167:
178:
31:
156:
69:
1166:
51:
1138:
239:
The
Eudaminae have been recent subject to significant taxonomic revisions based on genome analysis, including by Brower & Warren 2009, and Li et al. 2019.
1207:
1125:
1236:
166:
1231:
1200:
1130:
959:
Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; Hallwachs, Winnie; Janzen, Daniel H.; Grishin, Nick V. (26 March 2019).
530:
1143:
1241:
68:
1193:
937:
177:
155:
1052:
882:
592:
30:
972:
582:
572:
197:
130:
380:
1226:
699:
63:
815:
450:
1032:
1000:
793:
562:
520:
510:
1177:
410:
990:
980:
679:
201:
251:
217:
205:
1090:
976:
995:
960:
212:
40:
1024:
1220:
420:
906:
further divided into four subtribes: Eudamina, Loboclina, Cephisina and
Telemiadina
490:
145:
1025:"Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera)"
1173:
1165:
1112:
1084:
709:
602:
552:
480:
460:
430:
353:
323:
282:
Except where otherwise noted, the classification below follows Li et al., 2019:
250:, based on perceived similarities with two of the tribes in that subfamily, the
120:
961:"Genomes of skipper butterflies reveal extensive convergence of wing patterns"
842:
783:
642:
470:
222:
1075:
985:
852:
632:
622:
612:
303:
293:
271:
263:
193:
100:
80:
1036:
1004:
1099:
1069:
825:
652:
370:
343:
267:
259:
247:
1104:
773:
756:
719:
689:
500:
440:
333:
255:
227:
1117:
50:
872:
739:
729:
400:
390:
313:
110:
90:
1046:
941:
258:. As of Li et al. 2019, the Eudaminae are divided into four tribes:
1023:
Zhang, Jing; Cong, Qian; Shen, Jinhui; Grishin, Nick (2022-03-14).
225:, with some extending into temperate North America, and one genus,
862:
669:
176:
165:
154:
1050:
1181:
242:
Historically, the subfamily has been included as tribe
915:
1059:
971:(13). Supporting Information: Appendix pp.10–24.
944:. Version of 2009-JUN-11. Retrieved 2009-DEC-24.
965:Proceedings of the National Academy of Sciences
1201:
8:
1208:
1194:
1047:
49:
29:
20:
994:
984:
954:
952:
950:
934:Brower, Andrew V.Z. & Warren, Andrew
1018:
1016:
1014:
923:
899:
929:
927:
7:
1162:
1160:
1180:. You can help Knowledge (XXG) by
14:
1164:
415:O. Mielke & Casagrande, 2016
221:. They are largely found in the
67:
278:Current status and subdivisions
1:
204:Hesperiidae). Their original
1258:
1237:Taxa named by Paul Mabille
1159:
58:Typical resting positions
495:Godman & Salvin, 1893
64:Scientific classification
62:
57:
48:
37:
28:
23:
938:Tree of Life Web Project
894:Footnotes and references
231:, endemic to East Asia.
16:Subfamily of butterflies
986:10.1073/pnas.1821304116
1176:-related article is a
557:Herrich-Schäffer, 1869
185:
174:
163:
1232:Butterfly subfamilies
180:
169:
158:
836:Subtribe Typhedanina
767:Subtribe Telemiadina
977:2019PNAS..116.6232L
198:skipper butterflies
809:Subtribe Oileidina
750:Subtribe Cephisina
663:Subtribe Loboclina
186:
175:
164:
1242:Hesperiidae stubs
1189:
1188:
1154:
1153:
1053:Taxon identifiers
888:
878:
868:
858:
848:
831:
821:
799:
789:
779:
762:
745:
735:
725:
715:
705:
695:
685:
675:
674:R. Williams, 1927
658:
648:
638:
628:
627:Steinhauser, 1986
618:
608:
598:
588:
578:
568:
558:
546:Subtribe Eudamina
536:
526:
516:
506:
496:
486:
476:
466:
456:
446:
436:
426:
416:
406:
396:
386:
376:
359:
349:
339:
329:
319:
309:
299:
153:
152:
149:
1249:
1210:
1203:
1196:
1168:
1161:
1147:
1146:
1134:
1133:
1121:
1120:
1108:
1107:
1095:
1094:
1093:
1080:
1079:
1078:
1048:
1041:
1040:
1020:
1009:
1008:
998:
988:
956:
945:
935:
931:
916:
913:
907:
904:
886:
876:
866:
856:
846:
829:
819:
797:
787:
777:
760:
743:
733:
723:
713:
703:
693:
683:
673:
656:
646:
636:
626:
616:
606:
596:
586:
576:
566:
556:
534:
524:
514:
504:
494:
484:
474:
464:
454:
444:
434:
424:
414:
404:
394:
384:
374:
357:
347:
337:
327:
317:
307:
297:
144:
72:
71:
53:
38:Brown longtail (
33:
21:
1257:
1256:
1252:
1251:
1250:
1248:
1247:
1246:
1217:
1216:
1215:
1214:
1157:
1155:
1150:
1142:
1137:
1129:
1124:
1116:
1111:
1103:
1098:
1089:
1088:
1083:
1074:
1073:
1068:
1055:
1045:
1044:
1022:
1021:
1012:
958:
957:
948:
933:
932:
925:
920:
919:
914:
910:
905:
901:
896:
891:
867:A. Butler, 1870
857:E. Watson, 1893
847:A. Butler, 1870
838:
811:
806:
804:Tribe Oileidini
769:
752:
665:
607:A. Butler, 1870
548:
543:
475:E. Watson, 1893
425:E. Watson, 1893
395:E. Watson, 1893
366:
364:Tribe Phocidini
358:E. Watson, 1893
308:A. Butler, 1870
289:
280:
252:Celaenorrhinini
237:
182:Spicauda teleus
143:
66:
17:
12:
11:
5:
1255:
1253:
1245:
1244:
1239:
1234:
1229:
1219:
1218:
1213:
1212:
1205:
1198:
1190:
1187:
1186:
1169:
1152:
1151:
1149:
1148:
1135:
1122:
1109:
1096:
1081:
1065:
1063:
1057:
1056:
1051:
1043:
1042:
1010:
946:
922:
921:
918:
917:
908:
898:
897:
895:
892:
890:
889:
879:
869:
859:
849:
837:
834:
833:
832:
822:
810:
807:
805:
802:
801:
800:
790:
780:
768:
765:
764:
763:
751:
748:
747:
746:
736:
726:
716:
706:
696:
686:
676:
664:
661:
660:
659:
649:
639:
629:
619:
609:
599:
589:
579:
569:
559:
547:
544:
542:
541:Tribe Eudamini
539:
538:
537:
527:
517:
507:
505:Westwood, 1852
497:
487:
477:
467:
457:
447:
437:
427:
417:
407:
397:
387:
377:
365:
362:
361:
360:
350:
340:
330:
320:
310:
300:
288:
287:Tribe Entheini
285:
284:
279:
276:
236:
233:
213:junior synonym
171:Spicauda tanna
160:Spicauda tanna
151:
150:
138:
134:
133:
128:
124:
123:
118:
114:
113:
108:
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103:
98:
94:
93:
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78:
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73:
60:
59:
55:
54:
46:
45:
41:Urbanus procne
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1254:
1243:
1240:
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1233:
1230:
1228:
1225:
1224:
1222:
1211:
1206:
1204:
1199:
1197:
1192:
1191:
1185:
1183:
1179:
1175:
1170:
1167:
1163:
1158:
1145:
1140:
1136:
1132:
1127:
1123:
1119:
1114:
1110:
1106:
1101:
1097:
1092:
1086:
1082:
1077:
1071:
1067:
1066:
1064:
1062:
1058:
1054:
1049:
1038:
1034:
1030:
1029:Insecta Mundi
1026:
1019:
1017:
1015:
1011:
1006:
1002:
997:
992:
987:
982:
978:
974:
970:
966:
962:
955:
953:
951:
947:
943:
939:
930:
928:
924:
912:
909:
903:
900:
893:
887:Mabille, 1903
885:
884:
880:
877:Mabille, 1888
875:
874:
870:
865:
864:
860:
855:
854:
850:
845:
844:
840:
839:
835:
828:
827:
823:
820:Grishin, 2019
818:
817:
813:
812:
808:
803:
796:
795:
791:
786:
785:
781:
778:Mabille, 1878
776:
775:
771:
770:
766:
759:
758:
754:
753:
749:
742:
741:
737:
732:
731:
727:
724:Lindsey, 1925
722:
721:
717:
714:Lindsey, 1921
712:
711:
707:
704:Grishin, 2019
702:
701:
697:
692:
691:
687:
684:Grishin, 2019
682:
681:
677:
672:
671:
667:
666:
662:
657:Lindsey, 1921
655:
654:
650:
645:
644:
640:
635:
634:
630:
625:
624:
620:
615:
614:
610:
605:
604:
600:
595:
594:
590:
585:
584:
580:
575:
574:
570:
567:Grishin, 2019
565:
564:
560:
555:
554:
550:
549:
545:
540:
535:Grishin, 2022
533:
532:
528:
525:Grishin, 2022
523:
522:
518:
515:Grishin, 2022
513:
512:
508:
503:
502:
498:
493:
492:
488:
483:
482:
478:
473:
472:
468:
463:
462:
458:
455:Grishin, 2019
453:
452:
448:
443:
442:
438:
433:
432:
428:
423:
422:
421:Porphyrogenes
418:
413:
412:
408:
403:
402:
398:
393:
392:
388:
383:
382:
378:
373:
372:
368:
367:
363:
356:
355:
351:
346:
345:
341:
336:
335:
331:
328:Mabille, 1878
326:
325:
321:
316:
315:
311:
306:
305:
301:
296:
295:
291:
290:
286:
283:
277:
275:
273:
269:
265:
261:
257:
253:
249:
246:in subfamily
245:
240:
234:
232:
230:
229:
224:
220:
219:
214:
210:
207:
203:
199:
195:
191:
183:
179:
172:
168:
161:
157:
147:
142:
139:
136:
135:
132:
129:
126:
125:
122:
119:
116:
115:
112:
109:
106:
105:
102:
99:
96:
95:
92:
89:
86:
85:
82:
79:
76:
75:
70:
65:
61:
56:
52:
47:
43:
42:
36:
32:
27:
22:
19:
1182:expanding it
1171:
1156:
1060:
1028:
968:
964:
911:
902:
881:
871:
861:
851:
841:
824:
814:
792:
782:
772:
755:
738:
728:
718:
708:
698:
688:
678:
668:
651:
641:
631:
621:
611:
601:
597:Hübner, 1823
591:
581:
571:
561:
551:
529:
519:
509:
499:
491:Dyscophellus
489:
485:Austin, 2008
479:
469:
459:
449:
439:
435:Austin, 2008
429:
419:
409:
399:
389:
385:Austin, 2008
381:Pseudonascus
379:
369:
352:
342:
332:
322:
312:
302:
298:Watson, 1893
292:
281:
243:
241:
238:
226:
216:
208:
189:
187:
181:
170:
159:
140:
39:
18:
1174:Hesperiidae
1113:iNaturalist
1085:Wikispecies
761:Evans, 1952
744:Evans, 1952
734:Evans, 1952
710:Codatractus
700:Lobotractus
694:Moore, 1884
603:Spathilepia
553:Cecropterus
481:Euriphellus
461:Sarmientoia
445:Evans, 1952
431:Nicephellus
405:Evans, 1937
354:Tarsoctenus
324:Hyalothyrus
211:is today a
137:Subfamily:
131:Hesperiidae
121:Lepidoptera
1221:Categories
843:Typhedanus
816:Flattoides
784:Telemiades
643:Epargyreus
471:Bungalotis
465:Berg, 1897
223:Neotropics
206:type genus
184:, Colombia
173:, Colombia
162:, Colombia
101:Arthropoda
24:Eudaminae
1227:Eudaminae
1091:Eudaminae
1061:Eudaminae
942:Eudaminae
853:Oechydrus
794:Polygonus
633:Proteides
623:Narcosius
613:Astraptes
593:Autochton
583:Telegonus
451:Salantoia
304:Udranomia
294:Drephalys
272:Oileidini
264:Phocidini
194:subfamily
190:Eudaminae
141:Eudaminae
87:Kingdom:
81:Eukaryota
1100:BugGuide
1070:Wikidata
1037:35370352
1005:30877254
940:–
936:(2009):
830:Hübner,
826:Oileides
798:Hübner,
788:Hübner,
653:Chioides
647:Hübner,
637:Hübner,
617:Hübner,
587:Hübner,
577:Hübner,
563:Spicauda
521:Fulvatis
511:Ornilius
375:Hübner,
371:Phocides
348:Hübner,
344:Augiades
338:Hübner,
318:Hübner,
268:Eudamini
260:Entheini
248:Pyrginae
244:Eudamini
235:Taxonomy
127:Family:
97:Phylum:
91:Animalia
77:Domain:
1144:1053962
1076:Q774220
996:6442542
973:Bibcode
774:Ectomis
757:Cephise
720:Zestusa
690:Lobocla
573:Urbanus
501:Phareas
441:Salatis
411:Emmelus
334:Entheus
256:Pyrgini
228:Lobocla
218:Urbanus
209:Eudamus
146:Mabille
117:Order:
111:Insecta
107:Class:
1131:777462
1118:320509
1105:261419
1035:
1003:
993:
883:Marela
873:Nerula
740:Venada
730:Ridens
680:Zeutus
401:Aurina
391:Nascus
314:Phanus
202:family
192:are a
148:, 1877
1172:This
863:Cogia
670:Aguna
531:Adina
1178:stub
1139:NCBI
1126:ITIS
1033:PMID
1001:PMID
270:and
254:and
188:The
991:PMC
981:doi
969:116
215:of
196:of
1223::
1141::
1128::
1115::
1102::
1087::
1072::
1031:.
1027:.
1013:^
999:.
989:.
979:.
967:.
963:.
949:^
926:^
274:.
266:,
262:,
44:)
1209:e
1202:t
1195:v
1184:.
1039:.
1007:.
983::
975::
200:(
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