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Comephorus

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462:, at more than 90% of its diet. They are easily identifiable, and are large enough to be easily seen. Golomyankas are unusual for occurring pelagically throughout the entire water column of Lake Baikal, ranging from depths of around 1.6 km (1.0 mi) to near the surface, but they primarily occur deeper than 100 m (330 ft). During the night they move up the water column to feed, often occurring as shallow as 10–25 m (33–82 ft), and in the winter they can occasionally be seen swimming just below the ice-covered surface. They are considered the world's most abyssal freshwater fish, together with certain Lake Baikal 487:, giving birth to a swarm of 2000–3000 larvae. Females containing developing embryos can be seen year-round, but there does appear to be a level of seasonality in the birth, with the largest number of larvae occurring from August to April (some differences in the peak timing between the two species). Most females die after giving birth. It has been claimed that their bellies burst open to release the young, killing the female in the process, but this is a myth. Golomyanka can reach an age of up to 6–8 years. 66: 525: 44: 53: 482:
are not known for this species, although groups of up to about 20 individuals have been recorded near the lake bottom. Females are more common than males, which only make up about 32% of the population in the little Baikal oilfish and 17% in the big Baikal oilfish. The females do not lay eggs;
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by native Siberians when storms tossed the fish up on shore. They are of no value as a food source for humans, dogs or cats, but are the primary food source for the Baikal seal, and also eaten by other fish in the lake. They are so numerous and spawn so rapidly that they represent the largest
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of the golomyanka population is estimated at about 150 thousand tons, making it the most populous fish in Lake Baikal. It is estimated that about 70% of the fish in the lake are golomyankas. Their juveniles are also the most abundant pelagic
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Despite being numerous in the lake, golomyankas are difficult to catch in large quantities as they do not gather in large schools. Golomyankas are not harvested commercially, although their fats and oils were used in lamps and as
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W. Leo Smith & Morgan S. Busby (2014). "Phylogeny and taxonomy of sculpins, sandfishes, and snailfishes (Perciformes: Cottoidei) with comments on the phylogenetic significance of their early-life-history specializations".
466:(Abyssocottidae). They move without much regard for changes in pressure, although they can exist only in cold temperatures, preferring water that is no more than 5 °C (41 °F) and dying at 10 °C (50 °F). 1067:
Jakubowski, Tugarina, and Żuwała (2003). Pectoral fin development in the Baikalian viviparous golomyankas (Comephoridae; Cottoidei), with a remark on eggs and embryos of Comephorus baicalensis (Pallas). J.Anat. 203(3):
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They are sluggish fish, and have relatively large mouths with several rows of somewhat brush-like teeth, allowing them to sift out fine organisms from the water. They primarily feed on the planktonic copepod
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Miyasaka, Dzyuba, Genkai-Kato and Wada (2006). Feeding ecology of two planktonic sculpins, Comephorus baicalensis and Comephorus dybowskii (Comephoridae), in Lake Baikal. Ichthyological Research 53(4):
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Dzyuba, E.V. (2006). Two coexisting species of Baikal golomyankas, Comephorus baicalensis and C. dybowski: seasonal dynamics of juveniles and their feeding. Hydrobiologia 568, Supplement 1: 111–114.
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Sideleva, V.G. (1996). Comparative character of the deep-water and inshore cottoid fishes endemic to Lake Baikal. Journal of Fish Biology 49(sA): 192–206.
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concentration of fish biomass within the entire lake, and would seriously unbalance the ecosystem of Lake Baikal if not constantly preyed upon by
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in sunlight, leaving behind only fatty oil and bones, although the accuracy of this claim has been questioned. Unusually, their eyes only contain
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A few golomyankas (for example, the pinkish fish on the upper-center part of the photo and another in the lower-left), as well as several
923: 951: 1085: 264: 161: 1337: 1284: 355:. The lateral line system on the head consists of large cavities linked by narrow, bony bridges with small external pores. High 1322: 1045:
Hunt, D. M., et al. (1997). Molecular evolution of the cottoid fish endemic to Lake Baikal deduced from nuclear DNA evidence.
603: 1240: 1342: 1157: 625: 65: 1253: 428: 1327: 1097: 831: 761: 479: 408: 1162: 567: 513: 493: 391:), indicating a high sensitivity to light. Some of the adaptions in golomyankas resemble those seen in 292:
studies which have found that Baikal sculpins that were classified in the subfamilies Comephorinae and
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have translucent bodies with no scales, but appear dull when dead. The big Baikal oilfish (
475: 278: 206: 117: 524: 293: 107: 43: 847: 333:) up to 16 cm (6.3 in). Males are smaller than females, only reaching about 1306: 985: 976: 947: 919: 877: 757: 392: 380: 302: 289: 329:) can reach up to 21 cm (8.3 in) in length and the little Baikal oilfish ( 1149: 368: 364: 352: 344: 311: 170: 1175: 836:
Comparative Biochemistry and Physiology Part B: Biochemistry and Molecular Biology
1209: 1129: 676: 459: 458:, which also eat small golomyankas, and represent a primary food source for the 241: 222: 367:
allows the fish to tolerate varying pressure extremes as they move through the
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and that the classification of the Baikal sculpins in a different taxon from
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from Baikal Web World, accessed May 5, 2006 (with a photo of the golomyanka).
529: 388: 230: 77: 17: 855: 684: 450:, meaning "far from the shore" in the local dialect and referring to their 1114: 960: 932: 770: 641: 555: 537: 498: 384: 285: 218: 127: 97: 706: 1201: 509: 470: 451: 237: 214: 210: 52: 1214: 87: 1091: 898:"Baikal "shark": Amazing fact about the Baikal oilfish (golomyanka)" 523: 356: 300:
radiated from an ancestor which was likely to be within the genus
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Irkutsk State University Lake Baikal FAQ, accessed May 5, 2006
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behavior. They are the principal ecological competitor to the
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There are currently two recognized species in this genus:
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contains far less (4.7%). It has been said that they
288:, the typical sculpins. Other authorities have used 1019: 1017: 624:
J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016).
1104: 830:Kozlova, T. A.; Khotimchenko, S. V. (August 2000). 791: 789: 787: 785: 783: 781: 709:
from Irkutsk State University, accessed May 5, 2006
347:, and although pelvic bones are present, they lack 592:; Fricke, Ron & van der Laan, Richard (eds.). 1000:10.2305/IUCN.UK.2016-1.RLTS.T41676A45231738.en 892: 890: 888: 886: 371:. Over a third of the body weight (38.9%) of 359:content and porous bones result in an almost 8: 1027:(in Russian). Fishwatching. 21 January 2016 799:(in Russian). Fishwatching. 29 January 2016 1092: 816: 814: 797:"ГОЛОМЯНКИ, ЧАСТЬ 2: ОСОБЕННОСТИ СТРОЕНИЯ" 752: 750: 51: 42: 31: 1333:Taxa named by Bernard Germain de Lacépède 998: 702: 700: 698: 696: 694: 630:(5th ed.). Wiley. pp. 467–495. 271:as its only species. The 5th edition of 240:fishes and the main food source for the 735:. Science First Hand. 30 September 2004 578: 873: 871: 869: 867: 865: 727: 725: 723: 721: 719: 717: 715: 584: 582: 1047:Molecular Phylogenetics and Evolution 665:Molecular Phylogenetics and Evolution 7: 1025:"ГОЛОМЯНКИ, ЧАСТЬ 1: ЭКОЛОГИЯ ВИДОВ" 986:IUCN Red List of Threatened Species 281:subfamily Comephorinae within the 25: 707:Animals and plants of Lake Baikal 733:"Mysterious Fish of Lake Baikal" 446:originates from the old-Russian 64: 594:"Genera in the family Cottidae" 604:California Academy of Sciences 1: 848:10.1016/S0305-0491(00)00207-8 900:. Key to Baikal. 19 May 2017 878:Ichthyofauna of Lake Baikal 677:10.1016/j.ympev.2014.06.028 343:the length. They have long 265:Bernard Germain de Lacépède 1359: 964:. September 2016 version. 936:. September 2016 version. 277:places this genus in the 229:is the only genus in the 176: 169: 61:Scientific classification 59: 50: 41: 34: 950:; Pauly, Daniel (eds.). 922:; Pauly, Daniel (eds.). 774:. December 2012 version. 760:; Pauly, Daniel (eds.). 255:was first proposed as a 1338:Endemic fauna of Russia 433:(Little Baikal oilfish) 269:Callionymus baikalensis 178:Callionymus baikalensis 1323:Scorpaeniformes genera 926:Comephorus baikalensis 545: 520:Relationship to humans 409:Comephorus baikalensis 259:in 1800 by the French 993:: e.T41676A45231738. 590:Eschmeyer, William N. 568:List of fish families 527: 512:, including those of 494:Epischura baikalensis 351:. They have a strong 975:Goodman, S. (2016). 954:Comephorus dybowskii 551:traditional medicine 424:Comephorus dybowskii 419:(Big Baikal oilfish) 1343:Fish of Lake Baikal 627:Fishes of the World 478:in the lake. Large 464:deep-water sculpins 438:Biology and ecology 298:Fishes of the World 274:Fishes of the World 762:"Species in genus 546: 257:monospecific genus 236:. Golomyankas are 1300: 1299: 1262:Open Tree of Life 1098:Taxon identifiers 637:978-1-118-34233-6 599:Catalog of Fishes 534:Baikal yellowfins 514:their own species 483:rather, they are 432: 418: 190: 189: 165: 146: 16:(Redirected from 1350: 1293: 1292: 1280: 1279: 1270: 1269: 1257: 1256: 1244: 1243: 1231: 1230: 1218: 1217: 1205: 1204: 1192: 1191: 1179: 1178: 1166: 1165: 1153: 1152: 1140: 1139: 1138: 1125: 1124: 1123: 1093: 1079: 1075: 1069: 1065: 1059: 1056: 1050: 1043: 1037: 1036: 1034: 1032: 1021: 1012: 1011: 1009: 1007: 1002: 972: 966: 965: 944: 938: 937: 916: 910: 909: 907: 905: 894: 881: 875: 860: 859: 827: 821: 818: 809: 808: 806: 804: 793: 776: 775: 754: 745: 744: 742: 740: 729: 710: 704: 689: 688: 659: 653: 652: 650: 649: 640:. Archived from 621: 615: 614: 612: 610: 586: 442:The common name 427: 412: 361:neutral buoyancy 342: 341: 337: 160: 141: 69: 68: 55: 46: 32: 21: 1358: 1357: 1353: 1352: 1351: 1349: 1348: 1347: 1328:Viviparous fish 1303: 1302: 1301: 1296: 1288: 1283: 1275: 1273: 1265: 1260: 1252: 1247: 1239: 1234: 1226: 1221: 1213: 1208: 1200: 1195: 1187: 1182: 1174: 1169: 1161: 1156: 1148: 1143: 1134: 1133: 1128: 1119: 1118: 1113: 1100: 1086:Lake Baikal FAQ 1082: 1076: 1072: 1066: 1062: 1057: 1053: 1044: 1040: 1030: 1028: 1023: 1022: 1015: 1005: 1003: 974: 973: 969: 946: 945: 941: 918: 917: 913: 903: 901: 896: 895: 884: 876: 863: 829: 828: 824: 819: 812: 802: 800: 795: 794: 779: 756: 755: 748: 738: 736: 731: 730: 713: 705: 692: 661: 660: 656: 647: 645: 638: 623: 622: 618: 608: 606: 588: 587: 580: 576: 564: 522: 440: 401: 339: 335: 334: 320: 263:and politician 250: 209:comprising two 197:, known as the 186: 180: 159: 140: 118:Scorpaeniformes 63: 28: 27:Genus of fishes 23: 22: 15: 12: 11: 5: 1356: 1354: 1346: 1345: 1340: 1335: 1330: 1325: 1320: 1315: 1305: 1304: 1298: 1297: 1295: 1294: 1281: 1271: 1258: 1245: 1232: 1219: 1206: 1193: 1180: 1167: 1154: 1141: 1126: 1110: 1108: 1102: 1101: 1096: 1090: 1089: 1081: 1080: 1070: 1060: 1051: 1038: 1013: 967: 948:Froese, Rainer 939: 920:Froese, Rainer 911: 882: 861: 842:(4): 477–485. 822: 810: 777: 758:Froese, Rainer 746: 711: 690: 654: 636: 616: 577: 575: 572: 571: 570: 563: 560: 540:caught during 521: 518: 508:and larvae of 439: 436: 435: 434: 420: 400: 397: 373:C. baikalensis 327:C. baikalensis 319: 316: 294:Abyssocottinae 249: 246: 203:Baikal oilfish 188: 187: 181: 174: 173: 167: 166: 152: 148: 147: 135: 131: 130: 125: 121: 120: 115: 111: 110: 108:Actinopterygii 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 57: 56: 48: 47: 39: 38: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 1355: 1344: 1341: 1339: 1336: 1334: 1331: 1329: 1326: 1324: 1321: 1319: 1316: 1314: 1311: 1310: 1308: 1291: 1286: 1282: 1278: 1272: 1268: 1263: 1259: 1255: 1250: 1246: 1242: 1237: 1233: 1229: 1224: 1220: 1216: 1211: 1207: 1203: 1198: 1194: 1190: 1185: 1181: 1177: 1172: 1168: 1164: 1159: 1155: 1151: 1146: 1142: 1137: 1131: 1127: 1122: 1116: 1112: 1111: 1109: 1107: 1103: 1099: 1094: 1087: 1084: 1083: 1074: 1071: 1064: 1061: 1055: 1052: 1049:8(3), 415-22. 1048: 1042: 1039: 1026: 1020: 1018: 1014: 1001: 996: 992: 988: 987: 982: 980: 979:Pusa sibirica 971: 968: 963: 962: 957: 955: 949: 943: 940: 935: 934: 929: 927: 921: 915: 912: 899: 893: 891: 889: 887: 883: 879: 874: 872: 870: 868: 866: 862: 857: 853: 849: 845: 841: 837: 833: 826: 823: 817: 815: 811: 798: 792: 790: 788: 786: 784: 782: 778: 773: 772: 767: 765: 759: 753: 751: 747: 734: 728: 726: 724: 722: 720: 718: 716: 712: 708: 703: 701: 699: 697: 695: 691: 686: 682: 678: 674: 670: 666: 658: 655: 644:on 2022-06-01 643: 639: 633: 629: 628: 620: 617: 605: 601: 600: 595: 591: 585: 583: 579: 573: 569: 566: 565: 561: 559: 557: 552: 543: 539: 535: 531: 526: 519: 517: 515: 511: 507: 505: 504:Macrohectopus 500: 496: 495: 488: 486: 481: 477: 472: 467: 465: 461: 457: 453: 449: 445: 437: 430: 426: 425: 421: 416: 411: 410: 406: 405: 404: 398: 396: 394: 393:deep sea fish 390: 386: 382: 378: 374: 370: 366: 362: 358: 354: 350: 346: 345:pectoral fins 332: 328: 324: 317: 315: 313: 309: 305: 304: 299: 295: 291: 287: 284: 280: 276: 275: 270: 266: 262: 258: 254: 247: 245: 243: 239: 235: 232: 228: 224: 220: 216: 213:of peculiar, 212: 208: 204: 200: 196: 195: 184: 179: 175: 172: 168: 163: 158: 157: 153: 150: 149: 144: 139: 136: 133: 132: 129: 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 89: 86: 83: 82: 79: 76: 73: 72: 67: 62: 58: 54: 49: 45: 40: 37: 33: 30: 19: 1105: 1073: 1063: 1054: 1046: 1041: 1029:. Retrieved 1004:. Retrieved 990: 984: 978: 970: 959: 953: 942: 931: 925: 914: 902:. Retrieved 839: 835: 825: 801:. Retrieved 769: 763: 737:. Retrieved 668: 664: 657: 646:. Retrieved 642:the original 626: 619: 607:. Retrieved 597: 547: 502: 492: 489: 468: 447: 443: 441: 422: 407: 402: 377:C. dybowskii 376: 375:is oil, but 372: 369:water column 365:swim bladder 363:and lack of 353:lateral line 331:C. dybowskii 330: 326: 322: 321: 312:paraphyletic 307: 301: 297: 290:phylogenetic 272: 268: 252: 251: 234:Comephorinae 233: 226: 202: 198: 193: 192: 191: 177: 171:Type species 155: 154: 138:Comephorinae 137: 35: 29: 18:Comephorinae 1210:iNaturalist 1130:Wikispecies 1006:13 November 671:: 332–352. 544:using a net 501:crustacean 476:fish larvae 460:Baikal seal 349:pelvic fins 318:Description 242:Baikal seal 225:in Russia. 223:Lake Baikal 199:golomyankas 134:Subfamily: 1313:Comephorus 1307:Categories 1150:Comephorus 1136:Comephorus 1106:Comephorus 764:Comephorus 648:2023-03-04 574:References 542:icefishing 485:viviparous 444:golomyanka 389:cone cells 323:Comephorus 261:naturalist 253:Comephorus 227:Comephorus 194:Comephorus 156:Comephorus 36:Comephorus 1318:Cottoidei 556:predators 538:amphipods 530:whitefish 506:branickii 429:Korotneff 385:rod cells 381:decompose 279:monotypic 231:subfamily 84:Kingdom: 78:Eukaryota 1115:Wikidata 1078:419-422. 1068:317-322. 961:FishBase 933:FishBase 856:11026659 771:FishBase 685:25014569 562:See also 510:sculpins 499:amphipod 286:Cottidae 248:Taxonomy 205:, are a 162:Lacepède 128:Cottidae 124:Family: 98:Chordata 94:Phylum: 88:Animalia 74:Domain: 1277:4122801 1228:1314784 1202:2334856 1121:Q803775 1031:5 March 803:5 March 609:4 March 471:biomass 452:pelagic 448:golomen 417:, 1776) 399:Species 338:⁄ 238:pelagic 219:endemic 217:fishes 215:sculpin 211:species 151:Genus: 143:Günther 114:Order: 104:Class: 1290:268821 1274:uBio: 1267:231446 1241:167400 1163:498738 904:31 May 854:  739:31 May 683:  634:  497:, the 480:shoals 431:, 1904 415:Pallas 308:Cottus 303:Cottus 283:family 185:, 1776 183:Pallas 164:, 1800 145:, 1861 1285:WoRMS 1254:61640 1223:IRMNG 1215:87581 1189:45558 387:(not 357:lipid 267:with 207:genus 1249:NCBI 1236:ITIS 1197:GBIF 1176:3SRM 1158:BOLD 1033:2018 1008:2021 991:2016 906:2017 852:PMID 805:2018 741:2017 681:PMID 632:ISBN 611:2023 536:and 469:The 456:omul 310:was 1184:EoL 1171:CoL 1145:ADW 995:doi 844:doi 840:126 673:doi 314:. 296:by 221:to 201:or 1309:: 1287:: 1264:: 1251:: 1238:: 1225:: 1212:: 1199:: 1186:: 1173:: 1160:: 1147:: 1132:: 1117:: 1016:^ 989:. 983:. 958:. 930:. 885:^ 864:^ 850:. 838:. 834:. 813:^ 780:^ 768:. 749:^ 714:^ 693:^ 679:. 669:79 667:. 602:. 596:. 581:^ 558:. 532:, 516:. 395:. 244:. 1035:. 1010:. 997:: 981:" 977:" 956:" 952:" 928:" 924:" 908:. 858:. 846:: 807:. 766:" 743:. 687:. 675:: 651:. 613:. 413:( 340:3 336:2 20:)

Index

Comephorinae


Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Actinopterygii
Scorpaeniformes
Cottidae
Comephorinae
Günther
Comephorus
Lacepède
Type species
Pallas
genus
species
sculpin
endemic
Lake Baikal
subfamily
pelagic
Baikal seal
monospecific genus
naturalist
Bernard Germain de Lacépède
Fishes of the World
monotypic

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