Knowledge (XXG)

Paraves

Source 📝

4786: 583:) had been known since the mid-20th century, their possible functions were the subject mainly of speculation, and few actual studies were published. Initial speculation regarded the claws as slashing implements used to disembowel large prey. In this scenario, the shortened upper foot would serve as an anchor point for powerful tendons to improve kicking ability. However, subsequent studies of the actual claw shape showed that the underside of the claw was only weakly keeled and would not have been an effective cutting instrument. Instead, it appeared to be more of a hooking implement. Manning et al. suggested in 2006 that the claws were similar to 431: 597:, which had slightly opposing first toes and strong tendons in the toes and foot. This makes it likely that advanced dromaeosaurids also used their claws to puncture and grip their prey to aid in pinning it to the ground, while using shallow wing beats and tail movements to stabilize themselves. Other lines of evidence for this behavior include teeth which had large, hooked serrations only on the back edge (useful in pulling flesh upward rather than slicing it) and large claws on the wings (for greater maneuvering of prey while mantling it with the wings). 7994: 562: 591:
species with modern birds to shed light on their actual function. Fowler and colleagues showed that many modern predatory birds also have enlarged claws on the second toes. In modern raptors, these claws are used to help grip and hold prey of sizes smaller than or equal to the predator, while the birds use their body weight to pin their prey to the ground and eat it alive. Fowler and colleagues suggested that this behavior is entirely consistent with the anatomy of advanced dromaeosaurids like
4793: 6176: 4030: 5737: 8404: 5742: 184: 4779: 4024: 132: 4018: 7821: 6236: 7845: 523:, have often been referred to as "four winged dinosaurs". Though it has been suggested that these hind wings would have prevented some paravians from getting around on the ground, and that they must have lived in trees, there is very little evidence that any of the earliest paravians were capable of climbing. This apparent paradox was addressed by later studies which showed that early paravians like 6182: 8392: 476:. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research. 7857: 8380: 7833: 8368: 1451:("true maniraptorans") was named by Padian, Hutchinson, & Holtz (1997). They defined their clade to include only avialans and deinonychosaurs. Paraves and Eumaniraptora are generally considered to be synonyms, though some phylogenetic studies suggest that the two groups have a similar but not identical content; Agnolín and Novas (2011) recovered 1472:, initially based primarily on the presence of a retractable second toe with sickle-claw (now also known to be present in some avialans). The name Deinonychosauria was coined by Ned Colbert and Dale Russell in 1969, and defined as a clade (all theropods closer to dromaeosaurids than to birds) by Jacques Gauthier in 1986. 590:
A larger study of sickle-claw function, published in 2011 by Fowler and colleagues, concluded that the earlier study by Manning and colleagues was correct and that the "sickle claws" would have been ineffective as cutting weapons. They compared the claw and overall foot anatomy of various primitive
539:
Most theropods walked with three toes contacting the ground, but fossilized footprint tracks confirm that many basal paravians, including dromaeosaurids, troodontids, and some early avialans, held the second toe off the ground in a hyperextended position, with only the third and fourth toes bearing
509:
Paravians generally have long, winged forelimbs, though these have become smaller in many flightless species and some extinct lineages that evolved before flight. The wings usually bore three large, flexible, clawed fingers in early forms. The fingers became fused and stiffened and the claws highly
496:
Most of the earliest paravian groups were carnivorous, though some smaller species (especially among the troodontids and early avialans) are known to have been omnivores, and it has been suggested that an omnivorous diet was the ancestral state for this group, with strict carnivory evolving in some
574:
One of the best-known features of paravians is the presence of an enlarged and strongly curved "sickle claw" on a hyper-extendible second toe, modified to hold the sickle claw clear of the ground when walking, most notably developed in the dromaeosaurids and troodontids. While this characteristic
1475:
However, several more recent studies have cast doubt on the hypothesis that dromaeosaurids and troodontids were more closely related to each other than either was to birds, instead finding that troodontids were more closely related to birds than to dromaeosaurids. Because Deinonychosauria was
613:
are found in Paraves. The ancestors of Paraves first started to shrink in size in the early Jurassic 200 million years ago, and fossil evidence shows that this theropod line evolved new adaptations four times faster than other groups of dinosaurs, and was shrinking 160 times faster than other
1577:
are often considered to be synonyms, depending on the interpretation of paravian systematics. Deinonychosauria will become a synonym of Dromaeosauridae when troodontids are found to form a clade with Avialae, to the exclusion of Dromaeosauridae. Tetrapterygidae is a polyphyletic grouping of
600:
In more primitive dromaeosaurids and in troodontids, the feet were not as specialized and the claws were not as large or as hooked. Additionally, the toe joints allowed more range of motion than the simple up-down movements of advanced dromaeosaurids. This makes it likely that these species
540:
the weight of the animal. This is called functional didactyly. The enlarged second toe bore an unusually large, curved sickle-shaped claw (held off the ground or 'retracted' when walking). This claw was especially large and flattened from side to side in the large-bodied predatory
1476:
originally defined as all animals closer to dromaeosaurids than to birds without specific reference to troodontids, this would render Deinonychosauria a synonym of Dromaeosauridae. The clade containing avialans, microraptorians, unenlagiids,
2589:
Lefèvre, Ulysse; Cau, Andrea; Cincotta, Aude; Hu, Dongyu; Chinsamy, Anusuya; Escuillié, François; Godefroit, Pascal (2017). "A new Jurassic theropod from China documents a transitional step in the macrostructure of feathers".
618:. (2007) suggested that extreme miniaturization was ancestral for the clade, whose common ancestor has been estimated to have been around 65 centimetres (26 in) long and 600–700 grams (21–25 oz) in mass. In 2646:
Cau, Andrea; Beyrand, Vincent; Voeten, Dennis F.A.E.; Fernandez, Vincent; Tafforeau, Paul; Stein, Koen; Barsbold, Rinchen; Tsogtbaatar, Khishigjav; Currie, Philip J.; Godefroit, Pascal (2017-12-06).
1781:
Li, Rihui; Lockley, M.G.; Makovicky, P.J.; Matsukawa, M.; Norell, M.A.; Harris, J.D.; Liu, M. (2007). "Behavioral and faunal implications of Early Cretaceous deinonychosaur trackways from China".
517:
Many early members of Paraves had both well-developed wings and long feathers on the hind legs, which in some cases, formed a second set of airfoils. These species, most famously represented by
2824:
Godefroit, Pascal; Cau, Andrea; Hu, Dong-Yu; Escuillié, François; Wu, Wenhao; Dyke, Gareth (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds".
634:. Fossils show that all the earliest members of Paraves found to date started out as small, while Troodontidae and Dromaeosauridae gradually increased in size during the Cretaceous period. 4785: 2047: 630:
went later through four independent events of gigantism, three times in dromaeosaurids and once in troodontids, while the body mass continued to decrease in many forms within
1733: 1687:
Zhang, H.; Wang, M.; Liu, X. (2008). "Constraints on the upper boundary age of the Tiaojishan Formation volcanic rocks in West Liaoning-North Hebei by LA-ICP-MS dating".
493:. The serrations on the front edge of dromaeosaurid and troodontid teeth were very small and fine, while the back edge had serrations which were very large and hooked. 6733: 3103:
Foth, Christian; Tischlinger, Helmut; Rauhut, Oliver W.M. (2 July 2014). "New specimen of Archaeopteryx provides insights into the evolution of pennaceous feathers".
8439: 7895: 548:) was usually small and angled inward toward the center of the body, but only became fully reversed in more specialized members of the bird lineage. One species, 2718: 2523: 8251: 2070:
Hu, Dongyu; Lianhi, Hou; Zhang, Lijun; Xu, Xing (2009). "A pre-Archaeopteryx troodontid theropod from China with long feathers on the metatarsus".
6728: 497:
specialized lineages. Fossils also suggest that legs and feet covered with feathers was an ancestral condition, possibly having originated in the
8256: 3176: 2915: 7888: 8449: 1895: 1637: 643: 514:, allowed the forelimbs to elongate and an elaboration of their plumage, traits that made the evolution of flapping flight possible. 4029: 430: 6175: 8429: 510:
reduced or lost in some advanced lineages. An increasingly asymmetric wrist joint, a trend that can be traced back to primitive
6272: 2753:
Sereno, P. C. (1998). "A rationale for phylogenetic definitions, with application to the higher level taxonomy of Dinosauria".
1655: 601:
specialized in smaller prey that could be pinned using only the inner toes, not requiring the feet to be as strong or sturdy.
554:, possessed a first toe which was highly modified in parallel with the second. Both the first and second toes on each foot of 7881: 5736: 4792: 7861: 7993: 8358: 587:
and were used for climbing, and in the case of larger species or individuals, climbing up the flanks of very large prey.
4778: 8444: 8396: 8015: 6718: 2390: 2058: 1468:
Since the 1960s, the dromaeosaurids and troodontids have often been classified together in a group or clade named the
487:
The teeth of basal paravians were curved and serrated, but not blade-like except in some specialized species, such as
5741: 8434: 8107: 7943: 6500: 2513: 1660: 1610: 4023: 1835:
Agnolin, Federico L.; Motta, Matias J.; Brissón Egli, Federico; Lo Coco, Gastón; Novas, Fernando E. (2019-02-12).
531:
in particular also seems to represent a case of flight evolving independently of the bird lineage within Paraves.
6415: 6405: 3969: 3169: 1598: 1444: 1354: 489: 527:
were capable of flapping flight and powered launching from the ground into the air without relying on climbing.
183: 6741: 6723: 4322: 3267: 2048:
125-Million-Year-Old Biplanes: New Evidence Suggests the Earliest Bird Species Had Feathers on their Hind Limbs
4017: 6746: 5861: 4726: 1590:. Then, around 110–90 Mya, the ancestors of Neornithes (modern birds) split from the other paravians. 561: 8278: 8246: 8190: 8185: 8170: 6713: 6525: 6475: 6239: 5838: 5782: 2896:
Avian ancestors. A review of the phylogenetic relationships of the theropods Unenlagiidae, Microraptoria,
1627: 168: 2179:
Xu, X.; Zhang, F. (2005). "A new maniraptoran dinosaur from China with long feathers on the metatarsus".
8338: 8261: 6550: 6098: 4348: 3913: 6069: 2232:"The wings before the bird: an evaluation of flapping-based locomotory hypotheses in bird antecedents" 2059:
The asymmetry of the carpal joint and the evolution of wing folding in maniraptoran theropod dinosaurs
8325: 8200: 8112: 7178: 6181: 3162: 3112: 3010: 2948: 2833: 2727: 2659: 2599: 2553: 2460: 2353: 2294: 2188: 2137: 2079: 2002: 1943: 1790: 1696: 1518:. (2017) and other phylogenies in which find Eudromaeosauria to be an outgroup to a clade including 8408: 8195: 8122: 6708: 6600: 6540: 6485: 6265: 5130: 4625: 4491: 4126: 2518: 1452: 1218: 706: 465: 371: 7844: 1514:
and modern birds. Averaptora additionally contains troodontids according to Cau, Beyrand, Voeten
8424: 8210: 8175: 7928: 7491: 6555: 6515: 6480: 6126: 5258: 5207: 4854: 4594: 3817: 3294: 3136: 2964: 2921: 2857: 2691: 2623: 2484: 2212: 2161: 2103: 1814: 1760: 1632: 1459:
as paravians that were not eumaniraptorans, while Turner, Makovicky, and Norell (2012) recovered
402: 178: 5035: 4955: 8343: 7837: 6026: 5410: 3054:
Fountaine, Toby M.R.; Benton, Michael J.; Dyke, Gareth J.; Nudds, Robert L. (4 February 2005).
8226: 8205: 8165: 8045: 7963: 7617: 7564: 7514: 6657: 6647: 6590: 6390: 6385: 5978: 5237: 4965: 4480: 4265: 3869: 3236: 3128: 3085: 3036: 2911: 2849: 2806: 2683: 2675: 2615: 2571: 2476: 2433: 2381: 2322: 2263: 2204: 2153: 2095: 2030: 1971: 1891: 1868: 1806: 550: 355: 2648:"Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs" 8384: 8236: 8010: 7483: 6625: 6297: 6209: 6134: 6076: 5669: 5560: 5267: 4945: 4669: 4648: 3958: 3892: 3799: 3436: 3120: 3075: 3067: 3026: 3018: 2956: 2903: 2841: 2796: 2762: 2735: 2667: 2607: 2561: 2468: 2423: 2415: 2371: 2361: 2312: 2302: 2253: 2243: 2196: 2145: 2087: 2020: 2010: 1961: 1951: 1858: 1848: 1798: 1750: 1742: 1704: 1601:
of Paraves, there are no extant survivors or genetic material, so their entire phylogeny is
1469: 1366: 1088: 675: 2939:
Senter, P' (2007). "A new look at the phylogeny of Coelurosauria (Dinosauria: Theropoda)".
8180: 8084: 8067: 7938: 7715: 7543: 7013: 6535: 6520: 6458: 6357: 6285: 6156: 6149: 6112: 6033: 6017: 5597: 5569: 5216: 4844: 4703: 4634: 4506: 4469: 4311: 4296: 4215: 4171: 4150: 3927: 3806: 3763: 3588: 3519: 3252: 1643: 1622: 1530: 1485: 1456: 1165: 1114: 968: 623: 580: 541: 519: 461: 453: 449: 380: 363: 47: 2878: 2785:"Unenlagiid theropods: are they members of the Dromaeosauridae (Theropoda, Maniraptora)?" 2283:"A microraptorine (Dinosauria–Dromaeosauridae) from the Late Cretaceous of North America" 3116: 3014: 2952: 2837: 2731: 2663: 2603: 2557: 2540:
Turner, Alan H.; Pol, Diego; Clarke, Julia A.; Erickson, Gregory M.; Norell, M. (2007).
2464: 2357: 2298: 2192: 2141: 2083: 2006: 1947: 1794: 1700: 1426:
in 1997. The clade was defined by Sereno in 1998 as a branch-based clade containing all
131: 8241: 8132: 8055: 8002: 7973: 7933: 7908: 7825: 7755: 7707: 7641: 7580: 7535: 7522: 7396: 7359: 7210: 6984: 6821: 6575: 6570: 6565: 6545: 6530: 6425: 6420: 6400: 6395: 6377: 6325: 6258: 6199: 6141: 6105: 6002: 5934: 5909: 5882: 5850: 5494: 5464: 5329: 5168: 5086: 4975: 4715: 4641: 4604: 4102: 3899: 3658: 3554: 3493: 3472: 3080: 3055: 3031: 2998: 2428: 2403: 2376: 2341: 2317: 2282: 2258: 2231: 2025: 1990: 1966: 1927: 1378: 336: 141: 2710: 2122: 1991:"Herbivorous ecomorphology and specialization patterns in theropod dinosaur evolution" 575:
claw and its associated modifications to the anatomy of the foot (such as a shortened
8418: 8316: 8062: 7978: 7968: 7763: 7747: 7670: 7662: 7496: 7294: 7144: 7031: 6989: 6789: 6781: 6703: 6667: 6585: 6509: 6219: 6214: 6083: 5943: 5827: 5771: 5450: 5369: 5313: 5283: 5230: 5223: 5109: 5100: 4884: 4864: 4407: 4396: 4282: 4251: 4208: 4157: 4143: 3837: 3569: 3547: 3532: 3450: 2925: 2107: 1602: 1510: 1500: 1461: 1392: 1387: 1370: 1192: 931: 748: 619: 498: 147: 89: 27:
Clade of all dinosaurs that are more closely related to birds than to oviraptorosaurs
2968: 2801: 2784: 2766: 2627: 1818: 1764: 8372: 8127: 7849: 7593: 7438: 7254: 7092: 7040: 6912: 6873: 6642: 6630: 6620: 6610: 6307: 6229: 6091: 5818: 5692: 5685: 5624: 5542: 5275: 5154: 5066: 5015: 4985: 4934: 4874: 4692: 4545: 4530: 4515: 4449: 4337: 4289: 4201: 4091: 3946: 3906: 3885: 3878: 3858: 3851: 3844: 3791: 3776: 3712: 3669: 3630: 3500: 3479: 3410: 3388: 3140: 2861: 2739: 2695: 2488: 2342:"An aberrant island-dwelling theropod dinosaur from the Late Cretaceous of Romania" 2165: 1519: 1400: 1140: 1072: 1054:
Cladogram following the results of a phylogenetic study by Lefèvre, Cau, Cincotta,
952: 875: 661: 627: 511: 457: 415: 388: 326: 267: 2999:"A new time tree reveals Earth history's imprint on the evolution of modern birds" 2216: 472:, which include diverse extinct taxa as well as the over 10,000 species of living 1956: 484:
Like other theropods, all paravians are bipedal, walking on their two hind legs.
8310: 8139: 8089: 8030: 8025: 8020: 7654: 7501: 7307: 7286: 7275: 7240: 7107: 7097: 7055: 6946: 6813: 6615: 6440: 6435: 6367: 6352: 6320: 6054: 5964: 5899: 5640: 5590: 5521: 5473: 5298: 5244: 5192: 5139: 5025: 4995: 4914: 4678: 4428: 4418: 4081: 3920: 3685: 3644: 3615: 3608: 3576: 3486: 3420: 3360: 3347: 3306: 3185: 1888:
Avian Evolution: The Fossil Record of Birds and its Paleobiological Significance
1594: 1539: 1427: 1423: 1340:
Cladogram following the results of a phylogenetic study by Cau, Beyrand, Voeten
593: 566: 315: 305: 285: 153: 64: 2346:
Proceedings of the National Academy of Sciences of the United States of America
8040: 7983: 7953: 7948: 7771: 7633: 7572: 7551: 7383: 7367: 7302: 7247: 7219: 7151: 7116: 6994: 6842: 6677: 6652: 6410: 6119: 5987: 5794: 5678: 5648: 5615: 5583: 5549: 5535: 5528: 5352: 5322: 5116: 5076: 5005: 4894: 4823: 4810: 4738: 4685: 4561: 4538: 4523: 4459: 4386: 4274: 4244: 4237: 4180: 4136: 3830: 3719: 3703: 3692: 3651: 3637: 3601: 3562: 3540: 2960: 2907: 2647: 2611: 2451:
Bhullar, B.A.; et al. (2012). "Birds have paedomorphic dinosaur skulls".
2200: 1802: 1708: 1587: 1557: 1545: 1431: 1396: 1374: 1260: 918: 857: 730: 610: 576: 241: 171: 159: 109: 74: 8301: 2679: 1872: 1853: 1836: 17: 8144: 8117: 7731: 7686: 7475: 7268: 7185: 7169: 7158: 7068: 7004: 6963: 6662: 6463: 6450: 6430: 6330: 6224: 6204: 6040: 5995: 5957: 5950: 5507: 5440: 5420: 5400: 5390: 5380: 5184: 5177: 5146: 4611: 4438: 4230: 4112: 3784: 3726: 3678: 3622: 3508: 3377: 3279: 3227: 2566: 2541: 2402:
Manning, P.L.; Payne, D.; Pennicott, J.; Barrett, P.M.; Ennos, R.A. (2006).
2366: 2307: 2015: 1666: 1436: 1244: 805: 346: 254: 195: 114: 58: 3132: 3089: 3071: 3040: 3022: 2853: 2810: 2687: 2619: 2575: 2480: 2437: 2419: 2385: 2326: 2267: 2208: 2157: 2099: 2034: 1975: 1810: 8391: 1508:, which Agnolín and Novas redefine as the least inclusive clade including 8295: 8149: 8072: 8050: 7958: 7699: 7429: 7404: 7350: 7261: 7233: 7123: 7087: 7079: 7061: 6956: 6941: 6925: 6672: 6635: 6468: 6047: 5919: 5576: 5514: 5336: 5290: 4904: 4655: 4579: 4223: 4187: 3733: 3465: 3218: 3209: 1551: 1358: 823: 584: 445: 442: 228: 215: 104: 99: 84: 79: 69: 51: 7873: 3124: 2845: 2671: 2501: 2472: 2149: 2091: 1863: 8077: 7609: 7462: 7341: 7226: 7192: 7130: 6951: 6805: 6797: 6490: 6362: 6347: 6337: 6061: 5807: 5655: 5480: 5055: 4365: 4194: 2248: 1649: 1562: 1523: 1505: 1382: 992: 631: 469: 393: 119: 94: 31: 6250: 8035: 7739: 7678: 7454: 7446: 7375: 7315: 7199: 7137: 7020: 6687: 6682: 6560: 5632: 5430: 5045: 4924: 3399: 3203: 1755: 1494: 545: 205: 8272: 1926:
Fowler, D.W.; Freedman, E.A.; Scannella, J.B.; Kambic, R.E. (2011).
1728: 2121:
Xing, X.; Zhou, Z.; Wang, X.; Kuang, X.; Zhang, F.; Du, X. (2003).
1369:. The ancestral paravian is the earliest common ancestor of birds, 7625: 7412: 6975: 3457: 1837:"Paravian Phylogeny and the Dinosaur-Bird Transition: An Overview" 1746: 560: 429: 8330: 2542:"A basal dromaeosaurid and size evolution preceding avian flight" 1727:
Turner, Alan Hamilton; Makovicky, Peter J.; Norell, Mark (2012).
1434:(which includes all the birds living in the world today) than to 8231: 8099: 7912: 7601: 6968: 6931: 6901: 6315: 6281: 5759: 5704: 2986:. Baltimore, MD: Johns Hopkins University Press. pp. 45–48. 1362: 473: 164: 8276: 7877: 6765: 6254: 5757: 4808: 4045: 3345: 3196: 3158: 7920: 7904: 7723: 6580: 6342: 3154: 2340:
Z., Csiki; Vremir, M.; Brusatte, S. L.; Norell, M. A. (2010).
1729:"A review of dromaeosaurid systematics and paravian phylogeny" 1492:
by Agnolín and Novas (2013), defined as all animals closer to
434:
Birds, members of the paraves, are the only living dinosaurs.
1586:
Paravians diverged from other maniraptorans around 165 
1561:; together they were proposed to be the sister group of the 558:
were held retracted and bore enlarged, sickle-shaped claws.
501:, even if this trait was later lost in more advanced birds. 2902:. SpringerBriefs in Earth System Sciences. pp. 1–96. 1605:
only from the fossil record. The prototypical fossil is
2984:
The Rise of Birds: 225 million years of evolution
8356: 2778: 2776: 2641: 2639: 2637: 2230:
Dececchi, T.A.; Larsson, H.C.E.; Habib, M.B. (2016).
1597:
of modern birds, which are direct descendants in the
6014: 5975: 5931: 5915: 5905: 5895: 5666: 5612: 5491: 5461: 5446: 5436: 5426: 5416: 5406: 5396: 5386: 5376: 5365: 5310: 5255: 5204: 5165: 5127: 5097: 5082: 5072: 5062: 5051: 5041: 5031: 5021: 5011: 5001: 4991: 4981: 4971: 4961: 4951: 4941: 4930: 4920: 4910: 4900: 4890: 4880: 4870: 4860: 4850: 4840: 4712: 4666: 4622: 4591: 4574: 4503: 4487: 4476: 4465: 4455: 4445: 4434: 4424: 4414: 4403: 4392: 4382: 4334: 4308: 4262: 4168: 4123: 4108: 4098: 4087: 4077: 3955: 3760: 3433: 3416: 3406: 3395: 3384: 3373: 3056:"The quality of the fossil record of Mesozoic birds" 1890:(1 ed.). John Wiley & Sons. p. Ch. 2. 1240: 1215: 1189: 1162: 1137: 1111: 1085: 385: 377: 368: 360: 352: 342: 332: 322: 311: 301: 139:Six paravian dinosaurs (top left to bottom right): 8285: 8219: 8158: 8098: 8001: 7919: 7698: 7653: 7592: 7563: 7534: 7513: 7474: 7437: 7428: 7395: 7358: 7349: 7340: 7285: 7209: 7168: 7106: 7078: 7054: 7030: 7003: 6924: 6911: 6872: 6841: 6780: 6696: 6599: 6499: 6449: 6376: 6306: 6013: 5974: 5930: 5880: 5849: 5826: 5817: 5793: 5770: 5703: 5665: 5611: 5559: 5490: 5460: 5350: 5309: 5254: 5203: 5164: 5126: 5096: 4821: 4737: 4711: 4702: 4665: 4621: 4590: 4559: 4502: 4363: 4333: 4307: 4261: 4167: 4122: 4058: 3980: 3954: 3945: 3868: 3816: 3759: 3702: 3668: 3587: 3518: 3431: 3358: 3305: 3278: 3251: 2873: 2871: 1734:Bulletin of the American Museum of Natural History 1381:. Paraves often comprises three major sub-groups: 2894:Agnolín, Federico L.; Novas, Fernando E. (2013). 2783:Agnolín, Federico L.; Novas, Fernando E. (2011). 1722: 1720: 1718: 1535:The Rise of Birds: 225 Million Years of Evolution 2997:Claramunt, S.; Cracraft, J. (11 December 2015). 2889: 2887: 1921: 1919: 1917: 1915: 1913: 1911: 1909: 1907: 1670:, a putative but possibly chimeric Triassic bird 2535: 2533: 1465:as the only known non-eumaniraptoran paravian. 7889: 6266: 3170: 2755:Neues Jahrbuch für Geologie und Paläontologie 2719:Annual Review of Earth and Planetary Sciences 2404:"Dinosaur killer claws or climbing crampons?" 8: 2877:Mortimer, M. (2012): The Theropod Database: 2502:Dinosaurs 'shrank' regularly to become birds 1776: 1774: 1504:. Most studies use a similar definition for 1652:, a hypothetical dinosaur-bird intermediary 8273: 7896: 7882: 7874: 7434: 7355: 7346: 7337: 7075: 6921: 6908: 6869: 6865: 6838: 6777: 6773: 6762: 6758: 6273: 6259: 6251: 5888: 5823: 5799: 5767: 5754: 5499: 5358: 4833: 4829: 4818: 4805: 4708: 4567: 4375: 4371: 4070: 4066: 4055: 4042: 3951: 3822: 3768: 3756: 3593: 3524: 3442: 3366: 3355: 3342: 3284: 3257: 3248: 3193: 3177: 3163: 3155: 130: 38: 8252:Tradeoffs for locomotion in air and water 3079: 3030: 2800: 2565: 2427: 2375: 2365: 2316: 2306: 2257: 2247: 2024: 2014: 1965: 1955: 1862: 1852: 1754: 544:. In these early species, the first toe ( 2789:Anais da Academia Brasileira de Ciências 609:Extreme examples of miniaturization and 8363: 2711:"The origin and evolution of dinosaurs" 2514:"How Dinosaurs Shrank and Became Birds" 1679: 614:dinosaur lineages were growing. Turner 8440:Extant Late Jurassic first appearances 2281:Longrich, N.R.; Currie, P.J. (2009). 1989:Zanno, L.E.; Makovicky, P.J. (2011). 7: 7832: 1932:and the Origin of Flapping in Birds" 1830: 1828: 7856: 2941:Journal of Systematic Palaeontology 1578:four-winged basal paravian genera. 1233: 1208: 1181: 1155: 1130: 1104: 1078: 1068: 944: 937: 909: 848: 796: 789: 782: 721: 698: 691: 667: 657: 452:period. In addition to the extinct 2123:"Four-winged dinosaurs from China" 1638:Biogeography of paravian dinosaurs 1533:in the second edition of his book 1361:which are more closely related to 644:Biogeography of paravian dinosaurs 25: 6734:Late Quaternary prehistoric birds 1391:and modern birds, as well as the 8402: 8390: 8378: 8366: 7992: 7855: 7843: 7831: 7820: 7819: 7710:(New World vultures and condors) 6235: 6234: 6180: 6174: 5740: 5735: 4791: 4784: 4777: 4028: 4022: 4016: 2526:from the original on 2023-07-20. 1377:which was not also ancestral to 182: 2802:10.1590/S0001-37652011000100008 1656:Temporal paradox (paleontology) 2740:10.1146/annurev.earth.25.1.435 1399:, which may or may not form a 468:, the group also contains the 1: 1613:, both complete and partial. 1609:, of which 11 specimens 1957:10.1371/journal.pone.0028964 7774:(woodpeckers and relatives) 1529:In 2015 Chatterjee created 8466: 8108:Flying and gliding animals 7944:Fin and flipper locomotion 7636:(cormorants and relatives) 6729:Extinct species since 1500 1928:"The Predatory Ecology of 1841:Frontiers in Earth Science 1661:List of fossil bird genera 1582:Origin and early evolution 1406:The name 'Paraves' (Greek 641: 441:are a widespread group of 29: 8450:Taxa named by Paul Sereno 7990: 7815: 7766:(kingfishers and rollers) 7620:(albatrosses and petrels) 7504:(swifts and hummingbirds) 7486:(nightjars and relatives) 7336: 7310:(pheasants and relatives) 6868: 6837: 6776: 6772: 6761: 6757: 6293: 6194: 6172: 5891: 5802: 5766: 5753: 5733: 5502: 5361: 4836: 4832: 4817: 4804: 4775: 4570: 4378: 4374: 4073: 4069: 4054: 4041: 4014: 3825: 3771: 3755: 3596: 3527: 3445: 3369: 3354: 3341: 3287: 3260: 3247: 3192: 2961:10.1017/S1477201907002143 2908:10.1007/978-94-007-5637-3 2767:10.1127/njgpa/210/1998/41 2612:10.1007/s00114-017-1496-y 2201:10.1007/s00114-004-0604-y 1803:10.1007/s00114-007-0310-7 1709:10.1007/s11434-008-0287-4 1258: 1238: 1231: 1213: 1206: 1186: 1179: 1160: 1153: 1135: 1128: 1109: 1102: 1083: 1076: 990: 965: 949: 942: 935: 914: 907: 871: 853: 846: 819: 801: 794: 787: 780: 744: 726: 719: 703: 696: 689: 672: 665: 490:Dromaeosaurus albertensis 408: 401: 298: 293: 179:Scientific classification 177: 138: 129: 41: 7665:(seriemas and relatives) 7644:(pelicans and relatives) 2881:. Retrieved 2013-AUG-15. 1854:10.3389/feart.2018.00252 1689:Chinese Science Bulletin 30:Not to be confused with 8430:Late Jurassic dinosaurs 7758:(hornbills and hoopoes) 7673:(falcons and relatives) 2982:Chatterjee, S. (2015). 2900:and Scansoriopterygidae 2567:10.1126/science.1144066 2367:10.1073/pnas.1006970107 2308:10.1073/pnas.0811664106 2016:10.1073/pnas.1011924108 1414:'beside, near' + Latin 1357:defined to include all 605:Dichotomy in body sizes 448:that originated in the 8247:Terrestrial locomotion 8191:Evolution of cetaceans 8186:Origin of avian flight 8171:Evolution of tetrapods 7742:(trogons and quetzals) 7554:(cranes and relatives) 6824:(emus and cassowaries) 6714:Glossary of bird terms 6526:Confuciusornithiformes 3072:10.1098/rspb.2004.2923 3023:10.1126/sciadv.1501005 2879:Phylogeny of Theropoda 2420:10.1098/rsbl.2005.0395 2391:Supporting Information 1995:Proc Natl Acad Sci USA 1628:Origin of avian flight 1422:'bird') was coined by 571: 435: 8339:Paleobiology Database 8262:Undulatory locomotion 8211:Homologous structures 7583:(kagu and sunbittern) 7546:(gulls and relatives) 6551:Songlingornithiformes 6516:Omnivoropterygiformes 3914:Paralitherizinosaurus 2709:Sereno, P.C. (1997). 2592:The Science of Nature 1886:Mayr, G. (Oct 2016). 564: 433: 8397:Evolutionary biology 8206:Analogous structures 8201:Convergent evolution 7405:Phoenicopteriformes 1644:thecodont hypothesis 1537:, where he included 1484:to the exclusion of 535:Feet and sickle claw 8257:Rotating locomotion 8196:Comparative anatomy 7376:Mesitornithiformes 7370:(doves and pigeons) 6704:Families and orders 6566:Ichthyornithiformes 6541:Patagopterygiformes 5131:Patagopterygiformes 4626:Confuciusornithidae 4492:Scansoriopterygidae 4127:Scansoriopterygidae 3125:10.1038/nature13467 3117:2014Natur.511...79F 3015:2015SciA....1E1005C 2953:2007JSPal...5..429S 2846:10.1038/nature12168 2838:2013Natur.498..359G 2732:1997AREPS..25..435S 2672:10.1038/nature24679 2664:2017Natur.552..395C 2604:2017SciNa.104...74L 2558:2007Sci...317.1378T 2552:(5843): 1378–1381. 2519:Scientific American 2473:10.1038/nature11146 2465:2012Natur.487..223B 2358:2010PNAS..10715357C 2299:2009PNAS..106.5002L 2193:2005NW.....92..173X 2181:Naturwissenschaften 2150:10.1038/nature01342 2142:2003Natur.421..335X 2092:10.1038/nature08322 2084:2009Natur.461..640H 2007:2011PNAS..108..232Z 1948:2011PLoSO...628964F 1795:2008NW.....95..185L 1783:Naturwissenschaften 1701:2008SciBu..53.3574Z 1219:Scansoriopterygidae 707:Scansoriopterygidae 464:, and possibly the 372:Scansoriopterygidae 8445:Evolution of birds 8176:Evolution of birds 7929:Aquatic locomotion 7718:(eagles and hawks) 7618:Procellariiformes 7523:Opisthocomiformes 7492:Steatornithiformes 6719:List by population 6556:Hongshanornithidae 6481:Evolution of birds 5259:Songlingornithidae 5208:Hongshanornithidae 4595:Omnivoropterygidae 4507:Jeholornithiformes 3818:Therizinosauroidea 2249:10.7717/peerj.2159 1667:Protoavis texensis 1633:Evolution of birds 1453:scansoriopterygids 1355:branch-based clade 572: 466:scansoriopterygids 436: 8435:Prehistoric birds 8354: 8353: 8279:Taxon identifiers 8270: 8269: 8227:Animal locomotion 8166:Evolution of fish 8046:facultative biped 7871: 7870: 7811: 7810: 7807: 7806: 7803: 7802: 7799: 7798: 7795: 7794: 7791: 7790: 7787: 7786: 7783: 7782: 7604:(loons or divers) 7573:Phaethontiformes 7565:Phaethontimorphae 7515:Opisthocomiformes 7484:Caprimulgiformes 7424: 7423: 7413:Podicipediformes 7332: 7331: 7328: 7327: 7324: 7323: 7050: 7049: 6833: 6832: 6790:Struthioniformes 6658:Waterfowl hunting 6591:Gastornithiformes 6586:Aepyornithiformes 6576:Lithornithiformes 6248: 6247: 6190: 6189: 6170: 6169: 6166: 6165: 6070:Lutetodontopteris 5979:Gastornithiformes 5876: 5875: 5872: 5871: 5760:Aves / Neornithes 5749: 5748: 5731: 5730: 5727: 5726: 5723: 5722: 5705:Aves / Neornithes 5607: 5606: 5346: 5345: 5238:Parahongshanornis 4800: 4799: 4773: 4772: 4769: 4768: 4765: 4764: 4761: 4760: 4757: 4756: 4555: 4554: 4481:Archaeopterygidae 4359: 4358: 4266:Archaeopterygidae 4037: 4036: 4012: 4011: 4008: 4007: 4004: 4003: 4000: 3999: 3941: 3940: 3937: 3936: 3870:Therizinosauridae 3751: 3750: 3747: 3746: 3743: 3742: 3337: 3336: 3333: 3332: 3329: 3328: 3325: 3324: 3237:Maniraptoriformes 3066:(1560): 289–294. 2917:978-94-007-5636-6 2832:(7454): 359–362. 2658:(7685): 395–399. 2459:(7406): 223–226. 2136:(6921): 335–340. 2078:(7264): 640–643. 1695:(22): 3574–3584. 1348: 1347: 1334: 1333: 1325: 1324: 1316: 1315: 1307: 1306: 1298: 1297: 1289: 1288: 1280: 1279: 1271: 1270: 1062: 1061: 1048: 1047: 1039: 1038: 1030: 1029: 1021: 1020: 1012: 1011: 1003: 1002: 979: 978: 896: 895: 887: 886: 835: 834: 769: 768: 760: 759: 428: 427: 422: 356:Archaeopterygidae 289: 16:(Redirected from 8457: 8407: 8406: 8405: 8395: 8394: 8383: 8382: 8381: 8371: 8370: 8369: 8362: 8347: 8346: 8334: 8333: 8321: 8320: 8319: 8306: 8305: 8304: 8274: 8237:Robot locomotion 8011:Limb development 7996: 7969:Lobe-finned fish 7898: 7891: 7884: 7875: 7859: 7858: 7847: 7835: 7834: 7823: 7822: 7775: 7767: 7759: 7751: 7748:Leptosomiformes 7743: 7735: 7727: 7719: 7716:Accipitriformes 7711: 7690: 7689:(perching birds) 7682: 7674: 7666: 7645: 7637: 7629: 7621: 7613: 7610:Sphenisciformes 7605: 7584: 7576: 7555: 7547: 7544:Charadriiformes 7526: 7505: 7487: 7466: 7458: 7455:Musophagiformes 7450: 7435: 7416: 7408: 7387: 7379: 7371: 7356: 7347: 7338: 7311: 7298: 7076: 7072: 7065: 6978: 6971: 6935: 6922: 6918: 6909: 6905: 6870: 6866: 6839: 6825: 6817: 6809: 6801: 6793: 6778: 6774: 6763: 6759: 6626:Bird collections 6581:Dinornithiformes 6486:Darwin's finches 6476:Origin of flight 6416:Seabird breeding 6406:Sexual selection 6275: 6268: 6261: 6252: 6238: 6237: 6210:Deinonychosauria 6185: 6184: 6178: 6135:Protodontopteryx 6077:Macrodontopteryx 6016: 5977: 5933: 5917: 5907: 5897: 5889: 5824: 5800: 5768: 5755: 5744: 5739: 5670:Cimolopterygidae 5668: 5614: 5561:Hesperornithidae 5500: 5493: 5463: 5448: 5438: 5428: 5418: 5408: 5398: 5388: 5378: 5367: 5359: 5312: 5257: 5206: 5167: 5129: 5099: 5084: 5074: 5064: 5053: 5043: 5033: 5023: 5013: 5003: 4993: 4983: 4973: 4963: 4953: 4943: 4932: 4922: 4912: 4902: 4892: 4882: 4872: 4862: 4852: 4842: 4834: 4830: 4819: 4806: 4795: 4789: 4788: 4781: 4714: 4709: 4670:Jinguofortisidae 4668: 4649:Eoconfuciusornis 4624: 4593: 4576: 4568: 4505: 4489: 4478: 4467: 4457: 4447: 4436: 4426: 4416: 4405: 4394: 4384: 4376: 4372: 4336: 4310: 4264: 4170: 4125: 4110: 4100: 4089: 4079: 4071: 4067: 4056: 4043: 4032: 4026: 4020: 3970:Oviraptorosauria 3959:Oviraptorosauria 3957: 3952: 3893:Nanshiungosaurus 3823: 3800:Jianchangosaurus 3769: 3762: 3757: 3594: 3525: 3443: 3437:Alvarezsauroidea 3435: 3418: 3408: 3397: 3386: 3375: 3367: 3356: 3343: 3285: 3258: 3249: 3242: 3241: 3194: 3179: 3172: 3165: 3156: 3145: 3144: 3100: 3094: 3093: 3083: 3051: 3045: 3044: 3034: 3009:(11): e1501005. 3003:Science Advances 2994: 2988: 2987: 2979: 2973: 2972: 2936: 2930: 2929: 2891: 2882: 2875: 2866: 2865: 2821: 2815: 2814: 2804: 2780: 2771: 2770: 2750: 2744: 2743: 2715: 2706: 2700: 2699: 2643: 2632: 2631: 2586: 2580: 2579: 2569: 2537: 2528: 2527: 2510: 2504: 2499: 2493: 2492: 2448: 2442: 2441: 2431: 2399: 2393: 2389: 2379: 2369: 2352:(35): 15357–61. 2337: 2331: 2330: 2320: 2310: 2278: 2272: 2271: 2261: 2251: 2227: 2221: 2220: 2176: 2170: 2169: 2127: 2118: 2112: 2111: 2067: 2061: 2056: 2050: 2045: 2039: 2038: 2028: 2018: 1986: 1980: 1979: 1969: 1959: 1923: 1902: 1901: 1883: 1877: 1876: 1866: 1856: 1832: 1823: 1822: 1778: 1769: 1768: 1758: 1724: 1713: 1712: 1684: 1470:Deinonychosauria 1445:node-based clade 1242: 1234: 1217: 1209: 1191: 1182: 1164: 1156: 1139: 1131: 1113: 1105: 1089:Oviraptorosauria 1087: 1079: 1069: 1064: 1063: 945: 938: 910: 849: 797: 790: 783: 722: 699: 692: 676:Oviraptorosauria 668: 658: 653: 652: 414: 387: 379: 370: 362: 354: 344: 334: 324: 313: 303: 284: 187: 186: 134: 124: 61: 45:Temporal range: 39: 21: 8465: 8464: 8460: 8459: 8458: 8456: 8455: 8454: 8415: 8414: 8413: 8403: 8401: 8389: 8379: 8377: 8367: 8365: 8357: 8355: 8350: 8342: 8337: 8329: 8324: 8315: 8314: 8309: 8300: 8299: 8294: 8281: 8271: 8266: 8215: 8181:Origin of birds 8154: 8094: 8016:Limb morphology 7997: 7988: 7974:Ray-finned fish 7939:Fish locomotion 7915: 7902: 7872: 7867: 7779: 7773: 7765: 7757: 7756:Bucerotiformes 7750:(cuckoo-roller) 7749: 7741: 7733: 7725: 7717: 7709: 7708:Cathartiformes 7694: 7688: 7680: 7679:Psittaciformes 7672: 7664: 7649: 7643: 7642:Pelecaniformes 7635: 7627: 7619: 7611: 7603: 7588: 7582: 7581:Eurypygiformes 7574: 7559: 7553: 7545: 7530: 7524: 7509: 7503: 7485: 7470: 7464: 7456: 7448: 7420: 7414: 7406: 7391: 7385: 7384:Pterocliformes 7377: 7369: 7320: 7309: 7296: 7281: 7205: 7164: 7102: 7067: 7066: 7059: 7058: 7046: 7026: 6999: 6976: 6969: 6929: 6928: 6916: 6915: 6899: 6898: 6895: 6893: 6891: 6889: 6887: 6885: 6883: 6881: 6879: 6877: 6875: 6860: 6858: 6856: 6854: 6852: 6850: 6848: 6846: 6844: 6829: 6823: 6822:Casuariiformes 6815: 6814:Apterygiformes 6807: 6799: 6791: 6768: 6753: 6724:Lists by region 6692: 6602: 6595: 6536:Chaoyangiformes 6521:Jeholornithidae 6495: 6459:Origin of birds 6445: 6426:Brood parasites 6372: 6302: 6289: 6279: 6249: 6244: 6186: 6179: 6162: 6157:Tympanonesiotes 6150:Pseudodontornis 6113:Palaeochenoides 6034:Caspiodontornis 6018:Pelagornithidae 6009: 5970: 5926: 5868: 5845: 5813: 5789: 5762: 5745: 5719: 5699: 5661: 5603: 5598:Parahesperornis 5570:Asiahesperornis 5555: 5486: 5456: 5355: 5342: 5305: 5250: 5217:Archaeornithura 5199: 5160: 5122: 5092: 4845:Archaeorhynchus 4826: 4813: 4796: 4783: 4753: 4733: 4704:Ornithothoraces 4698: 4661: 4635:Changchengornis 4617: 4586: 4564: 4551: 4498: 4470:Anchiornithidae 4368: 4355: 4329: 4323:Dromaeosauridae 4312:Dromaeosauridae 4303: 4297:Anchiornithidae 4257: 4216:Liaoningvenator 4172:Anchiornithidae 4163: 4151:Scansoriopteryx 4118: 4063: 4050: 4033: 3996: 3976: 3933: 3928:Therizinosaurus 3864: 3812: 3807:Lingyuanosaurus 3764:Therizinosauria 3739: 3698: 3664: 3589:Parvicursorinae 3583: 3520:Alvarezsauridae 3514: 3439: 3427: 3363: 3350: 3321: 3301: 3274: 3268:Avemetatarsalia 3253:Avemetatarsalia 3243: 3199: 3198: 3188: 3183: 3153: 3148: 3111:(7507): 79–82. 3102: 3101: 3097: 3053: 3052: 3048: 2996: 2995: 2991: 2981: 2980: 2976: 2938: 2937: 2933: 2918: 2893: 2892: 2885: 2876: 2869: 2823: 2822: 2818: 2782: 2781: 2774: 2752: 2751: 2747: 2713: 2708: 2707: 2703: 2645: 2644: 2635: 2588: 2587: 2583: 2539: 2538: 2531: 2512: 2511: 2507: 2500: 2496: 2450: 2449: 2445: 2408:Biology Letters 2401: 2400: 2396: 2339: 2338: 2334: 2280: 2279: 2275: 2229: 2228: 2224: 2178: 2177: 2173: 2125: 2120: 2119: 2115: 2069: 2068: 2064: 2057: 2053: 2046: 2042: 1988: 1987: 1983: 1925: 1924: 1905: 1898: 1885: 1884: 1880: 1834: 1833: 1826: 1780: 1779: 1772: 1726: 1725: 1716: 1686: 1685: 1681: 1677: 1646:of bird origins 1623:Origin of birds 1619: 1611:have been found 1593:Other than the 1584: 1531:Tetrapterygidae 1486:Eudromaeosauria 1379:oviraptorosaurs 1367:oviraptorosaurs 1335: 1326: 1317: 1308: 1299: 1290: 1281: 1272: 1166:Anchiornithinae 1115:Dromaeosauridae 1049: 1040: 1031: 1022: 1013: 1004: 980: 969:Dromaeosauridae 897: 888: 836: 770: 761: 651: 646: 640: 624:Dromaeosauridae 607: 537: 520:Microraptor gui 507: 482: 450:Middle Jurassic 381:Dromaeosauridae 364:Anchiornithidae 283: 181: 125: 123: 122: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 56: 55: 48:Middle Jurassic 46: 43: 35: 28: 23: 22: 15: 12: 11: 5: 8463: 8461: 8453: 8452: 8447: 8442: 8437: 8432: 8427: 8417: 8416: 8412: 8411: 8399: 8387: 8375: 8352: 8351: 8349: 8348: 8335: 8322: 8307: 8291: 8289: 8283: 8282: 8277: 8268: 8267: 8265: 8264: 8259: 8254: 8249: 8244: 8239: 8234: 8229: 8223: 8221: 8217: 8216: 8214: 8213: 8208: 8203: 8198: 8193: 8188: 8183: 8178: 8173: 8168: 8162: 8160: 8156: 8155: 8153: 8152: 8147: 8145:Pterosaur wing 8142: 8137: 8136: 8135: 8130: 8125: 8115: 8110: 8104: 8102: 8096: 8095: 8093: 8092: 8087: 8082: 8081: 8080: 8070: 8065: 8060: 8059: 8058: 8053: 8048: 8043: 8038: 8033: 8028: 8023: 8013: 8007: 8005: 7999: 7998: 7991: 7989: 7987: 7986: 7981: 7976: 7971: 7966: 7961: 7956: 7951: 7946: 7941: 7936: 7934:Cephalopod fin 7931: 7925: 7923: 7917: 7916: 7903: 7901: 7900: 7893: 7886: 7878: 7869: 7868: 7866: 7865: 7853: 7841: 7829: 7816: 7813: 7812: 7809: 7808: 7805: 7804: 7801: 7800: 7797: 7796: 7793: 7792: 7789: 7788: 7785: 7784: 7781: 7780: 7778: 7777: 7769: 7764:Coraciiformes 7761: 7753: 7745: 7740:Trogoniformes 7737: 7729: 7721: 7713: 7704: 7702: 7696: 7695: 7693: 7692: 7687:Passeriformes 7684: 7676: 7671:Falconiformes 7668: 7663:Cariamiformes 7659: 7657: 7651: 7650: 7648: 7647: 7639: 7631: 7626:Ciconiiformes 7623: 7615: 7607: 7598: 7596: 7590: 7589: 7587: 7586: 7578: 7569: 7567: 7561: 7560: 7558: 7557: 7549: 7540: 7538: 7536:Cursorimorphae 7532: 7531: 7529: 7528: 7519: 7517: 7511: 7510: 7508: 7507: 7499: 7494: 7489: 7480: 7478: 7472: 7471: 7469: 7468: 7460: 7452: 7443: 7441: 7432: 7426: 7425: 7422: 7421: 7419: 7418: 7410: 7401: 7399: 7397:Mirandornithes 7393: 7392: 7390: 7389: 7381: 7373: 7368:Columbiformes 7364: 7362: 7360:Columbimorphae 7353: 7344: 7334: 7333: 7330: 7329: 7326: 7325: 7322: 7321: 7319: 7318: 7313: 7305: 7300: 7295:Meleagridinae 7291: 7289: 7283: 7282: 7280: 7279: 7272: 7265: 7258: 7251: 7244: 7237: 7230: 7223: 7215: 7213: 7211:Odontophoridae 7207: 7206: 7204: 7203: 7196: 7189: 7182: 7174: 7172: 7166: 7165: 7163: 7162: 7155: 7148: 7141: 7134: 7127: 7120: 7112: 7110: 7104: 7103: 7101: 7100: 7095: 7090: 7084: 7082: 7073: 7052: 7051: 7048: 7047: 7045: 7044: 7036: 7034: 7028: 7027: 7025: 7024: 7017: 7009: 7007: 7001: 7000: 6998: 6997: 6992: 6990:Stictonettinae 6987: 6985:Dendrocygninae 6982: 6981: 6980: 6973: 6961: 6960: 6959: 6954: 6949: 6938: 6936: 6919: 6906: 6863: 6835: 6834: 6831: 6830: 6828: 6827: 6819: 6811: 6803: 6795: 6786: 6784: 6770: 6769: 6766: 6755: 6754: 6752: 6751: 6750: 6749: 6744: 6738:Notable birds 6736: 6731: 6726: 6721: 6716: 6711: 6706: 6700: 6698: 6694: 6693: 6691: 6690: 6688:Egg collecting 6685: 6680: 6675: 6670: 6665: 6660: 6655: 6650: 6645: 6640: 6639: 6638: 6628: 6623: 6618: 6613: 6607: 6605: 6597: 6596: 6594: 6593: 6588: 6583: 6578: 6573: 6571:Hesperornithes 6568: 6563: 6558: 6553: 6548: 6546:Ambiortiformes 6543: 6538: 6533: 6531:Enantiornithes 6528: 6523: 6518: 6513: 6505: 6503: 6497: 6496: 6494: 6493: 6488: 6483: 6478: 6473: 6472: 6471: 6466: 6455: 6453: 6447: 6446: 6444: 6443: 6438: 6433: 6428: 6423: 6418: 6413: 6408: 6403: 6398: 6393: 6388: 6382: 6380: 6374: 6373: 6371: 6370: 6365: 6360: 6355: 6350: 6345: 6340: 6335: 6334: 6333: 6323: 6318: 6312: 6310: 6304: 6303: 6301: 6300: 6294: 6291: 6290: 6280: 6278: 6277: 6270: 6263: 6255: 6246: 6245: 6243: 6242: 6232: 6227: 6222: 6217: 6212: 6207: 6202: 6200:Archaeornithes 6195: 6192: 6191: 6188: 6187: 6173: 6171: 6168: 6167: 6164: 6163: 6161: 6160: 6153: 6146: 6142:Protopelicanus 6138: 6131: 6123: 6116: 6109: 6106:Osteodontornis 6102: 6095: 6088: 6080: 6073: 6066: 6058: 6051: 6044: 6037: 6030: 6022: 6020: 6011: 6010: 6008: 6007: 6003:Dromornithidae 5999: 5992: 5983: 5981: 5972: 5971: 5969: 5968: 5961: 5954: 5947: 5939: 5937: 5935:Dromornithidae 5928: 5927: 5925: 5924: 5913: 5910:Perplexicervix 5903: 5892: 5886: 5883:Incertae sedis 5878: 5877: 5874: 5873: 5870: 5869: 5867: 5866: 5865: 5864: 5862:Pangalliformes 5855: 5853: 5851:Pangalliformes 5847: 5846: 5844: 5843: 5842: 5841: 5832: 5830: 5821: 5815: 5814: 5812: 5811: 5803: 5797: 5791: 5790: 5788: 5787: 5786: 5785: 5776: 5774: 5764: 5763: 5758: 5751: 5750: 5747: 5746: 5734: 5732: 5729: 5728: 5725: 5724: 5721: 5720: 5718: 5717: 5716: 5715: 5709: 5707: 5701: 5700: 5698: 5697: 5689: 5682: 5674: 5672: 5663: 5662: 5660: 5659: 5652: 5645: 5637: 5629: 5620: 5618: 5609: 5608: 5605: 5604: 5602: 5601: 5594: 5587: 5580: 5573: 5565: 5563: 5557: 5556: 5554: 5553: 5546: 5539: 5532: 5525: 5518: 5511: 5503: 5497: 5495:Hesperornithes 5488: 5487: 5485: 5484: 5477: 5469: 5467: 5465:Ichthyornithes 5458: 5457: 5455: 5454: 5444: 5434: 5424: 5414: 5404: 5394: 5384: 5374: 5362: 5356: 5351: 5348: 5347: 5344: 5343: 5341: 5340: 5333: 5330:Similiyanornis 5326: 5318: 5316: 5307: 5306: 5304: 5303: 5295: 5287: 5280: 5272: 5263: 5261: 5252: 5251: 5249: 5248: 5241: 5234: 5227: 5220: 5212: 5210: 5201: 5200: 5198: 5197: 5189: 5181: 5173: 5171: 5169:Ambiortiformes 5162: 5161: 5159: 5158: 5151: 5143: 5135: 5133: 5124: 5123: 5121: 5120: 5113: 5105: 5103: 5094: 5093: 5091: 5090: 5087:Zhongjianornis 5080: 5070: 5060: 5049: 5039: 5029: 5019: 5009: 4999: 4989: 4979: 4976:Jianchangornis 4969: 4959: 4949: 4939: 4928: 4918: 4908: 4898: 4888: 4878: 4868: 4858: 4848: 4837: 4827: 4822: 4815: 4814: 4809: 4802: 4801: 4798: 4797: 4776: 4774: 4771: 4770: 4767: 4766: 4763: 4762: 4759: 4758: 4755: 4754: 4752: 4751: 4750: 4749: 4743: 4741: 4735: 4734: 4732: 4731: 4730: 4729: 4727:Enantiornithes 4720: 4718: 4716:Enantiornithes 4706: 4700: 4699: 4697: 4696: 4689: 4682: 4674: 4672: 4663: 4662: 4660: 4659: 4652: 4645: 4642:Confuciusornis 4638: 4630: 4628: 4619: 4618: 4616: 4615: 4608: 4605:Omnivoropteryx 4600: 4598: 4588: 4587: 4585: 4584: 4571: 4565: 4560: 4557: 4556: 4553: 4552: 4550: 4549: 4542: 4535: 4527: 4520: 4511: 4509: 4500: 4499: 4497: 4496: 4485: 4474: 4463: 4453: 4443: 4432: 4422: 4412: 4401: 4390: 4379: 4369: 4364: 4361: 4360: 4357: 4356: 4354: 4353: 4352: 4351: 4342: 4340: 4331: 4330: 4328: 4327: 4326: 4325: 4316: 4314: 4305: 4304: 4302: 4301: 4293: 4286: 4279: 4270: 4268: 4259: 4258: 4256: 4255: 4248: 4241: 4234: 4227: 4220: 4212: 4205: 4198: 4191: 4184: 4176: 4174: 4165: 4164: 4162: 4161: 4154: 4147: 4140: 4132: 4130: 4120: 4119: 4117: 4116: 4106: 4103:Pneumatoraptor 4096: 4085: 4074: 4064: 4059: 4052: 4051: 4046: 4039: 4038: 4035: 4034: 4015: 4013: 4010: 4009: 4006: 4005: 4002: 4001: 3998: 3997: 3995: 3994: 3993: 3992: 3986: 3984: 3978: 3977: 3975: 3974: 3973: 3972: 3963: 3961: 3949: 3943: 3942: 3939: 3938: 3935: 3934: 3932: 3931: 3924: 3917: 3910: 3903: 3900:Neimongosaurus 3896: 3889: 3882: 3874: 3872: 3866: 3865: 3863: 3862: 3855: 3848: 3841: 3834: 3826: 3820: 3814: 3813: 3811: 3810: 3803: 3796: 3788: 3781: 3772: 3766: 3753: 3752: 3749: 3748: 3745: 3744: 3741: 3740: 3738: 3737: 3730: 3723: 3716: 3708: 3706: 3700: 3699: 3697: 3696: 3689: 3682: 3674: 3672: 3666: 3665: 3663: 3662: 3659:Trierarchuncus 3655: 3648: 3641: 3634: 3627: 3619: 3612: 3605: 3597: 3591: 3585: 3584: 3582: 3581: 3573: 3566: 3559: 3555:Bonapartenykus 3551: 3544: 3537: 3528: 3522: 3516: 3515: 3513: 3512: 3505: 3497: 3494:Shishugounykus 3490: 3483: 3476: 3473:Bonapartenykus 3469: 3462: 3454: 3446: 3440: 3432: 3429: 3428: 3426: 3425: 3414: 3404: 3393: 3382: 3370: 3364: 3359: 3352: 3351: 3346: 3339: 3338: 3335: 3334: 3331: 3330: 3327: 3326: 3323: 3322: 3320: 3319: 3318: 3317: 3311: 3309: 3303: 3302: 3300: 3299: 3298: 3297: 3288: 3282: 3276: 3275: 3273: 3272: 3271: 3270: 3261: 3255: 3245: 3244: 3240: 3239: 3230: 3221: 3212: 3206: 3197: 3190: 3189: 3184: 3182: 3181: 3174: 3167: 3159: 3152: 3151:External links 3149: 3147: 3146: 3095: 3046: 2989: 2974: 2947:(4): 429–463. 2931: 2916: 2883: 2867: 2816: 2795:(1): 117–162. 2772: 2745: 2701: 2633: 2581: 2529: 2505: 2494: 2443: 2414:(1): 110–112. 2394: 2332: 2293:(13): 5002–7. 2273: 2222: 2187:(4): 173–177. 2171: 2113: 2062: 2051: 2040: 2001:(1): 232–237. 1981: 1942:(12): e28964. 1903: 1897:978-1119020769 1896: 1878: 1824: 1770: 1714: 1678: 1676: 1673: 1672: 1671: 1663: 1658: 1653: 1647: 1640: 1635: 1630: 1625: 1618: 1615: 1583: 1580: 1457:alvarezsaurids 1393:dromaeosaurids 1371:dromaeosaurids 1346: 1345: 1337: 1336: 1332: 1331: 1328: 1327: 1323: 1322: 1319: 1318: 1314: 1313: 1310: 1309: 1305: 1304: 1301: 1300: 1296: 1295: 1292: 1291: 1287: 1286: 1283: 1282: 1278: 1277: 1274: 1273: 1269: 1268: 1265: 1264: 1257: 1254: 1253: 1250: 1249: 1237: 1232: 1230: 1227: 1226: 1223: 1222: 1212: 1207: 1205: 1202: 1201: 1198: 1197: 1185: 1180: 1178: 1174: 1173: 1170: 1169: 1159: 1154: 1152: 1149: 1148: 1145: 1144: 1134: 1129: 1127: 1123: 1122: 1119: 1118: 1108: 1103: 1101: 1097: 1096: 1093: 1092: 1082: 1077: 1075: 1067: 1060: 1059: 1051: 1050: 1046: 1045: 1042: 1041: 1037: 1036: 1033: 1032: 1028: 1027: 1024: 1023: 1019: 1018: 1015: 1014: 1010: 1009: 1006: 1005: 1001: 1000: 997: 996: 989: 986: 985: 982: 981: 977: 976: 973: 972: 964: 961: 960: 957: 956: 948: 943: 941: 936: 934: 928: 927: 924: 923: 913: 908: 906: 903: 902: 899: 898: 894: 893: 890: 889: 885: 884: 881: 880: 870: 867: 866: 863: 862: 852: 847: 845: 842: 841: 838: 837: 833: 832: 829: 828: 818: 815: 814: 811: 810: 800: 795: 793: 788: 786: 781: 779: 776: 775: 772: 771: 767: 766: 763: 762: 758: 757: 754: 753: 743: 740: 739: 736: 735: 725: 720: 718: 715: 714: 711: 710: 702: 697: 695: 690: 688: 684: 683: 680: 679: 671: 666: 664: 656: 650: 647: 639: 636: 606: 603: 581:eudromaeosaurs 542:eudromaeosaurs 536: 533: 506: 503: 481: 478: 462:anchiornithids 454:dromaeosaurids 426: 425: 424: 423: 406: 405: 399: 398: 397: 396: 391: 383: 375: 366: 358: 350: 340: 337:Pneumatoraptor 330: 320: 309: 296: 295: 291: 290: 278: 271: 270: 265: 258: 257: 252: 245: 244: 239: 232: 231: 226: 219: 218: 213: 209: 208: 203: 199: 198: 193: 189: 188: 175: 174: 142:Confuciusornis 136: 135: 127: 126: 118: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 62: 44: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 8462: 8451: 8448: 8446: 8443: 8441: 8438: 8436: 8433: 8431: 8428: 8426: 8423: 8422: 8420: 8410: 8400: 8398: 8393: 8388: 8386: 8376: 8374: 8364: 8360: 8345: 8340: 8336: 8332: 8327: 8323: 8318: 8312: 8308: 8303: 8297: 8293: 8292: 8290: 8288: 8284: 8280: 8275: 8263: 8260: 8258: 8255: 8253: 8250: 8248: 8245: 8243: 8240: 8238: 8235: 8233: 8230: 8228: 8225: 8224: 8222: 8218: 8212: 8209: 8207: 8204: 8202: 8199: 8197: 8194: 8192: 8189: 8187: 8184: 8182: 8179: 8177: 8174: 8172: 8169: 8167: 8164: 8163: 8161: 8157: 8151: 8148: 8146: 8143: 8141: 8138: 8134: 8131: 8129: 8126: 8124: 8121: 8120: 8119: 8116: 8114: 8111: 8109: 8106: 8105: 8103: 8101: 8097: 8091: 8088: 8086: 8083: 8079: 8076: 8075: 8074: 8071: 8069: 8066: 8064: 8061: 8057: 8054: 8052: 8049: 8047: 8044: 8042: 8039: 8037: 8034: 8032: 8029: 8027: 8024: 8022: 8019: 8018: 8017: 8014: 8012: 8009: 8008: 8006: 8004: 8000: 7995: 7985: 7982: 7980: 7979:Pectoral fins 7977: 7975: 7972: 7970: 7967: 7965: 7962: 7960: 7957: 7955: 7952: 7950: 7947: 7945: 7942: 7940: 7937: 7935: 7932: 7930: 7927: 7926: 7924: 7922: 7918: 7914: 7910: 7906: 7899: 7894: 7892: 7887: 7885: 7880: 7879: 7876: 7864: 7863: 7854: 7852: 7851: 7846: 7842: 7840: 7839: 7830: 7828: 7827: 7818: 7817: 7814: 7776: 7770: 7768: 7762: 7760: 7754: 7752: 7746: 7744: 7738: 7736: 7730: 7728: 7724:Strigiformes 7722: 7720: 7714: 7712: 7706: 7705: 7703: 7701: 7697: 7691: 7685: 7683: 7677: 7675: 7669: 7667: 7661: 7660: 7658: 7656: 7652: 7646: 7640: 7638: 7632: 7630: 7624: 7622: 7616: 7614: 7608: 7606: 7600: 7599: 7597: 7595: 7591: 7585: 7579: 7577: 7575:(tropicbirds) 7571: 7570: 7568: 7566: 7562: 7556: 7550: 7548: 7542: 7541: 7539: 7537: 7533: 7527: 7521: 7520: 7518: 7516: 7512: 7506: 7500: 7498: 7497:Podargiformes 7495: 7493: 7490: 7488: 7482: 7481: 7479: 7477: 7473: 7467: 7461: 7459: 7453: 7451: 7447:Cuculiformes 7445: 7444: 7442: 7440: 7436: 7433: 7431: 7427: 7417: 7411: 7409: 7403: 7402: 7400: 7398: 7394: 7388: 7382: 7380: 7374: 7372: 7366: 7365: 7363: 7361: 7357: 7354: 7352: 7348: 7345: 7343: 7339: 7335: 7317: 7314: 7312: 7306: 7304: 7301: 7299: 7293: 7292: 7290: 7288: 7284: 7278: 7277: 7273: 7271: 7270: 7266: 7264: 7263: 7259: 7257: 7256: 7252: 7250: 7249: 7245: 7243: 7242: 7238: 7236: 7235: 7231: 7229: 7228: 7224: 7222: 7221: 7217: 7216: 7214: 7212: 7208: 7202: 7201: 7197: 7195: 7194: 7190: 7188: 7187: 7183: 7181: 7180: 7176: 7175: 7173: 7171: 7167: 7161: 7160: 7156: 7154: 7153: 7149: 7147: 7146: 7145:Macrocephalon 7142: 7140: 7139: 7135: 7133: 7132: 7128: 7126: 7125: 7121: 7119: 7118: 7114: 7113: 7111: 7109: 7105: 7099: 7096: 7094: 7091: 7089: 7086: 7085: 7083: 7081: 7077: 7074: 7070: 7063: 7057: 7053: 7043: 7042: 7038: 7037: 7035: 7033: 7032:Anseranatidae 7029: 7023: 7022: 7018: 7016: 7015: 7011: 7010: 7008: 7006: 7002: 6996: 6993: 6991: 6988: 6986: 6983: 6979: 6974: 6972: 6967: 6966: 6965: 6962: 6958: 6955: 6953: 6950: 6948: 6945: 6944: 6943: 6940: 6939: 6937: 6933: 6927: 6923: 6920: 6914: 6910: 6907: 6903: 6897: 6871: 6867: 6864: 6862: 6840: 6836: 6826: 6820: 6818: 6812: 6810: 6806:Tinamiformes 6804: 6802: 6796: 6794: 6788: 6787: 6785: 6783: 6782:Palaeognathae 6779: 6775: 6771: 6764: 6760: 6756: 6748: 6745: 6743: 6740: 6739: 6737: 6735: 6732: 6730: 6727: 6725: 6722: 6720: 6717: 6715: 6712: 6710: 6707: 6705: 6702: 6701: 6699: 6695: 6689: 6686: 6684: 6681: 6679: 6676: 6674: 6671: 6669: 6668:Pigeon racing 6666: 6664: 6661: 6659: 6656: 6654: 6651: 6649: 6646: 6644: 6641: 6637: 6634: 6633: 6632: 6629: 6627: 6624: 6622: 6619: 6617: 6614: 6612: 6609: 6608: 6606: 6604: 6598: 6592: 6589: 6587: 6584: 6582: 6579: 6577: 6574: 6572: 6569: 6567: 6564: 6562: 6559: 6557: 6554: 6552: 6549: 6547: 6544: 6542: 6539: 6537: 6534: 6532: 6529: 6527: 6524: 6522: 6519: 6517: 6514: 6512: 6511: 6510:Archaeopteryx 6507: 6506: 6504: 6502: 6498: 6492: 6489: 6487: 6484: 6482: 6479: 6477: 6474: 6470: 6467: 6465: 6462: 6461: 6460: 6457: 6456: 6454: 6452: 6448: 6442: 6439: 6437: 6434: 6432: 6429: 6427: 6424: 6422: 6419: 6417: 6414: 6412: 6409: 6407: 6404: 6402: 6399: 6397: 6394: 6392: 6389: 6387: 6384: 6383: 6381: 6379: 6375: 6369: 6366: 6364: 6361: 6359: 6356: 6354: 6351: 6349: 6346: 6344: 6341: 6339: 6336: 6332: 6329: 6328: 6327: 6324: 6322: 6319: 6317: 6314: 6313: 6311: 6309: 6305: 6299: 6296: 6295: 6292: 6287: 6283: 6276: 6271: 6269: 6264: 6262: 6257: 6256: 6253: 6241: 6233: 6231: 6228: 6226: 6223: 6221: 6220:Odontornithes 6218: 6216: 6215:Odontognathae 6213: 6211: 6208: 6206: 6203: 6201: 6197: 6196: 6193: 6183: 6177: 6159: 6158: 6154: 6152: 6151: 6147: 6144: 6143: 6139: 6137: 6136: 6132: 6129: 6128: 6124: 6122: 6121: 6117: 6115: 6114: 6110: 6108: 6107: 6103: 6101: 6100: 6096: 6094: 6093: 6089: 6086: 6085: 6081: 6079: 6078: 6074: 6072: 6071: 6067: 6064: 6063: 6059: 6057: 6056: 6052: 6050: 6049: 6045: 6043: 6042: 6038: 6036: 6035: 6031: 6029: 6028: 6024: 6023: 6021: 6019: 6012: 6005: 6004: 6000: 5998: 5997: 5993: 5990: 5989: 5985: 5984: 5982: 5980: 5973: 5967: 5966: 5962: 5960: 5959: 5955: 5953: 5952: 5948: 5946: 5945: 5944:Barawertornis 5941: 5940: 5938: 5936: 5929: 5922: 5921: 5914: 5912: 5911: 5904: 5902: 5901: 5894: 5893: 5890: 5887: 5885: 5884: 5879: 5863: 5859: 5858: 5857: 5856: 5854: 5852: 5848: 5840: 5839:Anserimorphae 5836: 5835: 5834: 5833: 5831: 5829: 5828:Anserimorphae 5825: 5822: 5820: 5816: 5810: 5809: 5805: 5804: 5801: 5798: 5796: 5792: 5784: 5783:Palaeognathae 5780: 5779: 5778: 5777: 5775: 5773: 5772:Palaeognathae 5769: 5765: 5761: 5756: 5752: 5743: 5738: 5713: 5712: 5711: 5710: 5708: 5706: 5702: 5695: 5694: 5690: 5688: 5687: 5683: 5681: 5680: 5676: 5675: 5673: 5671: 5664: 5658: 5657: 5653: 5651: 5650: 5646: 5643: 5642: 5638: 5635: 5634: 5630: 5627: 5626: 5622: 5621: 5619: 5617: 5610: 5600: 5599: 5595: 5593: 5592: 5588: 5586: 5585: 5581: 5579: 5578: 5574: 5572: 5571: 5567: 5566: 5564: 5562: 5558: 5552: 5551: 5547: 5545: 5544: 5540: 5538: 5537: 5533: 5531: 5530: 5526: 5524: 5523: 5519: 5517: 5516: 5512: 5510: 5509: 5505: 5504: 5501: 5498: 5496: 5489: 5483: 5482: 5478: 5476: 5475: 5471: 5470: 5468: 5466: 5459: 5453: 5452: 5451:Tingmiatornis 5445: 5443: 5442: 5435: 5433: 5432: 5425: 5423: 5422: 5415: 5413: 5412: 5405: 5403: 5402: 5395: 5393: 5392: 5385: 5383: 5382: 5375: 5372: 5371: 5370:Antarcticavis 5364: 5363: 5360: 5357: 5354: 5349: 5339: 5338: 5334: 5332: 5331: 5327: 5325: 5324: 5320: 5319: 5317: 5315: 5314:Yanornithidae 5308: 5301: 5300: 5296: 5293: 5292: 5288: 5286: 5285: 5284:Songlingornis 5281: 5278: 5277: 5273: 5270: 5269: 5265: 5264: 5262: 5260: 5253: 5247: 5246: 5242: 5240: 5239: 5235: 5233: 5232: 5231:Longicrusavis 5228: 5226: 5225: 5224:Hongshanornis 5221: 5219: 5218: 5214: 5213: 5211: 5209: 5202: 5195: 5194: 5190: 5187: 5186: 5182: 5180: 5179: 5175: 5174: 5172: 5170: 5163: 5157: 5156: 5152: 5149: 5148: 5144: 5142: 5141: 5137: 5136: 5134: 5132: 5125: 5119: 5118: 5114: 5112: 5111: 5110:Mengciusornis 5107: 5106: 5104: 5102: 5101:Schizoouridae 5095: 5089: 5088: 5081: 5079: 5078: 5071: 5069: 5068: 5061: 5058: 5057: 5050: 5048: 5047: 5040: 5038: 5037: 5030: 5028: 5027: 5020: 5018: 5017: 5010: 5008: 5007: 5000: 4998: 4997: 4990: 4988: 4987: 4980: 4978: 4977: 4970: 4968: 4967: 4960: 4958: 4957: 4950: 4948: 4947: 4940: 4937: 4936: 4929: 4927: 4926: 4919: 4917: 4916: 4909: 4907: 4906: 4899: 4897: 4896: 4889: 4887: 4886: 4885:Changzuiornis 4879: 4877: 4876: 4869: 4867: 4866: 4865:Brevidentavis 4859: 4857: 4856: 4849: 4847: 4846: 4839: 4838: 4835: 4831: 4828: 4825: 4820: 4816: 4812: 4807: 4803: 4794: 4790: 4787: 4780: 4747: 4746: 4745: 4744: 4742: 4740: 4736: 4728: 4724: 4723: 4722: 4721: 4719: 4717: 4710: 4707: 4705: 4701: 4695: 4694: 4690: 4688: 4687: 4683: 4681: 4680: 4676: 4675: 4673: 4671: 4664: 4658: 4657: 4653: 4651: 4650: 4646: 4644: 4643: 4639: 4637: 4636: 4632: 4631: 4629: 4627: 4620: 4614: 4613: 4609: 4607: 4606: 4602: 4601: 4599: 4596: 4589: 4583: 4581: 4573: 4572: 4569: 4566: 4563: 4558: 4548: 4547: 4543: 4541: 4540: 4536: 4533: 4532: 4528: 4526: 4525: 4521: 4518: 4517: 4513: 4512: 4510: 4508: 4501: 4494: 4493: 4486: 4483: 4482: 4475: 4472: 4471: 4464: 4462: 4461: 4454: 4452: 4451: 4444: 4441: 4440: 4433: 4431: 4430: 4423: 4421: 4420: 4413: 4410: 4409: 4408:Cretaaviculus 4402: 4399: 4398: 4391: 4389: 4388: 4381: 4380: 4377: 4373: 4370: 4367: 4362: 4350: 4346: 4345: 4344: 4343: 4341: 4339: 4332: 4324: 4320: 4319: 4318: 4317: 4315: 4313: 4306: 4299: 4298: 4294: 4292: 4291: 4287: 4285: 4284: 4283:Archaeopteryx 4280: 4277: 4276: 4272: 4271: 4269: 4267: 4260: 4254: 4253: 4252:Yixianosaurus 4249: 4247: 4246: 4242: 4240: 4239: 4235: 4233: 4232: 4228: 4226: 4225: 4221: 4218: 4217: 4213: 4211: 4210: 4209:Fujianvenator 4206: 4204: 4203: 4199: 4197: 4196: 4192: 4190: 4189: 4185: 4183: 4182: 4178: 4177: 4175: 4173: 4166: 4160: 4159: 4155: 4153: 4152: 4148: 4146: 4145: 4144:Epidexipteryx 4141: 4139: 4138: 4134: 4133: 4131: 4128: 4121: 4115: 4114: 4107: 4105: 4104: 4097: 4094: 4093: 4086: 4084: 4083: 4076: 4075: 4072: 4068: 4065: 4062: 4057: 4053: 4049: 4044: 4040: 4031: 4027: 4025: 4019: 3990: 3989: 3988: 3987: 3985: 3983: 3979: 3971: 3967: 3966: 3965: 3964: 3962: 3960: 3953: 3950: 3948: 3944: 3930: 3929: 3925: 3923: 3922: 3918: 3916: 3915: 3911: 3909: 3908: 3904: 3902: 3901: 3897: 3895: 3894: 3890: 3888: 3887: 3883: 3881: 3880: 3876: 3875: 3873: 3871: 3867: 3861: 3860: 3856: 3854: 3853: 3849: 3847: 3846: 3842: 3840: 3839: 3838:Beipiaosaurus 3835: 3833: 3832: 3828: 3827: 3824: 3821: 3819: 3815: 3809: 3808: 3804: 3802: 3801: 3797: 3794: 3793: 3789: 3787: 3786: 3782: 3779: 3778: 3774: 3773: 3770: 3767: 3765: 3758: 3754: 3736: 3735: 3731: 3729: 3728: 3724: 3722: 3721: 3717: 3715: 3714: 3710: 3709: 3707: 3705: 3701: 3695: 3694: 3690: 3688: 3687: 3683: 3681: 3680: 3676: 3675: 3673: 3671: 3667: 3661: 3660: 3656: 3654: 3653: 3649: 3647: 3646: 3642: 3640: 3639: 3635: 3633: 3632: 3628: 3625: 3624: 3620: 3618: 3617: 3613: 3611: 3610: 3606: 3604: 3603: 3599: 3598: 3595: 3592: 3590: 3586: 3579: 3578: 3574: 3572: 3571: 3570:Heptasteornis 3567: 3565: 3564: 3560: 3557: 3556: 3552: 3550: 3549: 3548:Alvarezsaurus 3545: 3543: 3542: 3538: 3535: 3534: 3533:Achillesaurus 3530: 3529: 3526: 3523: 3521: 3517: 3511: 3510: 3506: 3503: 3502: 3498: 3496: 3495: 3491: 3489: 3488: 3484: 3482: 3481: 3477: 3475: 3474: 3470: 3468: 3467: 3463: 3460: 3459: 3455: 3453: 3452: 3451:Achillesaurus 3448: 3447: 3444: 3441: 3438: 3430: 3423: 3422: 3415: 3413: 3412: 3405: 3402: 3401: 3394: 3391: 3390: 3383: 3380: 3379: 3372: 3371: 3368: 3365: 3362: 3357: 3353: 3349: 3344: 3340: 3315: 3314: 3313: 3312: 3310: 3308: 3304: 3296: 3292: 3291: 3290: 3289: 3286: 3283: 3281: 3277: 3269: 3265: 3264: 3263: 3262: 3259: 3256: 3254: 3250: 3246: 3238: 3234: 3231: 3229: 3225: 3222: 3220: 3216: 3213: 3211: 3207: 3205: 3201: 3200: 3195: 3191: 3187: 3180: 3175: 3173: 3168: 3166: 3161: 3160: 3157: 3150: 3142: 3138: 3134: 3130: 3126: 3122: 3118: 3114: 3110: 3106: 3099: 3096: 3091: 3087: 3082: 3077: 3073: 3069: 3065: 3061: 3060:Proc Biol Sci 3057: 3050: 3047: 3042: 3038: 3033: 3028: 3024: 3020: 3016: 3012: 3008: 3004: 3000: 2993: 2990: 2985: 2978: 2975: 2970: 2966: 2962: 2958: 2954: 2950: 2946: 2942: 2935: 2932: 2927: 2923: 2919: 2913: 2909: 2905: 2901: 2897: 2890: 2888: 2884: 2880: 2874: 2872: 2868: 2863: 2859: 2855: 2851: 2847: 2843: 2839: 2835: 2831: 2827: 2820: 2817: 2812: 2808: 2803: 2798: 2794: 2790: 2786: 2779: 2777: 2773: 2768: 2764: 2760: 2756: 2749: 2746: 2741: 2737: 2733: 2729: 2725: 2721: 2720: 2712: 2705: 2702: 2697: 2693: 2689: 2685: 2681: 2677: 2673: 2669: 2665: 2661: 2657: 2653: 2649: 2642: 2640: 2638: 2634: 2629: 2625: 2621: 2617: 2613: 2609: 2605: 2601: 2597: 2593: 2585: 2582: 2577: 2573: 2568: 2563: 2559: 2555: 2551: 2547: 2543: 2536: 2534: 2530: 2525: 2521: 2520: 2515: 2509: 2506: 2503: 2498: 2495: 2490: 2486: 2482: 2478: 2474: 2470: 2466: 2462: 2458: 2454: 2447: 2444: 2439: 2435: 2430: 2425: 2421: 2417: 2413: 2409: 2405: 2398: 2395: 2392: 2387: 2383: 2378: 2373: 2368: 2363: 2359: 2355: 2351: 2347: 2343: 2336: 2333: 2328: 2324: 2319: 2314: 2309: 2304: 2300: 2296: 2292: 2288: 2284: 2277: 2274: 2269: 2265: 2260: 2255: 2250: 2245: 2241: 2237: 2233: 2226: 2223: 2218: 2214: 2210: 2206: 2202: 2198: 2194: 2190: 2186: 2182: 2175: 2172: 2167: 2163: 2159: 2155: 2151: 2147: 2143: 2139: 2135: 2131: 2124: 2117: 2114: 2109: 2105: 2101: 2097: 2093: 2089: 2085: 2081: 2077: 2073: 2066: 2063: 2060: 2055: 2052: 2049: 2044: 2041: 2036: 2032: 2027: 2022: 2017: 2012: 2008: 2004: 2000: 1996: 1992: 1985: 1982: 1977: 1973: 1968: 1963: 1958: 1953: 1949: 1945: 1941: 1937: 1933: 1931: 1922: 1920: 1918: 1916: 1914: 1912: 1910: 1908: 1904: 1899: 1893: 1889: 1882: 1879: 1874: 1870: 1865: 1860: 1855: 1850: 1846: 1842: 1838: 1831: 1829: 1825: 1820: 1816: 1812: 1808: 1804: 1800: 1796: 1792: 1789:(3): 185–91. 1788: 1784: 1777: 1775: 1771: 1766: 1762: 1757: 1752: 1748: 1747:10.1206/748.1 1744: 1740: 1736: 1735: 1730: 1723: 1721: 1719: 1715: 1710: 1706: 1702: 1698: 1694: 1690: 1683: 1680: 1674: 1669: 1668: 1664: 1662: 1659: 1657: 1654: 1651: 1648: 1645: 1641: 1639: 1636: 1634: 1631: 1629: 1626: 1624: 1621: 1620: 1616: 1614: 1612: 1608: 1607:Archaeopteryx 1604: 1600: 1596: 1591: 1589: 1581: 1579: 1576: 1572: 1571:Eumaniraptora 1568: 1564: 1560: 1559: 1554: 1553: 1548: 1547: 1542: 1541: 1536: 1532: 1527: 1525: 1521: 1517: 1513: 1512: 1511:Archaeopteryx 1507: 1503: 1502: 1501:Dromaeosaurus 1497: 1496: 1491: 1487: 1483: 1479: 1473: 1471: 1466: 1464: 1463: 1462:Epidexipteryx 1458: 1454: 1450: 1449:Eumaniraptora 1446: 1441: 1439: 1438: 1433: 1429: 1425: 1421: 1417: 1413: 1409: 1404: 1402: 1401:natural group 1398: 1394: 1390: 1389: 1388:Archaeopteryx 1384: 1380: 1376: 1372: 1368: 1364: 1360: 1356: 1352: 1343: 1339: 1338: 1330: 1329: 1321: 1320: 1312: 1311: 1303: 1302: 1294: 1293: 1285: 1284: 1276: 1275: 1267: 1266: 1263: 1262: 1256: 1255: 1252: 1251: 1248: 1247: 1246: 1236: 1235: 1229: 1228: 1225: 1224: 1221: 1220: 1211: 1210: 1204: 1203: 1200: 1199: 1196: 1195: 1194: 1193:Archaeopteryx 1184: 1183: 1176: 1175: 1172: 1171: 1168: 1167: 1158: 1157: 1151: 1150: 1147: 1146: 1143: 1142: 1133: 1132: 1125: 1124: 1121: 1120: 1117: 1116: 1107: 1106: 1099: 1098: 1095: 1094: 1091: 1090: 1081: 1080: 1074: 1071: 1070: 1066: 1065: 1057: 1053: 1052: 1044: 1043: 1035: 1034: 1026: 1025: 1017: 1016: 1008: 1007: 999: 998: 995: 994: 988: 987: 984: 983: 975: 974: 971: 970: 963: 962: 959: 958: 955: 954: 947: 946: 940: 939: 933: 932:Eumaniraptora 930: 929: 926: 925: 922: 921: 920: 912: 911: 905: 904: 901: 900: 892: 891: 883: 882: 879: 878: 877: 869: 868: 865: 864: 861: 860: 859: 851: 850: 844: 843: 840: 839: 831: 830: 827: 826: 825: 817: 816: 813: 812: 809: 808: 807: 799: 798: 792: 791: 785: 784: 778: 777: 774: 773: 765: 764: 756: 755: 752: 751: 750: 749:Yixianosaurus 742: 741: 738: 737: 734: 733: 732: 724: 723: 717: 716: 713: 712: 709: 708: 701: 700: 694: 693: 686: 685: 682: 681: 678: 677: 670: 669: 663: 660: 659: 655: 654: 649:Relationships 648: 645: 637: 635: 633: 629: 625: 621: 620:Eumaniraptora 617: 612: 604: 602: 598: 596: 595: 588: 586: 582: 578: 570:"sickle claw" 569: 568: 563: 559: 557: 553: 552: 551:Balaur bondoc 547: 543: 534: 532: 530: 526: 522: 521: 515: 513: 504: 502: 500: 499:Coelurosauria 494: 492: 491: 485: 479: 477: 475: 471: 467: 463: 459: 455: 451: 447: 444: 440: 432: 420: 417: 413: 412:Eumaniraptora 410: 409: 407: 404: 400: 395: 392: 390: 384: 382: 376: 373: 367: 365: 359: 357: 351: 349: 348: 341: 339: 338: 331: 329: 328: 321: 318: 317: 310: 308: 307: 300: 299: 297: 292: 287: 282: 279: 276: 273: 272: 269: 266: 263: 260: 259: 256: 253: 250: 247: 246: 243: 240: 237: 234: 233: 230: 227: 224: 221: 220: 217: 214: 211: 210: 207: 204: 201: 200: 197: 194: 191: 190: 185: 180: 176: 173: 170: 166: 162: 161: 156: 155: 150: 149: 148:Dromaeosaurus 144: 143: 137: 133: 128: 121: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 60: 53: 49: 40: 37: 33: 19: 18:Eumaniraptora 8409:Paleontology 8286: 7860: 7848: 7836: 7824: 7734:(mousebirds) 7732:Coliiformes 7602:Gaviiformes 7594:Aequornithes 7502:Apodiformes 7463:Otidiformes 7439:Otidimorphae 7386:(sandgrouse) 7308:Phasianinae 7274: 7267: 7260: 7255:Odontophorus 7253: 7246: 7239: 7232: 7225: 7218: 7198: 7191: 7184: 7177: 7157: 7150: 7143: 7136: 7129: 7122: 7115: 7093:Oreophasinae 7039: 7019: 7012: 6917:(waterfowls) 6913:Anseriformes 6663:Cockfighting 6648:Conservation 6643:Bird feeding 6631:Birdwatching 6621:Ornithomancy 6561:Gansuiformes 6508: 6501:Fossil birds 6391:Intelligence 6230:Unenlagiidae 6155: 6148: 6140: 6133: 6125: 6118: 6111: 6104: 6097: 6092:Odontopteryx 6090: 6082: 6075: 6068: 6060: 6053: 6046: 6039: 6032: 6025: 6001: 5994: 5986: 5963: 5956: 5949: 5942: 5918: 5908: 5898: 5881: 5819:Galloanserae 5806: 5693:Lamarqueavis 5691: 5686:Cimolopteryx 5684: 5677: 5654: 5647: 5639: 5631: 5625:Australornis 5623: 5596: 5589: 5582: 5575: 5568: 5548: 5543:Pasquiaornis 5541: 5534: 5527: 5520: 5513: 5506: 5479: 5472: 5449: 5439: 5429: 5419: 5409: 5399: 5389: 5379: 5368: 5335: 5328: 5321: 5297: 5289: 5282: 5276:Piscivoravis 5274: 5266: 5243: 5236: 5229: 5222: 5215: 5191: 5183: 5176: 5155:Patagopteryx 5153: 5145: 5138: 5115: 5108: 5085: 5075: 5067:Xinghaiornis 5065: 5054: 5044: 5034: 5024: 5016:Khinganornis 5014: 5004: 4994: 4986:Jiuquanornis 4984: 4974: 4964: 4954: 4944: 4935:Gargantuavis 4933: 4923: 4913: 4903: 4893: 4883: 4875:Changmaornis 4873: 4863: 4853: 4843: 4782: 4693:Jinguofortis 4691: 4684: 4677: 4654: 4647: 4640: 4633: 4610: 4603: 4577: 4546:Neimengornis 4544: 4537: 4531:Jixiangornis 4529: 4522: 4516:Dalianraptor 4514: 4490: 4479: 4468: 4458: 4450:Yandangornis 4448: 4437: 4427: 4417: 4406: 4395: 4385: 4349:Troodontidae 4338:Troodontidae 4295: 4290:Wellnhoferia 4288: 4281: 4273: 4250: 4243: 4236: 4229: 4222: 4214: 4207: 4202:Eosinopteryx 4200: 4193: 4186: 4179: 4156: 4149: 4142: 4135: 4111: 4101: 4092:Palaeopteryx 4090: 4080: 4060: 4047: 4021: 3981: 3947:Pennaraptora 3926: 3919: 3912: 3907:Nothronychus 3905: 3898: 3891: 3886:Erlikosaurus 3884: 3879:Erliansaurus 3877: 3859:Suzhousaurus 3857: 3852:Martharaptor 3850: 3845:Enigmosaurus 3843: 3836: 3829: 3805: 3798: 3792:Fukuivenator 3790: 3783: 3777:Eshanosaurus 3775: 3732: 3725: 3718: 3713:Albertonykus 3711: 3691: 3684: 3677: 3670:Ceratonykini 3657: 3650: 3643: 3636: 3631:Nemegtonykus 3629: 3621: 3614: 3607: 3600: 3575: 3568: 3561: 3553: 3546: 3539: 3531: 3507: 3501:Tugulusaurus 3499: 3492: 3485: 3480:Haplocheirus 3478: 3471: 3464: 3456: 3449: 3419: 3411:Migmanychion 3409: 3398: 3389:Fukuivenator 3387: 3376: 3232: 3223: 3214: 3108: 3104: 3098: 3063: 3059: 3049: 3006: 3002: 2992: 2983: 2977: 2944: 2940: 2934: 2899: 2895: 2829: 2825: 2819: 2792: 2788: 2758: 2754: 2748: 2723: 2717: 2704: 2655: 2651: 2598:(9–10): 74. 2595: 2591: 2584: 2549: 2545: 2517: 2508: 2497: 2456: 2452: 2446: 2411: 2407: 2397: 2349: 2345: 2335: 2290: 2286: 2276: 2239: 2235: 2225: 2184: 2180: 2174: 2133: 2129: 2116: 2075: 2071: 2065: 2054: 2043: 1998: 1994: 1984: 1939: 1935: 1929: 1887: 1881: 1864:11336/130197 1844: 1840: 1786: 1782: 1738: 1732: 1692: 1688: 1682: 1665: 1606: 1599:stem lineage 1592: 1585: 1574: 1570: 1566: 1556: 1550: 1544: 1538: 1534: 1528: 1520:Troodontidae 1515: 1509: 1499: 1493: 1489: 1481: 1477: 1474: 1467: 1460: 1448: 1442: 1435: 1419: 1418:, plural of 1415: 1411: 1407: 1405: 1386: 1385:, including 1350: 1349: 1341: 1259: 1243: 1239: 1214: 1188: 1187: 1161: 1141:Troodontidae 1136: 1110: 1084: 1073:Pennaraptora 1055: 991: 966: 953:Troodontidae 950: 917: 915: 876:Eosinopteryx 874: 872: 856: 854: 822: 820: 804: 802: 747: 745: 729: 727: 704: 673: 662:Pennaraptora 628:Troodontidae 615: 608: 599: 592: 589: 573: 565: 555: 549: 538: 528: 524: 518: 516: 512:coelurosaurs 508: 495: 488: 486: 483: 438: 437: 418: 411: 389:Troodontidae 345: 335: 327:Palaeopteryx 325: 314: 304: 280: 274: 268:Pennaraptora 261: 248: 235: 222: 169:Prince Creek 158: 152: 146: 140: 36: 8311:Wikispecies 8140:Insect wing 8090:Webbed foot 8031:unguligrade 8026:plantigrade 8021:digitigrade 7862:WikiProject 7772:Piciformes 7655:Australaves 7634:Suliformes 7552:Gruiformes 7407:(flamingos) 7316:Tetraoninae 7287:Phasianidae 7276:Rhynchortyx 7241:Dactylortyx 7108:Megapodidae 7098:Penelopinae 7056:Galliformes 6798:Rheiformes 6792:(ostriches) 6742:individuals 6616:Ornithology 6603:interaction 6358:Preen gland 6127:Proceriavis 6099:Odontoptila 6055:Gigantornis 5965:Ilbandornis 5900:Asteriornis 5641:Neogaeornis 5591:Hesperornis 5522:Chupkaornis 5474:Ichthyornis 5299:Yixianornis 5245:Tianyuornis 5193:Palintropus 5140:Alamitornis 5026:Meemannavis 4996:Juehuaornis 4915:Eogranivora 4855:Bellulornis 4679:Chongmingia 4429:Overoraptor 4419:Fukuipteryx 4082:Imperobator 3921:Segnosaurus 3686:Ceratonykus 3645:Parvicursor 3616:Khulsanurus 3609:Jaculinykus 3577:Patagonykus 3487:Patagonykus 3421:Yaverlandia 3361:Maniraptora 3348:Maniraptora 3307:Maniraptora 3186:Maniraptora 2726:: 435–489. 1930:Deinonychus 1595:crown group 1540:Microraptor 1428:Maniraptora 1397:troodontids 1375:troodontids 1126:Averaptora 594:Deinonychus 567:Deinonychus 529:Microraptor 525:Microraptor 480:Description 458:troodontids 316:Overoraptor 306:Imperobator 154:Microraptor 57:165–0  8419:Categories 8068:Cephalopod 7984:Pelvic fin 7954:Dorsal fin 7949:Caudal fin 7612:(penguins) 7465:(bustards) 7303:Perdicinae 7248:Dendrortyx 7220:Callipepla 7152:Megapodius 7117:Aepypodius 6995:Tadorninae 6977:true geese 6808:(tinamous) 6767:Neornithes 6678:Pheasantry 6653:Aviculture 6421:Incubation 6411:Lek mating 6198:See also: 6120:Pelagornis 5988:Brontornis 5795:Neognathae 5714:see below↓ 5679:Ceramornis 5649:Polarornis 5616:Vegaviidae 5584:Fumicollis 5550:Potamornis 5536:Judinornis 5529:Enaliornis 5353:Ornithurae 5323:Abitusavis 5117:Schizooura 5077:Yumenornis 5036:Platanavis 5006:Kaririavis 4956:Horezmavis 4895:Chaoyangia 4824:Euornithes 4811:Euornithes 4748:see below↓ 4739:Euornithes 4686:Cratonavis 4562:Pygostylia 4539:Kompsornis 4524:Jeholornis 4460:Zhongornis 4387:Alcmonavis 4275:Alcmonavis 4245:Xiaotingia 4238:Serikornis 4181:Anchiornis 4137:Ambopteryx 3991:see below↓ 3831:Alxasaurus 3720:Linhenykus 3704:Mononykini 3693:Xixianykus 3652:Qiupanykus 3638:Ondogurvel 3602:Dzharaonyx 3563:Bradycneme 3541:Alnashetri 3316:see below↓ 3219:Dinosauria 2898:Anchiornis 1675:References 1575:Averaptora 1558:Anchiornis 1546:Xiaotingia 1490:Averaptora 1488:was named 1482:Xiaotingia 1478:Anchiornis 1432:Neornithes 1430:closer to 1261:Pygostylia 919:Anchiornis 858:Serikornis 731:Xiaotingia 642:See also: 611:progenesis 577:metatarsus 294:Subgroups 242:Saurischia 229:Dinosauria 172:troodontid 167:, and the 160:Anchiornis 8425:Paravians 8385:Dinosaurs 8159:Evolution 8118:Bird wing 8063:Arthropod 8056:quadruped 7681:(parrots) 7525:(hoatzin) 7476:Strisores 7457:(turacos) 7449:(cuckoos) 7378:(mesites) 7297:(turkeys) 7269:Philortyx 7186:Agelastes 7179:Acryllium 7170:Numididae 7159:Talegalla 7069:gamebirds 7062:landfowls 7041:Anseranas 7005:Anhimidae 6964:Anserinae 6747:fictional 6469:dinosaurs 6464:Theropoda 6451:Evolution 6396:Migration 6378:Behaviour 6225:Sauriurae 6205:Carinatae 6041:Cyphornis 6027:Aequornis 5996:Gastornis 5958:Genyornis 5951:Dromornis 5508:Baptornis 5441:Limenavis 5421:Iaceornis 5411:Guildavis 5401:Gallornis 5391:Cerebavis 5381:Apatornis 5185:Apsaravis 5178:Ambiortus 5147:Kuszholia 4612:Sapeornis 4439:Rahonavis 4231:Pedopenna 4113:Rahonavis 3785:Falcarius 3727:Mononykus 3679:Albinykus 3509:Xiyunykus 3378:Elopteryx 3295:Theropoda 3280:Theropoda 3228:Theropoda 3202:Kingdom: 2926:199493087 2761:: 41–83. 2680:1476-4687 2242:: e2159. 2108:205218015 1873:2296-6463 1756:2246/6352 1741:: 1–206. 1437:Oviraptor 1359:dinosaurs 1344:. (2017) 1245:Rahonavis 806:Pedopenna 638:Evolution 556:B. bondoc 446:dinosaurs 347:Rahonavis 255:Theropoda 202:Kingdom: 196:Eukaryota 42:Paravians 8331:42329848 8296:Wikidata 8150:Wingspan 8133:feathers 8128:skeleton 8113:Bat wing 8073:Tetrapod 7959:Fish fin 7826:Category 7700:Afroaves 7628:(storks) 7430:Passerea 7415:(grebes) 7351:Columbea 7262:Oreortyx 7234:Cyrtonyx 7124:Alectura 7088:Cracinae 7080:Cracidae 6957:Oxyurini 6947:Aythyini 6942:Anatinae 6926:Anatidae 6673:Falconry 6636:big year 6491:Seabirds 6401:Foraging 6348:Feathers 6240:Category 6048:Dasornis 5920:Qinornis 5577:Canadaga 5515:Brodavis 5337:Yanornis 5291:Yanornis 5268:Hollanda 4966:Iteravis 4946:Hollanda 4905:Dingavis 4656:Yangavis 4580:Proornis 4224:Ostromia 4188:Aurornis 3734:Shuvuuia 3466:Bannykus 3210:Chordata 3208:Phylum: 3204:Animalia 3133:24990749 3090:15705554 3041:26824065 2969:83726237 2854:23719374 2811:21437379 2688:29211712 2628:32780661 2620:28831510 2576:17823350 2524:Archived 2481:22722850 2438:17148340 2386:20805514 2327:19289829 2268:27441115 2209:15685441 2158:12540892 2100:19794491 2035:21173263 1976:22194962 1936:PLOS ONE 1819:16380823 1811:17952398 1765:83572446 1617:See also 1603:inferred 1552:Aurornis 1498:than to 1365:than to 1177:Avialae 1100:Paraves 824:Aurornis 687:Paraves 585:crampons 470:avialans 443:theropod 403:Synonyms 216:Chordata 212:Phylum: 206:Animalia 192:Domain: 8359:Portals 8317:Paraves 8302:Q136586 8287:Paraves 8220:Related 8078:dactyly 7964:Flipper 7838:Commons 7342:Neoaves 7227:Colinus 7193:Guttera 7131:Eulipoa 6952:Mergini 6816:(kiwis) 6800:(rheas) 6611:Ringing 6436:Hybrids 6431:Nesting 6386:Singing 6363:Plumage 6338:Dactyly 6308:Anatomy 6298:Outline 6288:: Aves) 6062:Laornis 5808:Neoaves 5656:Vegavis 5481:Janavis 5056:Wyleyia 4366:Avialae 4195:Caihong 4061:Paraves 4048:Paraves 3982:Paraves 3141:4464659 3113:Bibcode 3081:1634967 3032:4730849 3011:Bibcode 2949:Bibcode 2862:4364892 2834:Bibcode 2728:Bibcode 2696:4471941 2660:Bibcode 2600:Bibcode 2554:Bibcode 2546:Science 2489:4370675 2461:Bibcode 2429:1617199 2377:2932599 2354:Bibcode 2318:2664043 2295:Bibcode 2259:4941780 2189:Bibcode 2166:1160118 2138:Bibcode 2080:Bibcode 2026:3017133 2003:Bibcode 1967:3237572 1944:Bibcode 1791:Bibcode 1697:Bibcode 1650:Proavis 1567:Paraves 1563:Avialae 1524:Avialae 1506:Avialae 1447:called 1383:Avialae 1351:Paraves 1058:(2017) 993:Avialae 632:Avialae 622:, both 439:Paraves 394:Avialae 281:Paraves 52:Present 32:Panaves 8242:Samara 8051:triped 8036:uniped 7850:Portal 7726:(owls) 7200:Numida 7138:Leipoa 7021:Chauna 7014:Anhima 6709:Genera 6683:Imping 6601:Human 6441:Colony 6368:Vision 6353:Flight 5633:Maaqwi 5431:Kookne 5046:Vorona 4925:Gansus 4397:Balaur 3400:Kakuru 3139:  3131:  3105:Nature 3088:  3078:  3039:  3029:  2967:  2924:  2914:  2860:  2852:  2826:Nature 2809:  2694:  2686:  2678:  2652:Nature 2626:  2618:  2574:  2487:  2479:  2453:Nature 2436:  2426:  2384:  2374:  2325:  2315:  2266:  2256:  2217:789908 2215:  2207:  2164:  2156:  2130:Nature 2106:  2098:  2072:Nature 2033:  2023:  1974:  1964:  1894:  1871:  1817:  1809:  1763:  1573:, and 1555:, and 1495:Passer 1480:, and 1424:Sereno 1373:, and 1056:et al. 546:hallux 421:. 1999 416:Padian 288:, 1997 286:Sereno 8373:Birds 8344:64113 8100:Wings 8085:Digit 8041:biped 8003:Limbs 7913:wings 7909:limbs 6970:swans 6932:ducks 6902:fowls 6697:Lists 6321:Brain 6286:class 6282:Birds 3458:Aorun 3233:Clade 3224:Clade 3215:Clade 3137:S2CID 2965:S2CID 2922:S2CID 2858:S2CID 2714:(PDF) 2692:S2CID 2624:S2CID 2485:S2CID 2236:PeerJ 2213:S2CID 2162:S2CID 2126:(PDF) 2104:S2CID 1815:S2CID 1761:S2CID 1516:et al 1363:birds 1353:is a 1342:et al 616:et al 505:Wings 474:birds 419:et al 275:Clade 262:Clade 249:Clade 236:Clade 223:Clade 8232:Gait 8123:keel 7921:Fins 7911:and 7905:Fins 6343:Eggs 6331:milk 6326:Crop 6316:Beak 6084:Manu 5860:see 5837:see 5781:see 4725:see 4347:see 4321:see 3968:see 3293:see 3266:see 3129:PMID 3086:PMID 3037:PMID 2912:ISBN 2850:PMID 2807:PMID 2684:PMID 2676:ISSN 2616:PMID 2572:PMID 2477:PMID 2434:PMID 2382:PMID 2323:PMID 2287:PNAS 2264:PMID 2205:PMID 2154:PMID 2096:PMID 2031:PMID 1972:PMID 1892:ISBN 1869:ISSN 1807:PMID 1642:The 1522:and 1455:and 1420:avis 1416:aves 1412:par' 1408:pará 1395:and 626:and 165:crow 163:, a 65:PreꞒ 8326:EoL 3623:Kol 3121:doi 3109:511 3076:PMC 3068:doi 3064:272 3027:PMC 3019:doi 2957:doi 2904:doi 2842:doi 2830:498 2797:doi 2763:doi 2759:210 2736:doi 2668:doi 2656:552 2608:doi 2596:104 2562:doi 2550:317 2469:doi 2457:487 2424:PMC 2416:doi 2372:PMC 2362:doi 2350:107 2313:PMC 2303:doi 2291:106 2254:PMC 2244:doi 2197:doi 2146:doi 2134:421 2088:doi 2076:461 2021:PMC 2011:doi 1999:108 1962:PMC 1952:doi 1859:hdl 1849:doi 1799:doi 1751:hdl 1743:doi 1739:371 1705:doi 1588:Mya 579:in 8421:: 8341:: 8328:: 8313:: 8298:: 7907:, 4158:Yi 3235:: 3226:: 3217:: 3135:. 3127:. 3119:. 3107:. 3084:. 3074:. 3062:. 3058:. 3035:. 3025:. 3017:. 3005:. 3001:. 2963:. 2955:. 2943:. 2920:. 2910:. 2886:^ 2870:^ 2856:. 2848:. 2840:. 2828:. 2805:. 2793:83 2791:. 2787:. 2775:^ 2757:. 2734:. 2724:25 2722:. 2716:. 2690:. 2682:. 2674:. 2666:. 2654:. 2650:. 2636:^ 2622:. 2614:. 2606:. 2594:. 2570:. 2560:. 2548:. 2544:. 2532:^ 2522:. 2516:. 2483:. 2475:. 2467:. 2455:. 2432:. 2422:. 2410:. 2406:. 2380:. 2370:. 2360:. 2348:. 2344:. 2321:. 2311:. 2301:. 2289:. 2285:. 2262:. 2252:. 2238:. 2234:. 2211:. 2203:. 2195:. 2185:92 2183:. 2160:. 2152:. 2144:. 2132:. 2128:. 2102:. 2094:. 2086:. 2074:. 2029:. 2019:. 2009:. 1997:. 1993:. 1970:. 1960:. 1950:. 1938:. 1934:. 1906:^ 1867:. 1857:. 1847:. 1843:. 1839:. 1827:^ 1813:. 1805:. 1797:. 1787:95 1785:. 1773:^ 1759:. 1749:. 1737:. 1731:. 1717:^ 1703:. 1693:53 1691:. 1569:, 1565:. 1549:, 1543:, 1526:. 1443:A 1440:. 1410:, 1403:. 460:, 456:, 277:: 264:: 251:: 238:: 225:: 157:, 151:, 145:, 115:Pg 59:Ma 54:, 8361:: 7897:e 7890:t 7883:v 7071:) 7064:- 7060:( 6934:) 6930:( 6904:) 6900:( 6896:e 6894:a 6892:r 6890:e 6888:s 6886:n 6884:a 6882:o 6880:l 6878:l 6876:a 6874:G 6861:e 6859:a 6857:h 6855:t 6853:a 6851:n 6849:g 6847:o 6845:e 6843:N 6284:( 6274:e 6267:t 6260:v 6145:? 6130:? 6087:? 6065:? 6015:† 6006:? 5991:? 5976:† 5932:† 5923:? 5916:† 5906:† 5896:† 5696:? 5667:† 5644:? 5636:? 5628:? 5613:† 5492:† 5462:† 5447:† 5437:† 5427:† 5417:† 5407:† 5397:† 5387:† 5377:† 5373:? 5366:† 5311:† 5302:? 5294:? 5279:? 5271:? 5256:† 5205:† 5196:? 5188:? 5166:† 5150:? 5128:† 5098:† 5083:† 5073:† 5063:† 5059:? 5052:† 5042:† 5032:† 5022:† 5012:† 5002:† 4992:† 4982:† 4972:† 4962:† 4952:† 4942:† 4938:? 4931:† 4921:† 4911:† 4901:† 4891:† 4881:† 4871:† 4861:† 4851:† 4841:† 4713:† 4667:† 4623:† 4597:? 4592:† 4582:" 4578:" 4575:† 4534:? 4519:? 4504:† 4495:? 4488:† 4484:? 4477:† 4473:? 4466:† 4456:† 4446:† 4442:? 4435:† 4425:† 4415:† 4411:? 4404:† 4400:? 4393:† 4383:† 4335:† 4309:† 4300:? 4278:? 4263:† 4219:? 4169:† 4129:? 4124:† 4109:† 4099:† 4095:? 4088:† 4078:† 3956:† 3795:? 3780:? 3761:† 3626:? 3580:? 3558:? 3536:? 3504:? 3461:? 3434:† 3424:? 3417:† 3407:† 3403:? 3396:† 3392:? 3385:† 3381:? 3374:† 3178:e 3171:t 3164:v 3143:. 3123:: 3115:: 3092:. 3070:: 3043:. 3021:: 3013:: 3007:1 2971:. 2959:: 2951:: 2945:5 2928:. 2906:: 2864:. 2844:: 2836:: 2813:. 2799:: 2769:. 2765:: 2742:. 2738:: 2730:: 2698:. 2670:: 2662:: 2630:. 2610:: 2602:: 2578:. 2564:: 2556:: 2491:. 2471:: 2463:: 2440:. 2418:: 2412:2 2388:. 2364:: 2356:: 2329:. 2305:: 2297:: 2270:. 2246:: 2240:4 2219:. 2199:: 2191:: 2168:. 2148:: 2140:: 2110:. 2090:: 2082:: 2037:. 2013:: 2005:: 1978:. 1954:: 1946:: 1940:6 1900:. 1875:. 1861:: 1851:: 1845:6 1821:. 1801:: 1793:: 1767:. 1753:: 1745:: 1711:. 1707:: 1699:: 1241:† 1216:† 1190:† 1163:† 1138:† 1112:† 1086:† 967:† 951:† 916:† 873:† 855:† 821:† 803:† 746:† 728:† 705:† 674:† 386:† 378:† 374:? 369:† 361:† 353:† 343:† 333:† 323:† 319:? 312:† 302:† 120:N 110:K 105:J 100:T 95:P 90:C 85:D 80:S 75:O 70:Ꞓ 50:– 34:. 20:)

Index

Eumaniraptora
Panaves
Middle Jurassic
Present
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Confuciusornis
Dromaeosaurus
Microraptor
Anchiornis
crow
Prince Creek
troodontid
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata

Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.