1606:
2273:
1629:
1417:
123:
144:
1279:
1864:
2405:, which involves raking, would have become significantly less effective the larger the animal grew since a larger and larger portion of its prey would be small enough to pass between its sweep-feeding spines. Any specimen over the size of a metre (3.2 ft) which continued to feed on small invertebrates would need modified sweep-feeding appendages or would need to employ a different feeding method altogether. As such, it is more than possible that later
169:
4385:
676:
2617:
754:
4688:
2705:. The tracks indicate a lumbering, jerky and dragging movement. Scarps with crescent-shapes were left by the outer limbs, inner markings were made by the keeled belly and the telson carved a central groove. The slow progression and dragging of the tail indicate that the animal responsible was moving out of water. The presence of terrestrial tracks indicate that
1896:, ornamentation consisting of scales or other similar structures on the exoskeleton, the fourth pair of appendages possessing spines, the more posterior tergites of the abdomen possessing tongue-shaped scales near their edges and there being lobes positioned posterolaterally (posteriorly on both sides) on the prosoma. Historically, the morphology of
1320:(segments from the back of the animal), of a large and strange arthropod discovered in deposits in Scotland of Lower Carboniferous age, but did not assign a name to the fossils. Through Scouler's examination, the fossils represent the second eurypterid to be scientifically studied, just six years after the 1825 description of
1526:
represented yet another name applied to some scattered segments, a practice they deemed "taxonomically unsound". Though they suggested that further research was required to determine whether or not the taxon was valid at all, they did note that the presence of a fringe to the segments formed by their
2814:
has been discovered, also preserves a diverse
Carboniferous fauna and some species of plants. Interpreted as having been a large and open fresh to brackish water lake, with possibly occasional influences by storms and glacial processes, fossil remains recovered is most commonly that of various types
741:
was very deep-bodied and compact in comparison to other eurypterids and the mass of the specimen in question would likely have rivalled that of other giant eurypterids (and other giant arthropods), if not surpassed them. In addition to fossil finds of large specimens, fossil trackways attributed to
835:
The status of the 10 species listed below follow a 2018 survey by German paleontologists Jason A. Dunlop and Denise Jekel and
British paleontologist David Penney and size- and temporal ranges follow a 2009 study by American paleontologists James Lamsdell and Simon J. Braddy unless otherwise noted.
2635:
were sweep-feeders, having modified spines on their forward-facing prosomal appendages that allowed them to rake through the substrate of their living environments. Though sweep-feeding was used as a strategy by many genera within the
Stylonurina, it was most developed within the hibbertopterids,
622:
of the environments in which it lived in search for small invertebrates to eat, which it could then push towards its mouth. Though long hypothesised, the fact that eurypterids were capable of terrestrial locomotion was definitely proven through the discovery of a fossil trackway made by
822:
had extensions at their base and lacked longitudinal posterior grooves in all of its podomeres (leg segments). Some of these characteristics, in particular the shape of the telson, are thought to have been shared by other hibbertopterids, which are much less well preserved than
1839:, is remarkably complete, preserving not only the prosoma, the telson and several tergites, but also coxae and even part of the digestive system. The discovery was also important for eurypterid research in general, since it represents one of the few eurypterids known from the
852:
follow the 1968 description of these species. The descriptors, Norwegian paleontologist Leif Størmer and
British paleontologist Charles D. Waterston, did not consider these species to represent eurypterids, though any emended diagnosis of them is yet to be published.
3019:
2677:
could sweep up small animals from the soft sediments of shallow bodies of water, presumably small crustaceans and other arthropods, and could then sweep them into its mouth when it detected them. Through the different adaptations of juveniles and adults
788:
of its environment in search for food. The fourth pair of appendages, though used in feeding like the second and third pairs, was also used for locomotion and the two final pairs of legs (pairs five and six overall) were solely locomotory. As such,
2469:, a more primitid mycteropoid eurypterid, large-scale changes in the developments of the appendages over the course of the life of a single animal have been proven to have happened in some eurypterids. One of the key features distinguishing
3592:
724:
was unusually wide relative to its length for a eurypterid, the animal in question would probably have measured around 180–200 centimetres (5.9–6.6 ft) in length. Even though there were eurypterids of greater length (such as
2701:. The trackway found was roughly 6 metres (20 ft) long and 1 metre (3.3 ft) wide, and suggests that the eurypterid responsible was 160 centimetres (5.2 ft) long, consistent with other giant sizes attributed to
666:
changed significantly over the course of its life, from simpler raking organs present in younger specimens to specialised comb-like organs capable of trapping prey (rather than simply pushing it towards the mouth) in adults.
1572:
was more narrow, had a subelliptical (almost elliptical) shape and had its widest point in the middle rather than at the base. Further differences were noted in the position and shape of the animal's compound eyes, which in
2709:
was able to survive on land at least briefly, possible due to the probability that their gills could function in air as long as they remained wet. Additionally, some studies suggest that eurypterids possessed a dual
2668:
wasn't preying on large animals is also supported by the complete lack of adaptations towards any organs used for trapping prey in younger specimens (though they are present on adult specimens once referred to
2652:
animal, as modern filter feeding crustaceans, but the robust and massive nature of the genus (in contrast to modern filter feeding crustaceans which are typically very small) makes such a conclusion unlikely.
1803:
was also known only from a single specimen, a fragment of an appendage described in 1951. No distinguishing features were given for the species, and the authors noted that it was possibly synonymous with
2378:
would have had further adaptations towards sweep-feeding than any other hibbertopterid, with its blades modified into comb-like rachis that could entrap smaller prey or other organic food particles.
1413:
but since the parts of the body preserved in the fossils described don't completely overlap it is impossible to say if Peach's diagnostic characteristics actually apply to the two original species.
635:
could survive on land, however briefly, is unknown but it might have been possible through either its gills being able to function in air as long as they were wet or by the animal possessing a dual
2852:. Plant fossils in the Waaiport Formation are notably less diverse than those of preceding ages in the same location, possibly because of climate reasons. Among the genera present are the common
1405:
is highly problematic; some of the diagnostic characteristics used when describing it are either questionable or outright meaningless. For instance, the original description had been based on
2729:
contains deposits that were once a freshwater lake near a volcano. The locality has preserved a diverse fauna of the Viséan age of the
Carboniferous (about 335 million years ago). Other than
2640:
and other sweep-feeders was probably composed of what they could find raking through its living environment, likely primarily small invertebrates. This method of feeding is quite similar to
2905:
Tetlie, O. E. (2008). "Hallipterus excelsior, a
Stylonurid (Chelicerata: Eurypterida) from the Late Devonian Catskill Delta Complex, and Its Phylogenetic Position in the Hardieopteridae".
1518:. Scottish paleontologists Lyall I. Anderson and Nigel H. Trewin and German paleontologist Jason A. Dunlop noted in 2000 that Kjellesvig-Waerings acception of the original designation for
3545:"Redescription of Drepanopterus pentlandicus Laurie, 1892, the earliest known mycteropoid (Chelicerata: Eurypterida) from the early Silurian (Llandovery) of the Pentland Hills, Scotland"
1510:
by
American paleontologist Erik N. Kjellesvig-Waering in 1964 as part of a greater re-examination of the various species assigned to the family Pterygotidae. Kjellesvig-Waering retained
2588:. The same conclusions and suggestions were also published in a later 2020 conference abstract, co-authored by Hughes and James Lamsdell. However, a 2023 study describing a new species
983:
is too limited to confidently establish any distinguishing features, though it can be confidently be assigned to the genus based on its "striking similarities" to the other species.
1915:
makes it clear that both genera represent hibbertopterid eurypterids, but the incomplete nature of all fossil specimens referred to them make any further study of the precise
1667:
prompted
Norwegian paleontologist Leif Størmer and British paleontologist Charles D. Waterston to in 1968 re-examine the various species that had been referred to it. Because
2840:. Among the palaeoniscoids, eight distinct genera have been identified. Several of these palaeoniscoid genera also occur in deposits of similar age in Scotland. Other than
3868:
3766:
2447:. The three fragmentary genera were suggested to by synonyms of each other by American paleontologist James Lamsdell in 2010, which would have meant the oldest name,
2381:
It was suggested as early as 1993 by
American paleontologist Paul Selden and British paleontologist Andrew J. Jeram that these adaptations might not have been due to
1843:, where eurypterid finds are rare and usually fragmentary. The presence of the gut in the fossil proves that the specimen represents a dead individual, and not only
780:(leg segments) of these three pairs of limbs were covered with long spines, and the end of each limb was covered with sensory organs. These adaptations suggest that
4751:
2682:"), individuals of different ages would possibly have preferred different types of prey, which would have reduced competition between members of the same genus.
2421:
4792:
627:
in
Scotland. The trackway showed that an animal measuring around 160 centimetres (5.2 ft) had slowly lumbered across a stretch of land, dragging its
2272:
1117:
are the large number filaments on its first pair of appendages and the longer filaments on the second pair of appendages, which also have large fulcra.
2489:
is not surprising as movements of the eyes through ontogeny has been described in other eurypterid genera. Lamsdell considered it almost certain that
4738:
1605:
1628:
800:
Although not enough fossil material is known of the other hibbertopterid eurypterids to discuss the differences between them with full confidence,
3932:
Evans, Francis John (1999). "Palaeobiology of Early Carboniferous lacustrine biota of the Waaipoort Formation (Witteberg Group), South Africa".
746:
from South Africa indicates an animal around 250 centimetres (8.2 ft) in length (the same size attributed to the largest known eurypterid,
4464:
4454:
3976:
3588:
Laurie, 1892, the earliest known mycteropoid (Chelicerata: Eurypterida) from the early Silurian (Llandovery) of the Pentland Hills, Scotland"
1185:
is the precise arrangement of lunules (crescent-shaped markings) on said leg segments, forming rows. This is different from, for an example,
585:
could reach lengths around 180–200 centimetres (5.9–6.6 ft). Though this is significantly smaller than the largest eurypterid overall,
3039:
2481:
suggest might have been a feature which appeared late in an animal's life cycle. Differences in the positions of the eyes in specimens of
4807:
3145:
Waterston, Charles D.; Størmer, Leif (1968). "IV. Cyrtoctenus gen. nov., a large late Palaeozoic Arthropod with pectinate Appendages*".
3075:
4812:
4416:
1709:(named in honour of Ben Peach), as its type. Both of these species were based on fragmentary fossil remains. Furthermore, the species
2572:. In particular, she noted that though the feeding appendages were different, the ornamentation and form of the raking tools seen in
1427:
Though only represented by two small, jointed and vaguely cylindrical fossil fragments (both discovered in the Portage sandstones of
4431:
3916:
1435:
has had a complicated taxonomic history. Originally described in 1881 as a species of plant, the fragmentary fossil referred to as "
750:), though the largest known fossil specimens of the species only appear to have reached lengths of 135 centimetres (4.43 ft).
3903:. Geological Society of America Special Papers. Vol. 244. Geological Society of America Special Publication. pp. 13–24.
3895:
Rolfe, W. D. I.; Durant, G. P.; Fallick, A. E.; Hall, A. J.; Large, D. J.; Scott, A. C.; Smithson, T. R.; Walkden, G. M. (1990).
1892:. The hibbertopterids are united as a group by being large mycteropoids with broad prosomas, a hastate telson similar to that of
2636:
which possessed blades on the second, third and fourth pair of appendages. Inhabiting freshwater swamps and rivers, the diet of
4675:
4459:
4401:
2886:
1225:
can be distinguished from other species by the oval shape of its ocellar node (the raised portion of the carapace on which the
1147:
Size estimate is based on a complete carapace. As the type species, no particular distinguishing features are established for
4822:
4817:
4449:
4439:
4393:
3330:"Cyrtoctenus wittebergensis sp. nov. (Chelicerata: Eurypterida), a large sweep-feeder from the Carboniferous of South Africa"
784:, like other hibbertopterids, would have fed by a method referred to as sweep-feeding, using its limbs to sweep through the
3277:
818:
sword) and had a keel running down the middle, with in turn had a small indentation in its own centre. The walking legs of
4411:
4406:
2831:. Among the acanthodians, at least three genera have been identified from fossil scales and spines, including the derived
1877:
is classified as part of the family Hibbertopteridae, which it also lends its name to, a family of eurypterids within the
168:
4444:
4421:
2389:
representing more derived genera of hibbertopterids, but rather due to both genera perhaps representing adult forms of
1795:
other than the unique fragmentary type specimen, which at this point had been plastically preserved in sandstone. Like
3241:
Kjellesvig-Waering, Erik N. (1964). "A Synopsis of the Family Pterygotidae Clarke and Ruedemann, 1912 (Eurypterida)".
1443:
915:) of its second pair of appendages and the presence of small cone-shaped fulcra (supportive tissue) along the limbs.
2827:. Though shark material is too fragmentary to be identifiable, at least some fossils might represent the remains of
1522:
was "burdensome" as it is based on highly fragmentary material. They noted that like many other pterygotid species,
2358:(various hardened body parts) which have little diagnostic potential and are poorly known in fossils attributed to
1900:
and the other hibbertopterids has been seen as so unusual that they have been thought to be an order separate from
1620:
1451:
1283:
3896:
1181:
Fragmentary fossils known from Scotland. Known primarily from leg segments, the primary distinguishing feature of
4802:
4077:
3969:
3031:
2881:
1847:, and scientists examining it could conclude that it had been preserved as lying on its back. The description of
1923:
itself, though the highly incomplete nature of their remains again makes that hypothesis impossible to confirm.
1779:
fossils represented remains of a new order of aquatic arthropods which they dubbed "Cyrtoctenida". The species
1775:, respectively. Despite noting the presence of eurypterid-type tergites, Størmer and Waterston thought that the
1471:
was probably based on the slightly spinose surface of the fossils, but in 1888 Hall and American paleontologist
1221:
The lower size estimate follows known fossil evidence. The higher size estimate derives from trackway evidence.
2945:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates"
4384:
3506:
Tollerton, V P (1989). "Morphology, Taxonomy, and Classification of the Order Eurypterida Burmeister, 1843".
2918:
1479:
actually had spines similar to what Woodward and Jones suggested and as such, reassigned the species back to
1353:
by later researchers although it had not originally been designated as such) in 1882. The genus was based on
1259:
Isolated podomeres of a prosomal appendage have been discovered in the United States. The fossil material of
3508:
34:
2420:
1416:
4706:
2698:
1926:
The cladogram below is adapted from Lamsdell (2012), collapsed to only show the superfamily Mycteropoidea.
1878:
3842:
2350:
were congeneric (e.g. synonymous) was first suggested by British geologist Charles D. Waterston in 1985.
1824:
1823:
known from reasonably complete remains other than the type species itself. The fossil, discovered in the
4764:
3760:
1503:
2864:
122:
3631:
Discerning the Diets of Sweep-Feeding Eurypterids Through Analyses of Mesh-Modified Appendage Armature
1721:
was its comb-like first appendages. Waterston remarked in another 1968 paper that the "controversial"
1495:
in 1912, though no distinguishing features of the fossils were given due to their fragmentary nature.
1263:
is too limited to confidently establish any distinguishing features, but it is noted to be similar to
4797:
4691:
3962:
3795:
2992:
1242:
1202:
1164:
1130:
1092:
1058:
996:
962:
928:
890:
642:
Though sometimes, and often historically, treated as distinct genera, the hibbertopterid eurypterids
3370:
Waterston, Charles D. (1968). "I.—Further Observations on the Scottish Carboniferous Eurypterids*".
2854:
143:
2828:
2779:
2753:. Several terrestrial invertebrates are also known from the location, including several species of
2726:
1840:
1366:
1327:
1278:
785:
619:
556:
1919:
relationships within the Hibbertopteridae difficult. Both genera could even represent synonyms of
662:. The features of fossils associated with these genera suggest that the sweep-feeding strategy of
3941:
3821:
3697:
3609:
3564:
3525:
3488:
3430:
3395:
3349:
3305:
3258:
3220:
3170:
3094:
2922:
2711:
1783:
had originally been described as the fragmentary remains of a eurypterid in 1889 was assigned to
1456:
1437:
636:
456:
163:
2287:
as composing three separate, but closely related, hibbertopterid genera. In these arrangements,
1863:
1013:
The only known specimen, discovered in Ireland, is probably lost and was originally assigned to
697:
was a large, broad-bodied and heavy animal. It was the largest known eurypterid of the suborder
4769:
3676:
Braddy, Simon J.; Lerner, Allan J.; Lucas, Spencer G. (2023). "A new species of the eurypterid
3449:
4756:
4280:
4213:
4205:
3912:
3813:
3748:
3480:
3422:
3387:
3297:
3250:
3212:
3162:
2984:
2966:
2816:
2406:
1472:
655:
3471:
Kjellesvig-Waering, Erik N. (1959). "A Taxonomic Review of Some Late Paleozoic Eurypterida".
3035:
1229:
are located) and by the many crenulations present on the distal margins of its leg segments.
4620:
4334:
4326:
4316:
4308:
4288:
4259:
4175:
4124:
3904:
3877:
3803:
3786:
3738:
3730:
3689:
3658:
3601:
3556:
3517:
3379:
3341:
3289:
3204:
3154:
3084:
2974:
2958:
2914:
2718:
2155:
1488:
687:
249:
3117:
4649:
4495:
4352:
4293:
4249:
4152:
4139:
3721:
2949:
2549:
2460:
2169:
2050:
1943:
1556:
1428:
1151:. Its latest published diagnosis is simply the same as the diagnosis of the genus itself.
596:
30:
3071:"The systematics and phylogeny of the Stylonurina (Arthropoda: Chelicerata: Eurypterida)"
3799:
3413:
Kjellesvig-Waering, Erik N. (1961). "The Silurian Eurypterida of the Welsh Borderland".
675:
4565:
4481:
4344:
4271:
4231:
4223:
4147:
3866:
Tetlie, O E (2007). "Distribution and dispersal history of Eurypterida (Chelicerata)".
3743:
3716:
2979:
2869:
2832:
2804:
2759:
2734:
2641:
1787:
on the basis of the perceived filaments present on its appendages, similar to those of
1663:
1442:" was noted to represent the fossil remains of a eurypterid by American paleontologist
538:
3193:"A Middle Devonian chasmataspid arthropod from Achanarras Quarry, Caithness, Scotland"
2616:
1369:
in 1863. This designation was reinforced with more fossil fragments discovered in the
4786:
4572:
4544:
4537:
4509:
4488:
4241:
4180:
4170:
4162:
4116:
4106:
4098:
4054:
4018:
3701:
3648:
3583:
3544:
3399:
3353:
3329:
3309:
3224:
3174:
2859:
2836:
2773:
2746:
2664:
would probably have been incapable of preying on larger animals. The conclusion that
2465:
2036:
1998:
1981:
1957:
1933:
1916:
1881:
1832:
1808:, but they chose to maintain it as distinct due to the very limited fossil material.
1685:
1295:
766:
727:
714:
615:
587:
564:
530:
526:
236:
80:
3945:
3613:
3568:
3529:
3098:
2944:
2926:
2660:
were weak and they would not have been able to grasp any potential prey which means
595:
is likely to have been the heaviest due to its broad and compact body. Furthermore,
4530:
4362:
4298:
4254:
4129:
3825:
3192:
2824:
2791:
2649:
1836:
1483:, interpreting the fossils as fragments of the long walking legs. An assignment to
1461:
1382:
1309:
815:
753:
542:
275:
3693:
3043:
614:
fed through a method called sweep-feeding. It used its specialised forward-facing
3881:
4627:
4604:
4502:
4196:
4089:
4027:
4009:
3662:
2800:
2576:
were probably the precursors of the more moveable finger-like organs present in
2393:. In this case, the development of the more specialized sweep-feeding method of
1901:
1828:
1762:
1215:
1103:
1007:
949:
is the presence of ~110 filaments in every row of its comb-like feeding organs.
733:
706:
698:
578:
210:
55:
4593:
4579:
4558:
4551:
4516:
4357:
4045:
4036:
3985:
3630:
3605:
3593:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
3560:
3549:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
3521:
3383:
3372:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
3345:
3293:
3158:
3147:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
3089:
3070:
3024:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
2742:
2686:
2115:
1464:
1370:
1322:
1300:
1189:, in which these lunules are more in number and not as well sorted into rows.
1069:
690:
551:
502:
223:
100:
65:
42:
4729:
3680:
from the Carboniferous of New Mexico, and a review of the Hibbertopteridae".
3650:
Evidence for Synonymy and Ontogenetic Shifts Among Hibbertopterid Eurypterids
3484:
3426:
3391:
3301:
3254:
3216:
3166:
2970:
4670:
4654:
4644:
4003:
2849:
2785:
2768:
2754:
2738:
2722:
2542:
2098:
2074:
1342:
1253:
939:
506:
200:
180:
105:
49:
3817:
3752:
3734:
2988:
2962:
1175:
1141:
1107:
1075:
Fossil isolated rachis found in the Czech Republic. The fossil material of
901:
2397:
can directly be explained by the larger size of the specimens referred to
4723:
4613:
4586:
2845:
2764:
2750:
2621:
2355:
1612:
1398:
1394:
1317:
1113:
Fossil isolated rachis found in Scotland. The distinguishing features of
973:
808:(the posteriormost division of the body) was hastate (e.g. shaped like a
794:
777:
717:
702:
631:(the posteriormost division of its body) across the ground after it. How
522:
514:
131:
95:
90:
75:
70:
60:
38:
3897:"An early terrestrial biota preserved by Visean vulcanicity in Scotland"
3208:
4743:
3908:
3492:
3434:
3262:
2714:, which would allow short periods of time in terrestrial environments.
2584:, due to its distinct ornamentation, represented a genus distinct from
1844:
1635:
1386:
1313:
945:
Fossil isolated rachis found in Belgium. The distinguishing feature of
810:
534:
518:
152:
110:
85:
1577:
are surrounded by a ring-like shape of hardened integument (absent in
3997:
3191:
Trewin, Nigel H.; Dunlop, Jason A.; Anderson, Lyall I. (2000-11-01).
3020:"Eurypterids from the Viséan of East Kirkton, West Lothian, Scotland"
1855:-type eurypterids, not representatives of a new order of arthropods.
1683:
had been designated the type species of a distinct eurypterid genus,
1291:
1226:
1079:
is too limited to confidently establish any distinguishing features.
805:
774:
628:
591:, which could reach lengths of around 250 centimetres (8.2 ft),
190:
4700:
3808:
3781:
2771:. The site also preserves abundant plant life, including the genera
2560:
due to the "strong morphological similarities" between them, and as
761:, the largest species known from fossil remains, compared to a human
3328:
D. Waterston, C; W. Oelofsen, B; D. F. Oosthuizen, R (1985-01-01).
3278:"Chelicerata from the Dinantian of Foulden, Berwickshire, Scotland"
1554:
had already been referred to (considered a species of) the related
979:
Fossil isolated tergites found in Scotland. The fossil material of
3717:"The true identity of the supposed giant fossil spider Megarachne"
2820:
2767:
proven to have been terrestrial) and early representatives of the
2615:
2552:, American geologist Emily Hughes suggested the synonymization of
2419:
2271:
1862:
1415:
1277:
804:
is defined based on a collection of definite characteristics. The
752:
674:
3069:
James C. Lamsdell, Simon J. Braddy & O. Erik Tetlie (2010).
2690:
2374:, particularly in their adaptations to sweep-feeding. If valid,
1446:
in 1884, three years later. Though Hall assigned the species to
701:, composed of those eurypterids that lacked swimming paddles. A
4704:
3958:
639:, theorised to have been present in at least some eurypterids.
3954:
3334:
Transactions of the Royal Society of Edinburgh: Earth Sciences
3282:
Transactions of the Royal Society of Edinburgh: Earth Sciences
2477:
is the presence of grooves on its podomeres, which studies on
2283:
Many analyses and overviews treat the ten species assigned to
1585:
are also located near the center of the head whereas those of
563:, coined more than a century later, combines his name and the
549:, was first named as a species of the significantly different
2673:) and a lack of swimming adaptations. Through sweep-feeding,
2529:
specimens are more complete than the known fossil remains of
1725:
was similar to the unusual and massive prosomal appendage of
1304:(with swimming paddles and unspecialised walking appendages).
911:
is distinguished by the long filaments (longer than those of
1502:(a species described from Ireland by British paleontologist
1330:
redescribed the same fossil specimens, giving them the name
773:(pairs 2, 3 and 4) were specialised for gathering food. The
581:
suborder, with the largest fossil specimens suggesting that
2541:
specimens represent the actual mortalities, susceptible to
1312:
described the remains, consisting of a massive and unusual
618:(limbs), equipped with several spines, to rake through the
2733:, the fauna includes several terrestrial animals, such as
1791:. Størmer and Waterston disregarded specimens referred to
3715:
Selden, P.A., Corronca, J.A. & Hünicken, M.A (2005).
1454:
and Thomas Rupert Jones assigned the fossil to the genus
2919:
10.3374/0079-032X(2008)49[19:HEASCE]2.0.CO;2
2424:
More fossil spines and fossil ornamentation referred to
2844:, the only known invertebrates are two rare species of
2721:
of Scotland, 27 kilometres (16.8 miles) to the west of
3120:. In World Spider Catalog. Natural History Museum Bern
3782:"Palaeoecology: A gigantic fossil arthropod trackway"
2435:
frequently occur together with fragments referred to
2417:
to be able to continue to feed at larger body sizes.
1326:
itself. Five years later, in 1836, British geologist
3323:
3321:
3319:
3118:
A summary list of fossil spiders and their relatives
2749:, representing some of the oldest known terrestrial
1653:
was questionable at best and that its type species,
4713:
4663:
4637:
4603:
4473:
4430:
4392:
4343:
4325:
4307:
4279:
4268:
4240:
4222:
4204:
4193:
4161:
4138:
4115:
4097:
4086:
4070:
4063:
3635:
Graduate Theses, Dissertations, and Problem Reports
1487:was affirmed by Clarke and American paleontologist
720:, measures 65 centimetres (26 in) wide. Since
3454:Memoir (New York State Museum and Science Service)
2370:does suggest that they were more specialised than
1531:, but "strikingly similar" to what was present in
513:have been discovered in deposits ranging from the
3869:Palaeogeography, Palaeoclimatology, Palaeoecology
2907:Bulletin of the Peabody Museum of Natural History
1851:affirmed that the "cyrtoctenids" were definitely
1593:was selected to honor the original descriptor of
1564:as distinct from the type species of that genus,
1527:ornamentation was absent in all other species of
3116:Dunlop, J. A., Penney, D. & Jekel, D. 2018.
2644:. This has led some researchers to suggest that
2354:is known from very fragmentary material, mainly
1811:Known from a single specimen described in 1985,
1697:was to be part of a new genus, which they named
709:which covered the head) referred to the species
1713:, named in 1936, was referred to the new genus
1705:, a curved comb) and they named a new species,
3448:Clarke, John Mason; Ruedemann, Rudolf (1912).
2537:specimens would then represent exuviae whilst
3970:
2943:Lamsdell, James C.; Braddy, Simon J. (2009).
2303:being the type and only species of the genus
1560:for some time. Kjellesvig-Waering recognised
1546:When Kjellesvig-Waering designated the genus
1498:Though no specification was given as to why,
8:
3899:. In Lockley, Martin G.; Rice, Alan (eds.).
3765:: CS1 maint: multiple names: authors list (
411:(Waterston, Oelofsen & Oosthuizen, 1985)
2795:. Locally, the strange fossil carapaces of
2564:was found to be paraphyletic in regards to
1689:. Størmer and Waterston concluded that the
1589:are located further back. The generic name
4701:
4276:
4201:
4094:
4067:
3977:
3963:
3955:
3018:Jeram, Andrew J.; Selden, Paul A. (1993).
2362:. The morphology of fossils attributed to
1675:, the genus itself became synonymous with
1661:) had recently been referred to the genus
907:Fossil isolated rachis found in Scotland.
855:
142:
121:
20:
3807:
3742:
3088:
2978:
1450:, that same year British paleontologists
1045:Tail spine, ventral keels, and trackways
1021:in 1964 with no specification as to why.
610:Like many other stylonurine eurypterids,
599:indicates that the South African species
3647:Hughes, Emily; Lamsdell, James (2020).
3064:
3062:
3060:
3058:
3056:
3054:
3052:
2897:
2505:in turn represented junior synonyms of
2463:. Following studies on the ontogeny of
2291:is typically restricted to the species
654:have been suggested to represent adult
438:
347:
3758:
1460:, believing the fossils represented a
603:might have reached lengths similar to
577:was the largest eurypterid within the
3843:"Giant Water Scorpion Walked on Land"
3837:
3835:
3625:
3623:
3365:
3363:
3236:
3234:
3186:
3184:
3112:
3110:
3108:
2533:, often fragmentary. The majority of
2509:, which would subsume all three into
2451:, taking priority and subsuming both
1420:The two fossil fragments referred to
1199:Waterston, Oelofsen & Oosthuizen
7:
3140:
3138:
3136:
3134:
3132:
3130:
3128:
3126:
3014:
3012:
3010:
3008:
3006:
3004:
3002:
2938:
2936:
2568:, all three were subsumed into just
1361:, previously described as the plant
3076:Journal of Systematic Palaeontology
2185:
2161:
2090:
2066:
2042:
2032:
1973:
1949:
1939:
1929:
1431:), the species today recognised as
4793:Carboniferous arthropods of Europe
2685:A fossil trackway discovered near
1701:(the name deriving from the Greek
14:
2413:developed the structures seen in
4687:
4686:
4383:
3653:. Geological Society of America
3629:Hughes, Emily Samantha (2019), "
2803:"Scouler's heids" ("heid" being
2697:was capable of at least limited
1752:, further species were added to
1717:. The key diagnostic feature of
1627:
1604:
1349:(treated as the type species of
1282:Outdated 1872 reconstruction by
167:
4676:Timeline of eurypterid research
2887:Timeline of eurypterid research
2815:of fish. Among these types are
2810:The Waaipoort Formation, where
2525:would also explain why smaller
1341:was named by British geologist
836:The distinguishing features of
400:(Størmer & Waterston, 1968)
3276:Waterston, Charles D. (1985).
2620:Fossil trackway discovered in
1679:. That same year, the species
1657:, (and other species, such as
1535:. Subsequent research treated
1475:pointed out that no described
1:
3694:10.1080/08912963.2022.2032690
3450:"The Eurypterida of New York"
1345:, who also named the species
1308:In 1831, Scottish naturalist
1195:Hibbertopterus wittebergensis
505:, a group of extinct aquatic
470:Størmer & Waterston, 1968
3882:10.1016/j.palaeo.2007.05.011
2656:The chelicerae (pincers) of
2331:being referred to the genus
1767:, described in 1951, became
1357:and the fragmentary species
389:(Augusta & Přibyl, 1951)
3582:Lamsdell, James C. (2012).
3197:Scottish Journal of Geology
2600:should be synonymized with
1999:Drepanopterus odontospathus
1506:in 1872) was reassigned to
1493:The Eurypterida of New York
1051:Hibbertopterus ostraviensis
16:Extinct genus of arthropods
4839:
4808:Prehistoric life of Europe
3901:Volcanism and Fossil Biota
3586:Drepanopterus pentlandicus
2211:Cyrtoctenus wittebergensis
1958:Drepanopterus pentlandicus
1869:Cyrtoctenus wittebergensis
1817:Cyrtoctenus wittebergensis
1756:by Størmer and Waterston;
1377:. In 1887 Peach described
1375:Glyptoscorpius caledonicus
1373:, which Peach referred to
1365:by English paleontologist
1235:Hibbertopterus wrightianus
883:Hibbertopterus caledonicus
559:in 1836. The generic name
4813:Carboniferous eurypterids
4684:
4381:
4078:List of eurypterid genera
3992:
3663:10.1130/abs/2020AM-353747
3606:10.1017/S1755691012000072
3561:10.1017/S1755691012000072
3522:10.1017/S0022336000041275
3384:10.1017/S0080456800014472
3346:10.1017/S0263593300010555
3294:10.1017/S0263593300010269
3159:10.1017/S0080456800014563
3090:10.1080/14772011003603564
3036:10.1017/S0263593300006118
2882:List of eurypterid genera
2858:(possibly representing a
2693:, Scotland, reveals that
2207:
2190:
2183:
2166:
2159:
2112:
2095:
2088:
2071:
2064:
2051:Woodwardopterus scabrosus
2047:
2040:
2030:
1995:
1978:
1971:
1954:
1947:
1937:
1819:) is the only species of
1760:, described in 1889, and
1568:, in that the prosoma of
1157:Hibbertopterus stevensoni
989:Hibbertopterus hibernicus
876:
873:
462:
455:
300:
295:
281:
274:
164:Scientific classification
162:
150:
141:
129:
120:
23:
3934:Palaeontologica Africana
3543:Lamsdell, James (2012).
2848:, possibly representing
2631:Hibbertopterids such as
2580:. Hughes suggested that
2493:was a junior synonym of
2170:Campylocephalus oculatus
1671:was the type species of
1562:Campylocephalus scouleri
1089:Størmer & Waterston
1027:Hibbertopterus lamsdelli
921:Hibbertopterus dewalquei
877:Notes & description
861:
269:Kjellesvig-Waering, 1959
3655:Abstracts with Programs
3509:Journal of Paleontology
3473:Journal of Paleontology
3415:Journal of Paleontology
3243:Journal of Paleontology
2194:Hibbertopterus scouleri
1982:Drepanopterus abonensis
1884:, alongside the genera
1739:Dunsopterus wrightianus
1729:and as such reassigned
1422:Cyrtoctenus wrightianus
1123:Hibbertopterus scouleri
3735:10.1098/rsbl.2004.0272
2963:10.1098/rsbl.2009.0700
2699:terrestrial locomotion
2628:
2428:
2426:Dunsopterus stevensoni
2339:is the type species).
2280:
2278:Dunsopterus stevensoni
1871:
1723:Stylonurus wrightianus
1424:
1305:
762:
683:
4823:Eurypterids of Africa
4818:Eurypterids of Europe
4765:Paleobiology Database
3780:Whyte, M. A. (2005).
2993:Supplemental material
2763:(one of the earliest
2619:
2608:is distinct from it.
2423:
2275:
1866:
1769:Cyrtoctenus dewalquei
1504:William Hellier Baily
1500:Pterygotus hibernicus
1491:in their influential
1419:
1363:Cycadites caledonicus
1337:The eurypterid genus
1281:
1085:Hibbertopterus peachi
1055:Augusta & Přibyl
1017:. It was referred to
756:
678:
2799:have been given the
2431:Fossil specimens of
1758:Eurypterus dewalquei
1467:. The assignment to
955:Hibbertopterus dicki
545:. The type species,
533:period in Scotland,
447:(Etheridge Jr, 1877)
3800:2005Natur.438..576W
3209:10.1144/sjg36020151
2727:East Kirkton Quarry
2299:, with the species
2116:Megarachne servinei
1841:southern hemisphere
1825:Waaipoort Formation
1642:, drawn by Woodward
1552:Eurypterus scouleri
1367:John William Salter
1332:Eurypterus scouleri
1316:(head) and several
1274:History of research
1042:Late Carboniferous
797:(six-legged) gait.
765:The forward-facing
285:Eurypterus scouleri
4206:Onychopterelloidea
3909:10.1130/SPE244-p13
3682:Historical Biology
3584:"Redescription of
2712:respiratory system
2648:would have been a
2629:
2429:
2407:ontogenetic stages
2281:
2075:Mycterops mathieui
1872:
1597:, Samuel Hibbert.
1425:
1306:
1298:based on those of
793:would have used a
763:
684:
656:ontogenetic stages
637:respiratory system
571:) meaning "wing".
407:H. wittebergensis
4780:
4779:
4707:Taxon identifiers
4698:
4697:
4379:
4378:
4375:
4374:
4371:
4370:
4327:Adelophthalmoidea
4309:Waeringopteroidea
4281:Carcinosomatoidea
4214:Onychopterellidae
4189:
4188:
2868:(a small type of
2865:Archaeosigillaria
2842:H. wittebergensis
2812:H. wittebergensis
2329:H. wittebergensis
2276:Fossil spines of
2258:
2257:
2249:
2248:
2240:
2239:
2231:
2230:
2222:
2221:
2145:
2144:
2136:
2135:
2127:
2126:
2019:
2018:
2010:
2009:
1849:H. wittebergensis
1813:H. wittebergensis
1681:G. minutisculptus
1473:John Mason Clarke
1379:G. minutisculptus
1271:
1270:
1223:H. wittebergensis
744:H. wittebergensis
705:(the part of the
686:Like other known
601:H. wittebergensis
597:trackway evidence
494:
493:
488:
479:
471:
448:
435:
423:
412:
401:
390:
379:
368:
357:
344:
333:
322:
311:
270:
48:387.7–323.2
4830:
4803:Fossil trackways
4773:
4772:
4760:
4759:
4747:
4746:
4734:
4733:
4732:
4702:
4690:
4689:
4664:Related articles
4621:Merostomichnites
4387:
4335:Adelophthalmidae
4317:Waeringopteridae
4289:Carcinosomatidae
4277:
4260:Strobilopteridae
4202:
4176:Hibbertopteridae
4125:Parastylonuridae
4095:
4068:
3979:
3972:
3965:
3956:
3950:
3949:
3929:
3923:
3922:
3892:
3886:
3885:
3876:(3–4): 557–574.
3863:
3857:
3856:
3854:
3853:
3839:
3830:
3829:
3811:
3777:
3771:
3770:
3764:
3756:
3746:
3712:
3706:
3705:
3673:
3667:
3666:
3657:. Vol. 52.
3644:
3638:
3627:
3618:
3617:
3579:
3573:
3572:
3540:
3534:
3533:
3503:
3497:
3496:
3468:
3462:
3461:
3445:
3439:
3438:
3410:
3404:
3403:
3367:
3358:
3357:
3340:(2–3): 339–358.
3325:
3314:
3313:
3273:
3267:
3266:
3238:
3229:
3228:
3188:
3179:
3178:
3142:
3121:
3114:
3103:
3102:
3092:
3066:
3047:
3030:(3-4): 301–308.
3016:
2997:
2996:
2982:
2940:
2931:
2930:
2902:
2829:protacrodontoids
2624:, attributed to
2401:. The method of
2307:and the species
2186:
2162:
2156:Hibbertopteridae
2099:Hastimima whitei
2091:
2067:
2043:
2033:
1974:
1950:
1940:
1930:
1907:The features of
1631:
1608:
1539:as a species of
1520:Pterygotus dicki
1489:Rudolf Ruedemann
1218:(Carboniferous)
1178:(Carboniferous)
1144:(Carboniferous)
1110:(Carboniferous)
1072:(Carboniferous)
1010:(Carboniferous)
904:(Carboniferous)
856:
831:Table of species
486:
477:
469:
446:
430:
421:
410:
399:
388:
385:H. ostraviensis
377:
367:(Fraipont, 1889)
366:
355:
339:
331:
320:
309:
268:
261:
250:Hibbertopteridae
248:
235:
222:
172:
171:
146:
125:
115:
52:
29:Temporal range:
21:
4838:
4837:
4833:
4832:
4831:
4829:
4828:
4827:
4783:
4782:
4781:
4776:
4768:
4763:
4755:
4750:
4742:
4737:
4728:
4727:
4722:
4709:
4699:
4694:
4680:
4659:
4650:Chasmataspidida
4633:
4599:
4496:Campylocephalus
4469:
4426:
4388:
4367:
4353:Hughmilleriidae
4339:
4321:
4303:
4294:Megalograptidae
4270:
4264:
4250:Dolichopteridae
4236:
4224:Moselopteroidea
4218:
4195:
4185:
4171:Drepanopteridae
4157:
4153:Hardieopteridae
4140:Kokomopteroidea
4134:
4111:
4088:
4082:
4059:
3988:
3983:
3953:
3931:
3930:
3926:
3919:
3894:
3893:
3889:
3865:
3864:
3860:
3851:
3849:
3841:
3840:
3833:
3809:10.1038/438576a
3779:
3778:
3774:
3757:
3722:Biology Letters
3714:
3713:
3709:
3675:
3674:
3670:
3646:
3645:
3641:
3628:
3621:
3581:
3580:
3576:
3542:
3541:
3537:
3505:
3504:
3500:
3470:
3469:
3465:
3447:
3446:
3442:
3412:
3411:
3407:
3369:
3368:
3361:
3327:
3326:
3317:
3275:
3274:
3270:
3240:
3239:
3232:
3190:
3189:
3182:
3144:
3143:
3124:
3115:
3106:
3068:
3067:
3050:
3017:
3000:
2950:Biology Letters
2942:
2941:
2934:
2904:
2903:
2899:
2895:
2878:
2614:
2550:graduate thesis
2513:. Synonymizing
2461:junior synonyms
2321:H. ostraviensis
2270:
2259:
2250:
2241:
2232:
2223:
2146:
2137:
2128:
2020:
2011:
1944:Drepanopteridae
1909:Campylocephalus
1886:Campylocephalus
1867:Restoration of
1861:
1801:C. ostraviensis
1773:C. ostraviensis
1647:
1646:
1645:
1644:
1643:
1632:
1624:
1623:
1611:Drawing of the
1609:
1587:Campylocephalus
1581:). The eyes of
1579:Campylocephalus
1570:Campylocephalus
1557:Campylocephalus
1429:Italy, New York
1276:
1211:
1161:Etheridge, Jr.
1077:H. ostraviensis
874:Temporal range
846:H. ostraviensis
833:
679:Restoration of
673:
478:Waterston, 1968
451:
437:
436:
426:
352:H. caledonicus
346:
345:
321:(Hibbert, 1836)
291:
288:
267:
259:
246:
233:
220:
166:
116:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
31:Middle Devonian
27:
17:
12:
11:
5:
4836:
4834:
4826:
4825:
4820:
4815:
4810:
4805:
4800:
4795:
4785:
4784:
4778:
4777:
4775:
4774:
4761:
4748:
4735:
4719:
4717:
4715:Hibbertopterus
4711:
4710:
4705:
4696:
4695:
4685:
4682:
4681:
4679:
4678:
4673:
4667:
4665:
4661:
4660:
4658:
4657:
4652:
4647:
4641:
4639:
4638:Related groups
4635:
4634:
4632:
4631:
4624:
4617:
4609:
4607:
4601:
4600:
4598:
4597:
4590:
4583:
4576:
4569:
4566:Onychopterella
4562:
4555:
4548:
4541:
4534:
4527:
4524:Hibbertopterus
4520:
4513:
4506:
4499:
4492:
4485:
4482:Adelophthalmus
4477:
4475:
4474:Notable genera
4471:
4470:
4468:
4467:
4462:
4457:
4452:
4447:
4442:
4436:
4434:
4428:
4427:
4425:
4424:
4419:
4414:
4409:
4404:
4398:
4396:
4390:
4389:
4382:
4380:
4377:
4376:
4373:
4372:
4369:
4368:
4366:
4365:
4360:
4355:
4349:
4347:
4345:Pterygotioidea
4341:
4340:
4338:
4337:
4331:
4329:
4323:
4322:
4320:
4319:
4313:
4311:
4305:
4304:
4302:
4301:
4296:
4291:
4285:
4283:
4274:
4272:Diploperculata
4266:
4265:
4263:
4262:
4257:
4252:
4246:
4244:
4238:
4237:
4235:
4234:
4232:Moselopteridae
4228:
4226:
4220:
4219:
4217:
4216:
4210:
4208:
4199:
4191:
4190:
4187:
4186:
4184:
4183:
4178:
4173:
4167:
4165:
4159:
4158:
4156:
4155:
4150:
4148:Kokomopteridae
4144:
4142:
4136:
4135:
4133:
4132:
4127:
4121:
4119:
4113:
4112:
4110:
4109:
4103:
4101:
4099:Rhenopteroidea
4092:
4084:
4083:
4081:
4080:
4074:
4072:
4065:
4061:
4060:
4058:
4057:
4048:
4039:
4030:
4021:
4012:
4006:
4000:
3993:
3990:
3989:
3984:
3982:
3981:
3974:
3967:
3959:
3952:
3951:
3924:
3917:
3887:
3858:
3831:
3772:
3707:
3688:(2): 257–263.
3678:Hibbertopterus
3668:
3639:
3619:
3574:
3535:
3516:(5): 642–657.
3498:
3479:(2): 251–256.
3463:
3440:
3421:(4): 789–835.
3405:
3359:
3315:
3268:
3249:(2): 331–361.
3230:
3203:(2): 151–158.
3180:
3122:
3104:
3048:
2998:
2957:(2): 265–269.
2932:
2896:
2894:
2891:
2890:
2889:
2884:
2877:
2874:
2855:Praeramunculus
2817:palaeoniscoids
2760:Gigantoscorpio
2719:Midland Valley
2707:Hibbertopterus
2703:Hibbertopterus
2695:Hibbertopterus
2675:Hibbertopterus
2666:Hibbertopterus
2662:Hibbertopterus
2658:Hibbertopterus
2646:Hibbertopterus
2642:filter feeding
2638:Hibbertopterus
2633:Hibbertopterus
2626:Hibbertopterus
2613:
2610:
2602:Hibbertopterus
2586:Hibbertopterus
2574:Hibbertopterus
2570:Hibbertopterus
2554:Hibbertopterus
2535:Hibbertopterus
2527:Hibbertopterus
2515:Hibbertopterus
2511:Hibbertopterus
2495:Hibbertopterus
2483:Hibbertopterus
2475:Hibbertopterus
2433:Hibbertopterus
2411:Hibbertopterus
2403:Hibbertopterus
2391:Hibbertopterus
2342:The idea that
2309:H. caledonicus
2289:Hibbertopterus
2285:Hibbertopterus
2269:
2260:
2256:
2255:
2252:
2251:
2247:
2246:
2243:
2242:
2238:
2237:
2234:
2233:
2229:
2228:
2225:
2224:
2220:
2219:
2216:
2215:
2206:
2203:
2202:
2199:
2198:
2189:
2184:
2182:
2179:
2178:
2175:
2174:
2165:
2160:
2158:
2152:
2151:
2148:
2147:
2143:
2142:
2139:
2138:
2134:
2133:
2130:
2129:
2125:
2124:
2121:
2120:
2111:
2108:
2107:
2104:
2103:
2094:
2089:
2087:
2084:
2083:
2080:
2079:
2070:
2065:
2063:
2060:
2059:
2056:
2055:
2046:
2041:
2039:
2031:
2029:
2026:
2025:
2022:
2021:
2017:
2016:
2013:
2012:
2008:
2007:
2004:
2003:
1994:
1991:
1990:
1987:
1986:
1977:
1972:
1970:
1967:
1966:
1963:
1962:
1953:
1948:
1946:
1938:
1936:
1928:
1921:Hibbertopterus
1898:Hibbertopterus
1894:Hibbertopterus
1875:Hibbertopterus
1860:
1859:Classification
1857:
1853:Hibbertopterus
1821:Hibbertopterus
1815:(described as
1797:C. caledonicus
1793:C. caledonicus
1750:C. caledonicus
1731:S. wrightianus
1695:G. caledonicus
1691:Glyptoscorpius
1677:Adelophthalmus
1673:Glyptoscorpius
1664:Adelophthalmus
1651:Glyptoscorpius
1649:The fact that
1633:
1626:
1625:
1621:Henry Woodward
1610:
1603:
1602:
1601:
1600:
1599:
1591:Hibbertopterus
1583:Hibbertopterus
1575:Hibbertopterus
1548:Hibbertopterus
1508:Hibbertopterus
1452:Henry Woodward
1433:H. wrightianus
1407:G. caledonicus
1403:Glyptoscorpius
1359:G. caledonicus
1351:Glyptoscorpius
1339:Glyptoscorpius
1328:Samuel Hibbert
1284:Henry Woodward
1275:
1272:
1269:
1268:
1261:H. wrightianus
1257:
1251:
1248:
1245:
1240:
1237:
1231:
1230:
1219:
1213:
1208:
1205:
1200:
1197:
1191:
1190:
1179:
1173:
1170:
1167:
1162:
1159:
1153:
1152:
1145:
1139:
1136:
1133:
1128:
1125:
1119:
1118:
1111:
1101:
1098:
1095:
1090:
1087:
1081:
1080:
1073:
1067:
1064:
1061:
1056:
1053:
1047:
1046:
1043:
1040:
1038:
1035:
1032:
1031:Braddy et al.
1029:
1023:
1022:
1019:Hibbertopterus
1011:
1005:
1002:
999:
994:
991:
985:
984:
977:
971:
968:
965:
960:
957:
951:
950:
943:
937:
934:
931:
926:
923:
917:
916:
909:H. caledonicus
905:
899:
896:
893:
888:
885:
879:
878:
875:
872:
869:
866:
863:
860:
838:H. caledonicus
832:
829:
825:Hibbertopterus
820:Hibbertopterus
802:Hibbertopterus
791:Hibbertopterus
782:Hibbertopterus
771:Hibbertopterus
739:Hibbertopterus
722:Hibbertopterus
695:Hibbertopterus
688:hibbertopterid
672:
669:
664:Hibbertopterus
660:Hibbertopterus
633:Hibbertopterus
625:Hibbertopterus
612:Hibbertopterus
593:Hibbertopterus
575:Hibbertopterus
561:Hibbertopterus
557:Samuel Hibbert
539:Czech Republic
511:Hibbertopterus
501:is a genus of
498:Hibbertopterus
492:
491:
490:
489:
480:
472:
460:
459:
453:
452:
450:
449:
429:
428:
427:
425:
424:
422:(Dawson, 1881)
418:H. wrightianus
413:
402:
391:
380:
369:
358:
356:(Salter, 1863)
338:
337:
336:
335:
334:
332:(Braddy, 2023)
323:
312:
298:
297:
293:
292:
289:
279:
278:
272:
271:
264:Hibbertopterus
257:
253:
252:
244:
240:
239:
231:
227:
226:
218:
214:
213:
208:
204:
203:
198:
194:
193:
188:
184:
183:
178:
174:
173:
160:
159:
148:
147:
139:
138:
127:
126:
118:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
28:
25:Hibbertopterus
15:
13:
10:
9:
6:
4:
3:
2:
4835:
4824:
4821:
4819:
4816:
4814:
4811:
4809:
4806:
4804:
4801:
4799:
4796:
4794:
4791:
4790:
4788:
4771:
4766:
4762:
4758:
4753:
4749:
4745:
4740:
4736:
4731:
4725:
4721:
4720:
4718:
4716:
4712:
4708:
4703:
4693:
4683:
4677:
4674:
4672:
4669:
4668:
4666:
4662:
4656:
4653:
4651:
4648:
4646:
4643:
4642:
4640:
4636:
4630:
4629:
4625:
4623:
4622:
4618:
4616:
4615:
4611:
4610:
4608:
4606:
4602:
4596:
4595:
4591:
4589:
4588:
4584:
4582:
4581:
4577:
4575:
4574:
4573:Pentecopterus
4570:
4568:
4567:
4563:
4561:
4560:
4556:
4554:
4553:
4549:
4547:
4546:
4545:Megalograptus
4542:
4540:
4539:
4538:Jaekelopterus
4535:
4533:
4532:
4528:
4526:
4525:
4521:
4519:
4518:
4514:
4512:
4511:
4510:Drepanopterus
4507:
4505:
4504:
4500:
4498:
4497:
4493:
4491:
4490:
4489:Brachyopterus
4486:
4484:
4483:
4479:
4478:
4476:
4472:
4466:
4465:South America
4463:
4461:
4458:
4456:
4455:North America
4453:
4451:
4448:
4446:
4443:
4441:
4438:
4437:
4435:
4433:
4429:
4423:
4420:
4418:
4417:Carboniferous
4415:
4413:
4410:
4408:
4405:
4403:
4400:
4399:
4397:
4395:
4394:Geochronology
4391:
4386:
4364:
4361:
4359:
4356:
4354:
4351:
4350:
4348:
4346:
4342:
4336:
4333:
4332:
4330:
4328:
4324:
4318:
4315:
4314:
4312:
4310:
4306:
4300:
4297:
4295:
4292:
4290:
4287:
4286:
4284:
4282:
4278:
4275:
4273:
4267:
4261:
4258:
4256:
4253:
4251:
4248:
4247:
4245:
4243:
4242:Eurypteroidea
4239:
4233:
4230:
4229:
4227:
4225:
4221:
4215:
4212:
4211:
4209:
4207:
4203:
4200:
4198:
4192:
4182:
4181:Mycteroptidae
4179:
4177:
4174:
4172:
4169:
4168:
4166:
4164:
4163:Mycteropoidea
4160:
4154:
4151:
4149:
4146:
4145:
4143:
4141:
4137:
4131:
4128:
4126:
4123:
4122:
4120:
4118:
4117:Stylonuroidea
4114:
4108:
4107:Rhenopteridae
4105:
4104:
4102:
4100:
4096:
4093:
4091:
4085:
4079:
4076:
4075:
4073:
4069:
4066:
4062:
4056:
4055:Sclerophorata
4052:
4049:
4047:
4043:
4040:
4038:
4034:
4031:
4029:
4025:
4022:
4020:
4019:Euchelicerata
4016:
4013:
4011:
4007:
4005:
4001:
3999:
3995:
3994:
3991:
3987:
3980:
3975:
3973:
3968:
3966:
3961:
3960:
3957:
3947:
3943:
3939:
3935:
3928:
3925:
3920:
3918:9780813722443
3914:
3910:
3906:
3902:
3898:
3891:
3888:
3883:
3879:
3875:
3871:
3870:
3862:
3859:
3848:
3844:
3838:
3836:
3832:
3827:
3823:
3819:
3815:
3810:
3805:
3801:
3797:
3794:(7068): 576.
3793:
3789:
3788:
3783:
3776:
3773:
3768:
3762:
3754:
3750:
3745:
3740:
3736:
3732:
3728:
3724:
3723:
3718:
3711:
3708:
3703:
3699:
3695:
3691:
3687:
3683:
3679:
3672:
3669:
3664:
3660:
3656:
3652:
3651:
3643:
3640:
3636:
3632:
3626:
3624:
3620:
3615:
3611:
3607:
3603:
3599:
3595:
3594:
3589:
3587:
3578:
3575:
3570:
3566:
3562:
3558:
3554:
3550:
3546:
3539:
3536:
3531:
3527:
3523:
3519:
3515:
3511:
3510:
3502:
3499:
3494:
3490:
3486:
3482:
3478:
3474:
3467:
3464:
3459:
3455:
3451:
3444:
3441:
3436:
3432:
3428:
3424:
3420:
3416:
3409:
3406:
3401:
3397:
3393:
3389:
3385:
3381:
3377:
3373:
3366:
3364:
3360:
3355:
3351:
3347:
3343:
3339:
3335:
3331:
3324:
3322:
3320:
3316:
3311:
3307:
3303:
3299:
3295:
3291:
3287:
3283:
3279:
3272:
3269:
3264:
3260:
3256:
3252:
3248:
3244:
3237:
3235:
3231:
3226:
3222:
3218:
3214:
3210:
3206:
3202:
3198:
3194:
3187:
3185:
3181:
3176:
3172:
3168:
3164:
3160:
3156:
3153:(4): 63–104.
3152:
3148:
3141:
3139:
3137:
3135:
3133:
3131:
3129:
3127:
3123:
3119:
3113:
3111:
3109:
3105:
3100:
3096:
3091:
3086:
3082:
3078:
3077:
3072:
3065:
3063:
3061:
3059:
3057:
3055:
3053:
3049:
3045:
3041:
3037:
3033:
3029:
3025:
3021:
3015:
3013:
3011:
3009:
3007:
3005:
3003:
2999:
2994:
2990:
2986:
2981:
2976:
2972:
2968:
2964:
2960:
2956:
2952:
2951:
2946:
2939:
2937:
2933:
2928:
2924:
2920:
2916:
2912:
2908:
2901:
2898:
2892:
2888:
2885:
2883:
2880:
2879:
2875:
2873:
2871:
2867:
2866:
2861:
2860:progymnosperm
2857:
2856:
2851:
2847:
2843:
2839:
2838:
2837:Gyracanthides
2834:
2830:
2826:
2822:
2818:
2813:
2808:
2807:for "head").
2806:
2802:
2798:
2794:
2793:
2788:
2787:
2782:
2781:
2780:Lepidophloios
2776:
2775:
2774:Lepidodendron
2770:
2766:
2762:
2761:
2756:
2752:
2748:
2747:temnospondyls
2744:
2740:
2736:
2735:anthracosaurs
2732:
2728:
2724:
2720:
2715:
2713:
2708:
2704:
2700:
2696:
2692:
2688:
2683:
2681:
2676:
2672:
2667:
2663:
2659:
2654:
2651:
2647:
2643:
2639:
2634:
2627:
2623:
2618:
2612:Palaeoecology
2611:
2609:
2607:
2603:
2599:
2598:Vernonopterus
2595:
2591:
2587:
2583:
2582:Vernonopterus
2579:
2575:
2571:
2567:
2563:
2559:
2555:
2551:
2546:
2544:
2540:
2536:
2532:
2528:
2524:
2520:
2516:
2512:
2508:
2504:
2503:Vernonopterus
2500:
2496:
2492:
2488:
2484:
2480:
2479:Drepanopterus
2476:
2472:
2468:
2467:
2466:Drepanopterus
2462:
2458:
2457:Vernonopterus
2454:
2450:
2446:
2445:Vernonopterus
2442:
2438:
2434:
2427:
2422:
2418:
2416:
2412:
2408:
2404:
2400:
2396:
2392:
2388:
2384:
2379:
2377:
2373:
2369:
2365:
2361:
2357:
2353:
2349:
2345:
2340:
2338:
2334:
2330:
2326:
2322:
2318:
2314:
2310:
2306:
2302:
2301:H. stevensoni
2298:
2297:H. hibernicus
2294:
2290:
2286:
2279:
2274:
2268:
2264:
2261:
2254:
2253:
2245:
2244:
2236:
2235:
2227:
2226:
2218:
2217:
2214:
2213:
2212:
2205:
2204:
2201:
2200:
2197:
2196:
2195:
2188:
2187:
2181:
2180:
2177:
2176:
2173:
2172:
2171:
2164:
2163:
2157:
2154:
2153:
2150:
2149:
2141:
2140:
2132:
2131:
2123:
2122:
2119:
2118:
2117:
2110:
2109:
2106:
2105:
2102:
2101:
2100:
2093:
2092:
2086:
2085:
2082:
2081:
2078:
2077:
2076:
2069:
2068:
2062:
2061:
2058:
2057:
2054:
2053:
2052:
2045:
2044:
2038:
2037:Mycteroptidae
2035:
2034:
2028:
2027:
2024:
2023:
2015:
2014:
2006:
2005:
2002:
2001:
2000:
1993:
1992:
1989:
1988:
1985:
1984:
1983:
1976:
1975:
1969:
1968:
1965:
1964:
1961:
1960:
1959:
1952:
1951:
1945:
1942:
1941:
1935:
1934:Mycteropoidea
1932:
1931:
1927:
1924:
1922:
1918:
1914:
1913:Vernonopterus
1910:
1905:
1903:
1899:
1895:
1891:
1890:Vernonopterus
1887:
1883:
1882:Mycteropoidea
1880:
1876:
1870:
1865:
1858:
1856:
1854:
1850:
1846:
1842:
1838:
1834:
1833:Cape Province
1830:
1826:
1822:
1818:
1814:
1809:
1807:
1802:
1798:
1794:
1790:
1786:
1782:
1778:
1774:
1770:
1766:
1764:
1759:
1755:
1751:
1747:
1742:
1740:
1736:
1732:
1728:
1724:
1720:
1716:
1712:
1711:G. stevensoni
1708:
1704:
1700:
1696:
1692:
1688:
1687:
1686:Vernonopterus
1682:
1678:
1674:
1670:
1669:G. perornatus
1666:
1665:
1660:
1656:
1655:G. perornatus
1652:
1641:
1637:
1630:
1622:
1618:
1614:
1607:
1598:
1596:
1592:
1588:
1584:
1580:
1576:
1571:
1567:
1563:
1559:
1558:
1553:
1549:
1544:
1542:
1538:
1534:
1530:
1525:
1521:
1517:
1513:
1509:
1505:
1501:
1496:
1494:
1490:
1486:
1482:
1478:
1474:
1470:
1466:
1463:
1459:
1458:
1453:
1449:
1445:
1441:
1439:
1434:
1430:
1423:
1418:
1414:
1412:
1411:G. perornatus
1408:
1404:
1400:
1396:
1392:
1388:
1384:
1380:
1376:
1372:
1371:Coomsdon Burn
1368:
1364:
1360:
1356:
1355:G. perornatus
1352:
1348:
1347:G. perornatus
1344:
1340:
1335:
1333:
1329:
1325:
1324:
1319:
1315:
1311:
1303:
1302:
1297:
1293:
1289:
1285:
1280:
1273:
1266:
1265:H. stevensoni
1262:
1258:
1255:
1252:
1249:
1246:
1244:
1241:
1238:
1236:
1233:
1232:
1228:
1224:
1220:
1217:
1214:
1209:
1206:
1204:
1201:
1198:
1196:
1193:
1192:
1188:
1184:
1180:
1177:
1174:
1171:
1168:
1166:
1163:
1160:
1158:
1155:
1154:
1150:
1146:
1143:
1140:
1137:
1134:
1132:
1129:
1126:
1124:
1121:
1120:
1116:
1112:
1109:
1105:
1102:
1099:
1096:
1094:
1091:
1088:
1086:
1083:
1082:
1078:
1074:
1071:
1068:
1065:
1062:
1060:
1057:
1054:
1052:
1049:
1048:
1044:
1041:
1039:
1036:
1033:
1030:
1028:
1025:
1024:
1020:
1016:
1012:
1009:
1006:
1003:
1000:
998:
995:
992:
990:
987:
986:
982:
978:
975:
972:
969:
966:
964:
961:
958:
956:
953:
952:
948:
944:
941:
938:
935:
932:
930:
927:
924:
922:
919:
918:
914:
910:
906:
903:
900:
897:
894:
892:
889:
886:
884:
881:
880:
870:
867:
864:
858:
857:
854:
851:
847:
843:
839:
830:
828:
826:
821:
817:
813:
812:
807:
803:
798:
796:
792:
787:
783:
779:
776:
772:
768:
760:
755:
751:
749:
748:Jaekelopterus
745:
740:
736:
735:
730:
729:
728:Jaekelopterus
723:
719:
716:
715:Carboniferous
712:
708:
704:
700:
696:
692:
689:
682:
677:
670:
668:
665:
661:
657:
653:
652:
647:
646:
640:
638:
634:
630:
626:
621:
617:
613:
608:
606:
605:Jaekelopterus
602:
598:
594:
590:
589:
588:Jaekelopterus
584:
580:
576:
572:
570:
567:word πτερόν (
566:
562:
558:
554:
553:
548:
544:
540:
536:
532:
531:Carboniferous
528:
527:United States
524:
520:
516:
512:
509:. Fossils of
508:
504:
500:
499:
484:
483:Vernonopterus
481:
476:
473:
467:
464:
463:
461:
458:
454:
445:
444:
443:H. stevensoni
439:
434:
420:
419:
414:
409:
408:
403:
398:
397:
392:
387:
386:
381:
378:(Peach, 1883)
376:
375:
370:
365:
364:
363:H. dewalquei
359:
354:
353:
348:
343:
330:
329:
324:
319:
318:
313:
310:(Baily, 1872)
308:
307:
306:H. hibernicus
302:
301:
299:
294:
290:Hibbert, 1836
287:
286:
280:
277:
273:
266:
265:
258:
255:
254:
251:
245:
242:
241:
238:
237:Mycteropoidea
232:
230:Superfamily:
229:
228:
225:
219:
216:
215:
212:
209:
206:
205:
202:
199:
196:
195:
192:
189:
186:
185:
182:
179:
176:
175:
170:
165:
161:
158:
154:
149:
145:
140:
137:
133:
128:
124:
119:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
44:
40:
36:
35:Pennsylvanian
32:
26:
22:
19:
4714:
4626:
4619:
4612:
4592:
4585:
4578:
4571:
4564:
4557:
4550:
4543:
4536:
4531:Hughmilleria
4529:
4523:
4522:
4515:
4508:
4501:
4494:
4487:
4480:
4363:Pterygotidae
4299:Mixopteridae
4255:Eurypteridae
4130:Stylonuridae
4050:
4041:
4032:
4023:
4014:
3937:
3933:
3927:
3900:
3890:
3873:
3867:
3861:
3850:. Retrieved
3847:Live Science
3846:
3791:
3785:
3775:
3761:cite journal
3729:(1): 44–48.
3726:
3720:
3710:
3685:
3681:
3677:
3671:
3654:
3649:
3642:
3634:
3597:
3591:
3585:
3577:
3552:
3548:
3538:
3513:
3507:
3501:
3476:
3472:
3466:
3457:
3453:
3443:
3418:
3414:
3408:
3375:
3371:
3337:
3333:
3288:(1): 25–33.
3285:
3281:
3271:
3246:
3242:
3200:
3196:
3150:
3146:
3083:(1): 49–61.
3080:
3074:
3027:
3023:
2954:
2948:
2910:
2906:
2900:
2863:
2853:
2841:
2835:
2833:climatiiform
2825:acanthodians
2811:
2809:
2796:
2792:Sphenopteris
2790:
2784:
2778:
2772:
2758:
2730:
2716:
2706:
2702:
2694:
2684:
2679:
2674:
2670:
2665:
2661:
2657:
2655:
2645:
2637:
2632:
2630:
2625:
2605:
2601:
2597:
2593:
2592:argued that
2590:H. lamsdelli
2589:
2585:
2581:
2577:
2573:
2569:
2565:
2561:
2557:
2553:
2547:
2538:
2534:
2530:
2526:
2522:
2518:
2514:
2510:
2506:
2502:
2498:
2494:
2490:
2486:
2482:
2478:
2474:
2470:
2464:
2456:
2452:
2448:
2444:
2440:
2436:
2432:
2430:
2425:
2414:
2410:
2402:
2398:
2394:
2390:
2386:
2382:
2380:
2375:
2371:
2367:
2363:
2359:
2351:
2347:
2343:
2341:
2336:
2332:
2328:
2324:
2320:
2316:
2313:H. dewalquei
2312:
2308:
2304:
2300:
2296:
2292:
2288:
2284:
2282:
2277:
2266:
2262:
2210:
2209:
2208:
2193:
2192:
2191:
2168:
2167:
2114:
2113:
2097:
2096:
2073:
2072:
2049:
2048:
1997:
1996:
1980:
1979:
1956:
1955:
1925:
1920:
1917:phylogenetic
1912:
1908:
1906:
1897:
1893:
1889:
1885:
1874:
1873:
1868:
1852:
1848:
1837:South Africa
1820:
1816:
1812:
1810:
1805:
1800:
1796:
1792:
1788:
1784:
1781:C. dewalquei
1780:
1776:
1772:
1768:
1765:ostraviensis
1761:
1757:
1753:
1749:
1745:
1743:
1738:
1734:
1730:
1726:
1722:
1718:
1714:
1710:
1706:
1702:
1698:
1694:
1690:
1684:
1680:
1676:
1672:
1668:
1662:
1658:
1654:
1650:
1648:
1639:
1616:
1594:
1590:
1586:
1582:
1578:
1574:
1569:
1565:
1561:
1555:
1551:
1547:
1545:
1540:
1536:
1532:
1528:
1523:
1519:
1515:
1511:
1507:
1499:
1497:
1492:
1484:
1480:
1476:
1468:
1455:
1447:
1436:
1432:
1426:
1421:
1410:
1406:
1402:
1390:
1383:Mount Vernon
1378:
1374:
1362:
1358:
1354:
1350:
1346:
1338:
1336:
1331:
1321:
1310:John Scouler
1307:
1299:
1287:
1264:
1260:
1234:
1222:
1194:
1186:
1183:H. stevesoni
1182:
1156:
1148:
1122:
1114:
1084:
1076:
1050:
1026:
1018:
1014:
988:
980:
954:
947:H. dewalquei
946:
920:
912:
908:
882:
849:
845:
842:H. dewalquei
841:
837:
834:
824:
819:
809:
801:
799:
790:
781:
770:
764:
758:
757:The size of
747:
743:
742:the species
738:
732:
726:
721:
710:
694:
685:
680:
663:
659:
650:
649:
644:
643:
641:
632:
624:
611:
609:
604:
600:
592:
586:
582:
574:
573:
568:
560:
550:
546:
543:South Africa
510:
497:
496:
495:
482:
474:
465:
442:
441:
432:
431:Also called
417:
416:
406:
405:
395:
394:
384:
383:
373:
372:
362:
361:
351:
350:
341:
340:Also called
328:H. lamsdelli
327:
326:
316:
315:
305:
304:
284:
283:
276:Type species
263:
262:
156:
135:
24:
18:
4798:Stylonurina
4628:Palmichnium
4605:Ichnogenera
4503:Carcinosoma
4197:Eurypterina
4090:Stylonurina
4028:Prosomapoda
4010:Chelicerata
4008:Subphylum:
3986:Eurypterids
3378:(1): 1–20.
2801:common name
2797:H. scouleri
2731:H. scouleri
2680:Cyrtoctenus
2671:Cyrtoctenus
2606:Cyrtoctenus
2594:Dunsopterus
2578:Cyrtoctenus
2566:Cyrtoctenus
2562:Dunsopterus
2558:Dunsopterus
2539:Cyrtoctenus
2531:Cyrtoctenus
2523:Dunsopterus
2519:Cyrtoctenus
2507:Dunsopterus
2499:Cyrtoctenus
2491:Dunsopterus
2487:Cyrtoctenus
2471:Cyrtoctenus
2453:Cyrtoctenus
2449:Dunsopterus
2441:Dunsopterus
2437:Cyrtoctenus
2415:Cyrtoctenus
2399:Cyrtoctenus
2395:Cyrtoctenus
2387:Cyrtoctenus
2383:Dunsopterus
2376:Cyrtoctenus
2372:H. scouleri
2368:Cyrtoctenus
2364:Dunsopterus
2360:Cyrtoctenus
2352:Dunsopterus
2348:Cyrtoctenus
2344:Dunsopterus
2333:Cyrtoctenus
2305:Dunsopterus
2293:H. scouleri
2267:Dunsopterus
2263:Cyrtoctenus
1902:Eurypterida
1879:superfamily
1829:Klaarstroom
1785:Cyrtoctenus
1777:Cyrtoctenus
1763:Ctenopterus
1754:Cyrtoctenus
1744:Other than
1737:, creating
1735:Dunsopterus
1727:Dunsopterus
1719:Cyrtoctenus
1715:Dunsopterus
1703:Cyrtoctenos
1699:Cyrtoctenus
1659:G. kidstoni
1640:H. scouleri
1617:H. scouleri
1595:H. scouleri
1566:C. oculatus
1541:Cyrtoctenus
1533:Cyrtoctenus
1514:as part of
1477:Echinocaris
1469:Echinocaris
1462:phyllocarid
1457:Echinocaris
1438:Equisetides
1391:G. kidstoni
1288:H. scouleri
1256:(Devonian)
1216:Tournaisian
1149:H. scouleri
1104:Tournaisian
1008:Tournaisian
976:(Devonian)
942:(Devonian)
769:(limbs) of
759:H. scouleri
734:Carcinosoma
711:H. scouleri
707:exoskeleton
699:Stylonurina
691:eurypterids
681:H. scouleri
671:Description
651:Dunsopterus
645:Cyrtoctenus
583:H. scouleri
579:stylonurine
547:H. scouleri
487:Peach, 1905
475:Dunsopterus
466:Cyrtoctenus
433:Dunsopterus
342:Cyrtoctenus
317:H. scouleri
224:Eurypterida
211:Chelicerata
207:Subphylum:
157:H. scouleri
136:H. scouleri
4787:Categories
4594:Stylonurus
4580:Pterygotus
4559:Mixopterus
4552:Megarachne
4517:Eurypterus
4402:Ordovician
4358:Slimonidae
4269:Infraorder
4046:Dekatriata
4037:Planaterga
4004:Arthropoda
3852:2017-12-28
3600:: 77–103.
3555:: 77–103.
2893:References
2755:millipedes
2687:St Andrews
2548:In a 2019
2543:scavengers
1529:Pterygotus
1516:Pterygotus
1485:Stylonurus
1481:Stylonurus
1465:crustacean
1448:Stylonurus
1444:James Hall
1440:wrightiana
1401:. Peach's
1323:Eurypterus
1301:Eurypterus
1296:appendages
1247:Uncertain
1070:Bashkirian
1015:Pterygotus
1001:Uncertain
767:appendages
616:appendages
552:Eurypterus
517:period in
507:arthropods
503:eurypterid
201:Arthropoda
43:Bashkirian
4671:Metastoma
4655:Xiphosura
4645:Arachnida
4432:Geography
3996:Kingdom:
3702:246819305
3485:0022-3360
3427:0022-3360
3400:130931651
3392:2053-5945
3354:129589593
3310:128557225
3302:0263-5933
3255:0022-3360
3225:140167776
3217:0036-9276
3175:131694288
3167:2053-5945
3044:1755-6929
2971:1744-9561
2913:: 19–99.
2786:Stigmaria
2769:Opiliones
2765:scorpions
2751:tetrapods
2743:baphetids
2739:aistopods
2723:Edinburgh
2497:and that
2356:sclerites
2337:H. peachi
2325:H. peachi
1806:C. peachi
1789:C. peachi
1746:C. peachi
1707:C. peachi
1550:in 1959,
1343:Ben Peach
1254:Famennian
1212:(250 cm)
1187:H. peachi
1115:H. peachi
940:Famennian
925:Fraipont
913:H. peachi
850:H. peachi
795:hexapodal
786:substrate
778:podomeres
620:substrate
396:H. peachi
374:H. dicki
187:Kingdom:
181:Eukaryota
4724:Wikidata
4692:Category
4614:Arcuites
4587:Slimonia
4412:Devonian
4407:Silurian
4194:Suborder
4087:Suborder
4064:Taxonomy
4002:Phylum:
3998:Animalia
3946:56354233
3818:16319874
3753:17148124
3614:84151264
3569:84151264
3530:46953627
3099:85398946
2989:19828493
2927:85862868
2876:See also
2850:unionids
2846:bivalves
2622:Scotland
2604:, while
2317:H. dicki
1693:species
1636:tergites
1613:carapace
1537:P. dicki
1524:P. dicki
1512:P. dicki
1399:Somerset
1395:Radstock
1318:tergites
1127:Hibbert
981:H. dicki
974:Givetian
859:Species
827:itself.
718:Scotland
703:carapace
525:and the
523:Scotland
515:Devonian
457:Synonyms
296:Species
243:Family:
197:Phylum:
191:Animalia
177:Domain:
153:tergites
132:carapace
39:Givetian
4757:1055858
4744:4650096
4730:Q141251
4460:Oceania
4422:Permian
3940:: 1–6.
3826:4422644
3796:Bibcode
3744:1629066
3637:. 3890.
3493:1300755
3435:1301214
3263:1301554
2980:2865068
2870:lycopod
2717:In the
2650:pelagic
2335:(where
1845:exuviae
1634:Fossil
1387:Glasgow
1314:prosoma
1290:with a
1250:120 cm
1239:Dawson
1138:180 cm
887:Salter
871:Length
868:Status
862:Author
811:gladius
713:, from
535:Ireland
529:to the
519:Belgium
256:Genus:
217:Order:
151:Fossil
130:Fossil
4450:Europe
4440:Africa
4071:Genera
3944:
3915:
3824:
3816:
3787:Nature
3751:
3741:
3700:
3612:
3567:
3528:
3491:
3483:
3433:
3425:
3398:
3390:
3352:
3308:
3300:
3261:
3253:
3223:
3215:
3173:
3165:
3097:
3042:
2987:
2977:
2969:
2925:
2862:) and
2821:sharks
1389:, and
1292:telson
1227:ocelli
1210:135 cm
1207:Valid
1176:Viséan
1172:80 cm
1169:Valid
1142:Viséan
1135:Valid
1108:Viséan
1097:Valid
1063:Valid
1037:Valid
993:Baily
967:Valid
959:Peach
933:Valid
902:Viséan
895:Valid
806:telson
775:distal
629:telson
569:pteron
537:, the
4770:18973
4752:IRMNG
4051:Clade
4042:Clade
4033:Clade
4024:Clade
4015:Clade
3942:S2CID
3822:S2CID
3698:S2CID
3610:S2CID
3565:S2CID
3526:S2CID
3489:JSTOR
3431:JSTOR
3396:S2CID
3350:S2CID
3306:S2CID
3259:JSTOR
3221:S2CID
3171:S2CID
3095:S2CID
2923:S2CID
2805:Scots
2517:with
2473:from
1827:near
1393:from
1381:from
1034:2023
865:Year
816:Roman
565:Greek
4739:GBIF
4445:Asia
3913:ISBN
3814:PMID
3767:link
3749:PMID
3481:ISSN
3423:ISSN
3388:ISSN
3298:ISSN
3251:ISSN
3213:ISSN
3163:ISSN
3040:ISSN
2985:PMID
2967:ISSN
2823:and
2789:and
2745:and
2691:Fife
2596:and
2556:and
2521:and
2501:and
2485:and
2455:and
2443:and
2385:and
2366:and
2346:and
2327:and
2295:and
2265:and
1911:and
1888:and
1771:and
1748:and
1409:and
1294:and
1243:1881
1203:1985
1165:1877
1131:1836
1093:1968
1059:1951
997:1872
963:1883
929:1889
891:1863
848:and
814:, a
731:and
648:and
541:and
56:PreꞒ
3905:doi
3878:doi
3874:252
3804:doi
3792:438
3739:PMC
3731:doi
3690:doi
3659:doi
3633:".
3602:doi
3598:103
3557:doi
3553:103
3518:doi
3380:doi
3342:doi
3290:doi
3205:doi
3155:doi
3085:doi
3032:doi
2975:PMC
2959:doi
2915:doi
2872:).
2689:in
2459:as
2409:of
1733:to
1638:of
1619:by
1615:of
1397:in
1286:of
737:),
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