124:
3256:
507:
3245:
3251:
145:
3262:
728:
3285:
1571:(also analyzed in the study) likely included bone in their diet and were better adapted to handle the stresses associated with attacking struggling prey while troodontids, equipped with weaker jaws, preyed on softer animals and prey items such as invertebrates and carrion. This feeding strategy and ability to handle struggling prey was also a feature that the theropod also shared with tyrannosaurids such as
1482:
808:
1589:
1182:
1562:
likely preferred larger prey items than the troodontids it shared their environment with. Such differentiations in its diet likely allowed the theropod to inhabit the same environment as its more distant maniraptoran relatives. The same study also indicated that both
2061:
Choiniere, Jonah N.; Neenan, James M.; Schmitz, Lars; Ford, David P.; Chapelle, Kimberley E. J.; Balanoff, Amy M.; Sipla, Justin S.; Georgi, Justin A.; Walsh, Stig A.; Norell, Mark A.; Xu, Xing; Clark, James M.; Benson, Roger B. J. (7 May 2021).
1811:
Flaig, Peter P., Stephen T. Hasiotis, and
Anthony R. Fiorillo. "A paleopolar dinosaur track site in the Cretaceous (Maastrichtian) Prince Creek Formation of Arctic Alaska: Track characteristics and probable trackmakers." Ichnos 25.2-3 (2018):
1821:
Brown, Caleb
Marshall, et al. "Evidence for taphonomic size bias in the Dinosaur Park Formation (Campanian, Alberta), a model Mesozoic terrestrial alluvial‐paralic system." Palaeogeography, palaeoclimatology, palaeoecology 372 (2013):
745:, about 2 m (6 ft 7 in) in length and 16 kg (35 lb) in body mass. Its mouth was full of sharp teeth, and it probably would have had a sharply curved "sickle claw" on each foot. It lived during the
1549:
feeding habits were also discovered to be typical of coelurosaurian theropods, with a characteristic "puncture and pull" feeding method. Studies of wear patterns on the teeth of this animal by
Angelica Torices
775:
had a relatively robust skull with a deep snout. Its teeth were rather large and were shaped like a curved cone with a coat of enamel covering the crown. It had only nine teeth in each
571:
and some foot bones. The skull lacked most of the top of the snout. Several other skull fragments, and about thirty isolated teeth, are known from subsequent discoveries in
Alberta and
2379:
Therrien, Francois; Henderson, Donald M.; Ruff, Christopher B. (2005). "Bite Me: Biomechanical models of theropod mandibles and implications for feeding". In
Carpenter, Kenneth (ed.).
2168:
3419:
1656:"Comparative Taphonomy and Paleoecological Reconstruction of Two Microvertebrate Accumulations from the Late Cretaceous Hell Creek Formation (Maastrichtian), Eastern Montana"
1496:
differs from most of its relatives in having a short, massive skull, a deep mandible, and robust teeth. The teeth tend to be more heavily worn than those of its relative
3434:
514:
Despite receiving widespread attention in popular books on dinosaurs, and the usage of a complete mounted skeleton cast in museums throughout the world,
2438:
3365:
1554:
in a study regarding theropod feeding habits indicate that dromaeosaurid teeth share similar wear patterns to those seen in the
Tyrannosauridae and
873:
are somewhat unclear. Although its rugged build gives it a primitive appearance, it was actually a very specialized animal. In an analysis of the
3429:
3439:
709:
than other small theropods, although it was the first dromaeosaurid of which reasonably good cranial material was described. The genus
3278:
2200:
1796:
1602:
2410:
2392:
2348:
2253:
2234:
1737:
1631:
3444:
3454:
3255:
3244:
1514:
and suggested it relied more on its jaws than on the sickle claw to kill its prey. In a study predominantly centered around
1955:"New Dromaeosaurids (Dinosauria: Theropoda) from the Lower Cretaceous of Utah, and the Evolution of the Dromaeosaurid Tail"
2431:
3261:
144:
3250:
691:). Most of them were based on fragmentary material, some belonging to other genera, and far less complete than that of
681:
451:
123:
3449:
1502:, suggesting that its jaws were used for crushing and tearing rather than simply slicing through flesh. Therrien
132:
2535:
2424:
2384:
538:
753:. However, some fragmentary remains such as teeth that may belong to this genus have been found from the late
537:
on behalf of the
American Museum of Natural History. The area where these bones were collected is now part of
522:
was only made possible by knowledge gained from other dromaeosaurids that have been discovered more recently.
3424:
1715:
647:
623:
455:
3305:
2274:
579:
459:
435:
3391:
3288:
667:
1621:
815:
506:
3352:
2589:
2075:
2015:
DePalma, Robert A.; Burnham, David A.; Martin, Larry D.; Larson, Peter L.; Bakker, Robert T. (2015).
1966:
1670:
758:
447:
443:
3006:
2336:
2173:
792:
2266:(Theropoda: Allosauroidea) and its implications for theropod brain evolution. pp. 19–33. In:
3343:
2743:
2562:
2143:
2107:
2034:
1914:
1870:
1686:
139:
2169:"Scratches on dinosaur teeth reveal their fierce, efficient eating habits - The Washington Post"
1953:
Senter, P.; Kirkland, J. I.; Deblieux, D. D.; Madsen, S.; Toth, N. (2012). Dodson, Peter (ed.).
1704:
3396:
2281:(1922). "The family Deinodontidae, with notice of a new genus from the Cretaceous of Alberta".
3378:
3203:
3094:
3030:
2926:
2719:
2406:
2388:
2344:
2340:
2327:
2249:
2230:
2222:
2099:
2091:
1994:
1862:
1792:
1733:
1627:
1498:
1234:
1193:
The cladogram below follows a 2015 analysis by paleontologists Robert DePalma, David
Burnham,
3383:
1897:
Senter, Phil (2007). "A new look at the phylogeny of coelurosauria (Dlnosauria: Theropoda)".
3127:
3073:
2933:
2911:
2367:
2290:
2209:
2191:
2125:
2083:
2024:
1984:
1974:
1906:
1854:
1678:
1594:
964:
796:
315:
727:
3187:
3041:
2944:
2657:
2635:
2601:
2520:
2447:
2322:
1271:
1220:
1050:
937:
859:
835:
831:
766:
762:
750:
475:
471:
405:
395:
257:
244:
228:
30:
2079:
1970:
1674:
2845:
1989:
1954:
534:
519:
425:
2063:
1655:
3413:
3166:
2972:
2957:
2896:
2889:
2882:
2872:
2807:
2800:
2729:
2111:
1858:
1690:
1623:
Dinosaurs: The Most
Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
1202:
754:
711:
672:
413:
113:
76:
38:
2038:
1918:
1874:
1842:
3225:
3136:
3057:
3050:
3015:
2994:
2979:
2918:
2785:
2778:
2771:
2761:
2708:
2693:
2278:
2213:
2017:"The First Giant Raptor (Theropoda: Dromaeosauridae) from the Hell Creek Formation"
1555:
1381:
1333:
1309:
1258:
1198:
1194:
1088:
1019:
900:
885:
848:
697:
583:
530:
494:
484:
439:
283:
2016:
1481:
492:
was heavily built, more so than other dromaeosaurs that are similar in size, like
1979:
3337:
3180:
3158:
3109:
3065:
3022:
2987:
2965:
2904:
2815:
2750:
2574:
2302:
1573:
1285:
1139:
1064:
951:
891:
854:
839:
518:
is poorly known from actual fossils. The preparation of the popular cast by the
51:
1682:
838:) based on some similarities in the general proportions of the skull. In 1969,
3173:
3150:
3102:
2853:
2822:
2700:
2678:
2486:
2477:
2270:. Ed.s Tanke, D. H., Carpenter, K., Skrepnick, M. W. Indiana University Press.
2029:
1910:
1584:
1357:
1122:
879:
819:
568:
467:
202:
96:
61:
3328:
2095:
791:, that drained the front neck muscles through two long canals running to the
3211:
3143:
2838:
2830:
2736:
2547:
2495:
2223:"Theropod teeth from the Judith River Formation of southern Alberta, Canada"
2087:
1526:
978:
746:
742:
687:
409:
215:
156:
101:
45:
34:
2103:
1998:
1866:
1577:, which was also analyzed in said study alongside these smaller theropods.
807:
1181:
3322:
3195:
3116:
3087:
2861:
2671:
2471:
1516:
560:
542:
401:
398:
189:
176:
91:
86:
71:
66:
56:
1205:, using updated data from the Theropod Working Group. In this analysis,
3370:
3218:
2504:
1002:
776:
706:
653:
631:
572:
417:
106:
81:
2793:
2685:
2465:
2371:
2294:
1709:
907:, but more recent analysis do not recover such a close relationship.
421:
166:
3299:
765:, dating to 66 million years ago. Teeth have also been found in the
2401:
Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.):
2358:
Colbert, E.; Russell, D. A. (1969). "The small
Cretaceous dinosaur
2416:
2064:"Evolution of vision and hearing modalities in theropod dinosaurs"
1480:
1180:
874:
806:
564:
505:
391:
799:
of Dromaeosaurus is rather shallow and does not have much depth.
3357:
1558:, respectively. However, micro-wear on the teeth indicated that
784:
3303:
2458:
2420:
2405:, 2nd, Berkeley: University of California Press. 861 pp.
2309:, an unusual theropod from the Lower Cretaceous of Montana".
2194:(1995). "New information on the anatomy and relationships of
616:
609:
602:
595:
2144:"Dinosaurs' tooth wear sheds light on their predatory lives"
478:, which include many genera with similar characteristics to
345:
2328:"The Wonderful and Spectacular Dromaeosaurian Sickle-Claws"
2229:. Cambridge: Cambridge University Press. pp. 107–125.
366:
360:
348:
336:
333:
327:
695:, and those that haven't been reclassified are considered
1847:
Biological Reviews of the Cambridge Philosophical Society
458:. Teeth attributed to this genus have been found in the
1843:"The evolution of dinosaur tooth enamel microstructure"
2221:
Currie, P. J.; Rigby, K. J.; Sloan, Robert E. (1990).
1510:
had a bite nearly three times as powerful as that of
351:
342:
318:
545:
AMNH 5356, consisted of a partial skull 24 cm (
416:), sometime between 80 and 69 million years ago, in
324:
321:
290:
3312:
3126:
3040:
3005:
2943:
2871:
2760:
2718:
2655:
2600:
2573:
2546:
2519:
357:
339:
330:
2326:
2283:Bulletin of the American Museum of Natural History
2227:Dinosaur Systematics: Perspectives and Approaches
2010:
2008:
701:today. Nevertheless, it has grown apparent that
1753:
1751:
1749:
2126:"Shuvuuia: A dinosaur that hunted in the dark"
2432:
2262:Larsson, H.C.E. 2001. Endocranial anatomy of
594:. The generic name is derived from the Greek
8:
858:, and assigned these forms to a new family:
1773:
1771:
1769:
1767:
1765:
1763:
1730:So You Think You Know About...Velociraptor?
834:, within the "Deinodontidae" (now known as
375:
363:
3420:Late Cretaceous dinosaurs of North America
3300:
2949:
2663:
2652:
2579:
2552:
2525:
2516:
2455:
2439:
2425:
2417:
2311:Peabody Museum of Natural History Bulletin
1538:were discovered to be diurnal predators.
529:remains were discovered by paleontologist
122:
20:
2246:The Illustrated Encyclopedia of Dinosaurs
2225:. In Currie, P. J.; Carpenter, K (eds.).
2028:
1988:
1978:
1948:
1946:
726:
1887:"Description," Larsson (2001). Page 23.
1613:
1524:was compared to the former and also to
661:(Marsh 1888) Matthew & Brown 1922;
651:(Cope 1876) Matthew & Brown 1922 (
645:(Cope 1876) Matthew & Brown 1922;
482:such as possibly its closest relative
1209:is classified as the sister taxon of
715:was considered to be synonymous with
7:
3435:Taxa named by William Diller Matthew
1189:(4) compared with other dromaeosaurs
918:is recovered as the sister taxon to
862:. Since then, many new relatives of
826:Matthew and Brown originally placed
446:. Its fossils were unearthed in the
1899:Journal of Systematic Palaeontology
1787:Glut, Donald F. (30 January 2013).
1626:. Random House. 2007. p. 384.
1373:
1349:
1325:
1301:
1277:
1250:
1226:
1216:
1080:
1056:
994:
970:
943:
933:
424:and the western United States. The
2248:. Lorenz Books. pp. 200–201.
2201:Journal of Vertebrate Paleontology
1603:Timeline of dromaeosaurid research
582:and Brown named and described the
510:1922 diagram of the holotype skull
14:
877:Dromaeosaurinae, species such as
3284:
3283:
3260:
3254:
3249:
3243:
2333:Predatory Dinosaurs of the World
1859:10.1111/j.1469-185X.2010.00142.x
1587:
314:
143:
910:Below is a cladogram by Senter
895:have been recovered. The genus
679:(Cope 1876) Olshevsky 1979 and
2214:10.1080/02724634.1995.10011250
1841:Hwang, Sunny H. (2011-02-01).
671:(Marsh 1892) Russell 1972 (an
541:in Alberta, Canada. The find,
1:
3430:Fossil taxa described in 1922
2264:Carcharodontosaurus saharicus
2021:Paleontological Contributions
787:at the back of the head, the
1980:10.1371/journal.pone.0036790
617:
603:
533:during a 1914 expedition to
1789:Dinosaurs: The Encyclopedia
899:has been classified as the
869:The exact relationships of
685:(Barsbold 1983) Paul 1988 (
3471:
3440:Taxa named by Barnum Brown
2198:(Dinosauria: Theropoda)".
1683:10.2110/palo.2007.p07-006r
846:shared many features with
830:to its own subfamily, the
795:surface of the brain. The
682:Dromaeosaurus mongoliensis
610:
596:
452:Horseshoe Canyon Formation
16:Extinct genus of dinosaurs
3273:
3241:
2952:
2666:
2651:
2632:
2582:
2555:
2528:
2515:
2454:
2381:The Carnivorous Dinosaurs
2364:American Museum Novitates
2196:Dromaeosaurus albertensis
2030:10.17161/paleo.1808.18764
1911:10.1017/S1477201907002143
1757:Matthew and Brown (1922).
1395:
1378:
1371:
1354:
1347:
1330:
1323:
1306:
1299:
1282:
1275:
1255:
1248:
1231:
1224:
1136:
1119:
1102:
1085:
1078:
1061:
1054:
1016:
999:
992:
975:
968:
948:
941:
852:and the newly discovered
814:skull cast exhibited the
703:Dromaeosaurus albertensis
693:Dromaeosaurus albertensis
637:Another seven species of
592:Dromaeosaurus albertensis
434:, which was described by
431:Dromaeosaurus albertensis
300:Matthew & Brown, 1922
295:Dromaeosaurus albertensis
289:
282:
277:Matthew & Brown, 1922
140:Scientific classification
138:
133:Canadian Museum of Nature
130:
121:
23:
2385:Indiana University Press
2268:Mesozoic Vertebrate Life
2051:Therrien, et al. (2005).
1485:Cast of a reconstructed
731:Artistic restoration of
663:Dromaeosaurus explanatus
643:Dromaeosaurus laevifrons
622:) meaning 'lizard'. The
539:Dinosaur Provincial Park
131:Reconstructed skeleton,
3445:Paleontology in Alberta
2307:Deinonychus antirrhopus
2088:10.1126/science.abe7941
1716:Oxford University Press
665:(Cope 1876) Kuhn 1939;
659:Dromaeosaurus? gracilis
648:Dromaeosaurus cristatus
608:) meaning 'runner' and
559: in) in length, a
456:Dinosaur Park Formation
3455:Paleontology in Alaska
2305:(1969). "Osteology of
2244:Dixon, Dougal (2006).
1654:Wilson, L. E. (2008).
1506:(2005) estimated that
1490:
1190:
823:
735:
719:at one point in time.
677:Dromaeosaurus falculus
580:William Diller Matthew
511:
460:Prince Creek Formation
436:William Diller Matthew
404:that lived during the
3392:Paleobiology Database
1732:. Kane Miller. 2019.
1712:UK English Dictionary
1484:
1184:
810:
789:vena capitis dorsalis
730:
705:is even rarer in its
668:Dromaeosaurus minutus
509:
2387:. pp. 179–237.
2383:. Life of the Past.
2337:Simon & Schuster
502:Discovery and naming
448:Hell Creek Formation
408:period (middle late
3007:Saurornitholestinae
2275:Matthew, William D.
2174:The Washington Post
2080:2021Sci...372..610C
1971:2012PLoSO...736790S
1675:2008Palai..23..289W
741:was a medium-sized
1831:Weishampel, et al.
1491:
1191:
824:
736:
512:
3407:
3406:
3379:Open Tree of Life
3306:Taxon identifiers
3297:
3296:
3269:
3268:
3239:
3238:
3235:
3234:
3204:Saurornitholestes
3095:Saurornitholestes
3031:Saurornitholestes
2720:Halszkaraptorinae
2647:
2646:
2643:
2642:
2628:
2627:
2624:
2623:
2620:
2619:
2192:Currie, Philip J.
2074:(6542): 610–613.
1569:Saurornitholestes
1499:Saurornitholestes
1473:
1472:
1464:
1463:
1455:
1454:
1446:
1445:
1437:
1436:
1428:
1427:
1419:
1418:
1410:
1409:
1235:Saurornitholestes
1178:
1177:
1169:
1168:
1160:
1159:
1151:
1150:
1040:
1039:
1031:
1030:
866:have been found.
816:Geological museum
305:
304:
278:
3462:
3450:Campanian genera
3400:
3399:
3387:
3386:
3374:
3373:
3361:
3360:
3348:
3347:
3346:
3333:
3332:
3331:
3301:
3287:
3286:
3264:
3258:
3253:
3247:
3128:Velociraptorinae
3074:Dromaeosauroides
2950:
2934:Zhongjianosaurus
2912:Sinornithosaurus
2664:
2653:
2580:
2553:
2526:
2517:
2510:
2509:
2456:
2441:
2434:
2427:
2418:
2398:
2375:
2354:
2330:
2323:Paul, Gregory S.
2318:
2298:
2259:
2240:
2217:
2179:
2178:
2165:
2159:
2158:
2156:
2154:
2140:
2134:
2133:
2122:
2116:
2115:
2058:
2052:
2049:
2043:
2042:
2032:
2012:
2003:
2002:
1992:
1982:
1950:
1941:
1938:
1932:
1929:
1923:
1922:
1894:
1888:
1885:
1879:
1878:
1838:
1832:
1829:
1823:
1819:
1813:
1809:
1803:
1802:
1784:
1778:
1775:
1758:
1755:
1744:
1743:
1726:
1720:
1719:
1701:
1695:
1694:
1660:
1651:
1645:
1644:
1642:
1640:
1618:
1597:
1595:Dinosaurs portal
1592:
1591:
1590:
1542:Feeding behavior
1530:, in which both
1374:
1350:
1326:
1302:
1278:
1251:
1227:
1217:
1081:
1057:
995:
971:
965:Velociraptorinae
944:
934:
842:recognized that
797:Meckelian groove
763:Lance Formations
620:
613:
612:
606:
599:
598:
558:
557:
553:
550:
525:The first known
389:
386:
383:
380:
377:
373:
372:
369:
368:
365:
362:
359:
354:
353:
350:
347:
344:
341:
338:
335:
332:
329:
326:
323:
320:
292:
276:
269:
256:
243:
227:
148:
147:
126:
111:
48:
29:Temporal range:
21:
3470:
3469:
3465:
3464:
3463:
3461:
3460:
3459:
3410:
3409:
3408:
3403:
3395:
3390:
3382:
3377:
3369:
3364:
3356:
3351:
3342:
3341:
3336:
3327:
3326:
3321:
3308:
3298:
3293:
3265:
3231:
3188:Luanchuanraptor
3122:
3042:Dromaeosaurinae
3036:
3001:
2945:Eudromaeosauria
2939:
2867:
2756:
2714:
2660:
2658:Dromaeosauridae
2639:
2636:Dromaeosauridae
2616:
2602:Dromaeosauridae
2596:
2569:
2542:
2536:Avemetatarsalia
2521:Avemetatarsalia
2511:
2461:
2460:
2450:
2448:Dromaeosauridae
2445:
2395:
2378:
2357:
2351:
2321:
2303:Ostrom, John H.
2301:
2273:
2256:
2243:
2237:
2220:
2190:
2187:
2182:
2167:
2166:
2162:
2152:
2150:
2142:
2141:
2137:
2124:
2123:
2119:
2060:
2059:
2055:
2050:
2046:
2014:
2013:
2006:
1952:
1951:
1944:
1939:
1935:
1930:
1926:
1896:
1895:
1891:
1886:
1882:
1840:
1839:
1835:
1830:
1826:
1820:
1816:
1810:
1806:
1799:
1786:
1785:
1781:
1776:
1761:
1756:
1747:
1740:
1728:
1727:
1723:
1703:
1702:
1698:
1658:
1653:
1652:
1648:
1638:
1636:
1634:
1620:
1619:
1615:
1611:
1593:
1588:
1586:
1583:
1544:
1479:
1474:
1465:
1456:
1447:
1438:
1429:
1420:
1411:
1272:Dromaeosaurinae
1221:Eudromaeosauria
1179:
1170:
1161:
1152:
1051:Dromaeosaurinae
1041:
1032:
938:Eudromaeosauria
860:Dromaeosauridae
836:Tyrannosauridae
832:Dromaeosaurinae
805:
767:Aguja Formation
751:Late Cretaceous
725:
555:
551:
548:
546:
504:
476:Dromaeosaurinae
472:Dromaeosauridae
406:Late Cretaceous
387:
384:
381:
378:
356:
317:
313:
301:
298:
275:
267:
258:Dromaeosaurinae
254:
245:Eudromaeosauria
241:
229:Dromaeosauridae
225:
142:
117:
110:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
43:
42:
31:Late Cretaceous
27:
17:
12:
11:
5:
3468:
3466:
3458:
3457:
3452:
3447:
3442:
3437:
3432:
3427:
3425:Eudromaeosaurs
3422:
3412:
3411:
3405:
3404:
3402:
3401:
3388:
3375:
3362:
3349:
3334:
3318:
3316:
3310:
3309:
3304:
3295:
3294:
3292:
3291:
3281:
3274:
3271:
3270:
3267:
3266:
3242:
3240:
3237:
3236:
3233:
3232:
3230:
3229:
3222:
3215:
3208:
3200:
3192:
3184:
3177:
3170:
3163:
3155:
3147:
3140:
3132:
3130:
3124:
3123:
3121:
3120:
3113:
3106:
3099:
3091:
3084:
3077:
3070:
3062:
3054:
3046:
3044:
3038:
3037:
3035:
3034:
3027:
3019:
3011:
3009:
3003:
3002:
3000:
2999:
2991:
2984:
2976:
2969:
2962:
2953:
2947:
2941:
2940:
2938:
2937:
2930:
2923:
2915:
2908:
2901:
2893:
2886:
2878:
2876:
2869:
2868:
2866:
2865:
2858:
2850:
2846:Unquillosaurus
2842:
2835:
2827:
2819:
2812:
2804:
2797:
2790:
2782:
2775:
2767:
2765:
2758:
2757:
2755:
2754:
2747:
2740:
2733:
2725:
2723:
2716:
2715:
2713:
2712:
2705:
2697:
2690:
2682:
2675:
2667:
2661:
2656:
2649:
2648:
2645:
2644:
2641:
2640:
2633:
2630:
2629:
2626:
2625:
2622:
2621:
2618:
2617:
2615:
2614:
2613:
2612:
2606:
2604:
2598:
2597:
2595:
2594:
2593:
2592:
2583:
2577:
2571:
2570:
2568:
2567:
2566:
2565:
2556:
2550:
2544:
2543:
2541:
2540:
2539:
2538:
2529:
2523:
2513:
2512:
2508:
2507:
2498:
2489:
2480:
2474:
2468:
2459:
2452:
2451:
2446:
2444:
2443:
2436:
2429:
2421:
2415:
2414:
2403:The Dinosauria
2399:
2393:
2376:
2366:(2380): 1–49.
2355:
2349:
2319:
2299:
2271:
2260:
2254:
2241:
2235:
2218:
2208:(3): 576–591.
2186:
2183:
2181:
2180:
2160:
2135:
2117:
2053:
2044:
2004:
1942:
1933:
1931:Ostrom (1969).
1924:
1905:(4): 429–463.
1889:
1880:
1853:(1): 183–216.
1833:
1824:
1814:
1804:
1798:978-0786472222
1797:
1779:
1777:Currie (1995).
1759:
1745:
1738:
1721:
1696:
1669:(5): 289–297.
1646:
1632:
1612:
1610:
1607:
1606:
1605:
1599:
1598:
1582:
1579:
1547:Dromaeosaurus'
1543:
1540:
1487:D. albertensis
1478:
1475:
1471:
1470:
1467:
1466:
1462:
1461:
1458:
1457:
1453:
1452:
1449:
1448:
1444:
1443:
1440:
1439:
1435:
1434:
1431:
1430:
1426:
1425:
1422:
1421:
1417:
1416:
1413:
1412:
1408:
1407:
1404:
1403:
1394:
1391:
1390:
1387:
1386:
1377:
1372:
1370:
1367:
1366:
1363:
1362:
1353:
1348:
1346:
1343:
1342:
1339:
1338:
1329:
1324:
1322:
1319:
1318:
1315:
1314:
1305:
1300:
1298:
1295:
1294:
1291:
1290:
1281:
1276:
1274:
1268:
1267:
1264:
1263:
1254:
1249:
1247:
1244:
1243:
1240:
1239:
1230:
1225:
1223:
1215:
1176:
1175:
1172:
1171:
1167:
1166:
1163:
1162:
1158:
1157:
1154:
1153:
1149:
1148:
1145:
1144:
1135:
1132:
1131:
1128:
1127:
1118:
1115:
1114:
1111:
1110:
1101:
1098:
1097:
1094:
1093:
1084:
1079:
1077:
1074:
1073:
1070:
1069:
1060:
1055:
1053:
1047:
1046:
1043:
1042:
1038:
1037:
1034:
1033:
1029:
1028:
1025:
1024:
1015:
1012:
1011:
1008:
1007:
998:
993:
991:
988:
987:
984:
983:
974:
969:
967:
961:
960:
957:
956:
947:
942:
940:
932:
840:John H. Ostrom
812:D. albertensis
804:
803:Classification
801:
733:D. albertensis
724:
721:
535:Red Deer River
520:Tyrrell Museum
503:
500:
385:running lizard
303:
302:
299:
287:
286:
280:
279:
265:
261:
260:
252:
248:
247:
239:
232:
231:
223:
219:
218:
213:
206:
205:
200:
193:
192:
187:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
136:
135:
128:
127:
119:
118:
112:Possible Late
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
49:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3467:
3456:
3453:
3451:
3448:
3446:
3443:
3441:
3438:
3436:
3433:
3431:
3428:
3426:
3423:
3421:
3418:
3417:
3415:
3398:
3393:
3389:
3385:
3380:
3376:
3372:
3367:
3363:
3359:
3354:
3350:
3345:
3344:Dromaeosaurus
3339:
3335:
3330:
3324:
3320:
3319:
3317:
3315:
3314:Dromaeosaurus
3311:
3307:
3302:
3290:
3282:
3280:
3276:
3275:
3272:
3263:
3259:
3257:
3252:
3246:
3228:
3227:
3223:
3221:
3220:
3216:
3214:
3213:
3209:
3206:
3205:
3201:
3198:
3197:
3193:
3190:
3189:
3185:
3183:
3182:
3178:
3176:
3175:
3171:
3169:
3168:
3167:Kansaignathus
3164:
3161:
3160:
3156:
3153:
3152:
3148:
3146:
3145:
3141:
3139:
3138:
3134:
3133:
3131:
3129:
3125:
3119:
3118:
3114:
3112:
3111:
3107:
3105:
3104:
3100:
3097:
3096:
3092:
3090:
3089:
3085:
3083:
3082:
3081:Dromaeosaurus
3078:
3076:
3075:
3071:
3068:
3067:
3063:
3060:
3059:
3055:
3053:
3052:
3048:
3047:
3045:
3043:
3039:
3033:
3032:
3028:
3025:
3024:
3020:
3018:
3017:
3013:
3012:
3010:
3008:
3004:
2997:
2996:
2992:
2990:
2989:
2985:
2982:
2981:
2977:
2975:
2974:
2973:Dineobellator
2970:
2968:
2967:
2963:
2960:
2959:
2955:
2954:
2951:
2948:
2946:
2942:
2936:
2935:
2931:
2929:
2928:
2924:
2921:
2920:
2916:
2914:
2913:
2909:
2907:
2906:
2902:
2899:
2898:
2897:Hesperonychus
2894:
2892:
2891:
2890:Graciliraptor
2887:
2885:
2884:
2883:Changyuraptor
2880:
2879:
2877:
2874:
2873:Microraptoria
2870:
2864:
2863:
2859:
2856:
2855:
2851:
2848:
2847:
2843:
2841:
2840:
2836:
2833:
2832:
2828:
2825:
2824:
2820:
2818:
2817:
2813:
2810:
2809:
2808:Ornithodesmus
2805:
2803:
2802:
2801:Neuquenraptor
2798:
2796:
2795:
2791:
2788:
2787:
2783:
2781:
2780:
2776:
2774:
2773:
2769:
2768:
2766:
2763:
2759:
2753:
2752:
2748:
2746:
2745:
2741:
2739:
2738:
2734:
2732:
2731:
2730:Halszkaraptor
2727:
2726:
2724:
2721:
2717:
2711:
2710:
2706:
2703:
2702:
2698:
2696:
2695:
2691:
2688:
2687:
2683:
2681:
2680:
2676:
2674:
2673:
2669:
2668:
2665:
2662:
2659:
2654:
2650:
2638:
2637:
2631:
2610:
2609:
2608:
2607:
2605:
2603:
2599:
2591:
2587:
2586:
2585:
2584:
2581:
2578:
2576:
2572:
2564:
2560:
2559:
2558:
2557:
2554:
2551:
2549:
2545:
2537:
2533:
2532:
2531:
2530:
2527:
2524:
2522:
2518:
2514:
2506:
2502:
2499:
2497:
2493:
2490:
2488:
2484:
2481:
2479:
2475:
2473:
2469:
2467:
2463:
2462:
2457:
2453:
2449:
2442:
2437:
2435:
2430:
2428:
2423:
2422:
2419:
2412:
2411:0-520-24209-2
2408:
2404:
2400:
2396:
2394:0-253-34539-1
2390:
2386:
2382:
2377:
2373:
2369:
2365:
2361:
2360:Dromaeosaurus
2356:
2352:
2350:0-671-61946-2
2346:
2342:
2338:
2334:
2329:
2324:
2320:
2316:
2312:
2308:
2304:
2300:
2296:
2292:
2288:
2284:
2280:
2279:Brown, Barnum
2276:
2272:
2269:
2265:
2261:
2257:
2255:0-7548-1573-0
2251:
2247:
2242:
2238:
2236:0-521-43810-1
2232:
2228:
2224:
2219:
2215:
2211:
2207:
2203:
2202:
2197:
2193:
2189:
2188:
2184:
2176:
2175:
2170:
2164:
2161:
2149:
2145:
2139:
2136:
2131:
2127:
2121:
2118:
2113:
2109:
2105:
2101:
2097:
2093:
2089:
2085:
2081:
2077:
2073:
2069:
2065:
2057:
2054:
2048:
2045:
2040:
2036:
2031:
2026:
2022:
2018:
2011:
2009:
2005:
2000:
1996:
1991:
1986:
1981:
1976:
1972:
1968:
1965:(5): e36790.
1964:
1960:
1956:
1949:
1947:
1943:
1937:
1934:
1928:
1925:
1920:
1916:
1912:
1908:
1904:
1900:
1893:
1890:
1884:
1881:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1848:
1844:
1837:
1834:
1828:
1825:
1818:
1815:
1808:
1805:
1800:
1794:
1790:
1783:
1780:
1774:
1772:
1770:
1768:
1766:
1764:
1760:
1754:
1752:
1750:
1746:
1741:
1739:9781610678599
1735:
1731:
1725:
1722:
1717:
1713:
1711:
1706:
1705:"Dromaeosaur"
1700:
1697:
1692:
1688:
1684:
1680:
1676:
1672:
1668:
1664:
1657:
1650:
1647:
1635:
1633:9780375824197
1629:
1625:
1624:
1617:
1614:
1608:
1604:
1601:
1600:
1596:
1585:
1580:
1578:
1576:
1575:
1570:
1566:
1565:Dromaeosaurus
1561:
1560:Dromaeosaurus
1557:
1553:
1548:
1541:
1539:
1537:
1533:
1532:Dromaeosaurus
1529:
1528:
1523:
1522:Dromaeosaurus
1519:
1518:
1513:
1509:
1508:Dromaeosaurus
1505:
1501:
1500:
1495:
1494:Dromaeosaurus
1488:
1483:
1476:
1469:
1468:
1460:
1459:
1451:
1450:
1442:
1441:
1433:
1432:
1424:
1423:
1415:
1414:
1406:
1405:
1402:
1401:
1400:
1399:Dromaeosaurus
1393:
1392:
1389:
1388:
1385:
1384:
1383:
1376:
1375:
1369:
1368:
1365:
1364:
1361:
1360:
1359:
1352:
1351:
1345:
1344:
1341:
1340:
1337:
1336:
1335:
1328:
1327:
1321:
1320:
1317:
1316:
1313:
1312:
1311:
1304:
1303:
1297:
1296:
1293:
1292:
1289:
1288:
1287:
1280:
1279:
1273:
1270:
1269:
1266:
1265:
1262:
1261:
1260:
1253:
1252:
1246:
1245:
1242:
1241:
1238:
1237:
1236:
1229:
1228:
1222:
1219:
1218:
1214:
1212:
1208:
1207:Dromaeosaurus
1204:
1203:Robert Bakker
1200:
1196:
1188:
1187:Dromaeosaurus
1183:
1174:
1173:
1165:
1164:
1156:
1155:
1147:
1146:
1143:
1142:
1141:
1134:
1133:
1130:
1129:
1126:
1125:
1124:
1117:
1116:
1113:
1112:
1109:
1108:
1107:
1106:Dromaeosaurus
1100:
1099:
1096:
1095:
1092:
1091:
1090:
1083:
1082:
1076:
1075:
1072:
1071:
1068:
1067:
1066:
1059:
1058:
1052:
1049:
1048:
1045:
1044:
1036:
1035:
1027:
1026:
1023:
1022:
1021:
1014:
1013:
1010:
1009:
1006:
1005:
1004:
997:
996:
990:
989:
986:
985:
982:
981:
980:
973:
972:
966:
963:
962:
959:
958:
955:
954:
953:
946:
945:
939:
936:
935:
931:
929:
925:
921:
917:
916:Dromaeosaurus
913:
908:
906:
905:Dromaeosaurus
902:
898:
894:
893:
888:
887:
882:
881:
876:
872:
871:Dromaeosaurus
867:
865:
864:Dromaeosaurus
861:
857:
856:
851:
850:
845:
844:Dromaeosaurus
841:
837:
833:
829:
828:Dromaeosaurus
821:
817:
813:
809:
802:
800:
798:
794:
790:
786:
782:
781:Dromaeosaurus
778:
774:
773:Dromaeosaurus
770:
768:
764:
760:
756:
755:Maastrichtian
752:
749:stage of the
748:
744:
740:
739:Dromaeosaurus
734:
729:
722:
720:
718:
717:Dromaeosaurus
714:
713:
712:Chirostenotes
708:
704:
700:
699:
694:
690:
689:
684:
683:
678:
674:
673:alvarezsaurid
670:
669:
664:
660:
656:
655:
650:
649:
644:
640:
639:Dromaeosaurus
635:
633:
629:
628:"albertensis"
625:
624:specific name
621:
619:
607:
605:
593:
589:
588:Dromaeosaurus
585:
581:
576:
574:
570:
566:
562:
544:
540:
536:
532:
528:
527:Dromaeosaurus
523:
521:
517:
516:Dromaeosaurus
508:
501:
499:
497:
496:
491:
490:Dromaeosaurus
487:
486:
481:
480:Dromaeosaurus
477:
473:
469:
465:
464:Dromaeosaurus
461:
457:
453:
449:
445:
441:
437:
433:
432:
427:
423:
419:
415:
414:Maastrichtian
411:
407:
403:
400:
397:
396:dromaeosaurid
393:
371:
311:
310:
309:Dromaeosaurus
297:
296:
288:
285:
281:
274:
273:
272:Dromaeosaurus
266:
263:
262:
259:
253:
250:
249:
246:
240:
237:
234:
233:
230:
224:
221:
220:
217:
214:
211:
208:
207:
204:
201:
198:
195:
194:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
146:
141:
137:
134:
129:
125:
120:
115:
114:Maastrichtian
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
47:
44:80–69.1
40:
39:Maastrichtian
36:
32:
26:
25:Dromaeosaurus
22:
19:
3313:
3248:
3226:Velociraptor
3224:
3217:
3210:
3202:
3194:
3186:
3179:
3172:
3165:
3157:
3149:
3142:
3137:Acheroraptor
3135:
3115:
3108:
3101:
3093:
3086:
3080:
3079:
3072:
3064:
3058:Dakotaraptor
3056:
3051:Achillobator
3049:
3029:
3021:
3016:Atrociraptor
3014:
2995:Zhenyuanlong
2993:
2986:
2980:Tianyuraptor
2978:
2971:
2964:
2956:
2932:
2925:
2919:Tianyuraptor
2917:
2910:
2903:
2895:
2888:
2881:
2860:
2852:
2844:
2837:
2829:
2821:
2814:
2806:
2799:
2792:
2786:Dakotaraptor
2784:
2779:Buitreraptor
2777:
2772:Austroraptor
2770:
2762:Unenlagiinae
2749:
2742:
2735:
2728:
2709:Zhenyuanlong
2707:
2699:
2694:Tianyuraptor
2692:
2684:
2677:
2670:
2634:
2500:
2491:
2482:
2402:
2380:
2363:
2359:
2332:
2314:
2310:
2306:
2286:
2282:
2267:
2263:
2245:
2226:
2205:
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2195:
2172:
2163:
2151:. Retrieved
2148:ScienceDaily
2147:
2138:
2129:
2120:
2071:
2067:
2056:
2047:
2020:
1962:
1958:
1940:Paul (1988).
1936:
1927:
1902:
1898:
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1850:
1846:
1836:
1827:
1817:
1807:
1788:
1782:
1729:
1724:
1708:
1699:
1666:
1662:
1649:
1637:. Retrieved
1622:
1616:
1572:
1568:
1564:
1559:
1556:Troodontidae
1551:
1546:
1545:
1535:
1531:
1525:
1521:
1515:
1512:Velociraptor
1511:
1507:
1503:
1497:
1493:
1492:
1486:
1477:Paleobiology
1398:
1397:
1396:
1382:Dakotaraptor
1380:
1379:
1356:
1355:
1334:Achillobator
1332:
1331:
1310:Atrociraptor
1308:
1307:
1284:
1283:
1259:Velociraptor
1257:
1256:
1233:
1232:
1211:Dakotaraptor
1210:
1206:
1199:Peter Larson
1195:Larry Martin
1192:
1186:
1138:
1137:
1121:
1120:
1105:
1104:
1103:
1089:Achillobator
1087:
1086:
1063:
1062:
1020:Velociraptor
1018:
1017:
1001:
1000:
977:
976:
950:
949:
928:Achillobator
927:
923:
919:
915:
911:
909:
904:
901:sister taxon
897:Dakotaraptor
896:
890:
886:Achillobator
884:
878:
870:
868:
863:
853:
849:Velociraptor
847:
843:
827:
825:
811:
788:
780:
772:
771:
738:
737:
732:
716:
710:
702:
698:nomina dubia
696:
692:
686:
680:
676:
666:
662:
658:
652:
646:
642:
641:were named:
638:
636:
630:, refers to
627:
615:
601:
591:
587:
584:type species
577:
531:Barnum Brown
526:
524:
515:
513:
495:Velociraptor
493:
489:
485:Dakotaraptor
483:
479:
463:
440:Barnum Brown
430:
429:
426:type species
308:
307:
306:
294:
293:
284:Type species
271:
270:
235:
209:
196:
183:
24:
18:
3338:Wikispecies
3181:Linheraptor
3159:Deinonychus
3110:Yurgovuchia
3066:Deinonychus
3023:Bambiraptor
2988:Vectiraptor
2966:Bambiraptor
2905:Microraptor
2816:Pamparaptor
2751:Natovenator
2590:Maniraptora
2575:Maniraptora
2339:. pp.
2289:: 367–385.
2153:11 February
1574:Gorgosaurus
1286:Deinonychus
1140:Yurgovuchia
1065:Deinonychus
952:Bambiraptor
920:Yurgovuchia
892:Yurgovuchia
855:Deinonychus
783:also had a
723:Description
251:Subfamily:
3414:Categories
3277:See also:
3151:Boreonykus
3103:Utahraptor
2854:Variraptor
2823:Pyroraptor
2701:Variraptor
2679:Pyroraptor
2611:see below↓
2487:Dinosauria
2478:Sauropsida
2185:References
1358:Utahraptor
1123:Utahraptor
924:Utahraptor
880:Utahraptor
820:Copenhagen
759:Hell Creek
569:metacarpal
567:, a first
468:type genus
203:Saurischia
190:Dinosauria
3144:Adasaurus
2839:Unenlagia
2831:Rahonavis
2737:Hulsanpes
2563:Theropoda
2548:Theropoda
2496:Theropoda
2464:Kingdom:
2372:2246/2590
2295:2246/1300
2112:233872840
2096:0036-8075
1691:140668860
1609:Footnotes
1536:Alioramus
1527:Alioramus
979:Adasaurus
914:in 2012.
822:, Denmark
793:posterior
747:Campanian
743:carnivore
688:Adasaurus
410:Campanian
216:Theropoda
163:Kingdom:
157:Eukaryota
35:Campanian
3323:Wikidata
3289:Category
3279:Timeline
3196:Nuthetes
3117:Zapsalis
3088:Itemirus
2862:Ypupiara
2744:Mahakala
2672:Daurlong
2472:Chordata
2470:Phylum:
2466:Animalia
2325:(1988).
2317:: 1–165.
2130:phys.org
2104:33958472
2039:17099603
1999:22615813
1959:PLOS ONE
1919:83726237
1875:34613996
1867:20518758
1822:108-122.
1812:208-220.
1639:5 August
1581:See also
1517:Shuvuuia
1185:Size of
578:In 1922
561:mandible
543:holotype
470:of both
402:dinosaur
399:theropod
222:Family:
177:Chordata
173:Phylum:
167:Animalia
153:Domain:
3384:4946055
3371:4822955
3358:4433583
3329:Q131082
3219:Tsaagan
2505:Paraves
2476:Class:
2341:357–370
2076:Bibcode
2068:Science
1990:3352940
1967:Bibcode
1671:Bibcode
1663:PALAIOS
1003:Tsaagan
777:maxilla
707:habitat
654:Troodon
632:Alberta
604:dromeus
597:δρομεύς
573:Montana
554:⁄
466:is the
418:Alberta
390:) is a
379:
264:Genus:
2958:Balaur
2927:Wulong
2794:Diuqin
2686:Shanag
2409:
2391:
2347:
2252:
2233:
2110:
2102:
2094:
2037:
2023:(14).
1997:
1987:
1917:
1873:
1865:
1795:
1736:
1718:. n.d.
1710:Lexico
1689:
1630:
1552:et al.
1504:et al.
1201:, and
912:et al.
618:sauros
611:σαύρος
565:hyoids
563:, two
422:Canada
116:record
3397:38566
2501:Clade
2492:Clade
2483:Clade
2108:S2CID
2035:S2CID
1915:S2CID
1871:S2CID
1687:S2CID
1659:(PDF)
875:clade
392:genus
236:Clade
210:Clade
197:Clade
184:Clade
3366:GBIF
3212:Shri
3174:Kuru
2588:see
2561:see
2534:see
2407:ISBN
2389:ISBN
2345:ISBN
2250:ISBN
2231:ISBN
2155:2019
2100:PMID
2092:ISSN
1995:PMID
1863:PMID
1793:ISBN
1734:ISBN
1641:2020
1628:ISBN
1567:and
1534:and
1489:foot
926:and
889:and
785:vein
761:and
757:age
474:and
454:and
444:1922
438:and
412:and
376:lit.
52:PreꞒ
3353:EoL
2368:hdl
2362:".
2291:hdl
2210:doi
2084:doi
2072:372
2025:doi
1985:PMC
1975:doi
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1855:doi
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