:
1593:
1590:
1589:
1574:
1569:
1567:
1562:
1560:
1557:
1556:
1553:
1552:
1543:
1538:
1536:
1533:
1532:
1529:
1528:
1524:
1523:
1520:
1519:
1510:
1507:
1506:
1503:
1502:
1493:
1488:
1486:
1477:
1475:
1466:
1418:
1417:
1414:
1413:
1409:
1408:
1405:
1404:
1400:
1399:
1396:
1395:
1391:
1390:
1387:
1386:
1382:
1381:
1378:
1377:
1373:
1372:
1369:
1368:
1364:
1363:
1360:
1359:
1350:
1347:
1346:
1343:
1342:
1333:
1328:
1326:
1318:
1317:
1314:
1313:
1304:
1301:
1300:
1297:
1296:
1287:
1282:
1280:
1277:
1276:
1273:
1272:
1263:
1260:
1259:
1256:
1255:
1246:
1241:
1239:
1236:
1235:
1232:
1231:
1222:
1217:
1215:
1207:
1206:
1203:
1202:
1195:
1190:
1188:
1185:
1184:
1181:
1180:
1171:
1168:
1167:
1164:
1163:
1156:
1151:
1149:
1139:
1098:Baurusuchoidea
1091:
1090:
1087:
1086:
1078:
1077:
1074:
1073:
1069:
1068:
1065:
1064:
1060:
1059:
1056:
1055:
1051:
1050:
1047:
1046:
1042:
1041:
1038:
1037:
1033:
1032:
1029:
1028:
1019:
1016:
1015:
1012:
1011:
1006:
1001:
999:
996:
995:
992:
991:
982:
977:
975:
972:
971:
968:
967:
952:
947:
945:
942:
941:
938:
937:
922:
917:
915:
912:
911:
908:
907:
898:
893:
891:
888:
887:
884:
883:
874:
869:
867:
858:
748:paleontologist
740:
737:
587:
584:
499:
498:
497:
496:
495:
494:
484:
474:
454:
453:
452:
442:
432:
422:
402:
391:
381:
370:
367:Gondwanasuchus
360:
349:
339:
328:
324:Chimaerasuchus
314:
313:
309:
308:
293:
289:
288:
273:
266:
265:
257:
250:
249:
241:
234:
233:
228:
221:
220:
215:
208:
207:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
146:
142:
141:
128:
127:
118:
117:
109:
108:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
44:
26:
15:
13:
10:
9:
6:
4:
3:
2:
5170:
5159:
5156:
5154:
5151:
5149:
5146:
5144:
5141:
5139:
5136:
5134:
5131:
5130:
5128:
5111:
5106:
5102:
5098:
5093:
5089:
5085:
5080:
5076:
5072:
5067:
5063:
5059:
5054:
5050:
5045:
5044:Baurusuchidae
5039:
5035:
5030:
5024:
5020:
5019:
5017:
5015:
5014:Baurusuchidae
5011:
5007:
5002:
4986:
4983:
4982:
4979:
4972:
4968:
4958:
4953:
4950:
4943:
4936:
4906:
4905:
4901:
4899:
4898:
4897:Caipirasuchus
4894:
4893:
4891:
4889:
4885:
4879:
4878:
4874:
4872:
4871:
4870:Caryonosuchus
4867:
4865:
4864:
4860:
4859:
4857:
4855:
4851:
4845:
4844:
4840:
4838:
4837:
4836:Labidiosuchus
4833:
4831:
4830:
4826:
4825:
4822:
4819:
4817:
4813:
4807:
4806:
4802:
4799:
4798:
4797:Mariliasuchus
4794:
4792:
4791:
4787:
4784:
4783:
4782:Coronelsuchus
4779:
4778:
4776:
4774:
4770:
4764:
4763:
4759:
4757:
4756:
4755:Coronelsuchus
4752:
4750:
4749:
4748:Coringasuchus
4745:
4742:
4741:
4737:
4736:
4733:
4730:
4728:
4724:
4710:
4709:
4705:
4703:
4702:
4698:
4696:
4695:
4694:Campinasuchus
4691:
4690:
4688:
4686:
4682:
4676:
4675:
4671:
4669:
4668:
4664:
4662:
4661:
4660:Aplestosuchus
4657:
4655:
4654:
4653:Aphaurosuchus
4650:
4649:
4647:
4645:
4644:Baurusuchinae
4641:
4635:
4634:
4630:
4628:
4627:
4623:
4621:
4620:
4616:
4615:
4612:
4609:
4607:
4606:Baurusuchidae
4603:
4596:
4595:
4591:
4589:
4588:
4584:
4581:
4580:
4576:
4573:
4572:
4571:Comahuesuchus
4568:
4566:
4565:
4561:
4560:
4557:
4554:
4552:
4548:
4545:
4542:
4537:
4527:
4526:
4522:
4520:
4519:
4515:
4513:
4512:
4508:
4506:
4505:
4501:
4500:
4498:
4495:
4491:
4485:
4484:
4483:Uruguaysuchus
4480:
4478:
4477:
4473:
4470:
4469:
4465:
4463:
4462:
4458:
4455:
4454:
4453:Comahuesuchus
4450:
4448:
4447:
4446:Araripesuchus
4443:
4440:
4439:
4435:
4434:
4431:
4427:
4424:
4421:
4416:
4412:
4408:
4403:
4399:
4390:
4386:
4383:
4377:
4371:
4346:
4345:
4344:
4343:
4341:
4339:
4335:
4331:
4317:
4316:
4315:Uberabasuchus
4312:
4310:
4309:
4305:
4303:
4302:
4298:
4296:
4295:
4291:
4289:
4288:
4284:
4282:
4281:
4277:
4275:
4274:
4270:
4268:
4267:
4263:
4261:
4260:
4256:
4254:
4253:
4249:
4247:
4246:
4242:
4240:
4239:
4235:
4233:
4232:
4231:Barrosasuchus
4228:
4226:
4225:
4224:Barcinosuchus
4221:
4219:
4218:
4214:
4213:
4211:
4209:
4208:Peirosaurinae
4205:
4199:
4198:
4194:
4192:
4191:
4187:
4185:
4184:
4180:
4178:
4177:
4173:
4171:
4170:
4166:
4164:
4163:
4159:
4158:
4156:
4154:
4150:
4143:
4142:
4138:
4136:
4135:
4134:Miadanasuchus
4131:
4128:
4127:
4123:
4121:
4120:
4116:
4115:
4112:
4109:
4107:
4106:Peirosauridae
4103:
4097:
4096:
4092:
4090:
4089:
4085:
4083:
4082:
4078:
4076:
4075:
4071:
4069:
4068:
4064:
4062:
4061:
4057:
4055:
4054:
4050:
4048:
4047:
4043:
4041:
4040:
4036:
4034:
4033:
4029:
4027:
4026:
4022:
4020:
4019:
4015:
4013:
4012:
4008:
4007:
4005:
4003:
3999:
3993:
3992:
3988:
3986:
3985:
3981:
3980:
3978:
3976:
3972:
3966:
3965:
3961:
3958:
3957:
3953:
3951:
3950:
3946:
3944:
3943:
3939:
3938:
3935:
3932:
3929:
3924:
3913:
3912:
3908:
3907:
3904:
3900:
3897:
3894:
3889:
3885:
3881:
3876:
3872:
3851:
3850:
3849:
3848:
3846:
3844:
3840:
3832:
3828:
3827:
3826:
3825:
3822:
3819:
3817:
3813:
3809:
3801:
3797:
3794:
3792:
3788:
3786:
3782:
3780:
3776:
3774:
3770:
3769:
3764:
3760:
3756:
3749:
3744:
3742:
3737:
3735:
3730:
3729:
3726:
3714:
3709:
3705:
3701:
3697:
3693:
3692:Palaeontology
3689:
3682:
3679:
3674:
3670:
3666:
3662:
3655:
3653:
3643:
3640:
3635:
3631:
3627:
3623:
3619:
3615:
3611:
3607:
3603:
3601:
3600:Caipirasuchus
3592:
3589:
3578:on 2010-09-21
3577:
3572:
3567:
3564:(28): 1–143.
3563:
3559:
3552:
3550:
3546:
3541:
3537:
3533:
3529:
3522:
3519:
3514:
3507:
3504:
3499:
3495:
3491:
3487:
3483:
3479:
3472:
3469:
3461:
3457:
3453:
3449:
3445:
3441:
3437:
3433:
3429:
3422:
3420:
3411:
3408:
3403:
3399:
3395:
3391:
3387:
3383:
3379:
3378:Araripesuchus
3372:
3369:
3364:
3360:
3356:
3352:
3348:
3344:
3340:
3336:
3329:
3326:
3321:
3317:
3313:
3309:
3305:
3301:
3294:
3291:
3286:
3282:
3277:
3272:
3267:
3262:
3258:
3254:
3251:(7): e21916.
3250:
3246:
3242:
3235:
3233:
3231:
3229:
3225:
3214:on 2011-07-06
3210:
3206:
3202:
3198:
3194:
3190:
3186:
3182:
3178:
3171:
3169:
3160:
3158:
3154:
3149:
3145:
3141:
3134:
3131:
3126:
3122:
3115:
3112:
3106:
3101:
3097:
3094:
3090:
3086:
3082:
3080:
3071:
3068:
3062:
3057:
3053:
3049:
3045:
3043:
3034:
3032:
3030:
3026:
3021:
3017:
3010:
3007:
3002:
2998:
2994:
2990:
2986:
2982:
2975:
2972:
2967:
2963:
2959:
2955:
2948:
2945:
2940:
2936:
2932:
2928:
2924:
2920:
2913:
2906:
2903:
2898:
2894:
2889:
2884:
2880:
2876:
2872:
2868:
2864:
2860:
2856:
2849:
2847:
2845:
2843:
2839:
2834:
2830:
2825:
2820:
2816:
2812:
2807:
2802:
2798:
2794:
2790:
2786:
2782:
2780:
2770:
2767:
2760:
2758:
2755:
2754:
2749:
2745:
2741:
2737:
2729:
2722:
2721:
2713:
2712:
2704:
2703:
2695:
2694:
2686:
2685:
2677:
2676:
2668:
2667:
2659:
2658:
2650:
2649:
2646:
2645:
2644:
2637:
2636:
2633:
2632:
2629:
2628:
2627:
2620:
2619:
2613:
2612:
2609:
2608:
2605:
2604:
2603:
2596:
2595:
2589:
2588:
2585:
2584:
2581:
2580:
2579:
2572:
2571:
2565:
2564:
2561:
2560:
2557:
2556:
2555:
2548:
2547:
2541:
2540:
2537:
2536:
2533:
2532:
2531:
2524:
2523:
2517:
2516:Baurusuchinae
2514:
2513:
2510:
2509:
2501:
2500:
2492:
2491:
2488:
2487:
2486:
2479:
2478:
2475:
2474:
2471:
2470:
2469:
2462:
2461:
2455:
2454:
2451:
2450:
2447:
2446:
2445:
2438:
2437:
2431:
2428:
2427:
2421:
2420:
2417:
2416:
2413:
2412:
2411:
2404:
2403:
2397:
2396:
2393:
2392:
2389:
2388:
2387:
2380:
2379:
2373:
2372:Baurusuchidae
2370:
2369:
2365:
2363:
2359:
2358:Baurusuchinae
2355:
2353:
2352:Aphaurosuchus
2348:
2343:
2341:
2337:
2336:
2335:Campinasuchus
2325:
2324:
2316:
2315:
2307:
2306:
2298:
2297:
2289:
2288:
2280:
2279:
2271:
2270:
2262:
2261:
2253:
2252:
2249:
2248:
2247:
2240:
2239:
2236:
2235:
2232:
2231:
2230:
2223:
2222:
2216:
2215:
2212:
2211:
2208:
2207:
2206:
2199:
2198:
2192:
2191:
2187:
2186:
2183:
2182:
2179:
2178:
2177:
2170:
2169:
2162:
2161:Baurusuchinae
2158:
2157:
2154:
2153:
2145:
2144:
2141:
2140:
2139:
2132:
2131:
2128:
2127:
2124:
2123:
2122:
2115:
2114:
2107:
2103:
2102:
2096:
2095:
2092:
2091:
2088:
2087:
2086:
2079:
2078:
2071:
2070:Baurusuchidae
2067:
2066:
2063:
2062:
2059:
2058:
2057:
2050:
2049:
2043:
2042:
2039:
2038:
2035:
2034:
2033:
2032:Mariliasuchus
2026:
2025:
2019:
2018:
2015:
2014:
2011:
2010:
2009:
2002:
2001:
1994:
1991:
1990:
1986:
1984:
1979:
1977:
1973:
1969:
1965:
1961:
1957:
1953:
1949:
1945:
1941:
1940:Baurusuchinae
1937:
1933:
1928:
1926:
1925:
1924:Comahuesuchus
1920:
1919:
1914:
1912:
1905:
1904:
1902:
1897:
1896:
1891:
1890:
1885:
1884:
1879:
1878:
1873:
1872:
1867:
1863:
1861:
1856:
1854:
1850:
1845:
1841:
1837:
1833:
1829:
1825:
1821:
1817:
1813:
1809:
1808:
1803:
1793:
1792:
1789:Baurusuchids
1780:
1779:
1771:
1770:
1762:
1761:
1753:
1752:
1744:
1743:
1735:
1734:
1726:
1725:
1722:
1721:
1720:Peirosauridae
1715:
1714:
1711:
1710:
1702:
1701:
1698:
1697:
1696:
1689:
1688:
1685:
1684:
1681:
1680:
1679:
1672:
1671:
1664:
1660:
1659:
1653:
1652:
1649:
1648:
1645:
1644:
1643:
1641:
1640:
1630:
1629:
1622:
1618:
1617:
1614:
1613:
1610:
1609:
1603:
1602:
1596:
1595:
1592:
1591:
1588:
1587:
1586:
1584:
1583:
1573:
1572:
1566:
1565:
1559:
1558:
1555:
1554:
1551:
1550:
1549:
1548:Araripesuchus
1542:
1541:
1535:
1534:
1531:
1530:
1522:
1521:
1518:
1517:
1516:
1509:
1508:
1505:
1504:
1501:
1500:
1499:
1492:
1491:
1485:
1481:
1480:
1473:
1470:
1469:
1465:
1463:
1462:
1457:
1453:
1449:
1445:
1440:
1438:
1434:
1430:
1426:
1416:
1415:
1407:
1406:
1398:
1397:
1389:
1388:
1380:
1379:
1371:
1370:
1362:
1361:
1358:
1357:
1356:
1349:
1348:
1345:
1344:
1341:
1340:
1339:
1332:
1331:
1324:
1323:Baurusuchidae
1320:
1319:
1316:
1315:
1312:
1311:
1310:
1303:
1302:
1299:
1298:
1295:
1294:
1293:
1286:
1285:
1279:
1278:
1275:
1274:
1271:
1270:
1269:
1262:
1261:
1258:
1257:
1254:
1253:
1252:
1245:
1244:
1238:
1237:
1234:
1233:
1230:
1229:
1228:
1221:
1220:
1213:
1209:
1208:
1205:
1204:
1201:
1200:
1194:
1193:
1187:
1186:
1183:
1182:
1179:
1178:
1177:
1170:
1169:
1166:
1165:
1162:
1161:
1155:
1154:
1147:
1143:
1142:
1138:
1136:
1132:
1128:
1123:
1121:
1120:
1115:
1111:
1107:
1103:
1099:
1089:
1088:
1085:Baurusuchids
1076:
1075:
1067:
1066:
1058:
1057:
1049:
1048:
1040:
1039:
1031:
1030:
1027:
1026:
1025:
1018:
1017:
1014:
1013:
1010:
1005:
1004:
998:
997:
994:
993:
990:
989:
988:
981:
980:
974:
973:
970:
969:
966:
965:
964:
962:
961:
951:
950:
944:
943:
940:
939:
936:
935:
934:
932:
931:
921:
920:
914:
913:
910:
909:
906:
905:
904:
897:
896:
890:
889:
886:
885:
882:
881:
880:
873:
872:
865:
862:
861:
857:
855:
851:
847:
842:
841:
836:
835:
830:
826:
822:
818:
815:
811:
806:
804:
803:
798:
797:
792:
791:
786:
782:
778:
777:
772:
768:
764:
760:
756:
752:
749:
746:
738:
736:
734:
730:
726:
722:
721:Campinasuchus
719:A new genus,
717:
715:
711:
707:
706:Maastrichtian
703:
699:
691:
690:
684:
680:
678:
674:
670:
666:
662:
658:
654:
650:
649:
644:
643:
638:
637:
632:
631:
626:
625:
624:Campinasuchus
620:
616:
612:
608:
604:
601:
597:
593:
585:
583:
581:
577:
576:Baurusuchinae
573:
569:
566:
562:
560:
555:
553:
549:
544:
540:
536:
532:
528:
524:
523:South America
520:
516:
512:
509:
505:
504:Baurusuchidae
493:
492:
485:
483:
482:
475:
473:
472:
471:Campinasuchus
465:
464:
463:
462:
455:
451:
450:
443:
441:
440:
433:
431:
430:
429:Aplestosuchus
423:
421:
420:
419:Aphaurosuchus
413:
412:
411:
410:
409:Baurusuchinae
403:
400:
399:
392:
390:
389:
382:
379:
378:
371:
369:
368:
361:
358:
357:
350:
348:
347:
340:
337:
336:
335:Comahuesuchus
329:
326:
325:
318:
317:
315:
310:
306:
300:
299:Baurusuchidae
294:
291:
290:
286:
280:
274:
271:
268:
267:
264:
258:
255:
252:
251:
248:
242:
239:
236:
235:
232:
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200:
197:
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190:
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184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
150:
147:
144:
143:
138:
133:
129:
126:
125:
119:
115:
110:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
42:
34:
30:
22:
19:
5153:Baurusuchids
5013:
4938:
4902:
4895:
4877:Sphagesaurus
4875:
4868:
4861:
4841:
4834:
4827:
4803:
4795:
4788:
4780:
4773:Notosuchidae
4760:
4753:
4746:
4738:
4727:Sphagesauria
4706:
4699:
4692:
4672:
4665:
4658:
4651:
4631:
4624:
4617:
4605:
4592:
4585:
4577:
4569:
4562:
4523:
4518:Malawisuchus
4516:
4509:
4502:
4481:
4474:
4466:
4461:Libycosuchus
4459:
4451:
4444:
4436:
4373:
4313:
4306:
4299:
4292:
4287:Patagosuchus
4285:
4278:
4271:
4264:
4259:Hamadasuchus
4257:
4250:
4243:
4236:
4229:
4222:
4217:Antaeusuchus
4215:
4195:
4188:
4181:
4176:Epoidesuchus
4174:
4169:Caririsuchus
4167:
4160:
4153:Pepesuchinae
4139:
4132:
4126:Fortignathus
4124:
4119:Amargasuchus
4117:
4093:
4086:
4081:Sahitisuchus
4079:
4072:
4065:
4058:
4051:
4044:
4037:
4030:
4023:
4018:Barinasuchus
4016:
4009:
3989:
3982:
3962:
3954:
3947:
3940:
3909:
3831:Pseudosuchia
3816:Pseudosuchia
3795:
3695:
3691:
3681:
3664:
3660:
3651:
3642:
3609:
3605:
3599:
3591:
3580:. Retrieved
3576:the original
3561:
3531:
3527:
3521:
3512:
3506:
3484:(1): 87–98.
3481:
3477:
3471:
3460:the original
3431:
3427:
3418:
3410:
3388:(1): 57–76.
3385:
3381:
3377:
3371:
3346:
3342:
3338:
3334:
3328:
3303:
3299:
3293:
3248:
3244:
3216:. Retrieved
3209:the original
3183:(1): 11–30.
3180:
3176:
3167:
3147:
3143:
3139:
3133:
3124:
3120:
3114:
3088:
3084:
3078:
3070:
3051:
3047:
3041:
3019:
3015:
3009:
2984:
2980:
2974:
2965:
2961:
2957:
2953:
2947:
2922:
2918:
2905:
2862:
2858:
2788:
2784:
2778:
2769:
2751:
2743:
2733:
2730:Paleobiology
2641:
2640:
2624:
2623:
2600:
2599:
2576:
2575:
2552:
2551:
2528:
2527:
2483:
2482:
2466:
2465:
2442:
2441:
2408:
2407:
2384:
2383:
2371:
2350:
2346:
2344:
2339:
2333:
2331:
2244:
2243:
2227:
2226:
2203:
2202:
2188:
2174:
2173:
2136:
2135:
2119:
2118:
2083:
2082:
2069:
2054:
2053:
2030:
2029:
2006:
2005:
1982:
1980:
1975:
1971:
1967:
1959:
1955:
1951:
1947:
1943:
1931:
1930:Montefeltro
1929:
1922:
1916:
1909:
1906:
1899:
1893:
1887:
1881:
1875:
1869:
1858:
1847:
1831:
1827:
1823:
1819:
1815:
1811:
1805:
1801:
1799:
1718:
1693:
1692:
1676:
1675:
1638:
1637:
1634:
1633:
1606:
1581:
1580:
1577:
1576:
1546:
1545:
1515:Malawisuchus
1513:
1512:
1496:
1495:
1459:
1455:
1451:
1448:peirosaurids
1441:
1436:
1432:
1429:crocodylians
1422:
1353:
1352:
1336:
1335:
1322:
1307:
1306:
1290:
1289:
1266:
1265:
1249:
1248:
1225:
1224:
1212:Sebecosuchia
1197:
1176:Libycosaurus
1174:
1173:
1158:
1127:paraphyletic
1124:
1117:
1113:
1109:
1105:
1095:
1022:
1021:
1008:
985:
984:
959:
958:
955:
954:
929:
928:
925:
924:
903:Libycosuchus
901:
900:
877:
876:
853:
838:
832:
829:crocodylians
810:phylogenetic
807:
800:
794:
788:
784:
780:
774:
767:Sebecosuchia
754:
742:
728:
720:
718:
701:
697:
695:
687:
672:
668:
664:
652:
646:
640:
634:
628:
622:
618:
595:
591:
589:
567:
557:
546:
503:
502:
489:
479:
469:
459:
447:
437:
427:
417:
407:
396:
386:
375:
365:
354:
344:
333:
322:
298:
278:
269:
263:Sebecosuchia
253:
237:
224:
211:
205:Pseudosuchia
198:
185:
122:
24:Baurusuchids
18:
5038:Wikispecies
4790:Llanosuchus
4708:Wargosuchus
4667:Baurusuchus
4579:Eremosuchus
4551:Baurusuchia
4438:Anatosuchus
4308:Rukwasuchus
4294:Peirosaurus
4197:Roxochampsa
4095:Zulmasuchus
4046:Iberosuchus
4032:Bretesuchus
4025:Bergisuchus
4011:Ayllusuchus
3984:Kaprosuchus
3942:Baharijodon
3911:Microsuchus
3893:Ziphosuchia
3880:Ziphosuchia
3843:Ziphosuchia
3755:Ziphosuchia
3339:Iberosuchus
2744:Baurusuchus
2246:B. pachecoi
2205:B. albertoi
2190:Baurusuchus
2138:Wargosuchus
1976:Wargosuchus
1956:Baurusuchus
1948:Wargosuchus
1846:containing
1832:Wargosuchus
1816:Baurusuchus
1802:Baurusuchus
1695:Bretesuchus
1582:Baurusuchus
1456:Bretesuchus
1452:Baurusuchus
1437:Baurusuchus
1433:Baurusuchus
1338:Baurusuchus
1309:Bretesuchus
1292:Iberosuchus
1251:Eremosuchus
1114:Baurusuchus
1106:Baurusuchus
1024:Bretesuchus
960:Iberosuchus
930:Baurusuchus
846:Ziphosuchia
840:Eremosuchus
834:Iberosuchus
802:Wargosuchus
781:Baurusuchus
757:. In 1946,
755:Baurusuchus
673:Wargosuchus
665:Wargosuchus
648:Wargosuchus
619:Baurusuchus
603:Bauru Group
596:Baurusuchus
592:Baurusuchus
572:subfamilies
568:Baurusuchia
545:containing
491:Wargosuchus
439:Baurusuchus
356:Eremosuchus
279:Baurusuchia
192:Archosauria
39:86–62
5127:Categories
4843:Yacarerani
4805:Notosuchus
4525:Pakasuchus
4504:Candidodon
4476:Simosuchus
4420:Notosuchia
4407:Notosuchia
4347:see below↓
4338:Notosuchia
4301:Pepesuchus
4273:Lomasuchus
4266:Kinesuchus
4190:Pepesuchus
4074:Ogresuchus
4067:Lorosuchus
4060:Langstonia
3964:Wanosuchus
3852:see below↓
3800:Metasuchia
3785:Sauropsida
3582:2010-08-13
3218:2010-12-26
3150:: 217–270.
2761:References
2008:Notosuchus
1993:Notosuchia
1918:Pakasuchus
1498:Notosuchus
1484:Notosuchia
1472:Metasuchia
1199:Notosuchia
1146:Metasuchia
879:Notosuchus
864:Notosuchia
821:Notosuchia
817:Metasuchia
570:, and two
247:Notosuchia
4633:Pabwehshi
4183:Itasuchus
4053:Ilchunaia
4002:Sebecidae
3949:Doratodon
3771:Kingdom:
3667:: 19–42.
3634:235172623
3626:1477-2019
3456:1342-937X
3205:1342-937X
2879:0021-8782
2815:1932-6203
2748:oospecies
1911:Pabwehshi
1663:Sebecidae
1639:Pabwehshi
1608:Neosuchia
1461:Pabwehshi
1425:Neosuchia
1355:Pabwehshi
1160:Neosuchia
1135:cladogram
825:Neosuchia
771:Sebecidae
745:Brazilian
739:Phylogeny
702:Pabwehshi
686:Skull of
677:Argentina
615:Santonian
539:PhyloCode
527:Argentina
508:Gondwanan
377:Pabwehshi
155:Kingdom:
149:Eukaryota
121:Skull of
29:Santonian
5023:Wikidata
3779:Chordata
3777:Phylum:
3773:Animalia
3498:86257810
3402:85946263
3285:21765925
3245:PLOS ONE
2897:33870512
2833:30067779
2785:PLOS ONE
2746:. A new
1985:(2011):
1874:but not
779:). Both
759:American
710:Pakistan
611:Turonian
565:suborder
535:Pakistan
292:Family:
283:Warren,
179:Reptilia
169:Chordata
165:Phylum:
159:Animalia
145:Domain:
5097:3605935
5071:4822293
5058:4472863
5029:Q140104
4088:Sebecus
3928:Sebecia
3789:Order:
3783:Class:
3700:Bibcode
3661:Zootaxa
3562:ZooKeys
3436:Bibcode
3351:Bibcode
3343:Geobios
3308:Bibcode
3276:3135595
3253:Bibcode
3185:Bibcode
3140:Sebecus
3093:Preview
3085:Zootaxa
2989:Bibcode
2968:: 1–20.
2927:Bibcode
2888:8349455
2824:6070184
2793:Bibcode
2750:called
2345:Darlim
2163:
2108:
2072:
1995:
1678:Sebecus
1665:
1623:
1621:Sebecia
1474:
1444:Sebecia
1325:
1268:Sebecus
1214:
1148:
1110:Sebecus
987:Sebecus
866:
850:derived
785:Sebecus
776:Sebecus
714:Sebecia
657:English
312:Genera
303:Price,
175:Class:
5084:116722
3632:
3624:
3496:
3454:
3400:
3283:
3273:
3203:
3127:: 1–8.
2895:
2885:
2877:
2831:
2821:
2813:
2740:Brazil
2360:, and
2347:et al.
2159:
2104:
2068:
1983:et al.
1932:et al.
1830:, and
1812:et al.
1661:
1619:
1482:
1321:
1210:
1144:
854:et al.
799:, and
645:, and
607:Brazil
586:Genera
531:Brazil
511:family
270:Branch
33:Danian
5110:38393
5079:IRMNG
3796:Clade
3657:(PDF)
3630:S2CID
3494:S2CID
3463:(PDF)
3424:(PDF)
3398:S2CID
3212:(PDF)
3173:(PDF)
2915:(PDF)
1964:basal
1898:, or
1844:clade
1131:basal
814:clade
808:Many
543:clade
506:is a
254:Clade
238:Clade
225:Clade
212:Clade
199:Clade
186:Clade
5066:GBIF
3829:see
3665:2871
3622:ISSN
3452:ISSN
3281:PMID
3201:ISSN
3089:2871
2956:and
2893:PMID
2875:ISSN
2829:PMID
2811:ISSN
1954:and
1946:and
1938:and
1921:and
1857:and
1458:and
1116:and
1108:and
837:and
783:and
671:and
578:and
556:and
529:and
305:1945
285:1968
47:PreꞒ
5053:EoL
3708:doi
3669:doi
3614:doi
3566:doi
3536:doi
3486:doi
3444:doi
3390:doi
3359:doi
3316:doi
3271:PMC
3261:doi
3193:doi
3100:doi
3096:PDF
3056:doi
3052:149
2997:doi
2935:doi
2883:PMC
2867:doi
2863:239
2819:PMC
2801:doi
2738:of
1104:of
708:of
605:of
513:of
5129::
5107::
5094::
5081::
5068::
5055::
5040::
5025::
3798::
3706:.
3696:54
3694:.
3690:.
3663:.
3659:.
3628:.
3620:.
3610:19
3608:.
3604:.
3560:.
3548:^
3532:19
3530:.
3492:.
3482:25
3480:.
3450:.
3442:.
3430:.
3426:.
3396:.
3386:20
3384:.
3357:.
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3345:.
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3243:.
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3191:.
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3125:25
3123:.
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3050:.
3046:.
3028:^
3020:30
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2995:.
2983:.
2964:.
2933:.
2923:27
2921:.
2917:.
2891:.
2881:.
2873:.
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2857:.
2841:^
2827:.
2817:.
2809:.
2799:.
2789:13
2787:.
2783:.
1958:.
1927:)
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1886:,
1880:,
1851:,
1826:,
1822:,
1818:,
793:,
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679:.
651:.
639:,
633:,
627:,
613:-
582:.
550:,
272::
256::
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227::
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201::
188::
97:Pg
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102:N
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52:Ꞓ
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