3389:
1984:
137:
113:
567:
526:
3705:
2748:
2733:
3376:
2740:
3383:
665:
604:
378:, a ridge on the femur that serves as a muscle attachment in archosaurs, first appears in erythrosuchids. The triradiate pelvis and fourth trochanter are both features which indicate that erythrosuchids had an erect stance similar to later archosaurs. More
355:, the bone behind the premaxilla. This forms a characteristic "step" that makes erythrosuchids easily distinguishable from all other early archosauriformes, which have smooth jaw margins that are either straight or gradually curved.
1691:
doubtless constituted much of erythrosuchids' prey. However, the first erythrosuchids appear in the fossil record slightly earlier than the kannemeyeriids, so it must be assumed that they fed on other animals as well.
979:). The placement of fragmentary taxa that had to be removed to increase tree resolution is indicated by dashed lines (in the most derived position that they can be confidently assigned to). Taxa that are
2002:
344:(central parts of vertebrae) are deeply indented on either side, differing considerably from the usual cylindrical shape of the centra in early archosauriformes, but similar to later archosaurs.
441:, which lack these features. Thus the Erythrosuchidae occupy a transitional evolutionary position between the most primitive archisauriformes and more advanced Triassic archosaurs.
1808:
1762:
3798:
319:
Erythrosuchids were unusually large and robust archosauromorphs. Several features set them apart from other archosauriformes and are also seen in later, more
300:
of their day, with lengths of 2.5 m (8 ft 2 in) to almost 5 m (16 ft). Their fossil remains are known to date from South Africa (
2030:
3785:
3872:
3857:
3867:
3862:
1860:
1849:
1739:
1879:"The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms"
2023:
1997:
1818:
347:
The heads of erythrosuchids are generally disproportionately large and deep. In all erythrosuchids, the lower margin of the
136:
3388:
285:
222:
1729:
1395:
3708:
2016:
1547:
1088:
3307:
3198:
2732:
2682:
1530:
1281:
1191:
426:
359:
3375:
3852:
2631:
1489:
1148:
3382:
3725:
2747:
2657:
1513:
1305:
967:
Below is a cladogram from
Ezcurra (2016) that reexamined all historical members of the "Proterosuchia" (a
3824:
3336:
2739:
1264:
1131:
418:
379:
255:
3772:
3763:
3239:
1208:
1165:
2127:
1989:
1069:
1049:
3672:
1756:
1378:
985:
are indicated by the note "dubium". Bold terminal taxa are collapsed. Ezcurra (2016) recovered a
131:
437:
longer than the fourth, indicate that erythrosuchids are closer to the true archosaurs than the
3829:
3811:
2541:
2410:
2354:
1964:
1950:
1910:
1856:
1845:
1735:
1329:
1247:
430:
375:
3816:
3803:
2779:
2766:
2700:
2585:
2491:
2377:
2338:
2297:
2257:
2206:
2189:
2176:
2139:
2039:
1900:
1890:
1790:
1439:
1102:
745:
411:
204:
191:
112:
3667:
3601:
3466:
3296:
3187:
2989:
2875:
2861:
2519:
2454:
2438:
2391:
2226:
1685:
1456:
1224:
718:
713:
703:
689:
684:
651:
591:
549:
517:
512:
484:
474:
438:
313:
32:
1708:
D.M.S. Watson. 1917. "A sketch classification of the Pre-Jurassic tetrapod vertebrates".
566:
3682:
3261:
3166:
3116:
3025:
2891:
2620:
2596:
2484:
2399:
2267:
2247:
1958:
1905:
1878:
1115:
623:
613:
586:
545:
525:
403:
309:
301:
258:
251:
28:
3846:
3677:
3636:
3631:
3436:
3285:
3274:
3150:
3126:
3003:
2950:
2920:
2913:
2851:
2841:
2672:
2534:
2527:
2476:
2427:
2420:
2369:
2326:
2277:
2096:
1012:
976:
972:
827:
760:
535:
399:
383:
337:
297:
122:
70:
3657:
3498:
3456:
3446:
3426:
3325:
3106:
3032:
2927:
2468:
2361:
2287:
2216:
1794:
986:
981:
968:
851:
641:
595:
558:
422:
367:
363:
320:
989:
Erythrosuchidae, although the relationship of the Long Reef form (SAM P41754) and
1930:"The early evolution of archosaurs: Relationships and the origin of major clades"
1840:
Sues, H.-D.; Fraser, N.C. (2010). "Early and early Middle
Triassic in Gondwana".
3757:
3662:
3641:
3611:
3591:
3571:
3561:
3551:
3541:
3521:
3476:
3096:
3055:
3039:
2996:
2906:
2884:
2800:
2646:
2575:
2505:
2447:
1600:
991:
674:
502:
395:
45:
664:
3581:
3511:
3342:
3215:
3173:
3159:
3136:
3086:
3010:
2966:
2959:
2899:
2831:
2820:
2810:
2689:
2512:
2498:
2461:
2347:
2312:
2237:
2112:
2069:
1979:
1929:
1688:
912:
868:
434:
407:
351:, the bone at the tip of the upper jaw, is lower than the lower margin of the
348:
90:
55:
3748:
3622:
3486:
3251:
3228:
3076:
3018:
2935:
2606:
2558:
2384:
803:
735:
576:
332:
323:
269:
261:
148:
95:
39:
1914:
3742:
2063:
553:
358:
Erythrosuchids are notable for being the first archosauriforms to have a
341:
265:
168:
85:
80:
65:
60:
50:
603:
3790:
2078:
1895:
1813:
371:
352:
273:
178:
100:
75:
1731:
Anatomy, Phylogeny and
Palaeobiology of Early Archosaurs and Their Kin
448:
3531:
2087:
2057:
327:
158:
3719:
2008:
1728:
Sterling J. Nesbitt, Julia Brenda Desojo, Randall B. Irmis (2013).
656:
628:
489:
414:
305:
386:
lacked these features and probably had a more sprawling posture.
3777:
3313:
421:
to the
Archosauria proper. The presence of certain archosaurian
3723:
3404:
2764:
2174:
2050:
2012:
330:, which are found in other early archosauriformes, such as
1781:
362:
with three projecting areas formed from three bones: an
1844:. New York: Columbia University Press. pp. 19–36.
3607:
3597:
3587:
3577:
3567:
3557:
3547:
3537:
3527:
3517:
3507:
3482:
3472:
3462:
3452:
3442:
3432:
3422:
3281:
3270:
3225:
3184:
3147:
3132:
3122:
3112:
3102:
3092:
3082:
3072:
2977:
2947:
2872:
2857:
2847:
2837:
2827:
2816:
2806:
2796:
2669:
2643:
2617:
2602:
2592:
2581:
2571:
2309:
2293:
2283:
2273:
2263:
2253:
2243:
2233:
2222:
2212:
2202:
3732:
3650:
3621:
3496:
3415:
3324:
3295:
3259:
3250:
3224:
3213:
3183:
3146:
3053:
2976:
2946:
2871:
2777:
2699:
2668:
2642:
2616:
2556:
2437:
2409:
2337:
2307:
2187:
2138:
2111:
308:Basin), China, India and European Russia, from the
410:approach. In this, erythrosuchids constitute an
1710:Proceedings of the Zoological Society of London
1934:Columbia University (Open Access Dissertation)
1723:
1721:
2024:
2005:– list of species (Mikko's Phylogeny Archive)
1872:
1870:
8:
1761:: CS1 maint: multiple names: authors list (
1842:Triassic Life on Land: The Great Transition
402:. This classification is no longer used by
394:Erythrosuchids were formerly classified as
3720:
3412:
3401:
3256:
3221:
3065:
3061:
2789:
2785:
2774:
2761:
2564:
2318:
2195:
2184:
2171:
2117:
2108:
2047:
2031:
2017:
2009:
111:
20:
1904:
1894:
284:The family Erythrosuchidae was named by
1776:
1774:
1772:
1701:
466:
463:
1754:
326:. For example, they lack teeth on the
1969:Vertebrate Paleontology and Evolution
7:
1505:
1481:
1418:
1411:
1357:
1345:
1321:
1297:
1239:
1123:
1109:
1027:
1005:
998:
843:
819:
795:
788:
776:
752:
741:
1783:Journal of Vertebrate Paleontology
1734:. Geological Society. p. 22.
995:remain unknown within the family.
14:
3704:
3703:
3387:
3381:
3374:
2746:
2738:
2731:
1982:
663:
602:
565:
524:
272:) to the early Middle Triassic (
264:that lived from the later Early
135:
1795:10.1080/02724634.1992.10011434
1:
3873:Taxa named by D. M. S. Watson
250:(meaning "red crocodiles" in
3858:Prehistoric reptile families
1928:Sterling J. Nesbitt (2009).
374:. Although it is small, the
286:David Meredith Seares Watson
3868:Middle Triassic extinctions
3863:Olenekian first appearances
1877:Ezcurra, Martín D. (2016).
1396:Guchengosuchus shiguaiensis
971:historical group including
238:
3889:
1604:(=other archosauriformes)
3699:
3411:
3400:
3371:
3068:
3064:
2792:
2788:
2773:
2760:
2729:
2567:
2321:
2198:
2183:
2170:
2120:
2107:
2046:
1597:
1548:Shansisuchus shansisuchus
1544:
1527:
1510:
1503:
1486:
1479:
1453:
1436:
1423:
1416:
1409:
1392:
1375:
1362:
1355:
1343:
1326:
1319:
1302:
1295:
1278:
1261:
1244:
1237:
1222:
1205:
1188:
1162:
1145:
1128:
1121:
1107:
1089:Tasmaniosaurus triassicus
1085:
1065:
1045:
1032:
1025:
1010:
1003:
935:
909:
865:
848:
841:
824:
817:
800:
793:
786:
774:
757:
750:
340:. In erythrosuchids, the
235:
230:
132:Scientific classification
130:
119:
110:
23:
1531:Shansisuchus kuyeheensis
1282:Fugusuchus hejiapanensis
1192:Chasmatosuchus vjushkovi
427:triradiate pelvic girdle
382:archosauriforms such as
360:triradiate pelvic girdle
254:) are a family of large
1955:Vertebrate Paleontology
1684:The large contemporary
1490:Erythrosuchus africanus
1149:Chasmatosuchus rossicus
937:Other archosauriformes
1971:, WH Freeman & Co.
1514:Chalishevia cothurnata
1306:Sarmatosuchus otschevi
3825:Paleobiology Database
1959:Blackwell Science Ltd
1265:Kalisuchus rewanensis
1132:Chasmatosuchus magnus
738:from Parrish (1992):
1209:Vonhuenia friedrichi
1166:Gamosaurus lozovskii
1990:Paleontology portal
1070:Blomosuchus georgii
1050:Ankistrodon indicus
406:, who now employ a
3673:Ichthyosauromorpha
3663:Drepanosauromorpha
1961:(2004) 3rd edition
1896:10.7717/peerj.1778
1379:Uralosaurus magnus
16:Family of reptiles
3840:
3839:
3812:Open Tree of Life
3726:Taxon identifiers
3717:
3716:
3695:
3694:
3691:
3690:
3396:
3395:
3369:
3368:
3365:
3364:
3361:
3360:
3357:
3356:
3353:
3352:
3209:
3208:
3049:
3048:
2756:
2755:
2727:
2726:
2723:
2722:
2719:
2718:
2552:
2551:
2542:Sharovipterygidae
2411:Sharovipterygidae
2355:Dinocephalosaurus
2166:
2165:
2162:
2161:
2158:
2157:
1809:"Erythrosuchidae"
1676:
1675:
1667:
1666:
1658:
1657:
1649:
1648:
1640:
1639:
1631:
1630:
1622:
1621:
1613:
1612:
1586:
1585:
1577:
1576:
1568:
1567:
1559:
1558:
1468:
1467:
1330:Cuyosuchus huenei
1248:Eorasaurus olsoni
1177:
1176:
964:
963:
955:
954:
946:
945:
924:
923:
898:
897:
889:
888:
880:
879:
728:
727:
431:fourth trochanter
376:fourth trochanter
366:and an elongated
245:
244:
226:
3880:
3833:
3832:
3820:
3819:
3807:
3806:
3794:
3793:
3781:
3780:
3768:
3767:
3766:
3753:
3752:
3751:
3721:
3707:
3706:
3651:Possible members
3609:
3599:
3589:
3579:
3569:
3559:
3549:
3539:
3529:
3519:
3509:
3484:
3474:
3464:
3454:
3444:
3434:
3424:
3413:
3402:
3391:
3385:
3379:
3378:
3283:
3272:
3257:
3227:
3222:
3186:
3149:
3134:
3124:
3114:
3104:
3094:
3084:
3074:
3066:
3062:
2979:
2949:
2874:
2859:
2849:
2839:
2829:
2818:
2808:
2798:
2790:
2786:
2780:Archosauriformes
2775:
2767:Archosauriformes
2762:
2751:
2750:
2743:
2742:
2735:
2701:Archosauriformes
2671:
2645:
2619:
2604:
2594:
2586:Cryptovaranoides
2583:
2573:
2565:
2492:Langobardisaurus
2378:Protanystropheus
2339:Trachelosauridae
2319:
2311:
2298:Vritramimosaurus
2295:
2285:
2275:
2265:
2258:Mecistotrachelos
2255:
2245:
2235:
2224:
2214:
2207:Aenigmastropheus
2204:
2196:
2190:Archosauromorpha
2185:
2177:Archosauromorpha
2172:
2140:Archosauromorpha
2118:
2109:
2102:
2101:
2048:
2040:Archosauromorpha
2033:
2026:
2019:
2010:
1992:
1987:
1986:
1985:
1938:
1937:
1925:
1919:
1918:
1908:
1898:
1874:
1865:
1864:
1837:
1831:
1830:
1828:
1826:
1817:. Archived from
1805:
1799:
1798:
1778:
1767:
1766:
1760:
1752:
1750:
1748:
1725:
1716:
1706:
1506:
1482:
1440:Garjainia madiba
1419:
1412:
1358:
1346:
1322:
1298:
1240:
1124:
1110:
1103:Archosauriformes
1028:
1006:
999:
844:
820:
796:
789:
777:
753:
746:Archosauriformes
742:
667:
606:
569:
528:
449:
433:, and the third
412:Archosauriformes
398:of the suborder
221:
216:
205:Archosauriformes
192:Archosauromorpha
140:
139:
115:
105:
42:
27:Temporal range:
21:
3888:
3887:
3883:
3882:
3881:
3879:
3878:
3877:
3843:
3842:
3841:
3836:
3828:
3823:
3815:
3810:
3802:
3797:
3789:
3784:
3776:
3771:
3764:Erythrosuchidae
3762:
3761:
3756:
3747:
3746:
3741:
3734:Erythrosuchidae
3728:
3718:
3713:
3687:
3668:Kuehneosauridae
3646:
3617:
3602:Vjushkovisaurus
3492:
3467:Protecovasaurus
3407:
3392:
3373:
3349:
3320:
3308:Avemetatarsalia
3297:Avemetatarsalia
3291:
3246:
3218:
3205:
3199:Proterochampsia
3188:Proterochampsia
3179:
3142:
3058:
3045:
2990:Bharitalasuchus
2981:Erythrosuchidae
2972:
2942:
2876:Proterosuchidae
2867:
2862:Syntomiprosopus
2782:
2769:
2752:
2745:
2737:
2715:
2695:
2664:
2638:
2612:
2561:
2548:
2520:Sclerostropheus
2455:Augustaburiania
2439:Tanystropheidae
2433:
2405:
2392:Sclerostropheus
2333:
2315:
2303:
2227:Elachistosuchus
2192:
2179:
2154:
2134:
2103:
2053:
2052:
2042:
2037:
2003:Erythrosuchidae
1998:Erythrosuchidae
1988:
1983:
1981:
1978:
1947:
1942:
1941:
1927:
1926:
1922:
1876:
1875:
1868:
1852:
1839:
1838:
1834:
1824:
1822:
1821:on 9 April 2005
1807:
1806:
1802:
1780:
1779:
1770:
1753:
1746:
1744:
1742:
1727:
1726:
1719:
1707:
1703:
1698:
1682:
1677:
1668:
1659:
1650:
1641:
1632:
1623:
1614:
1587:
1578:
1569:
1560:
1469:
1457:Garjainia prima
1351:Erythrosuchidae
1225:Proterosuchidae
1178:
965:
956:
947:
925:
899:
890:
881:
782:Erythrosuchidae
733:
719:European Russia
714:Middle Triassic
704:Vjushkovisaurus
690:European Russia
685:Middle Triassic
652:Middle Triassic
592:European Russia
518:European Russia
513:Middle Triassic
485:Middle Triassic
475:Bharitalasuchus
447:
439:Proterosuchidae
404:paleontologists
392:
314:Middle Triassic
294:
282:
248:Erythrosuchidae
220:
218:Erythrosuchidae
214:
134:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
33:Middle Triassic
25:
17:
12:
11:
5:
3886:
3884:
3876:
3875:
3870:
3865:
3860:
3855:
3853:Erythrosuchids
3845:
3844:
3838:
3837:
3835:
3834:
3821:
3808:
3795:
3782:
3769:
3754:
3738:
3736:
3730:
3729:
3724:
3715:
3714:
3712:
3711:
3700:
3697:
3696:
3693:
3692:
3689:
3688:
3686:
3685:
3683:Thalattosauria
3680:
3675:
3670:
3665:
3660:
3654:
3652:
3648:
3647:
3645:
3644:
3639:
3634:
3628:
3626:
3619:
3618:
3616:
3615:
3605:
3595:
3585:
3575:
3565:
3555:
3545:
3535:
3525:
3515:
3504:
3502:
3494:
3493:
3491:
3490:
3480:
3470:
3460:
3450:
3440:
3430:
3419:
3417:
3409:
3408:
3406:Related topics
3405:
3398:
3397:
3394:
3393:
3372:
3370:
3367:
3366:
3363:
3362:
3359:
3358:
3355:
3354:
3351:
3350:
3348:
3347:
3346:
3345:
3339:
3330:
3328:
3322:
3321:
3319:
3318:
3317:
3316:
3310:
3301:
3299:
3293:
3292:
3290:
3289:
3279:
3267:
3265:
3262:Incertae sedis
3254:
3248:
3247:
3245:
3244:
3243:
3242:
3233:
3231:
3219:
3214:
3211:
3210:
3207:
3206:
3204:
3203:
3202:
3201:
3192:
3190:
3181:
3180:
3178:
3177:
3170:
3167:Halazhaisuchus
3163:
3155:
3153:
3144:
3143:
3141:
3140:
3130:
3120:
3117:Marcianosuchus
3110:
3100:
3090:
3080:
3069:
3059:
3054:
3051:
3050:
3047:
3046:
3044:
3043:
3036:
3029:
3026:Guchengosuchus
3022:
3015:
3007:
3000:
2993:
2985:
2983:
2974:
2973:
2971:
2970:
2963:
2955:
2953:
2944:
2943:
2941:
2940:
2932:
2924:
2917:
2910:
2903:
2896:
2892:Chasmatosuchus
2888:
2880:
2878:
2869:
2868:
2866:
2865:
2855:
2845:
2835:
2825:
2814:
2804:
2793:
2783:
2778:
2771:
2770:
2765:
2758:
2757:
2754:
2753:
2730:
2728:
2725:
2724:
2721:
2720:
2717:
2716:
2714:
2713:
2712:
2711:
2705:
2703:
2697:
2696:
2694:
2693:
2686:
2678:
2676:
2666:
2665:
2663:
2662:
2661:
2660:
2651:
2649:
2640:
2639:
2637:
2636:
2635:
2634:
2632:Allokotosauria
2625:
2623:
2621:Allokotosauria
2614:
2613:
2611:
2610:
2600:
2597:Tasmaniosaurus
2590:
2579:
2568:
2562:
2557:
2554:
2553:
2550:
2549:
2547:
2546:
2538:
2531:
2524:
2516:
2509:
2502:
2495:
2488:
2485:Gwyneddosaurus
2481:
2473:
2465:
2458:
2451:
2443:
2441:
2435:
2434:
2432:
2431:
2424:
2416:
2414:
2407:
2406:
2404:
2403:
2400:Trachelosaurus
2396:
2388:
2381:
2374:
2366:
2358:
2351:
2343:
2341:
2335:
2334:
2332:
2331:
2322:
2316:
2308:
2305:
2304:
2302:
2301:
2291:
2281:
2271:
2268:Prolacertoides
2261:
2251:
2248:Malutinisuchus
2241:
2231:
2220:
2210:
2199:
2193:
2188:
2181:
2180:
2175:
2168:
2167:
2164:
2163:
2160:
2159:
2156:
2155:
2153:
2152:
2151:
2150:
2144:
2142:
2136:
2135:
2133:
2132:
2131:
2130:
2121:
2115:
2105:
2104:
2100:
2099:
2090:
2081:
2072:
2066:
2060:
2051:
2044:
2043:
2038:
2036:
2035:
2028:
2021:
2013:
2007:
2006:
2000:
1994:
1993:
1977:
1976:External links
1974:
1973:
1972:
1965:Carroll, R. L.
1962:
1946:
1943:
1940:
1939:
1920:
1866:
1850:
1832:
1800:
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1157:
1154:
1153:
1144:
1141:
1140:
1137:
1136:
1127:
1122:
1120:
1116:Chasmatosuchus
1108:
1106:
1098:
1097:
1094:
1093:
1084:
1081:
1080:
1077:
1076:
1064:
1061:
1060:
1057:
1056:
1044:
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1040:
1037:
1036:
1031:
1026:
1024:
1021:
1020:
1017:
1016:
1009:
1004:
1002:
997:
977:erythrosuchids
973:proterosuchids
962:
961:
958:
957:
953:
952:
949:
948:
944:
943:
940:
939:
934:
931:
930:
927:
926:
922:
921:
918:
917:
908:
905:
904:
901:
900:
896:
895:
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891:
887:
886:
883:
882:
878:
877:
874:
873:
864:
861:
860:
857:
856:
847:
842:
840:
837:
836:
833:
832:
823:
818:
816:
813:
812:
809:
808:
799:
794:
792:
787:
785:
775:
773:
770:
769:
766:
765:
756:
751:
749:
740:
732:
729:
726:
725:
723:
721:
716:
711:
708:
697:
696:
694:
692:
687:
682:
679:
669:
668:
661:
659:
654:
649:
646:
636:
635:
633:
631:
626:
624:Early Triassic
621:
618:
614:Guchengosuchus
608:
607:
600:
598:
589:
587:Early Triassic
584:
581:
571:
570:
563:
561:
556:
543:
540:
530:
529:
522:
520:
515:
510:
507:
497:
496:
494:
492:
487:
482:
479:
469:
468:
465:
462:
459:
456:
453:
446:
443:
425:, such as the
391:
390:Classification
388:
384:proterosuchids
302:Beaufort Group
298:apex predators
296:They were the
293:
290:
281:
278:
259:archosauriform
243:
242:
233:
232:
228:
227:
212:
208:
207:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
146:
142:
141:
128:
127:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
26:
24:Erythrosuchids
15:
13:
10:
9:
6:
4:
3:
2:
3885:
3874:
3871:
3869:
3866:
3864:
3861:
3859:
3856:
3854:
3851:
3850:
3848:
3831:
3826:
3822:
3818:
3813:
3809:
3805:
3800:
3796:
3792:
3787:
3783:
3779:
3774:
3770:
3765:
3759:
3755:
3750:
3744:
3740:
3739:
3737:
3735:
3731:
3727:
3722:
3710:
3702:
3701:
3698:
3684:
3681:
3679:
3678:Sauropterygia
3676:
3674:
3671:
3669:
3666:
3664:
3661:
3659:
3656:
3655:
3653:
3649:
3643:
3640:
3638:
3637:Protorosauria
3635:
3633:
3632:Proterosuchia
3630:
3629:
3627:
3624:
3620:
3614:
3613:
3606:
3604:
3603:
3596:
3594:
3593:
3586:
3584:
3583:
3576:
3574:
3573:
3566:
3564:
3563:
3556:
3554:
3553:
3546:
3544:
3543:
3536:
3534:
3533:
3526:
3524:
3523:
3516:
3514:
3513:
3506:
3505:
3503:
3501:
3500:
3495:
3489:
3488:
3481:
3479:
3478:
3471:
3469:
3468:
3461:
3459:
3458:
3451:
3449:
3448:
3441:
3439:
3438:
3437:Lucianosaurus
3431:
3429:
3428:
3421:
3420:
3418:
3414:
3410:
3403:
3399:
3390:
3386:
3384:
3377:
3344:
3340:
3338:
3334:
3333:
3332:
3331:
3329:
3327:
3323:
3315:
3311:
3309:
3305:
3304:
3303:
3302:
3300:
3298:
3294:
3288:
3287:
3280:
3277:
3276:
3275:Sikannisuchus
3269:
3268:
3266:
3264:
3263:
3258:
3255:
3253:
3249:
3241:
3237:
3236:
3235:
3234:
3232:
3230:
3223:
3220:
3217:
3212:
3200:
3196:
3195:
3194:
3193:
3191:
3189:
3182:
3176:
3175:
3171:
3169:
3168:
3164:
3162:
3161:
3157:
3156:
3154:
3152:
3151:Euparkeriidae
3145:
3139:
3138:
3131:
3129:
3128:
3127:Polymorphodon
3121:
3119:
3118:
3111:
3109:
3108:
3101:
3099:
3098:
3091:
3089:
3088:
3081:
3079:
3078:
3071:
3070:
3067:
3063:
3060:
3057:
3052:
3042:
3041:
3037:
3035:
3034:
3030:
3028:
3027:
3023:
3021:
3020:
3016:
3013:
3012:
3008:
3006:
3005:
3004:Erythrosuchus
3001:
2999:
2998:
2994:
2992:
2991:
2987:
2986:
2984:
2982:
2975:
2969:
2968:
2964:
2962:
2961:
2957:
2956:
2954:
2952:
2951:Protopyknosia
2945:
2938:
2937:
2933:
2930:
2929:
2925:
2923:
2922:
2921:Samsarasuchus
2918:
2916:
2915:
2914:Proterosuchus
2911:
2909:
2908:
2904:
2902:
2901:
2897:
2894:
2893:
2889:
2887:
2886:
2882:
2881:
2879:
2877:
2870:
2864:
2863:
2856:
2854:
2853:
2852:Sarmatosuchus
2846:
2844:
2843:
2842:Koilamasuchus
2836:
2834:
2833:
2826:
2823:
2822:
2815:
2813:
2812:
2805:
2803:
2802:
2795:
2794:
2791:
2787:
2784:
2781:
2776:
2772:
2768:
2763:
2759:
2749:
2744:
2741:
2734:
2709:
2708:
2707:
2706:
2704:
2702:
2698:
2692:
2691:
2687:
2685:
2684:
2680:
2679:
2677:
2674:
2673:Prolacertidae
2667:
2659:
2658:Rhynchosauria
2655:
2654:
2653:
2652:
2650:
2648:
2647:Rhynchosauria
2641:
2633:
2629:
2628:
2627:
2626:
2624:
2622:
2615:
2609:
2608:
2601:
2599:
2598:
2591:
2588:
2587:
2580:
2578:
2577:
2570:
2569:
2566:
2563:
2560:
2555:
2544:
2543:
2539:
2537:
2536:
2535:Tanytrachelos
2532:
2530:
2529:
2528:Tanystropheus
2525:
2522:
2521:
2517:
2515:
2514:
2510:
2508:
2507:
2503:
2501:
2500:
2496:
2494:
2493:
2489:
2487:
2486:
2482:
2479:
2478:
2477:Gracilicollum
2474:
2471:
2470:
2466:
2464:
2463:
2459:
2457:
2456:
2452:
2450:
2449:
2445:
2444:
2442:
2440:
2436:
2430:
2429:
2428:Sharovipteryx
2425:
2423:
2422:
2418:
2417:
2415:
2412:
2408:
2402:
2401:
2397:
2394:
2393:
2389:
2387:
2386:
2382:
2380:
2379:
2375:
2372:
2371:
2370:Gracilicollum
2367:
2364:
2363:
2359:
2357:
2356:
2352:
2350:
2349:
2345:
2344:
2342:
2340:
2336:
2329:
2328:
2327:Jesairosaurus
2324:
2323:
2320:
2317:
2314:
2306:
2300:
2299:
2292:
2290:
2289:
2282:
2280:
2279:
2278:Protorosaurus
2272:
2270:
2269:
2262:
2260:
2259:
2252:
2250:
2249:
2242:
2240:
2239:
2232:
2229:
2228:
2221:
2219:
2218:
2211:
2209:
2208:
2201:
2200:
2197:
2194:
2191:
2186:
2182:
2178:
2173:
2169:
2148:
2147:
2146:
2145:
2143:
2141:
2137:
2129:
2125:
2124:
2123:
2122:
2119:
2116:
2114:
2110:
2106:
2098:
2097:Archelosauria
2094:
2091:
2089:
2085:
2082:
2080:
2076:
2073:
2071:
2067:
2065:
2061:
2059:
2055:
2054:
2049:
2045:
2041:
2034:
2029:
2027:
2022:
2020:
2015:
2014:
2011:
2004:
2001:
1999:
1996:
1995:
1991:
1980:
1975:
1970:
1966:
1963:
1960:
1956:
1952:
1951:Benton, M. J.
1949:
1948:
1944:
1935:
1931:
1924:
1921:
1916:
1912:
1907:
1902:
1897:
1892:
1888:
1884:
1880:
1873:
1871:
1867:
1863:
1862:
1861:0-231-13522-X
1858:
1853:
1851:9780231135221
1847:
1843:
1836:
1833:
1820:
1816:
1815:
1810:
1804:
1801:
1796:
1792:
1789:(1): 93–102.
1788:
1784:
1777:
1775:
1773:
1769:
1764:
1758:
1743:
1741:9781862393615
1737:
1733:
1732:
1724:
1722:
1718:
1714:
1711:
1705:
1702:
1695:
1693:
1690:
1687:
1686:kannemeyeriid
1679:
1672:
1671:
1663:
1662:
1654:
1653:
1645:
1644:
1636:
1635:
1627:
1626:
1618:
1617:
1609:
1608:
1605:
1603:
1602:
1595:
1594:
1591:
1590:
1582:
1581:
1573:
1572:
1564:
1563:
1555:
1554:
1551:
1550:
1549:
1542:
1541:
1538:
1537:
1534:
1533:
1532:
1525:
1524:
1521:
1520:
1517:
1516:
1515:
1508:
1507:
1501:
1500:
1497:
1496:
1493:
1492:
1491:
1484:
1483:
1477:
1476:
1473:
1472:
1464:
1463:
1460:
1459:
1458:
1451:
1450:
1447:
1446:
1443:
1442:
1441:
1434:
1433:
1430:
1429:
1426:
1421:
1420:
1414:
1413:
1407:
1406:
1403:
1402:
1399:
1398:
1397:
1390:
1389:
1386:
1385:
1382:
1381:
1380:
1373:
1372:
1369:
1368:
1365:
1360:
1359:
1352:
1348:
1347:
1341:
1340:
1337:
1336:
1333:
1332:
1331:
1324:
1323:
1317:
1316:
1313:
1312:
1309:
1308:
1307:
1300:
1299:
1293:
1292:
1289:
1288:
1285:
1284:
1283:
1276:
1275:
1272:
1271:
1268:
1267:
1266:
1259:
1258:
1255:
1254:
1251:
1250:
1249:
1242:
1241:
1235:
1234:
1231:
1230:
1227:
1226:
1220:
1219:
1216:
1215:
1212:
1211:
1210:
1203:
1202:
1199:
1198:
1195:
1194:
1193:
1186:
1185:
1182:
1181:
1173:
1172:
1169:
1168:
1167:
1160:
1159:
1156:
1155:
1152:
1151:
1150:
1143:
1142:
1139:
1138:
1135:
1134:
1133:
1126:
1125:
1118:
1117:
1112:
1111:
1104:
1100:
1099:
1096:
1095:
1092:
1091:
1090:
1083:
1082:
1079:
1078:
1075:
1073:
1071:
1063:
1062:
1059:
1058:
1055:
1053:
1051:
1043:
1042:
1039:
1038:
1035:
1030:
1029:
1023:
1022:
1019:
1018:
1015:
1014:
1013:Prolacertidae
1008:
1007:
1001:
1000:
996:
994:
993:
988:
984:
983:
978:
974:
970:
960:
959:
951:
950:
942:
941:
938:
933:
932:
929:
928:
920:
919:
916:
915:
914:
907:
906:
903:
902:
894:
893:
885:
884:
876:
875:
872:
871:
870:
863:
862:
859:
858:
855:
854:
853:
846:
845:
839:
838:
835:
834:
831:
830:
829:
828:Erythrosuchus
822:
821:
815:
814:
811:
810:
807:
806:
805:
798:
797:
791:
790:
783:
779:
778:
772:
771:
768:
767:
764:
763:
762:
761:Proterosuchus
755:
754:
747:
744:
743:
739:
737:
730:
724:
722:
720:
717:
715:
712:
709:
707:
706:
705:
699:
698:
695:
693:
691:
688:
686:
683:
680:
678:
677:
676:
671:
670:
666:
662:
660:
658:
655:
653:
650:
647:
645:
644:
643:
638:
637:
634:
632:
630:
627:
625:
622:
619:
617:
616:
615:
610:
609:
605:
601:
599:
597:
593:
590:
588:
585:
582:
580:
579:
578:
573:
572:
568:
564:
562:
560:
557:
555:
551:
547:
544:
541:
539:
538:
537:
536:Erythrosuchus
532:
531:
527:
523:
521:
519:
516:
514:
511:
508:
506:
505:
504:
499:
498:
495:
493:
491:
488:
486:
483:
480:
478:
477:
476:
471:
470:
460:
457:
454:
451:
450:
444:
442:
440:
436:
432:
428:
424:
420:
416:
413:
409:
405:
401:
400:Proterosuchia
397:
389:
387:
385:
381:
377:
373:
369:
365:
361:
356:
354:
350:
345:
343:
339:
335:
334:
329:
325:
322:
317:
315:
311:
307:
303:
299:
291:
289:
287:
279:
277:
275:
271:
267:
263:
260:
257:
253:
249:
241:
240:
234:
229:
224:
219:
213:
210:
209:
206:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
150:
147:
144:
143:
138:
133:
129:
125:
124:
123:Erythrosuchus
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
38:249–237
34:
30:
22:
19:
3733:
3658:Choristodera
3623:Paraphyletic
3610:
3600:
3590:
3580:
3570:
3560:
3550:
3540:
3530:
3520:
3510:
3499:Nomina dubia
3497:
3485:
3475:
3465:
3457:Palaeosaurus
3455:
3447:Tecovasaurus
3445:
3435:
3427:Crosbysaurus
3425:
3380:
3343:crocodilians
3337:Pseudosuchia
3326:Pseudosuchia
3284:
3273:
3260:
3172:
3165:
3158:
3135:
3125:
3115:
3107:Litorosuchus
3105:
3095:
3085:
3075:
3038:
3033:Shansisuchus
3031:
3024:
3017:
3009:
3002:
2995:
2988:
2980:
2965:
2958:
2934:
2928:Tsylmosuchus
2926:
2919:
2912:
2905:
2898:
2890:
2883:
2860:
2850:
2840:
2830:
2819:
2809:
2799:
2736:
2688:
2681:
2605:
2595:
2584:
2574:
2540:
2533:
2526:
2518:
2511:
2504:
2497:
2490:
2483:
2475:
2469:Fuyuansaurus
2467:
2460:
2453:
2446:
2426:
2419:
2398:
2390:
2383:
2376:
2368:
2362:Fuyuansaurus
2360:
2353:
2346:
2325:
2296:
2288:Rhombopholis
2286:
2276:
2266:
2256:
2246:
2236:
2225:
2217:Czatkowiella
2215:
2205:
2092:
2083:
2074:
1968:
1954:
1933:
1923:
1886:
1882:
1855:
1841:
1835:
1823:. Retrieved
1819:the original
1812:
1803:
1786:
1782:
1747:13 September
1745:. Retrieved
1730:
1712:
1709:
1704:
1683:
1680:Paleobiology
1599:
1598:
1546:
1545:
1529:
1528:
1512:
1511:
1488:
1487:
1455:
1454:
1438:
1437:
1424:
1394:
1393:
1377:
1376:
1363:
1350:
1328:
1327:
1304:
1303:
1280:
1279:
1263:
1262:
1246:
1245:
1223:
1207:
1206:
1190:
1189:
1164:
1163:
1147:
1146:
1130:
1129:
1114:
1087:
1086:
1067:
1066:
1047:
1046:
1033:
1011:
990:
987:monophyletic
982:nomina dubia
980:
969:polyphyletic
966:
936:
911:
910:
867:
866:
852:Shansisuchus
850:
849:
826:
825:
802:
801:
781:
759:
758:
734:
702:
700:
673:
672:
642:Shansisuchus
640:
639:
612:
611:
596:South Africa
575:
574:
559:South Africa
534:
533:
501:
500:
473:
472:
464:Description
393:
357:
346:
338:euparkeriids
331:
318:
295:
283:
247:
246:
236:
217:
198:
185:
121:
18:
3758:Wikispecies
3642:Thecodontia
3612:Wangisuchus
3592:Microcnemus
3572:Fenhosuchus
3562:Exilisuchus
3552:Collilongus
3542:Blomosuchus
3522:Ankistrodon
3477:Uatchitodon
3252:Archosauria
3240:Phytosauria
3229:Phytosauria
3097:Heteropelta
3056:Eucrocopoda
3040:Uralosaurus
2997:Chalishevia
2907:Jaikosuchus
2885:Archosaurus
2801:Antarctanax
2576:Boreopricea
2506:Macrocnemus
2448:Amotosaurus
1689:dicynodonts
1601:Eucrocopoda
1425:GHG 7433MI
1364:SAM P41754
1034:SAM-PK-591
992:Uralosaurus
675:Uralosaurus
503:Chalishevia
417:that is an
292:Description
3847:Categories
3582:Megacnemus
3512:Albisaurus
3416:Tooth taxa
3216:Crurotarsi
3174:Osmolskina
3160:Euparkeria
3137:Vancleavea
3087:Dorosuchus
3011:Fugusuchus
2967:Triopticus
2960:Kranosaura
2900:Gamosaurus
2832:Kalisuchus
2821:Eorasaurus
2811:Cuyosuchus
2710:see below↓
2690:Prolacerta
2683:Kadimakara
2513:Raibliania
2499:Luxisaurus
2462:Cosesaurus
2348:Austronaga
2313:Tanysauria
2238:Elessaurus
2149:see below↓
2128:Sauropsida
2113:Sauropsida
2070:Sauropsida
1957:, 2nd Ed.
1696:References
913:Fugusuchus
869:Vjushkovia
435:metatarsal
396:thecodonts
349:premaxilla
324:archosaurs
262:carnivores
3487:Zanclodon
3341:includes
3312:includes
3077:Asperoris
3019:Garjainia
2936:Vonhuenia
2607:Teyujagua
2559:Crocopoda
2385:Pectodens
2056:Kingdom:
1889:: e1778.
1757:cite book
1715:: 167–186
1074:(dubium)
1054:(dubium)
804:Garjainia
736:Cladogram
731:Phylogeny
577:Garjainia
461:Location
408:cladistic
333:Doswellia
288:in 1917.
270:Olenekian
155:Kingdom:
149:Eukaryota
120:Skull of
3743:Wikidata
3709:Category
2079:Diapsida
2064:Chordata
2062:Phylum:
2058:Animalia
1967:(1988),
1953:(2000),
1936:: 1–632.
1915:27162705
554:Triassic
423:features
419:outgroup
266:Triassic
211:Family:
179:Reptilia
169:Chordata
165:Phylum:
159:Animalia
145:Domain:
3817:4126953
3791:4817546
3778:4531083
3749:Q133077
2068:Class:
1945:Sources
1906:4860341
1814:Palaeos
1353:
1119:
1105:
784:
748:
467:Images
455:Status
372:ischium
353:maxilla
321:derived
304:of the
274:Anisian
231:Genera
175:Class:
3804:116289
3625:groups
3532:Avipes
2421:Ozimek
2088:Sauria
1913:
1903:
1859:
1848:
1825:1 July
1738:
1349:
1113:
1101:
780:
710:Valid
681:Valid
648:Valid
620:Valid
583:Valid
550:Middle
542:Valid
509:Valid
481:Valid
452:Genus
445:Genera
429:, the
342:centra
328:palate
280:Naming
225:, 1917
223:Watson
3830:38225
3799:IRMNG
3314:birds
2093:Clade
2084:Clade
2075:Clade
1883:PeerJ
657:China
629:China
546:Early
490:India
415:clade
380:basal
368:pubis
364:ilium
310:Early
306:Karoo
256:basal
252:Greek
239:below
199:Clade
186:Clade
29:Early
3786:GBIF
3335:see
3306:see
3286:Smok
3238:see
3197:see
2656:see
2630:see
2126:see
1911:PMID
1857:ISBN
1846:ISBN
1827:2010
1763:link
1749:2022
1736:ISBN
1713:1917
975:and
594:and
458:Age
370:and
336:and
237:see
46:PreꞒ
3773:EoL
1901:PMC
1891:doi
1791:doi
312:to
276:).
3849::
3827::
3814::
3801::
3788::
3775::
3760::
3745::
2095::
2086::
2077::
1932:.
1909:.
1899:.
1885:.
1881:.
1869:^
1854:.
1811:.
1787:12
1785:.
1771:^
1759:}}
1755:{{
1720:^
316:.
201::
188::
126:.
96:Pg
40:Ma
35:,
31:–
3608:†
3598:†
3588:†
3578:†
3568:†
3558:†
3548:†
3538:†
3528:†
3518:†
3508:†
3483:†
3473:†
3463:†
3453:†
3443:†
3433:†
3423:†
3282:†
3278:?
3271:†
3226:†
3185:†
3148:†
3133:†
3123:†
3113:†
3103:†
3093:†
3083:†
3073:†
3014:?
2978:†
2948:†
2939:?
2931:?
2895:?
2873:†
2858:†
2848:†
2838:†
2828:†
2824:?
2817:†
2807:†
2797:†
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2670:†
2644:†
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2603:†
2593:†
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2582:†
2572:†
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2523:?
2480:?
2472:?
2413:?
2395:?
2373:?
2365:?
2330:?
2310:†
2294:†
2284:†
2274:†
2264:†
2254:†
2244:†
2234:†
2230:?
2223:†
2213:†
2203:†
2032:e
2025:t
2018:v
1917:.
1893::
1887:4
1829:.
1797:.
1793::
1765:)
1751:.
1072:"
1068:"
1052:"
1048:"
701:?
552:-
548:-
268:(
215:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
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