792:
4371:
4366:
3456:
3470:
724:
4353:
884:
129:
147:
3463:
2410:
4377:
1522:
1210:
606:
4360:
2398:
1071:
2377:
1120:
and display an incorrect, more rectangular profile. The skull was in fact triangular with a pointed snout, equipped with a beak. Its teeth were more tightly packed in the jaw compared to other
Morrison euornithopods. Museum curator John Foster describes them as having "thick median ridges on their
1048:, consisting of an anterior cervical centrum; an anterior dorsal centrum; a dorsosacral centrum; four sacral centra; a caudosacral centrum; two anterior caudal centra; three medial caudal centra; and a distal fragment from the left humerus. The Fuentecillas specimen is seen to be closer to
826:
in 1909 based on the holotype and only known specimen USNM 4753, a fragmentary postcranium, by the "narrowness or depressed nature of the ilia". Carpenter and Wilson (2008) referred this species to
791:
2023:
Sachs, S.; Hornung, J.J. (2006). "Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper
Cretaceous (Lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria)".
1107:. The largest fragments from later strata indicate adult individuals around 6.79 metres (22.3 ft) long. The Quarry 13 individuals are smaller though. It is estimated that
1837:
Brill, K.; K. Carpenter (2007). "A description of a new ornithopod from the Lytle Member of the
Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of
1013:
by
Lydekker in 1889. However, Naish & Martill (2008), McDonald and colleagues (2010), and McDonald (2011) found that Seeley's original generic distinction was valid.
945:
dryosaurid, but a more recent analysis contends that the diagnostic features of
Dryosauridae do not overlap with its material, instead considering it an iguanodontian.
4862:
1724:
Bakker, R.T. (1998). "Dinosaur mid-life crisis: the
Jurassic-Cretaceous transition in Wyoming and Colorado". In Lucas, S.G.; Kirkland, J.I.; Estep, J.W. (eds.).
1536:), scientists believe they may have been able to achieve running speeds of 25 km per hour (15 mph). A tiny 9 inch fossilized embryo, referred to
4936:
4921:
1877:(Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb".
4951:
2242:"The largest ornithopod (Dinosauria: Ornithischia) from the Upper Jurassic of Europe sheds light on the evolutionary history of basal ankylopollexians"
4849:
4370:
1581:"Tracing Late Jurassic ornithopod diversity in the eastern Iberian Peninsula: Camptosaurus-like postcranial remains from Alpuente (Valencia, Spain)"
941:
consists of the holotype and only known specimen NHMUK R167, a poorly preserved left femur lacking the distal end. It was earlier believed to be a
3455:
4931:
4926:
4365:
2442:
2185:
2151:
1850:
3469:
1554:
2166:
Norman D.B. and
Barrett, P.M. 2002. Ornithischian dinosaurs from the Lower Cretaceous (Berriasian) of England. Palaeontology 68:161-189
2241:
2142:, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England". In Carpenter, Kenneth (ed.).
528:
4352:
1131:
teeth frequently exhibit extensive wear, which indicates that individuals in the genus had a diet of relatively tough vegetation.
899:
species in North
America, numerous species from Europe were also referred to the genus in the late 19th and early 20th centuries:
4941:
2091:
Fonseca, André O.; Reid, Iain J.; Venner, Alexander; Duncan, Ruairidh J.; Garcia, Mauricio S.; Müller, Rodrigo T. (2024-12-31).
1081:
is a relatively heavily built form, with robust hindlimbs and broad feet, still having four toes. Due to the separate status of
4731:
805:
1165:, of which group it would then be a basal member. This would mean that the genus is closely related to the ancestor of later
4754:
4741:
1705:
Galton, P.M.; Powell, H.P. (1980). "The ornithischian dinosaur
Camptosaurus prestwichii from the Upper Jurassic of England".
1041:
696:. Throughout the 1880 and 1890s, he continued to receive specimens from Quarry 13 and in 1894 named two additional species:
4946:
4746:
1121:
lateral sides and denticles along their edges," these features were similar to, but "more fully developed" than those in
866:
ending in a small "foot" among other features. An analysis by Andrew McDonald and colleagues in 2010 suggested that like
146:
4376:
638:, or "flexible back", from Greek κάμπτω, "to bend" and νῶτον, "back", in reference to the presumed flexibility of the
2388:
2314:
McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C. & Panzarin, L. (2010).
855:
840:. However, McDonald and colleagues (2010), and McDonald (2011) found that the horizontal postacetabular process of
3462:
2435:
2203:"Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinhã, Portugal"
819:
2542:
1917:
McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C.; Panzarin, L. (2010).
1890:
2381:
2316:"New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs"
1919:"New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs"
4956:
1143:
to a family of its own: the
Camptosauridae. Alternatively some authors considered it an early member of the
923:(Nopcsa, 1900) consists of fragmentary material, a dentary and articular from Upper Cretaceous rocks of the
4789:
2901:
2753:
723:
621:
1111:
typically reached 5–6 metres (16–20 ft) long and weighed 500–1,000 kilograms (1,100–2,200 lb).
4888:
4711:
3267:
2569:
629:
4916:
4836:
4131:
2428:
2414:
2327:
2214:
2104:
2032:
1930:
1752:
1647:
1592:
996:
326:
4359:
784:
617:
415:
883:
4483:
4094:
2679:
2202:
1894:
1768:
1663:
1616:
1185:
1088:
823:
705:
675:
401:
141:
4893:
4827:
128:
4875:
4867:
4490:
4229:
4150:
4110:
2994:
2880:
2826:
2797:
2355:
2181:
2147:
2120:
1958:
1846:
1608:
1549:
1104:
888:
4880:
4631:
4624:
4434:
4427:
3956:
3867:
3705:
3606:
3592:
3306:
3274:
2909:
2818:
2732:
2667:
2402:
2345:
2335:
2253:
2222:
2112:
2040:
2005:
1948:
1938:
1886:
1806:
1760:
1655:
1600:
1439:
815:
651:
639:
555:
484:
609:
Historical skeletal restoration by O.C. Marsh, with skull based on remains now referred to
4716:
4511:
4462:
4325:
4256:
4159:
3979:
3698:
3684:
3663:
3281:
3154:
2921:
2689:
2527:
1728:. Vol. 14. New Mexico Museum of Natural History and Science Bulletin. pp. 67–77.
2331:
2218:
2108:
2036:
1986:
1934:
1756:
1651:
1596:
1114:
Earlier reconstructions, such as those by Marsh and Gilmore, were based on the skull of
4726:
4721:
4659:
4596:
4469:
4318:
4281:
4263:
4208:
3840:
3656:
3649:
3578:
3500:
3487:
3415:
3403:
3372:
3347:
3211:
3133:
3094:
3080:
2957:
2868:
2790:
2768:
2761:
2502:
2350:
2315:
1953:
1918:
1580:
1351:
1229:
1191:
1162:
1144:
924:
844:
is more likely a product of distortion. Therefore, McDonald put it into its own genus,
271:
223:
2226:
1540:, was retrieved from Morrison Formation strata at Dinosaur National Monument in Utah.
4910:
4666:
4610:
4589:
4560:
4476:
4413:
4311:
4304:
4290:
4200:
4180:
4173:
4070:
4020:
4013:
3997:
3972:
3945:
3938:
3931:
3902:
3853:
3815:
3780:
3761:
3733:
3677:
3340:
3333:
3288:
3218:
3087:
2946:
2931:
2893:
2804:
2697:
2660:
2653:
2092:
1772:
1667:
1620:
1375:
1318:
1256:
1151:
1116:
1018:
951:
871:
800:
625:
567:
559:
552:
545:
86:
48:
2072:, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere".
1898:
1810:
1099:, from Quarry 13, have been recovered from very deep layers, probably dating to the
4574:
4550:
4538:
4504:
4497:
4448:
4441:
4247:
4191:
4166:
4138:
4027:
3829:
3794:
3740:
3670:
3642:
3528:
3521:
3361:
3326:
3253:
3246:
3109:
3064:
2966:
2840:
2833:
2782:
2746:
2704:
2554:
2493:
2061:
1795:, with a revision of the species of the genus, and descriptions of two new species"
1533:
1521:
1209:
1174:
1166:
988:
967:
942:
933:
846:
739:
683:
659:
443:
334:
207:
2116:
2064:(2009). "Notes on Neocomian (Late Cretaceous) ornithopod dinosaurs from England -
2044:
605:
2340:
2257:
2138:
Ruiz-Omeñaca, José Ignacio; Pereda Suberbiola, Xabier; Galton, Peter M. (2007). "
1943:
1790:
766:
was a valid species. They also considered a skull, YPM 1887, in 1886 referred to
4821:
4763:
4652:
4617:
4455:
4297:
4270:
4222:
4215:
4048:
4034:
4006:
3986:
3965:
3909:
3888:
3881:
3874:
3808:
3801:
3754:
3719:
3691:
3627:
3620:
3389:
3239:
3225:
3197:
3175:
3117:
2980:
2811:
2775:
2739:
2635:
2622:
2581:
2451:
1170:
992:
634:
428:
239:
61:
31:
1604:
4705:
4680:
4673:
4645:
4638:
4529:
4518:
4404:
4103:
4083:
4041:
3916:
3768:
3635:
3585:
3550:
3543:
3514:
3354:
3232:
3204:
3073:
3046:
3021:
2861:
2854:
2847:
2725:
2009:
1659:
1579:
Cubells-Alarcó, M.; Verdú, F. J.; Suñer, M.; de Santisteban, C. (2022-01-08).
1399:
1301:
1214:
1179:
1123:
778:, but the specimen was recently shown by Brill and Carpenter not to belong to
692:
667:
390:
255:
106:
71:
17:
4812:
2124:
1764:
1612:
716:
in his 1909 redescription of the Marsh specimens. In the Morrison Formation,
4687:
4603:
4582:
4420:
4117:
4055:
3564:
3536:
3396:
3260:
3190:
3161:
2987:
2973:
2716:
1199:
1100:
961:
828:
158:
111:
55:
2359:
1962:
931:. It is almost certainly a rhabdodontid and is no longer considered valid (
836:, on the basis of similarities between its ilium and the holotype ilium of
2376:
1845:. Bloomington and Indianapolis: Indiana University Press. pp. 49–67.
1070:
974:
by Norman and Barrett in 2002. It has since been transferred to the genus
4806:
4567:
4236:
4124:
3924:
3895:
3822:
3747:
3712:
3613:
3599:
3557:
3381:
3182:
3147:
3053:
3039:
3001:
2511:
2484:
2475:
2146:. Bloomington and Indianapolis: Indiana University Press. pp. 3–16.
1423:
1054:
1005:
999:
was driven into the side of a hill. It was described by Hulke in 1880 as
976:
643:
548:
436:
383:
191:
178:
101:
96:
81:
76:
66:
2093:"A comprehensive phylogenetic analysis on early ornithischian evolution"
1058:
from the Jurassic of Portugal, hence its tentative placement within the
4854:
3860:
3295:
1270:
1083:
928:
876:
863:
671:
460:
368:
355:
116:
91:
2289:
Jurassic West: The Dinosaurs of the Morrison Formation and Their World
1825:
Jurassic West: The Dinosaurs of the Morrison Formation and Their World
3726:
3319:
2469:
563:
168:
4783:
2240:
Sánchez-Fenollosa, Sergio; Verdú, Francisco; Cobos, Alberto (2023).
1740:
1635:
949:
is probably a valid dryosaurid that has been moved to the new genus
3571:
3101:
1520:
1208:
1159:
1069:
1045:
882:
790:
722:
650:
by him in 1885 because the original name was already in use for a
604:
541:
4841:
2939:
1040:. A single specimen was found in the Fuentecillas member of the
1032:
In 2022, the first confirmed European remains belonging to aff.
758:
to be different growth stages or different gender of the larger
4787:
4392:
3485:
2620:
2462:
2424:
870:
is actually more closely related to more advanced iguanodonts (
854:, was named by Carpenter and Wilson in 2008 for specimens from
2420:
774:
as well. Gilmore had used this skull to describe the skull of
662:
of the genus) for material he received from his collectors at
591:
584:
577:
570:
514:
798:
skeleton mounted in outdated quadrupedal posture, and with
517:
505:
731:
skeleton at the AMNH, now thought to be a growth stage of
1891:
10.2992/0097-4463(2008)76[227:ansoco]2.0.co;2
862:
in the lower jaw, shorter neck vertebrae, and straighter
493:
2201:
Mateus, Octávio; Telles Antunes, Miguel (January 2001).
1528:(blue) compared in size to a human and other ornithopods
646:
was YPM 1877, a partial skeleton. The genus was renamed
1021:, more closely related to advanced iguanodonts than to
782:. In 2007, they put it into its own genus and species,
2279:
2277:
2275:
2273:
2271:
2269:
2267:
2386:
2144:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
1843:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
529:
520:
511:
490:
487:
1873:
Carpenter, K.; Wilson, Y. (2008). "A new species of
502:
499:
379:
364:
4796:
4697:
4548:
4528:
4403:
4280:
4246:
4190:
4149:
4093:
4069:
3996:
3955:
3839:
3778:
3498:
3414:
3371:
3305:
3131:
3063:
3019:
2956:
2920:
2879:
2715:
2678:
2633:
2580:
2553:
2526:
770:by Marsh, later confirmed by Gilmore, to belong to
508:
496:
1868:
1866:
1864:
1862:
1741:"The typical Ornithopoda of the American Jurassic"
1726:Lower and Middle Cretaceous Terrestrial Ecosystems
1087:it has become problematic which material from the
1912:
1910:
1908:
1681:Marsh, O.C. (1885). "Names of extinct reptiles".
1799:Proceedings of the United States National Museum
720:fossils are present in stratigraphic zones 2–6.
682:based on the holotype YPM 1879, a foot found by
690:. The foot was later shown to have belonged to
1700:
1698:
1696:
1574:
1572:
1570:
2436:
1532:Based on studies of other iguanodonts (clade
8:
1095:. The specimens with certainty belonging to
1980:
1978:
1976:
1974:
1972:
1784:
1782:
4784:
4400:
4389:
4075:
3845:
3786:
3506:
3495:
3482:
3311:
3139:
3031:
3027:
2885:
2645:
2641:
2630:
2617:
2559:
2532:
2523:
2459:
2443:
2429:
2421:
1009:by Seeley in 1888, but was soon sunk into
742:and H.P. Powell in their redescription of
127:
38:
2349:
2339:
2246:Zoological Journal of the Linnean Society
1952:
1942:
2056:
2054:
1827:. Indiana University Press. pp. 327-329.
1036:.sp were described by Sánchez-Fenollosa
995:in England. The fossil was found when a
2393:
2291:. Indiana University Press. p. 219-221.
1566:
987:was recovered from Chawley Brick Pits,
2302:The Princeton Field Guide to Dinosaurs
874:). It has been moved to the new genus
1987:"The taxonomy of species assigned to
1202:analyses of McDonald and colleagues,
7:
1640:American Journal of Science and Arts
1555:Paleobiota of the Morrison Formation
566:. The name means 'flexible lizard' (
4937:Taxa named by Othniel Charles Marsh
4922:Dinosaurs of the Morrison Formation
2304:, Princeton University Press p. 284
2097:Journal of Systematic Palaeontology
1791:"Osteology of the Jurassic reptile
1415:
1391:
1367:
1343:
1293:
1286:
1262:
1235:
1225:
1003:, and then placed in its own genus
25:
4952:Late Jurassic dinosaurs of Europe
1636:"Notice of new Jurassic reptiles"
4375:
4369:
4364:
4358:
4351:
3468:
3461:
3454:
2408:
2396:
2375:
2178:Fossil Reptiles of Great Britain
1841:". In Carpenter, Kenneth (ed.).
632:described and named the find as
483:
145:
136:Reconstructed skeleton in Japan
4732:List of informal dinosaur names
1823:Foster, J. (2007). "Appendix."
1811:10.5479/si.00963801.36-1666.197
983:The remaining European species
806:Natural History Museum of Milan
4742:Timeline of hadrosaur research
2176:Benton MJ, Spencer PS (1995).
1042:Villar del Arzobispo Formation
708:named two additional species,
1:
4932:Fossil taxa described in 1885
4927:Fossil taxa described in 1879
2227:10.1016/S0753-3969(01)88003-4
2117:10.1080/14772019.2024.2346577
2045:10.1016/j.geobios.2005.01.003
1879:Annals of the Carnegie Museum
624:found the remains of a small
2341:10.1371/journal.pone.0014075
1944:10.1371/journal.pone.0014075
1177:than contemporaries such as
746:(see following), considered
592:
578:
375:Carpenter & Wilson, 2008
1985:Andrew T. McDonald (2011).
1745:American Journal of Science
1683:American Journal of Science
895:While Marsh was describing
628:. That same year Professor
4973:
2258:10.1093/zoolinnean/zlad076
1991:(Dinosauria: Ornithopoda)"
1605:10.1007/s41513-021-00182-z
1585:Journal of Iberian Geology
1158:by definition part of the
856:Dinosaur National Monument
585:
571:
29:
27:Extinct genus of dinosaurs
4399:
4388:
4349:
4078:
3848:
3789:
3509:
3494:
3481:
3452:
3314:
3142:
3034:
3030:
2888:
2648:
2644:
2629:
2616:
2562:
2535:
2522:
2458:
2010:10.11646/zootaxa.2783.1.4
1660:10.2475/ajs.s3-18.108.501
1437:
1420:
1413:
1396:
1389:
1372:
1365:
1348:
1341:
1315:
1298:
1291:
1284:
1267:
1260:
1240:
1233:
1025:, similar to the case of
970:in 1874 and was moved to
852:Camptosaurus aphanoecetes
850:. An additional species,
820:Hot Springs, South Dakota
704:, based in part on size.
407:
400:
361:
353:
340:
333:
142:Scientific classification
140:
135:
126:
41:
2207:Annales de Paléontologie
1765:10.2475/ajs.s3-48.283.85
1017:has been recovered as a
985:Camptosaurus prestwichii
544:of plant-eating, beaked
30:Not to be confused with
4942:Paleontology in Wyoming
1206:was placed as follows:
1173:dinosaurs and was more
1139:Marsh in 1885 assigned
814:was recovered from the
654:. In 1879, Marsh named
2180:. Chapman & Hall.
2074:Revue de Paléobiologie
1789:Gilmore, C.W. (1909).
1529:
1222:
1075:
892:
822:. It was described by
812:Camptosaurus depressus
808:
735:
622:Albany County, Wyoming
613:
4889:Paleobiology Database
4712:Basturs Poble bonebed
2384:at Wikimedia Commons
1524:
1217:dinosaurs, including
1212:
1198:In the 2010 and 2011
1073:
1001:Iguanodon prestwichii
959:was originally named
887:Cast of a skull from
886:
794:
762:, and therefore only
726:
630:Othniel Charles Marsh
616:On September 4, 1879
608:
597:) meaning 'lizard').
583:) meaning 'bent' and
4947:Ornithischian genera
4734:(contains hadrosaur
4132:Probrachylophosaurus
2283:Foster, J. (2007). "
2140:Callovosaurus leedsi
1739:Marsh, O.C. (1894).
1634:Marsh, O.C. (1879).
727:Outdated mount of a
601:History of discovery
2332:2010PLoSO...514075M
2285:Camptosaurus dispar
2219:2001AnPal..87...61M
2109:2024JSPal..2246577F
2037:2006Geobi..39..415S
1935:2010PLoSO...514075M
1757:1894AmJS...48...85M
1652:1879AmJS...18..501M
1597:2022JIbG...48...65S
1526:Camptosaurus dispar
1213:Restoration of six
1097:Camptosaurus dispar
1023:Camptosaurus dispar
785:Theiophytalia kerri
618:William Harlow Reed
4484:Ornithopodichnites
4095:Brachylophosaurini
2680:Hypsilophodontidae
2300:Paul, G.S., 2010,
1530:
1223:
1089:Morrison Formation
1076:
893:
858:. It differs from
838:Planicoxa venenica
809:
736:
706:Charles W. Gilmore
676:Morrison Formation
614:
562:and possibly also
343:Camptonotus dispar
4904:
4903:
4876:Open Tree of Life
4790:Taxon identifiers
4781:
4780:
4777:
4776:
4773:
4772:
4491:Shiraminesauropus
4384:
4383:
4347:
4346:
4343:
4342:
4339:
4338:
4335:
4334:
4230:Naashoibitosaurus
4151:Austrokritosauria
4111:Brachylophosaurus
4065:
4064:
3477:
3476:
3450:
3449:
3446:
3445:
3442:
3441:
3438:
3437:
3434:
3433:
3127:
3126:
3015:
3014:
3011:
3010:
2995:Transylvanosaurus
2881:Rhabdodontomorpha
2827:Notohypsilophodon
2798:Macrogryphosaurus
2612:
2611:
2608:
2607:
2604:
2603:
2600:
2599:
2380:Media related to
2187:978-0-412-62040-9
2153:978-0-253-34817-3
1852:978-0-253-34817-3
1805:(1666): 197–332.
1550:List of dinosaurs
1513:
1512:
1504:
1503:
1495:
1494:
1486:
1485:
1477:
1476:
1468:
1467:
1459:
1458:
1450:
1449:
1330:
1329:
889:Bone Cabin Quarry
818:near the town of
474:
473:
468:
456:
447:
432:
419:
394:
376:
329:
307:
290:
54:156.3–146.8
16:(Redirected from
4964:
4897:
4896:
4884:
4883:
4871:
4870:
4858:
4857:
4845:
4844:
4832:
4831:
4830:
4817:
4816:
4815:
4785:
4758:mummy S.M. R4036
4632:Microhadrosaurus
4625:Mandschurosaurus
4435:Camptosaurichnus
4428:Bonaparteichnium
4401:
4394:Related articles
4390:
4379:
4373:
4368:
4363:
4362:
4355:
4076:
3957:Parasaurolophini
3868:Angulomastacator
3846:
3787:
3706:Shuangmiaosaurus
3607:Huehuecanauhtlus
3593:Gongpoquansaurus
3507:
3496:
3483:
3472:
3466:
3465:
3458:
3312:
3307:Hadrosauriformes
3275:Ratchasimasaurus
3140:
3032:
3028:
2910:Muttaburrasaurus
2886:
2819:Muttaburrasaurus
2733:Atlascopcosaurus
2668:Thescelosauridae
2646:
2642:
2631:
2618:
2560:
2533:
2524:
2517:
2516:
2460:
2445:
2438:
2431:
2422:
2413:
2412:
2411:
2401:
2400:
2399:
2392:
2379:
2364:
2363:
2353:
2343:
2311:
2305:
2298:
2292:
2281:
2262:
2261:
2252:(4): 1013–1033.
2237:
2231:
2230:
2198:
2192:
2191:
2173:
2167:
2164:
2158:
2157:
2135:
2129:
2128:
2088:
2082:
2081:
2062:Galton, Peter M.
2058:
2049:
2048:
2020:
2014:
2013:
1995:
1982:
1967:
1966:
1956:
1946:
1914:
1903:
1902:
1870:
1857:
1856:
1834:
1828:
1821:
1815:
1814:
1786:
1777:
1776:
1736:
1730:
1729:
1721:
1715:
1714:
1702:
1691:
1690:
1678:
1672:
1671:
1646:(108): 501–505.
1631:
1625:
1624:
1576:
1440:Hadrosauriformes
1416:
1392:
1368:
1344:
1294:
1287:
1263:
1236:
1226:
816:Lakota Formation
640:sacral vertebrae
595:
588:
587:
581:
574:
573:
536:
532:
527:
526:
523:
522:
519:
516:
513:
510:
507:
504:
501:
498:
495:
492:
489:
466:
454:
442:
426:
414:
389:
381:
374:
366:
324:
317:
305:
300:
288:
283:
270:
254:
238:
222:
206:
150:
149:
131:
121:
58:
47:Temporal range:
39:
21:
4972:
4971:
4967:
4966:
4965:
4963:
4962:
4961:
4907:
4906:
4905:
4900:
4892:
4887:
4879:
4874:
4866:
4861:
4853:
4848:
4840:
4835:
4826:
4825:
4820:
4811:
4810:
4805:
4792:
4782:
4769:
4750:mummy AMNH 5060
4717:Dakota (fossil)
4693:
4544:
4524:
4512:Tetrapodosaurus
4463:Hadrosaurichnus
4395:
4380:
4357:
4331:
4326:Shantungosaurus
4276:
4257:Augustynolophus
4242:
4186:
4160:Bonapartesaurus
4145:
4089:
4061:
3992:
3980:Parasaurolophus
3951:
3835:
3783:
3774:
3699:Qianjiangsaurus
3685:Probactrosaurus
3664:Penelopognathus
3503:
3490:
3473:
3460:
3430:
3410:
3367:
3301:
3282:Riabininohadros
3155:Bayannurosaurus
3136:
3123:
3059:
3024:
3007:
2952:
2922:Tenontosauridae
2916:
2875:
2711:
2690:Gideonmantellia
2674:
2638:
2625:
2596:
2576:
2549:
2543:Avemetatarsalia
2528:Avemetatarsalia
2518:
2465:
2464:
2454:
2449:
2419:
2409:
2407:
2397:
2395:
2387:
2373:
2368:
2367:
2313:
2312:
2308:
2299:
2295:
2282:
2265:
2239:
2238:
2234:
2200:
2199:
2195:
2188:
2175:
2174:
2170:
2165:
2161:
2154:
2137:
2136:
2132:
2090:
2089:
2085:
2060:
2059:
2052:
2022:
2021:
2017:
1993:
1984:
1983:
1970:
1916:
1915:
1906:
1872:
1871:
1860:
1853:
1836:
1835:
1831:
1822:
1818:
1788:
1787:
1780:
1738:
1737:
1733:
1723:
1722:
1718:
1704:
1703:
1694:
1680:
1679:
1675:
1633:
1632:
1628:
1578:
1577:
1568:
1563:
1546:
1519:
1514:
1505:
1496:
1487:
1478:
1469:
1460:
1451:
1331:
1221:at the far left
1137:
1068:
868:C. aphanoecetes
824:Charles Gilmore
603:
534:
530:
486:
482:
465:
453:
441:
425:
413:
388:
373:
369:C. aphanoecetes
349:
346:
323:
315:
304:
298:
287:
285:Camptosauroidea
281:
268:
252:
236:
220:
204:
144:
122:
120:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
53:
52:
45:
35:
28:
23:
22:
15:
12:
11:
5:
4970:
4968:
4960:
4959:
4957:Jurassic Spain
4954:
4949:
4944:
4939:
4934:
4929:
4924:
4919:
4909:
4908:
4902:
4901:
4899:
4898:
4885:
4872:
4859:
4846:
4833:
4818:
4802:
4800:
4794:
4793:
4788:
4779:
4778:
4775:
4774:
4771:
4770:
4768:
4767:
4760:
4752:
4744:
4739:
4729:
4727:Hypsilophodont
4724:
4722:Hadrosaur diet
4719:
4714:
4709:
4701:
4699:
4698:Other articles
4695:
4694:
4692:
4691:
4684:
4677:
4670:
4663:
4660:Stephanosaurus
4656:
4649:
4642:
4635:
4628:
4621:
4614:
4607:
4600:
4597:Fulgurotherium
4593:
4586:
4579:
4571:
4564:
4556:
4554:
4546:
4545:
4543:
4542:
4534:
4532:
4526:
4525:
4523:
4522:
4515:
4508:
4501:
4494:
4487:
4480:
4473:
4470:Iguanodontipus
4466:
4459:
4452:
4445:
4438:
4431:
4424:
4417:
4409:
4407:
4397:
4396:
4393:
4386:
4385:
4382:
4381:
4350:
4348:
4345:
4344:
4341:
4340:
4337:
4336:
4333:
4332:
4330:
4329:
4322:
4319:Laiyangosaurus
4315:
4308:
4301:
4294:
4286:
4284:
4282:Edmontosaurini
4278:
4277:
4275:
4274:
4267:
4264:Prosaurolophus
4260:
4252:
4250:
4244:
4243:
4241:
4240:
4233:
4226:
4219:
4212:
4209:Coahuilasaurus
4205:
4196:
4194:
4188:
4187:
4185:
4184:
4177:
4170:
4163:
4155:
4153:
4147:
4146:
4144:
4143:
4135:
4128:
4121:
4114:
4107:
4099:
4097:
4091:
4090:
4088:
4087:
4079:
4073:
4067:
4066:
4063:
4062:
4060:
4059:
4052:
4045:
4038:
4031:
4024:
4017:
4010:
4002:
4000:
3994:
3993:
3991:
3990:
3983:
3976:
3969:
3961:
3959:
3953:
3952:
3950:
3949:
3942:
3935:
3928:
3921:
3913:
3906:
3899:
3892:
3885:
3878:
3871:
3864:
3857:
3849:
3843:
3841:Lambeosaurinae
3837:
3836:
3834:
3833:
3826:
3819:
3812:
3805:
3798:
3790:
3784:
3779:
3776:
3775:
3773:
3772:
3765:
3758:
3751:
3744:
3737:
3730:
3723:
3716:
3709:
3702:
3695:
3688:
3681:
3674:
3667:
3660:
3657:Nanyangosaurus
3653:
3650:Nanningosaurus
3646:
3639:
3632:
3624:
3617:
3610:
3603:
3596:
3589:
3582:
3579:Gilmoreosaurus
3575:
3568:
3561:
3554:
3547:
3540:
3533:
3525:
3518:
3510:
3504:
3501:Hadrosauroidea
3499:
3492:
3491:
3488:Hadrosauroidea
3486:
3479:
3478:
3475:
3474:
3453:
3451:
3448:
3447:
3444:
3443:
3440:
3439:
3436:
3435:
3432:
3431:
3429:
3428:
3427:
3426:
3420:
3418:
3416:Hadrosauroidea
3412:
3411:
3409:
3408:
3404:Mantellisaurus
3400:
3393:
3386:
3377:
3375:
3373:Iguanodontidae
3369:
3368:
3366:
3365:
3358:
3351:
3348:Mantellisaurus
3344:
3337:
3330:
3323:
3315:
3309:
3303:
3302:
3300:
3299:
3292:
3285:
3278:
3271:
3264:
3257:
3250:
3243:
3236:
3229:
3222:
3215:
3212:Iguanacolossus
3208:
3201:
3194:
3187:
3179:
3172:
3165:
3158:
3151:
3143:
3137:
3134:Ankylopollexia
3132:
3129:
3128:
3125:
3124:
3122:
3121:
3114:
3106:
3098:
3095:Eousdryosaurus
3091:
3084:
3081:Dysalotosaurus
3077:
3069:
3067:
3061:
3060:
3058:
3057:
3050:
3043:
3035:
3025:
3020:
3017:
3016:
3013:
3012:
3009:
3008:
3006:
3005:
2998:
2991:
2984:
2977:
2970:
2962:
2960:
2958:Rhabdodontidae
2954:
2953:
2951:
2950:
2943:
2936:
2927:
2925:
2918:
2917:
2915:
2914:
2906:
2898:
2889:
2883:
2877:
2876:
2874:
2873:
2869:Weewarrasaurus
2865:
2858:
2851:
2844:
2837:
2830:
2823:
2815:
2808:
2801:
2794:
2791:Leaellynasaura
2787:
2779:
2772:
2769:Gasparinisaura
2765:
2762:Galleonosaurus
2758:
2750:
2743:
2736:
2729:
2721:
2719:
2713:
2712:
2710:
2709:
2701:
2694:
2685:
2683:
2676:
2675:
2673:
2672:
2664:
2657:
2649:
2639:
2634:
2627:
2626:
2621:
2614:
2613:
2610:
2609:
2606:
2605:
2602:
2601:
2598:
2597:
2595:
2594:
2593:
2592:
2586:
2584:
2578:
2577:
2575:
2574:
2573:
2572:
2563:
2557:
2551:
2550:
2548:
2547:
2546:
2545:
2536:
2530:
2520:
2519:
2515:
2514:
2505:
2503:Neornithischia
2496:
2487:
2478:
2472:
2463:
2456:
2455:
2450:
2448:
2447:
2440:
2433:
2425:
2418:
2417:
2405:
2372:
2371:External links
2369:
2366:
2365:
2326:(11): e14075.
2306:
2293:
2263:
2232:
2193:
2186:
2168:
2159:
2152:
2130:
2083:
2050:
2031:(3): 415–425.
2015:
1968:
1929:(11): e14075.
1904:
1885:(4): 227–263.
1858:
1851:
1829:
1816:
1778:
1751:(283): 85–90.
1731:
1716:
1692:
1673:
1626:
1565:
1564:
1562:
1559:
1558:
1557:
1552:
1545:
1542:
1518:
1515:
1511:
1510:
1507:
1506:
1502:
1501:
1498:
1497:
1493:
1492:
1489:
1488:
1484:
1483:
1480:
1479:
1475:
1474:
1471:
1470:
1466:
1465:
1462:
1461:
1457:
1456:
1453:
1452:
1448:
1447:
1444:
1443:
1436:
1433:
1432:
1429:
1428:
1419:
1414:
1412:
1409:
1408:
1405:
1404:
1395:
1390:
1388:
1385:
1384:
1381:
1380:
1371:
1366:
1364:
1361:
1360:
1357:
1356:
1352:Iguanacolossus
1347:
1342:
1340:
1337:
1336:
1333:
1332:
1328:
1327:
1324:
1323:
1314:
1311:
1310:
1307:
1306:
1297:
1292:
1290:
1285:
1283:
1280:
1279:
1276:
1275:
1266:
1261:
1259:
1253:
1252:
1249:
1248:
1239:
1234:
1232:
1230:Ankylopollexia
1224:
1192:Othnielosaurus
1163:Ankylopollexia
1145:Iguanodontidae
1136:
1135:Classification
1133:
1067:
1064:
1019:styracosternan
738:Then in 1980,
602:
599:
472:
471:
470:
469:
467:McDonald, 2011
457:
448:
433:
420:
405:
404:
398:
397:
396:
395:
384:C. prestwichii
377:
359:
358:
351:
350:
347:
338:
337:
331:
330:
313:
309:
308:
302:Camptosauridae
296:
292:
291:
279:
275:
274:
272:Ankylopollexia
266:
259:
258:
250:
243:
242:
234:
227:
226:
224:Neornithischia
218:
211:
210:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
133:
132:
124:
123:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
59:
46:
26:
24:
18:Camptosauridae
14:
13:
10:
9:
6:
4:
3:
2:
4969:
4958:
4955:
4953:
4950:
4948:
4945:
4943:
4940:
4938:
4935:
4933:
4930:
4928:
4925:
4923:
4920:
4918:
4915:
4914:
4912:
4895:
4890:
4886:
4882:
4877:
4873:
4869:
4864:
4860:
4856:
4851:
4847:
4843:
4838:
4834:
4829:
4823:
4819:
4814:
4808:
4804:
4803:
4801:
4799:
4795:
4791:
4786:
4766:
4765:
4761:
4759:
4757:
4756:Edmontosaurus
4753:
4751:
4749:
4748:Edmontosaurus
4745:
4743:
4740:
4737:
4733:
4730:
4728:
4725:
4723:
4720:
4718:
4715:
4713:
4710:
4708:
4707:
4703:
4702:
4700:
4696:
4690:
4689:
4685:
4683:
4682:
4678:
4676:
4675:
4671:
4669:
4668:
4667:Syngonosaurus
4664:
4662:
4661:
4657:
4655:
4654:
4650:
4648:
4647:
4643:
4641:
4640:
4636:
4634:
4633:
4629:
4627:
4626:
4622:
4620:
4619:
4615:
4613:
4612:
4611:Koutalisaurus
4608:
4606:
4605:
4601:
4599:
4598:
4594:
4592:
4591:
4590:Eucercosaurus
4587:
4585:
4584:
4580:
4577:
4576:
4572:
4570:
4569:
4565:
4563:
4562:
4561:Arstanosaurus
4558:
4557:
4555:
4553:
4552:
4547:
4541:
4540:
4536:
4535:
4533:
4531:
4527:
4521:
4520:
4516:
4514:
4513:
4509:
4507:
4506:
4502:
4500:
4499:
4495:
4493:
4492:
4488:
4486:
4485:
4481:
4479:
4478:
4477:Orcauichnites
4474:
4472:
4471:
4467:
4465:
4464:
4460:
4458:
4457:
4453:
4451:
4450:
4446:
4444:
4443:
4439:
4437:
4436:
4432:
4430:
4429:
4425:
4423:
4422:
4418:
4416:
4415:
4414:Amblydactylus
4411:
4410:
4408:
4406:
4402:
4398:
4391:
4387:
4378:
4374:
4372:
4367:
4361:
4354:
4328:
4327:
4323:
4321:
4320:
4316:
4314:
4313:
4312:Kundurosaurus
4309:
4307:
4306:
4305:Kerberosaurus
4302:
4300:
4299:
4295:
4293:
4292:
4291:Edmontosaurus
4288:
4287:
4285:
4283:
4279:
4273:
4272:
4268:
4266:
4265:
4261:
4259:
4258:
4254:
4253:
4251:
4249:
4245:
4239:
4238:
4234:
4232:
4231:
4227:
4225:
4224:
4220:
4218:
4217:
4213:
4211:
4210:
4206:
4203:
4202:
4201:Anasazisaurus
4198:
4197:
4195:
4193:
4189:
4183:
4182:
4181:Secernosaurus
4178:
4176:
4175:
4174:Kelumapusaura
4171:
4169:
4168:
4164:
4162:
4161:
4157:
4156:
4154:
4152:
4148:
4141:
4140:
4136:
4134:
4133:
4129:
4127:
4126:
4122:
4120:
4119:
4115:
4113:
4112:
4108:
4106:
4105:
4101:
4100:
4098:
4096:
4092:
4086:
4085:
4081:
4080:
4077:
4074:
4072:
4071:Saurolophinae
4068:
4058:
4057:
4053:
4051:
4050:
4046:
4044:
4043:
4039:
4037:
4036:
4032:
4030:
4029:
4025:
4023:
4022:
4021:Hypacrosaurus
4018:
4016:
4015:
4014:Corythosaurus
4011:
4009:
4008:
4004:
4003:
4001:
3999:
3998:Lambeosaurini
3995:
3989:
3988:
3984:
3982:
3981:
3977:
3975:
3974:
3973:Charonosaurus
3970:
3968:
3967:
3963:
3962:
3960:
3958:
3954:
3948:
3947:
3946:Tsintaosaurus
3943:
3941:
3940:
3939:Pararhabdodon
3936:
3934:
3933:
3932:Nipponosaurus
3929:
3927:
3926:
3922:
3919:
3918:
3914:
3912:
3911:
3907:
3905:
3904:
3903:Jaxartosaurus
3900:
3898:
3897:
3893:
3891:
3890:
3886:
3884:
3883:
3879:
3877:
3876:
3872:
3870:
3869:
3865:
3863:
3862:
3858:
3856:
3855:
3854:Adynomosaurus
3851:
3850:
3847:
3844:
3842:
3838:
3832:
3831:
3827:
3825:
3824:
3820:
3818:
3817:
3816:Lapampasaurus
3813:
3811:
3810:
3806:
3804:
3803:
3799:
3797:
3796:
3792:
3791:
3788:
3785:
3782:
3781:Hadrosauridae
3777:
3771:
3770:
3766:
3764:
3763:
3762:Zhanghenglong
3759:
3757:
3756:
3752:
3750:
3749:
3745:
3743:
3742:
3738:
3736:
3735:
3734:Telmatosaurus
3731:
3729:
3728:
3724:
3722:
3721:
3717:
3715:
3714:
3710:
3708:
3707:
3703:
3701:
3700:
3696:
3694:
3693:
3689:
3687:
3686:
3682:
3680:
3679:
3678:Portellsaurus
3675:
3673:
3672:
3668:
3666:
3665:
3661:
3659:
3658:
3654:
3652:
3651:
3647:
3645:
3644:
3640:
3638:
3637:
3633:
3630:
3629:
3625:
3623:
3622:
3618:
3616:
3615:
3611:
3609:
3608:
3604:
3602:
3601:
3597:
3595:
3594:
3590:
3588:
3587:
3583:
3581:
3580:
3576:
3574:
3573:
3569:
3567:
3566:
3562:
3560:
3559:
3555:
3553:
3552:
3548:
3546:
3545:
3541:
3539:
3538:
3534:
3531:
3530:
3526:
3524:
3523:
3519:
3517:
3516:
3512:
3511:
3508:
3505:
3502:
3497:
3493:
3489:
3484:
3480:
3471:
3467:
3464:
3457:
3424:
3423:
3422:
3421:
3419:
3417:
3413:
3406:
3405:
3401:
3399:
3398:
3394:
3392:
3391:
3387:
3384:
3383:
3379:
3378:
3376:
3374:
3370:
3364:
3363:
3359:
3357:
3356:
3352:
3350:
3349:
3345:
3343:
3342:
3341:Jinzhousaurus
3338:
3336:
3335:
3334:Hypselospinus
3331:
3329:
3328:
3324:
3322:
3321:
3317:
3316:
3313:
3310:
3308:
3304:
3298:
3297:
3293:
3291:
3290:
3289:Theiophytalia
3286:
3284:
3283:
3279:
3277:
3276:
3272:
3270:
3269:
3265:
3263:
3262:
3258:
3256:
3255:
3251:
3249:
3248:
3244:
3242:
3241:
3237:
3235:
3234:
3230:
3228:
3227:
3223:
3221:
3220:
3219:Lanzhousaurus
3216:
3214:
3213:
3209:
3207:
3206:
3202:
3200:
3199:
3195:
3193:
3192:
3188:
3185:
3184:
3180:
3178:
3177:
3173:
3171:
3170:
3166:
3164:
3163:
3159:
3157:
3156:
3152:
3150:
3149:
3145:
3144:
3141:
3138:
3135:
3130:
3120:
3119:
3115:
3112:
3111:
3107:
3104:
3103:
3099:
3097:
3096:
3092:
3090:
3089:
3088:Elrhazosaurus
3085:
3083:
3082:
3078:
3076:
3075:
3071:
3070:
3068:
3066:
3062:
3056:
3055:
3051:
3049:
3048:
3044:
3042:
3041:
3037:
3036:
3033:
3029:
3026:
3023:
3018:
3004:
3003:
2999:
2997:
2996:
2992:
2990:
2989:
2985:
2983:
2982:
2978:
2976:
2975:
2971:
2969:
2968:
2964:
2963:
2961:
2959:
2955:
2949:
2948:
2947:Tenontosaurus
2944:
2942:
2941:
2937:
2934:
2933:
2932:Convolosaurus
2929:
2928:
2926:
2923:
2919:
2912:
2911:
2907:
2904:
2903:
2899:
2896:
2895:
2894:Ampelognathus
2891:
2890:
2887:
2884:
2882:
2878:
2871:
2870:
2866:
2864:
2863:
2859:
2857:
2856:
2852:
2850:
2849:
2845:
2843:
2842:
2838:
2836:
2835:
2831:
2829:
2828:
2824:
2821:
2820:
2816:
2814:
2813:
2809:
2807:
2806:
2805:Mahuidacursor
2802:
2800:
2799:
2795:
2793:
2792:
2788:
2785:
2784:
2780:
2778:
2777:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2756:
2755:
2751:
2749:
2748:
2744:
2742:
2741:
2737:
2735:
2734:
2730:
2728:
2727:
2723:
2722:
2720:
2718:
2714:
2707:
2706:
2702:
2700:
2699:
2698:Hypsilophodon
2695:
2692:
2691:
2687:
2686:
2684:
2681:
2677:
2670:
2669:
2665:
2663:
2662:
2661:Callovosaurus
2658:
2656:
2655:
2654:Burianosaurus
2651:
2650:
2647:
2643:
2640:
2637:
2632:
2628:
2624:
2619:
2615:
2590:
2589:
2588:
2587:
2585:
2583:
2579:
2571:
2567:
2566:
2565:
2564:
2561:
2558:
2556:
2552:
2544:
2540:
2539:
2538:
2537:
2534:
2531:
2529:
2525:
2521:
2513:
2509:
2506:
2504:
2500:
2497:
2495:
2491:
2488:
2486:
2482:
2479:
2477:
2473:
2471:
2467:
2466:
2461:
2457:
2453:
2446:
2441:
2439:
2434:
2432:
2427:
2426:
2423:
2416:
2415:United States
2406:
2404:
2394:
2390:
2385:
2383:
2378:
2370:
2361:
2357:
2352:
2347:
2342:
2337:
2333:
2329:
2325:
2321:
2317:
2310:
2307:
2303:
2297:
2294:
2290:
2286:
2280:
2278:
2276:
2274:
2272:
2270:
2268:
2264:
2259:
2255:
2251:
2247:
2243:
2236:
2233:
2228:
2224:
2220:
2216:
2212:
2208:
2204:
2197:
2194:
2189:
2183:
2179:
2172:
2169:
2163:
2160:
2155:
2149:
2145:
2141:
2134:
2131:
2126:
2122:
2118:
2114:
2110:
2106:
2102:
2098:
2094:
2087:
2084:
2080:(1): 211–273.
2079:
2075:
2071:
2067:
2066:Hypsilophodon
2063:
2057:
2055:
2051:
2046:
2042:
2038:
2034:
2030:
2026:
2019:
2016:
2011:
2007:
2003:
1999:
1992:
1990:
1981:
1979:
1977:
1975:
1973:
1969:
1964:
1960:
1955:
1950:
1945:
1940:
1936:
1932:
1928:
1924:
1920:
1913:
1911:
1909:
1905:
1900:
1896:
1892:
1888:
1884:
1880:
1876:
1869:
1867:
1865:
1863:
1859:
1854:
1848:
1844:
1840:
1833:
1830:
1826:
1820:
1817:
1812:
1808:
1804:
1800:
1796:
1794:
1785:
1783:
1779:
1774:
1770:
1766:
1762:
1758:
1754:
1750:
1746:
1742:
1735:
1732:
1727:
1720:
1717:
1712:
1708:
1707:Palaeontology
1701:
1699:
1697:
1693:
1688:
1684:
1677:
1674:
1669:
1665:
1661:
1657:
1653:
1649:
1645:
1641:
1637:
1630:
1627:
1622:
1618:
1614:
1610:
1606:
1602:
1598:
1594:
1590:
1586:
1582:
1575:
1573:
1571:
1567:
1560:
1556:
1553:
1551:
1548:
1547:
1543:
1541:
1539:
1535:
1527:
1523:
1516:
1509:
1508:
1500:
1499:
1491:
1490:
1482:
1481:
1473:
1472:
1464:
1463:
1455:
1454:
1446:
1445:
1442:
1441:
1435:
1434:
1431:
1430:
1427:
1426:
1425:
1418:
1417:
1411:
1410:
1407:
1406:
1403:
1402:
1401:
1394:
1393:
1387:
1386:
1383:
1382:
1379:
1378:
1377:
1376:Lanzhousaurus
1370:
1369:
1363:
1362:
1359:
1358:
1355:
1354:
1353:
1346:
1345:
1339:
1338:
1335:
1334:
1326:
1325:
1322:
1321:
1320:
1319:Theiophytalia
1313:
1312:
1309:
1308:
1305:
1304:
1303:
1296:
1295:
1289:
1288:
1282:
1281:
1278:
1277:
1274:
1273:
1272:
1265:
1264:
1258:
1257:Styracosterna
1255:
1254:
1251:
1250:
1247:
1246:
1245:
1238:
1237:
1231:
1228:
1227:
1220:
1216:
1211:
1207:
1205:
1201:
1196:
1194:
1193:
1188:
1187:
1182:
1181:
1176:
1172:
1168:
1164:
1161:
1157:
1153:
1152:phylogenetics
1148:
1146:
1142:
1134:
1132:
1130:
1126:
1125:
1119:
1118:
1117:Theiophytalia
1112:
1110:
1106:
1102:
1098:
1094:
1090:
1086:
1085:
1080:
1072:
1065:
1063:
1061:
1057:
1056:
1051:
1047:
1043:
1039:
1035:
1030:
1028:
1024:
1020:
1016:
1012:
1008:
1007:
1002:
998:
994:
990:
986:
981:
979:
978:
973:
969:
965:
963:
958:
954:
953:
952:Callovosaurus
948:
944:
940:
936:
935:
930:
926:
922:
918:
914:
913:C. prestwichi
910:
906:
902:
898:
891:West, Wyoming
890:
885:
881:
879:
878:
873:
872:Styracosterna
869:
865:
861:
857:
853:
849:
848:
843:
839:
835:
831:
830:
825:
821:
817:
813:
807:
803:
802:
801:Theiophytalia
797:
793:
789:
787:
786:
781:
777:
773:
769:
765:
761:
757:
753:
749:
745:
744:C. prestwichi
741:
734:
730:
725:
721:
719:
715:
711:
707:
703:
699:
695:
694:
689:
685:
681:
677:
673:
669:
665:
661:
657:
653:
649:
645:
641:
637:
636:
631:
627:
623:
619:
612:
611:Theiophytalia
607:
600:
598:
596:
594:
582:
580:
569:
565:
561:
560:North America
557:
554:
553:Late Jurassic
550:
547:
546:ornithischian
543:
539:
538:
525:
480:
479:
463:
462:
458:
452:
449:
445:
439:
438:
434:
430:
427:Marsh, 1879 (
424:
421:
417:
412:
409:
408:
406:
403:
399:
392:
386:
385:
378:
371:
370:
363:
362:
360:
357:
352:
345:
344:
339:
336:
332:
328:
322:
321:
314:
311:
310:
303:
297:
294:
293:
286:
280:
278:Superfamily:
277:
276:
273:
267:
264:
261:
260:
257:
251:
248:
245:
244:
241:
235:
232:
229:
228:
225:
219:
216:
213:
212:
209:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
134:
130:
125:
118:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
57:
50:
49:Late Jurassic
44:
40:
37:
33:
19:
4828:Camptosaurus
4798:Camptosaurus
4797:
4762:
4755:
4747:
4735:
4704:
4686:
4679:
4672:
4665:
4658:
4651:
4644:
4637:
4630:
4623:
4616:
4609:
4602:
4595:
4588:
4581:
4575:Claorhynchus
4573:
4566:
4559:
4551:Nomina dubia
4549:
4539:Guegoolithus
4537:
4517:
4510:
4505:Staurichnium
4503:
4498:Sousaichnium
4496:
4489:
4482:
4475:
4468:
4461:
4454:
4449:Dakotasaurus
4447:
4442:Caririchnium
4440:
4433:
4426:
4419:
4412:
4356:
4324:
4317:
4310:
4303:
4296:
4289:
4269:
4262:
4255:
4248:Saurolophini
4235:
4228:
4221:
4214:
4207:
4199:
4192:Kritosaurini
4179:
4172:
4167:Huallasaurus
4165:
4158:
4139:Wulagasaurus
4137:
4130:
4123:
4116:
4109:
4102:
4082:
4054:
4047:
4040:
4033:
4028:Lambeosaurus
4026:
4019:
4012:
4005:
3985:
3978:
3971:
3964:
3944:
3937:
3930:
3923:
3915:
3908:
3901:
3894:
3887:
3880:
3873:
3866:
3859:
3852:
3830:Yamatosaurus
3828:
3821:
3814:
3807:
3800:
3795:Aquilarhinus
3793:
3767:
3760:
3753:
3746:
3741:Tethyshadros
3739:
3732:
3725:
3718:
3711:
3704:
3697:
3690:
3683:
3676:
3671:Plesiohadros
3669:
3662:
3655:
3648:
3643:Lophorhothon
3641:
3634:
3626:
3619:
3612:
3605:
3598:
3591:
3584:
3577:
3570:
3563:
3556:
3549:
3542:
3535:
3529:Batyrosaurus
3527:
3522:Bactrosaurus
3520:
3513:
3459:
3402:
3395:
3388:
3380:
3362:Ouranosaurus
3360:
3353:
3346:
3339:
3332:
3327:Brighstoneus
3325:
3318:
3294:
3287:
3280:
3273:
3266:
3259:
3254:Osmakasaurus
3252:
3247:Oblitosaurus
3245:
3238:
3231:
3224:
3217:
3210:
3203:
3196:
3189:
3181:
3174:
3169:Camptosaurus
3168:
3167:
3160:
3153:
3146:
3116:
3110:Kangnasaurus
3108:
3100:
3093:
3086:
3079:
3072:
3065:Dryosauridae
3052:
3045:
3038:
3000:
2993:
2986:
2979:
2972:
2967:Matheronodon
2965:
2945:
2938:
2930:
2908:
2900:
2892:
2867:
2860:
2853:
2846:
2841:Sektensaurus
2839:
2834:Qantassaurus
2832:
2825:
2817:
2810:
2803:
2796:
2789:
2783:Kangnasaurus
2781:
2774:
2767:
2760:
2752:
2747:Diluvicursor
2745:
2738:
2731:
2724:
2705:Vectidromeus
2703:
2696:
2688:
2666:
2659:
2652:
2570:Ornithischia
2555:Ornithischia
2507:
2498:
2494:Ornithischia
2489:
2480:
2382:Camptosaurus
2374:
2323:
2319:
2309:
2301:
2296:
2288:
2284:
2249:
2245:
2235:
2213:(1): 61–73.
2210:
2206:
2196:
2177:
2171:
2162:
2143:
2139:
2133:
2100:
2096:
2086:
2077:
2073:
2069:
2065:
2028:
2024:
2018:
2001:
1997:
1989:Camptosaurus
1988:
1926:
1922:
1882:
1878:
1875:Camptosaurus
1874:
1842:
1839:Camptosaurus
1838:
1832:
1824:
1819:
1802:
1798:
1793:Camptosaurus
1792:
1748:
1744:
1734:
1725:
1719:
1710:
1706:
1686:
1682:
1676:
1643:
1639:
1629:
1591:(1): 65–78.
1588:
1584:
1538:Camptosaurus
1537:
1534:Iguanodontia
1531:
1525:
1517:Paleobiology
1438:
1422:
1421:
1398:
1397:
1374:
1373:
1350:
1349:
1317:
1316:
1300:
1299:
1269:
1268:
1244:Camptosaurus
1243:
1242:
1241:
1219:Camptosaurus
1218:
1204:Camptosaurus
1203:
1197:
1190:
1184:
1178:
1167:iguanodontid
1156:Camptosaurus
1155:
1149:
1141:Camptosaurus
1140:
1138:
1129:Camptosaurus
1128:
1122:
1115:
1113:
1109:Camptosaurus
1108:
1096:
1093:Camptosaurus
1092:
1082:
1079:Camptosaurus
1078:
1077:
1060:Camptosaurus
1059:
1053:
1050:Camptosaurus
1049:
1037:
1034:Camptosaurus
1033:
1031:
1026:
1022:
1014:
1011:Camptosaurus
1010:
1004:
1000:
989:Cumnor Hurst
984:
982:
975:
972:Camptosaurus
971:
968:Richard Owen
960:
956:
950:
946:
939:C. valdensis
938:
934:nomen dubium
932:
920:
917:C. valdensis
916:
912:
908:
904:
900:
897:Camptosaurus
896:
894:
875:
867:
859:
851:
847:Osmakasaurus
845:
842:C. depressus
841:
837:
833:
827:
811:
810:
799:
796:Camptosaurus
795:
783:
780:Camptosaurus
779:
776:Camptosaurus
775:
771:
767:
763:
759:
755:
751:
747:
743:
740:Peter Galton
737:
732:
728:
718:Camptosaurus
717:
714:C. depressus
713:
709:
701:
697:
691:
687:
684:Arthur Lakes
679:
663:
660:type species
655:
648:Camptosaurus
647:
633:
626:euornithopod
615:
610:
590:
576:
478:Camptosaurus
477:
476:
475:
459:
451:Symphyrophus
450:
435:
422:
411:Brachyrophus
410:
382:
367:
342:
341:
335:Type species
320:Camptosaurus
319:
318:
301:
284:
262:
246:
230:
214:
208:Ornithischia
198:
185:
43:Camptosaurus
42:
36:
4917:Iguanodonts
4822:Wikispecies
4764:Willinakaqe
4736:nomina nuda
4653:Siluosaurus
4618:Loncosaurus
4456:Dinehichnus
4298:Kamuysaurus
4271:Saurolophus
4223:Kritosaurus
4216:Gryposaurus
4049:Sahaliyania
4035:Magnapaulia
4007:Amurosaurus
3987:Tlatolophus
3966:Adelolophus
3910:Kazaklambia
3889:Blasisaurus
3882:Arenysaurus
3875:Aralosaurus
3809:Hadrosaurus
3802:Eotrachodon
3755:Yunganglong
3720:Sirindhorna
3692:Protohadros
3628:Koshisaurus
3621:Jintasaurus
3390:Comptonatus
3240:Napaisaurus
3226:Lurdusaurus
3198:Fukuisaurus
3176:Cedrorestes
3118:Valdosaurus
2981:Pareisactus
2812:Morrosaurus
2776:Isasicursor
2740:Chakisaurus
2636:Ornithopoda
2623:Ornithopoda
2582:Ornithopoda
2452:Ornithopoda
2070:Valdosaurus
1171:hadrosaurid
1091:belongs to
1074:Restoration
1066:Description
993:Oxfordshire
925:Haţeg Basin
834:P. depressa
635:Camptonotus
558:of western
429:preoccupied
423:Camptonotus
348:Marsh, 1879
306:Marsh, 1885
289:Marsh, 1885
240:Ornithopoda
32:Camposaurus
4911:Categories
4706:Arkharavia
4681:Tichosteus
4674:Thespesius
4646:Pteropelyx
4639:Orthomerus
4519:Wintonopus
4104:Acristavus
4084:Barsboldia
4042:Olorotitan
3917:Latirhinus
3769:Zuoyunlong
3636:Levnesovia
3586:Gobihadros
3551:Datonglong
3544:Claosaurus
3515:Altirhinus
3425:see below↓
3355:Morelladon
3233:Magnamanus
3205:Hippodraco
3074:Dryosaurus
3047:Hesperonyx
3022:Dryomorpha
2862:Trinisaura
2855:Tietasaura
2848:Talenkauen
2726:Anabisetia
2591:see below↓
2485:Dinosauria
1713:: 411–443.
1561:References
1400:Kukufeldia
1302:Hippodraco
1215:ornithopod
1180:Dryosaurus
1124:Dryosaurus
693:Allosaurus
668:Como Bluff
455:Cope, 1878
256:Dryomorpha
192:Dinosauria
4688:Trachodon
4604:Glishades
4583:Diclonius
4421:Anomoepus
4405:Ichnotaxa
4118:Maiasaura
4056:Velafrons
3565:Equijubus
3537:Choyrodon
3397:Iguanodon
3261:Planicoxa
3191:Dakotadon
3162:Calvarius
2988:Rhabdodon
2974:Mochlodon
2717:Elasmaria
2468:Kingdom:
2403:Dinosaurs
2125:1477-2019
2004:: 52–68.
1773:130777820
1668:131001110
1621:245804125
1613:1698-6180
1200:cladistic
1154:has made
1105:Oxfordian
1101:Callovian
962:Iguanodon
957:C. hoggii
947:C. leedsi
921:C. inkeyi
909:C. leedsi
905:C. hoggii
901:C. inkeyi
860:C. dispar
829:Planicoxa
772:C. dispar
768:C. amplus
764:C. dispar
760:C. dispar
756:C. browni
752:C. medius
733:C. dispar
710:C. browni
698:C. medius
688:Quarry 1A
680:C. amplus
664:Quarry 13
656:C. dispar
549:dinosaurs
165:Kingdom:
159:Eukaryota
4842:52571997
4807:Wikidata
4568:Cionodon
4237:Rhinorex
4125:Ornatops
3925:Minqaria
3896:Canardia
3823:Malefica
3748:Xuwulong
3713:Siamodon
3614:Jeyawati
3600:Gonkoken
3558:Eolambia
3382:Barilium
3183:Cumnoria
3148:Barilium
3054:Owenodon
3040:Draconyx
3002:Zalmoxes
2902:Fostoria
2754:Fostoria
2512:Cerapoda
2476:Chordata
2474:Phylum:
2470:Animalia
2360:21124919
2320:PLOS ONE
1963:21124919
1923:PLOS ONE
1899:86197297
1544:See also
1424:Barilium
1055:Draconyx
1052:than to
1015:Cumnoria
1006:Cumnoria
977:Owenodon
748:C. nanus
729:C. nanus
702:C. nanus
644:holotype
437:Cumnoria
402:Synonyms
295:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
4881:4946706
4868:1421899
4855:3238881
4813:Q133203
3861:Ajnabia
3296:Uteodon
2389:Portals
2351:2989904
2328:Bibcode
2215:Bibcode
2105:Bibcode
2033:Bibcode
2025:Geobios
1998:Zootaxa
1954:2989904
1931:Bibcode
1753:Bibcode
1648:Bibcode
1593:Bibcode
1271:Uteodon
1186:Drinker
1175:derived
1150:Modern
1084:Uteodon
1062:genus.
1027:Uteodon
997:tramway
943:dubious
929:Romania
877:Uteodon
864:ischium
804:skull,
674:in the
672:Wyoming
652:cricket
579:kamptos
572:καμπτος
551:of the
540:) is a
461:Uteodon
393:, 1880
356:species
325:Marsh,
312:Genus:
4530:Ootaxa
3727:Tanius
3320:Bolong
2358:
2348:
2184:
2150:
2123:
1961:
1951:
1897:
1849:
1771:
1689:: 169.
1666:
1619:
1611:
1189:, and
1038:et al.
964:hoggii
915:, and
642:. The
593:sauros
586:σαυρος
564:Europe
556:period
446:, 1888
444:Seeley
418:, 1878
354:Other
4894:38748
4863:IRMNG
3572:Fylax
3102:Iyuku
2508:Clade
2499:Clade
2490:Clade
2481:Clade
2103:(1).
1994:(PDF)
1895:S2CID
1769:S2CID
1747:. 3.
1664:S2CID
1617:S2CID
1160:clade
1046:Spain
832:, as
666:near
568:Greek
542:genus
391:Hulke
263:Clade
247:Clade
231:Clade
215:Clade
199:Clade
186:Clade
4850:GBIF
3268:Proa
2940:Iani
2568:see
2541:see
2356:PMID
2182:ISBN
2148:ISBN
2121:ISSN
2002:2783
1959:PMID
1847:ISBN
1609:ISSN
1169:and
754:and
712:and
700:and
678:and
533:-tə-
531:KAMP
416:Cope
327:1885
62:PreꞒ
4837:EoL
2346:PMC
2336:doi
2287:."
2254:doi
2250:199
2223:doi
2113:doi
2041:doi
2006:doi
1949:PMC
1939:doi
1887:doi
1807:doi
1761:doi
1656:doi
1601:doi
1044:in
991:in
966:by
937:).
927:in
686:at
620:in
537:-əs
535:SOR
515:ɔːr
4913::
4891::
4878::
4865::
4852::
4839::
4824::
4809::
2510::
2501::
2492::
2483::
2354:.
2344:.
2334:.
2322:.
2318:.
2266:^
2248:.
2244:.
2221:.
2211:87
2209:.
2205:.
2119:.
2111:.
2101:22
2099:.
2095:.
2078:28
2076:.
2068:,
2053:^
2039:.
2029:39
2027:.
2000:.
1996:.
1971:^
1957:.
1947:.
1937:.
1925:.
1921:.
1907:^
1893:.
1883:76
1881:.
1861:^
1803:36
1801:.
1797:.
1781:^
1767:.
1759:.
1749:48
1743:.
1711:23
1709:.
1695:^
1687:29
1685:.
1662:.
1654:.
1644:18
1642:.
1638:.
1615:.
1607:.
1599:.
1589:48
1587:.
1583:.
1569:^
1195:.
1183:,
1147:.
1127:.
1029:.
980:.
955:.
919:.
911:,
907:,
903:,
880:.
788:.
750:,
670:,
387:?
372:?
265::
249::
233::
217::
201::
188::
112:Pg
56:Ma
51:,
4738:)
4578:?
4204:?
4142:?
3920:?
3631:?
3532:?
3407:?
3385:?
3186:?
3113:?
3105:?
2935:?
2924:?
2913:?
2905:?
2897:?
2872:?
2822:?
2786:?
2757:?
2708:?
2693:?
2682:?
2671:?
2444:e
2437:t
2430:v
2391::
2362:.
2338::
2330::
2324:5
2260:.
2256::
2229:.
2225::
2217::
2190:.
2156:.
2127:.
2115::
2107::
2047:.
2043::
2035::
2012:.
2008::
1965:.
1941::
1933::
1927:5
1901:.
1889::
1855:.
1813:.
1809::
1775:.
1763::
1755::
1670:.
1658::
1650::
1623:.
1603::
1595::
1103:-
658:(
589:(
575:(
524:/
521:s
518:ə
512:s
509:ˈ
506:ə
503:t
500:p
497:m
494:æ
491:k
488:ˌ
485:/
481:(
464:?
440:?
431:)
380:†
365:†
316:†
299:†
282:†
269:†
253:†
237:†
221:†
205:†
117:N
107:K
102:J
97:T
92:P
87:C
82:D
77:S
72:O
67:Ꞓ
34:.
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
