Kambara - Knowledge (XXG)

3937:, the upper forelimb may have been able to swing with much greater force, while also providing greater stability against gravity and thus improving the animals ability to perform the crocodilian high walk, the mode of locomotion during which crocodiles lift their body off the ground. Additionally, these factors may also increase stride-length and walking speed. The change to the deltapectoral crest could have advantages for swimming, allowing the limbs to better resist the drag during the recovery stroke. Furthermore, the shortening of the humeral shaft may have also come with advantages, although this is less clear due to the fact that no material of the lower forelimb has been described yet, making it ambiguous how the musculature of the lower and upper arm would connect. Regardless, it is possible that the shortening could have brought the musculature into a more proximal position, which would mean that the lower limbs would have exerted less force while walking or swimming, allowing the animal to swing the lower limb faster. 3953:, based on material collected from Tingamarra, were described by Stein and colleagues in 2017 and feature a robust ilium similar to what is seen in modern alligators and gharials, but with notably shallower peduncles (the parts of the ilium that connect to the other bones of the hip). The joint for the femur is shallowly concave and the postacetabular process, the section of the ilium behind this joint, lacks the constriction seen in modern crocodiles. Otherwise the ilium does not differ significantly for what is seen in other Eusuchians. This morphotype is designated "pelvic form one" by Stein and colleagues and differs in its morphology from fossils found in the Rundle Formation that may pertain to another species of 3893:, shows several circular pits arranged in a linear fashion, which is in line with the bite marks left by a crocodilian. The amount of tooth marks present on the plastron indicates that the individual that tried to eat the turtle bit it several times, behaviour that would match the "juggling" performed by modern crocodiles. "Juggling" in this case describes the act of repeatedly biting prey like turtles in order to change its orientation, helping to align it either with the teeth or to make it easier to swallow. However, it is believed that in this instance the crocodile was overambitious, as the plastron's width and thickness would be too great for the average 4072:. Salisbury and Molnar suggest that, if not ecologically different, the two species may have differed in their habitat preferences and possibly wouldn't have crossed paths under normal circumstances. They point out that the fossil site near Murgon shows signs of having undergone both dry and wet periods, with the former greatly reducing the present bodies of water. A drought could have driven one of the species from their natural habitat and forced them to look for sanctuary in water that would otherwise be home to the other form. Somewhat similar circumstances may have led to the creation of the bonebed that preserves the fossils of 809: 4109: 4080:

large, adult males prefer deeper open waters. This could explain why the Murgon site primarily preserves large individuals alongside some remains of eggs and hatchlings, with only few animals of intermediate size. Such subadults may have lived largely separate from adults in so called "stockyards", as they would pose a threat to small hatchlings while themselves being threatened by cannibalistic adult males. This could explain why the Murgon site primarily preserves large individuals alongside some remains of eggs and hatchlings, with only few animals of intermediate size.

4053: 6735: 6730: 5890: 146: 3870: 990:, which had been studied and compared to the equivalent bones in saltwater and freshwater crocodiles. In both of these modern forms, the deltopectoral crest is offset medially from the lateral margin of the shaft. The apex of the crest is also directed medially, which essentially places it just above the midline of the shaft. The deltapectoral crest sits at a right angle relative to the ventral face of the humeral shaft. However, in 4045:. Both species are known from a single fossil site near Murgon, with the remains being intermingled with one another, suggesting they occurred in the same environment at the time of their deaths. Morphologically, both species share a variety of similarities and differences. Both are semi-aquatic animals with platyrostral snouts and both are believed to have reached similar lengths. The two however differ in that 820: 5895: 3962: 120: 592: 3806: 927:. Around the seventh and eight dentary teeth, as well as from the twelfth to seventeenth, the teeth appear to have interlocked. However, the tenth to twelfth teeth of the dentary are confluent, meaning they are too closely spaced to interlock with those of the upper jaw, necessitating an overbite in this area. This clearly sets it apart from all other species of 972: 1014:, the dorsal and ventral faces are compressed, which in turn lengthens the medial and lateral condyles, while also directing these surfaces towards the midline. The profile of the distal end is subsequently hexagonal. Profile and size of the medial and lateral condyles in saltwater and freshwater crocodiles are nearly identical, but very different in 4009:, is mentioned, but cannot be fully confirmed due to the poor preservation of that particular element. Simultaneously, Buchanan notes that the skeletal is not any more specialised in aquatic life than that of modern crocodilians either, suggesting a similarly generalized semi-aquatic mode of life. Finally, it is pointed out that the 892:

interlocking teeth in some areas and a slight overlap further back in the jaw. The enlarged fifth tooth of the maxilla does however occupy more space, with its pit being set much further laterally than any of the others. Regardless, this clearly sets not just the skulls but also the dentaries apart from those assigned to

548:, named in 2009 by Lucas A. Buchanan. Although the type description is limited to the skull material (two crania and a lower jaw), which is easily comparable to the previously established species, Buchanan takes note of a "considerable amount" of postcranial remains. These remains were partly described in a 4141:

attacked members of their own species. Such behavior is commonly seen in extant crocodilians and even recorded in a multitude of fossils. Crocodilians often attack limb bones, which would explain the fractures seen on the pathological humerus and metatarsals. The high number of individuals present at

3901:. A costal scute of what is thought to be the same individual has also been recovered, also showing signs of a crocodile bite, some healing and infection. While this suggests that turtles, which were a readily available food source, were fed on at least occasionally, it does not necessarily mean that 946:

All teeth are slightly compressed side to side and possess distinct carinae (cutting edges), but are not ziphodont like in some later mekosuchines. Both this compression and the development of the edges is strongest towards the front of the jaw and gradually grows weaker further back in the toothrow,

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differ significantly, as neither preserve fully overlapping dentition. Instead, distinct occlusal pits reveal that the teeth slid in between each other when the jaw was closed. This means that the teeth of these two species would interlock in a fashion much more similar to what is observed in species

3825:

was a semi-aquatic generalist. One method to determine the details of such a lifestyle may be found in the way the teeth of these crocodilians occlude with another, although the specifics of this are still poorly understood. One hypothesis by Charles C. Mook proposed that an overbite in crocodilians

4146:

bonebed lends itself well to this hypothesis as well, with the crowded nature of the area leading to increased aggression between its residents. In both cases, the injured animal survived and healed. The osteomyelitis of NMV P227802 meanwhile was caused by a bacterial infection causing necrosis and

4079:

It is also possible that the animals were typically kept apart not by preferences unique to the two species but through preferences based on size, age and sex. In modern saltwater crocodiles, the ranges of nesting females and juveniles often overlap upriver or in areas of denser vegetation, whereas

3845:

Another important factor that differentiates the ecology of the various species is the development of the retroarticular process. The retroarticular process serves as an attachment site for important jaw adductor and abductor muscles and is subsequently of great importance in capturing and subduing

4121:

and subsequently formed a callus, a growth of cancellous bone, during healing. Another specimen represents a series of metatarsals, toe bones, which were fractured during the animal's life and then fused into a single element as it healed, with the individual elements rotating due to the continued

4020:

seemingly displays a combination of adaptions for both semi-aquatic and somewhat terrestrial life, with the limbs suggesting a superior ability to walk on land relative to modern forms, whereas the shape and form of the skull are indicative of a semi-aquatic ambush hunting lifestyle not unlike the

994:

things are notably different. Neither the deltapectoral crest nor its apex are directed medially, with the crest as a whole being just above the lateral margin of the shaft and the apex being directed ventrally. The right angle formed by the crest and shaft however remains. This is associated with

431:

at Murgon. It is unclear how exactly these two species differed in their niche, but they did differ notably in their anatomy, with the latter being slightly more gracile and with interlocking teeth. It is also possible that the two species didn't coexist at all, but instead were driven together by

410:

is poorly understood. The general shape of the skull, which is flattened, and the environment the fossils were deposited in, all suggest that they were semi-aquatic ambush predators like modern crocodilians. However, subtle differences in the postcranial remains, especially the limb bones, suggest

3944:

is an additional point of consideration. The olecranon fossa and the articular surface of the humerus both suggest a wider range of motion than in saltwater and freshwater crocodiles, but especially the former would require the description of the lower arm to verify this conclusion. The increased

891:

clearly interlock with each other based on the presence of the aforementioned occlusal pits between the individual tooth sockets up to the ninth tooth of the upper jaw. After this point, the pits are located further medially. This is also confirmed by the pits present in the lower jaw, indicating

3957:

or possibly a different genus altogether. Assuming the former, the Rundle fossils ("pelvic form two") feature much deeper peduncles which are shared with younger mekosuchines from the Oligocene and Miocene, but lack the specific expansion of the pubic peduncle seen in later taxa. Additionally,

3779:

with its intermediate form, various undetermined crocodilians of the time with interlocking teeth and phylogenetic analysis all seem to suggest that the genus actually began with interlocking dentition, acquiring the overbite later. Recent phylogenies consistently point towards

849:

differs significantly and is used to set them apart. Generally, three different morphologies can be observed, with the difference being primarily expressed through how the upper and lower toothrow interact with one another. As in many other crocodilians, the dental margin of

500:

The mid 1980s saw the discovery of mammal remains in the area, leading to increased study and revised dating. This new-found attention led to the discovery of several additional crocodilian fossils, including a largely complete skull (QM F21115) collected in 1991. The genus

4064:, Salisbury and Willis argue that the difference in head shape was of little value to their respective ecologies. They suggest that the differences are so minor that they are effectively still the same ecomorph, which they determine may have been similar to that of today's 521:

seemingly showed a high degree of variation among its fossils, in particular regarding the better preserved lower jaws. For this reason, it was suggested that the fossils could belong to multiple species of a single genus. This was confirmed when a second species,

3830:

of mammalian predators, being used to break and slice. This would then suggest that interlocking dentition may be better suited when having to restrain large, struggling prey, requiring greater force that would be more evenly spread out across the toothrow.

423:. While this could suggest that it was faster on land than living crocodilians, it does not necessarily mean that it was terrestrial like some later mekosuchines are thought to have been. Another question regarding its ecology is posed by the coexistence of 3917:

is typically regarded as such a semi-aquatic animal, which would be supported by the environment it was found in. While the skull displays no particular adaptations that would suggest any degree of terrestriality, as is the case in mekosuchines like

4116:

In addition to the many well preserved bones that show no signs of non-taphonomic distortion, there is plentiful material that shows clear signs of injury. This material, covered in Buchanan's thesis, includes a femur that has suffered a

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and other extinct Australian crocodilians were noted since the description of the Murgon mandible in 1982. By the 1993 description, the idea of an endemic Australian crocodilian radiation was better established, at the time including

777:

was described, it was recognised as a highly variable genus, retaining certain key traits but differing notably in others. As noted by Buchanan, the most prominent of these differentiating traits centre around the anatomy of the

4029:. However it is also possible that this was no adaption towards movement on land, but rather used for bottom-walking in shallow waters, a habit that could have led to more terrestrial mekosuchines later during the Cenozoic. 995:

the torsion of the shaft. In the modern forms, the humerus is given a sigmoid shape by the fact that the articulating surfaces, the parts of the bone that connect to the joints, are at an angle to each other. The humerus of

2372:. In addition to the monophyletic Mekosuchinae recovered by the vast majority of their analysis, two more novel results were also found. These alternative topologies render Mekosuchinae paraphyletic, including the clade 4228:

may have been found across many of the inland waterways of Queensland, possibly even all of eastern Australia where temperatures allowed for the presence of crocodilians. If this was the case, then the distribution of

786:

stands out as having a long, high and wide process, offering a greater surface area for the musculature to attach to. By comparison, the retroarticular processes in older species are less well developed, with that of

4421:

Stein, M.; Salisbury, S. W.; Hand, S. J.; Archer, M.; Godthelp, H. (2012). "Humeral morphology of the early Eocene mekosuchine crocodylian Kambara from the Tingamarra Local Fauna southeastern Queensland, Australia".

4191:, although none of them were especially large, seemingly maxing out at 100 mm (3.9 in) in length. This would suggest that the waters of this locality were rather shallow, which could indicate that it was a 3981:

does indeed possess some traits associated with terrestrial locomotion, however, they are generally not as well developed as one would expect from a fully terrestrial animal. Among these is the development of the

887:, the first teeth of the lower jaw slide into very pronounced pits that pierce the bones of the upper jaw, emerging on its upper surface in front of the nostrils. The remaining premaxillary and maxillary teeth of 753:

appears typically crocodilian, being roughly triangular with a rostrum that begins relatively narrow and continues to widen towards the back. The precise degree to which the snout tapers differs between species.

4126:. Based on the advanced state of the infection, it is thought to have been chronic and to have been affecting the individual for years. The final pathological specimen noted by Buchanan is a highly deformed 629:

was not recognised as a distinct species until the discovery of nearly complete skulls in 1993, clearly showing major differences with the type species. Among these differences is the fact that the teeth of

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and their position in the lower jaw may have increased and focused the force of the bite in a smaller area, enhancing their ability to puncture and drawing further parallels to mammalian carnassials.

4130:, which in some areas is twice as wide as it should be. The reason for the extreme deformation of this particular specimen is not entirely clear, but could have been caused by one of several possible 4021:

one seen in most extant crocodilians, while certain adaptations of the arm are also advantageous for swimming. Buchanan interprets this to mean that while not being a true terrestrial crocodilian,

4333:

Salisbury, S. W.; Willis, P. M. A. (1996). "A new crocodylian from the early Eocene of south-eastern Queensland and a preliminary investigation of the phylogenetic relationships of crocodyloids".

3862:

show well defined attachment sites for musculature used to hold and crush prey, an activity the pterygoid would not be involved in. This collaboration between different muscle groups would allow

634:

interlock, rather than forming an overbite. It is this fact that the species name derives from. Additionally, the skull of this species was relatively more gracile, as are this species' teeth.

4122:

use of the limb. NMV P227802 on the other hand represents a heavily altered humerus, characterised by being swollen and containing numerous sinuses filled with small bone fragments, signs of

4167:

were a part of the Tingamarra Fauna, which also included a variety of terrestrial and aquatic animals. Mammals are represented through various marsupials, while the reptile fauna includes

1006:

There are also differences concerning the distal end, the end towards the toes rather than the hip. Here, modern Australian crocodiles have flat medial and lateral surfaces bordering the

3897:

individual to manage. Subsequently, the turtle was let go or escaped the crocodile, allowing the bite to somewhat heal. Despite this however, the turtle later died from infection and

3854:

stands out as having the most strongly developed processes, which correlates with larger pterygoid muscles and a stronger bite than that of its older relatives. This may suggest that

766:

was relatively flat, lacking the raised edges seen in today's crocodiles, but instead multiple specimens across species show the presence of well developed pits just before the

344:. It is generally thought to have been a semi-aquatic generalist, living a lifestyle similar to many of today's crocodiles. Four species are currently recognised, the sympatric 1080:, which may be a subadult, has been estimated to have been 2.5 m (8 ft 2 in), while a much larger mandible suggests lengths of up to 3.5 m (11 ft). 900:

the teeth clearly interlock in the premaxilla as well, however, while the first pair of dentary teeth also extend into the bone, they do not breach the dorsal surface as in

583:

is stated to derive from an Aboriginal word for crocodile, however, it is not specified which language in particular, only that it stems from the Queensland language group.

526:, was named three years later on the basis of new material collected from the same locality, with some of the previously recovered fossils now being assigned to this taxon. 2392:. However, these results are a minority within the team's analysis and even then nodal support for paraphyletic Mekosuchinae is considered to be poor by the authors. 6846: 3929:

Most postcranial material remains unpublished, the exception being the humerus, which is known from an undetermined species found at the Murgon locality (possibly

3775:

was the basal most species of the genus, which would have rendered the overbite characteristic for this species the ancestral condition. However, the discovery of

3958:"pelvic form two" displays a knob situated atop the postacetabular crest which appears to be convergently developed with the constriction of modern crocodylids. 702:. It was named for Murgon, the closest township to the type locality. It was a moderately robust animal with a skull much wider than that of the contemporary 4803: 947:

while also becoming more bulbous and peg-like. While the number of premaxillary teeth is consistent, the amount of teeth in the maxilla and dentary varies.

664:

species, but has enough differences to the other forms to be considered distinct. Most notably, the teeth appear to be intermediate between the overbite of

1049:

in articulation. Ribs and chevrons are likewise known. Most of this material however have not seen formal publication and are only described in Buchanan's

491:

in 1982, who reported a lower jaw bone he believed to belong to a Miocene crocodylid. While noting similarities to other Australian crocodilians, notably

4147:

bone death. While it is unclear how the animal was infected, it is possible that the area was exposed by an injury, possibly from the attack of another

3973:

Some additional discussion of other postcranial material can be found in Buchanan's PhD thesis, which also contained the later published description of

1129:

in 1993, but it was still recognised as the basalmost member of Mekosuchinae. Subsequent papers largely supported this suggestion, typically recovering

762:, although still platyrostral, is notably slender compared to its contemporary relative. Similarly, the teeth are also more slender. The skull table of 4683:"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem" 6995: 6926: 6833: 4013:

bonebed was likely the result of mass death caused by drought, something that would be less likely to affect an animal less dependent on water.

951:

has the lowest number of teeth in the maxillae (and thus throughout the upper jaw) with only 14 teeth on either side. The number is unknown for

4736:"Variation in the pelvic and pectoral girdles of Australian Oligo–Miocene mekosuchine crocodiles with implications for locomotion and habitus" 3858:

was capable of taking on larger, more terrestrial prey items, whereas older species fed on smaller animals. Furthermore, multiple species of

517:, Ralph Molnar and John D. Scanlon, who tentatively referred all Murgon crocodile fossils to this taxon. Even at this time it was noted that 3990:, the later of which only differing slightly from saltwater crocodiles and with only limited mobility of the ankle. The possibility that 1133:

as one of the earliest branching mekosuchines, although other taxa may be found as slightly more basal or on an even level as part of a

740:

people and simply means crocodile. This language was chosen as a proxy for that of the Bailai due to the links between the two cultures.

4631: 3821:

With its appearance generally resembling today's crocodiles, in particular the flat, platyrostral snout, it is generally thought that

537:

was known for a while, with the type specimen found in 1991. However the material was not closely examined until the description of

4096:, but also at least two hatchlings and some fragmentary egg shells. This may suggest that the Murgon site could have been used by 6990: 6729: 4796: 6734: 6985: 6062: 5889: 6108: 5977: 4108: 2841: 1766: 3949:

would have been capable of high walking at a greater speed than the crocodiles found in Australia now. The pelvic bones of

2342:

was still recovered as a mekosuchine, however, the overall make up of the clade was changed significantly. The study found

999:

meanwhile is much straighter, with both surfaces being level to each other. As a consequence, the shaft of the humerus in

808: 443:

analyses recover it as one of the earliest diverging mekosuchines, but recent studies have commonly found other taxa like

4557:

The systematics, palaeobiology and palaeoecology of Kambara taraina sp. nov. from the Eocene Rundle Formation, Queensland

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sp. nov. (Crocodylia, Crocodyloidea), a new Eocene mekosuchine from Queensland, Australia, and a revision of the genus".

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Ristevski, J.; Willis, P.M.A.; Yates, A.M.; White, M.A.; Hart, L.J.; Stein, M.D.; Price, G.J.; Salisbury, S.W. (2023).

4049:

was slightly more gracile both in regards to its skull and teeth in addition to the different ways the teeth occluded.

2368:

were not found to be mekosuchines in this analysis. Another, much more different placement was also shown in Ristevski

4789: 2666: 2044: 533:, although it stands out as the species with the least amount of available material. Like with the species before it, 5969: 5136: 3450: 415:

may have been better adapted at walking on land than the two crocodile species still inhabiting Australia today, the

372:

were medium-sized crocodilians, with mature specimens generally reaching lengths from 3–4 m (9.8–13.1 ft).

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and the dentition. Regarding the retroarticular process, an important attachment site for the abductor musculature,

6190: 4627:"Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil" 497:(named nine years later), Molnar refrained from naming the material due to the fragmentary nature of the dentary. 6162: 3019: 1803: 699: 476: 145: 5808: 1125:

in this group and redefining Mekosuchidae as Mekosuchinae. No detailed phylogenetic analysis was conducted for

736:

interlock and thus set it apart from the other known species. The species name derives from the dialect of the

625:

in the Boat Mountain area near Murgon, Queensland, indicating that the two species coexisted. Because of this,

5129: 3500: 5894: 758:

for instance is described as being moderately broad-snouted with a robust build. By comparison, the skull of

6183: 3324: 2700: 2625: 2091: 1990: 779: 767: 724:, it stems from the Kerosene Creek Member of the Rundle Formation, which means that it did not coexist with 6669: 6443: 6773: 6693: 6467: 6176: 4387:"A new species of mekosuchine crocodilian from the middle Palaeogene Rundle Formation, central Queensland" 4092:

other than the fact that the Murgon fossil site preserves not just the remains of large, presumably adult

3913:

As indicated by aspects of the skull shape, namely the resemblance to modern semi-aquatic ambush hunters,

3573: 2769: 1929: 1137:. For instance, a 2018 study by Lee and Yates, using genetic, morphological and stratigraphic data, found 22: 4578:"Migrations, diversifications and extinctions: the evolutionary history of crocodyliforms in Australasia" 4137:

At least some of these pathologies are explainable through intraspecific combat, meaning that individual

861:, the teeth of the upper jaw overlap those of the lower jaw, giving it an overbite similar to that of an 6980: 6952: 6872: 6653: 6646: 6459: 6451: 6335: 4972: 4892: 3419: 3307: 2017: 6677: 657: 361: 6913: 6820: 6169: 5858: 5851: 5801: 5599: 4589: 4505: 4431: 4342: 4234: 2330:

Although Mekosuchinae as a monophyletic clade became a well established group in the years following

439:

is considered an important piece in understanding the evolutionary history and origin of this group.

420: 3283: 2601: 1963: 1515: 5837: 5564: 4118: 3259: 3208: 2649: 416: 4056:

An example for modern sympatric crocodilians includes American alligators and American crocodiles.

676:. It was named in honour of Ralph Molnar for his consistent contributions to mekosuchine research. 6701: 6610: 6602: 6328: 5606: 5585: 5578: 5239: 5115: 4919: 4607: 4521: 4447: 4224:

may have been much more widespread than currently known. Holt, Salisbury and Willis propose that

3966: 2683: 2071: 1042: 140: 6957: 6877: 4217:

stemming from the younger deposits of the Kerosene Creek Member instead of the Brick Kiln Seam.

552:, but unlike the cranial material have not been formally published. An exception to this is the 4213:. Unlike with the species of the Tingamarra Fauna, the Rundle species were not sympatric, with 6939: 6859: 6851: 6617: 6116: 6083: 5844: 5571: 5376: 4767: 4716: 4660: 3869: 1084:

is thought to have been of similar size, albeit with a more robust build. The same applies to

6944: 6864: 4041:

was how two rather similar species of the same genus could coexist as closely as it did with

2334:

s description, it is not without competing ideas. This in particular affects the position of

6595: 5830: 5693: 5656: 5592: 5487: 5353: 4757: 4747: 4706: 4696: 4650: 4640: 4597: 4513: 4439: 4350: 4168: 3827: 3814: 2373: 1046: 695: 213: 200: 6685: 6661: 6624: 6544: 6292: 6019: 5778: 5763: 5728: 5671: 5541: 5397: 5342: 5159: 5143: 4195:. Furthermore, it is known that the locality was subject to cyclical wet and dry seasons. 3834:

Buchanan speculated on the potential implications for the intermediate dentition seen in

4593: 4509: 4435: 4346: 6558: 6536: 6278: 6270: 6228: 6026: 5961: 5947: 5820: 5533: 5517: 5446: 5404: 5362: 5334: 5312: 5269: 5204: 5191: 5166: 5151: 5122: 5053: 5009: 4994: 4762: 4735: 4711: 4682: 4655: 4626: 3945:

range of motion is more certain based on the articular surface, and would suggest that

2957: 2935: 1605: 1203: 1022: 493: 451: 6974: 6811: 6756: 6588: 6581: 6566: 6551: 6514: 6500: 6412: 6397: 6389: 6306: 6285: 6256: 6249: 6242: 6132: 6046: 6034: 5984: 5954: 5939: 5925: 5707: 5678: 5648: 5510: 5479: 5466: 5454: 5438: 5420: 5383: 5283: 5107: 5085: 4611: 4451: 4220:

Although so far only known from two sedimentary basins, each preserving two species,

4123: 3191: 2793: 1752: 1354: 933: 656:

is known from a partial mandible and several isolated remains all collected from the

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has been found, most of said material remains undescribed with the exception of the

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of Australia, with the first remains found in rock units of what may be part of the

379:

are easiest differentiated by the different occlusal patterns of their teeth. While

6751: 6574: 6507: 6420: 6405: 6372: 6361: 6313: 6095: 6005: 5992: 5912: 5791: 5742: 5735: 5663: 5525: 5502: 5327: 5319: 5290: 5232: 5225: 5100: 4985: 4877: 4852: 4286: 4172: 3998: 3983: 3961: 3679: 3467: 2987: 2568: 2551: 2493: 2389: 2348: 2283: 1789: 1716: 1417: 1390: 1189: 1034: 963:

possess 16 teeth in either maxilla. The lower jaw contains 17 or 18 dentary teeth.

939: 573: 510: 488: 330: 279: 239: 4602: 4577: 3889:

which appears to have been attacked by a crocodile. The fossil, a nearly complete

819: 487:. Although known for half a century, the material was first formally described by 6904: 4443: 6805: 6529: 6522: 6320: 6299: 6263: 6235: 6214: 5756: 5749: 5721: 5714: 5700: 5641: 5494: 5430: 5390: 5369: 5297: 5276: 5218: 5092: 5060: 5017: 4176: 4112:

Crocodiles will often attack their opponents' limbs during intraspecific combat.

3550: 3143: 1334: 1155: 1038: 1030: 514: 445: 52: 6796: 6483: 6347: 6152: 6076: 6069: 6055: 5634: 5251: 5031: 4931: 4843: 4813: 4354: 4180: 3987: 3341: 2867: 1578: 1121: 1088:, which was described as reaching lengths from 3–4 m (9.8–13.1 ft). 1050: 879: 484: 459:

was no mekosuchine at all, but said results are not supported by all authors.

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may have been much more adapt at leaving the water than members of the genus

6433: 5771: 5554: 5038: 4904: 4210: 4192: 4069: 4002: 3084: 2353: 2163: 1072: 862: 591: 565: 472: 396: 384: 341: 157: 102: 46: 4771: 4720: 4664: 4645: 3805: 4734:

Stein, Michael D.; Yates, Adam; Hand, Suzanne J.; Archer, Michael (2017).

4037:

One particular question that was raised in the wake of the description of

1064:

was a medium-sized crocodilian, smaller than the more recent mekosuchines

971: 854:

appears wave-like, though it is more gentle than some other mekosuchines.

6898: 6790: 6636: 6124: 5685: 5305: 5045: 4861: 4837: 4517: 4207: 4184: 3995: 3898: 3890: 3882: 3043: 2824: 2210: 1625: 1134: 1115: 1026: 737: 177: 92: 87: 72: 67: 57: 4701: 919:, which shows a somewhat intermediate condition between the overbite of 403:, although incompletely known, seems to represent an intermediate form. 6931: 6838: 4781: 4752: 4188: 1007: 987: 553: 323: 187: 107: 82: 5621: 4831: 4127: 4065: 3926:, some elements of the postcranium suggest a more complex lifestyle. 549: 480: 353: 337: 167: 39: 6767: 4278: 5067: 4687: 4279:"An early Eocene crocodilian from Murgon, southeastern Queensland" 4131: 4107: 4051: 3960: 3868: 3804: 1551: 1003:

is not lengthened as is the case in modern Australian crocodiles.

970: 590: 564:

has been recovered from the Rundle Formation, however, it was not

326: 6918: 6825: 6140: 5002: 4006: 3792:

is typically removed due to the limited nature of its material.

3067: 2190: 1437: 1066: 1053:. Their general proportions seem to resemble modern crocodiles. 6771: 5910: 5189: 4970: 4824: 4785: 4088:

Little has been published on the potential nesting behavior of

3933:). Based on the specific anatomy of the deltapectoral crest of 839:

had interlocking teeth similar to that of a crocodile (bottom).

3881:, but does exist. Specifically, Buchanan's thesis describes a 1119:. Willis and colleagues took this a step further by including 1021:

Additional known elements of the postcranial skeleton include

831:

had an overbite, much like modern alligators (top), whereas

3877:

Direct evidence for predator prey interaction is rare for

3866:

to attack large prey, before restraining and crushing it.

2388:

as an early diverging crocodyloid and immediate sister to

4681:

Rio, Jonathan P.; Mannion, Philip D. (6 September 2021).

4183:

environment. Fish are also known from this fauna, namely

21:

For an Indian poet in Kannada language by same name, see

4560:(PhD thesis). Melbourne: Monash University. p. 340. 455:

to be the oldest diverging. Other studies suggest that

4385:

Holt, T. R.; Salisbury, S. W.; Willis, P.M.A. (2005).

4277:

Willis, P. M. A.; Molnar, R.E.; Scanlon, J.D. (1993).

3838:. Buchanan argued that the confluent tooth sockets of 1010:, which makes the end look somewhat quadrilateral. In 395:

had interlocking dentition much more similar to true

4582:

Alcheringa: An Australasian Journal of Palaeontology

4424:

Alcheringa: An Australasian Journal of Palaeontology

2214: 2194: 2167: 2095: 2075: 2048: 2021: 1994: 1967: 1933: 1888: 1868: 1841: 1807: 1770: 1629: 1609: 1582: 1555: 1519: 1499: 1442: 1421: 1394: 1358: 1338: 1297: 1261: 1241: 1207: 303: 294: 285: 272: 6888: 6780: 6634: 6482: 6431: 6371: 6345: 6212: 6150: 6094: 6045: 6003: 5923: 5818: 5789: 5620: 5552: 5465: 5418: 5352: 5249: 5202: 5077: 4984: 4930: 4903: 4876: 4005:, which would allow for hinge-like movement of the 2338:. In a study dealing exclusively with morphology, 791:being described as long but narrow, while that of 4625:Michael S. Y. Lee; Adam M. Yates (27 June 2018). 923:and the more extensive interlocking dentition of 529:A third species was named in 2005 in the form of 2356:crocodyloid no longer believed to be related to 1045:, several dorsal and tail vertebrae as well as 560:and noted to differ greatly from modern forms. 435:As the oldest described and named mekosuchine, 845:The toothrow across all four known species of 4797: 4198:The Rundle Formation, which was home to both 1037:, hindlimbs as well as multiple parts of the 8: 4206:, on the other hand is thought to represent 4068:. One possible explanation may be found in 931:and more closely resembles animals such as 698:and have been found in areas overlying the 544:The most recent addition to this genus was 6768: 6492: 6381: 6353: 6220: 6100: 6011: 5931: 5920: 5907: 5626: 5471: 5462: 5261: 5257: 5210: 5199: 5186: 4981: 4967: 4909: 4882: 4873: 4821: 4804: 4790: 4782: 1167:to be the basalmost forms. In both cases, 982:Although a lot of postcranial material of 690:The type species of the genus, remains of 406:Despite plentiful fossils, the ecology of 118: 29: 4761: 4751: 4710: 4700: 4654: 4644: 4601: 4416: 4414: 4412: 4410: 4408: 4406: 4404: 3994:possessed a "fibular condyle" similar to 3817:. It broadly resembles modern crocodiles. 4272: 4270: 912:is characterised by being interlocking. 660:. It is the least well preserved of all 4676: 4674: 4328: 4326: 4324: 4322: 4320: 4318: 4316: 4268: 4266: 4264: 4262: 4260: 4258: 4256: 4254: 4252: 4250: 4246: 3873:A modern Alligator "juggling" a turtle. 3765: 2324: 4571: 4569: 4567: 4487: 4485: 4483: 4481: 4314: 4312: 4310: 4308: 4306: 4304: 4302: 4300: 4298: 4296: 3826:could serve a similar function as the 2918: 2397: 1739: 1176: 603:, scalebar = 10 cm (3.9 in). 4549: 4547: 4545: 4543: 4541: 4539: 4537: 4535: 4479: 4477: 4475: 4473: 4471: 4469: 4467: 4465: 4463: 4461: 4380: 4378: 4376: 4374: 4372: 4370: 4368: 4366: 4364: 3940:The range of motion for the limbs of 513:, was finally established in 1993 by 7: 4237:inhabit most of northern Australia. 1096:Similarities between a then unnamed 383:had an overbite similar to a modern 278:Willis, Molnar & Scanlon, 1993 ( 3620: 3596: 3589: 3565: 3543: 3492: 3442: 3435: 3411: 3299: 3275: 3251: 3183: 3159: 3135: 3127: 3059: 3035: 3011: 3004: 2979: 2972: 2950: 2928: 2920: 2816: 2809: 2786: 2761: 2641: 2617: 2593: 2543: 2536: 2468: 2444: 2420: 2413: 2406: 2399: 2182: 2155: 2063: 2036: 2009: 1982: 1955: 1948: 1921: 1856: 1829: 1822: 1795: 1785: 1758: 1748: 1741: 1597: 1570: 1543: 1491: 1484: 1409: 1382: 1326: 1319: 1312: 1285: 1229: 1222: 1195: 1185: 1178: 4632:Proceedings of the Royal Society B 4498:Journal of Vertebrate Paleontology 3905:was specialised for hunting them. 2346:in a basal position and including 668:and the interlocking dentition of 368:, also from the Rundle Formation. 300:Holt, Salisbury & Willis, 2005 259:Willis, Molnar & Scanlon, 1993 14: 904:. Regardless, much like those of 773:Ever since the first material of 579:In the type description the name 6996:Prehistoric pseudosuchian genera 6733: 6728: 5893: 5888: 4391:Memoirs of the Queensland Museum 818: 807: 144: 3771:Initially, it was thought that 2384:. Instead, these trees recover 1171:was found to be a mekosuchine. 471:are exclusively known from the 4233:may be similar to how today's 1041:, including nearly the entire 915:The third type is observed in 621:were found alongside those of 364:and the youngest of the four, 1: 4603:10.1080/03115518.2023.2201319 4060:In their 1996 description of 3809:The skull of an indetermined 3784:as the basal most species of 556:, which was covered by Stein 4444:10.1080/03115518.2012.671697 3977:. Here, Buchanan notes that 3518:Brachyuranochampsa eversolei 3168:Orientalosuchus naduongensis 291:Salisbury & Willis, 1996 2667:Mekosuchus whitehunterensis 2494:"Asiatosuchus" nanlingensis 2349:"Asiatosuchus" nanlingensis 2045:Mekosuchus whitehunterensis 1076:. The holotype specimen of 700:Oakdale Sandstone Formation 477:Oakdale Sandstone Formation 7012: 3885:from the same locality as 3468:"Crocodylus" depressifrons 3192:Jiangxisuchus nankangensis 1029:, bones of the lower arm, 20: 6746: 6726: 6495: 6384: 6356: 6223: 6103: 6014: 5934: 5919: 5906: 5886: 5629: 5474: 5264: 5260: 5213: 5198: 5185: 4980: 4966: 4912: 4885: 4872: 4820: 4355:10.1080/03115519608619189 3677: 3642: 3625: 3618: 3601: 3594: 3587: 3570: 3563: 3548: 3541: 3514: 3497: 3490: 3464: 3447: 3440: 3433: 3416: 3409: 3338: 3321: 3304: 3297: 3280: 3273: 3256: 3249: 3205: 3188: 3181: 3164: 3157: 3144:Krabisuchus siamogallicus 3140: 3133: 3125: 3081: 3064: 3057: 3040: 3033: 3020:Kalthifrons aurivellensis 3016: 3009: 3002: 2988:"Asiatosuchus" germanicus 2984: 2977: 2970: 2955: 2948: 2933: 2926: 2864: 2838: 2821: 2814: 2807: 2791: 2784: 2766: 2759: 2697: 2680: 2663: 2646: 2639: 2622: 2615: 2598: 2591: 2565: 2548: 2541: 2534: 2490: 2473: 2466: 2449: 2442: 2425: 2418: 2411: 2404: 2281: 2207: 2187: 2180: 2160: 2153: 2088: 2068: 2061: 2041: 2034: 2014: 2007: 1987: 1980: 1960: 1953: 1946: 1926: 1919: 1881: 1861: 1854: 1834: 1827: 1820: 1804:Kalthifrons aurivellensis 1800: 1793: 1783: 1763: 1756: 1746: 1714: 1622: 1602: 1595: 1575: 1568: 1548: 1541: 1512: 1496: 1489: 1482: 1434: 1414: 1407: 1387: 1380: 1351: 1331: 1324: 1317: 1310: 1290: 1283: 1254: 1234: 1227: 1220: 1200: 1193: 1183: 572:, which stems from older 269: 264: 141:Scientific classification 139: 126: 117: 32: 18:Extinct genus of reptiles 6063:"Crocodylus" gariepensis 4492:Buchanan, L.A. (2009). " 3969:performing the highwalk. 3909:Locomotion and lifestyle 720:The youngest species of 6991:Crocodiles of Australia 6336:"Tomistoma" lusitanicum 6109:"Crocodylus" megarhinus 5978:"Crocodylus" megarhinus 4554:Buchanan, L.A. (2008). 4285:(3): 27–33 – via 3325:Trilophosuchus rackhami 2842:"Crocodylus" megarhinus 2701:Mekosuchus inexpectatus 2626:Trilophosuchus rackhami 2092:Mekosuchus inexpectatus 1991:Trilophosuchus rackhami 1767:"Crocodylus" megarhinus 768:supratemporal fenestrae 356:, the poorly preserved 6986:Eocene crocodylomorphs 4646:10.1098/rspb.2018.1071 4113: 4057: 3970: 3874: 3818: 3574:Australosuchus clarkae 2770:Australosuchus clarkae 1930:Australosuchus clarkae 979: 799:Toothrow and dentition 780:retroarticular process 604: 336:that lived during the 23:Chandrashekhar Kambara 6953:Paleobiology Database 6873:Paleobiology Database 4235:freshwater crocodiles 4111: 4055: 3964: 3872: 3808: 3420:Prodiplocynodon langi 3308:Volia athollandersoni 2360:. Additionally, both 2018:Volia athollandersoni 1141:to be second only to 974: 648:The third species of 594: 479:near the township of 131:and the lower jaw of 6611:"Tomistoma" coppensi 6603:"Tomistoma" cairense 6329:"Tomistoma" cairense 5970:"Crocodylus" affinis 5137:"Crocodylus" affinis 4518:10.1671/039.029.0220 4179:, which inhabited a 3451:"Crocodylus" affinis 1149:alternatively found 1145:, whereas Ristevski 421:freshwater crocodile 352:from sediments near 6890:Kambara murgonensis 6618:"Tomistoma" dowsoni 4702:10.7717/peerj.12094 4594:2023Alch...47..370R 4510:2009JVPal..29..473B 4436:2012Alch...36..473S 4347:1996Alch...20..179S 4132:tumorous conditions 4119:comminuted fracture 4043:Kambara murgonensis 4039:Kambara implexidens 3773:Kambara murgonensis 3646:Kambara murgonensis 3605:Kambara implexidens 3260:Dongnanosuchus hsui 3209:Eoalligator chunyii 2650:Ultrastenos willisi 2477:Kambara murgonensis 2429:Kambara implexidens 1885:Kambara murgonensis 1838:Kambara implexidens 1498:Bullock Creek taxon 1294:Kambara murgonensis 1258:Kambara implexidens 1127:Kambara murgonensis 977:Kambara implexidens 975:Size comparison of 908:, the dentition of 795:was short and low. 601:Kambara murgonensis 597:Kambara implexidens 595:Comparison between 524:Kambara implexidens 350:Kambara implexidens 346:Kambara murgonensis 129:Kambara implexidens 6191:C. thorbjarnarsoni 5240:Listrognathosuchus 5116:Brachyuranochampsa 4753:10.7717/peerj.3501 4173:meiolaniid turtles 4114: 4058: 3971: 3967:American crocodile 3875: 3819: 2684:Mekosuchus sanderi 2072:Mekosuchus sanderi 980: 605: 463:History and naming 6968: 6967: 6940:Open Tree of Life 6860:Open Tree of Life 6774:Taxon identifiers 6765: 6764: 6742: 6741: 6724: 6723: 6720: 6719: 6716: 6715: 6712: 6711: 6478: 6477: 6208: 6207: 6204: 6203: 6200: 6199: 6163:C. anthropophagus 5902: 5901: 5884: 5883: 5880: 5879: 5876: 5875: 5872: 5871: 5868: 5867: 5616: 5615: 5414: 5413: 5377:Eurycephalosuchus 5181: 5180: 5177: 5176: 5130:"Crocodylus" acer 4962: 4961: 4958: 4957: 4954: 4953: 4950: 4949: 4169:softshell turtles 3769: 3768: 3762: 3761: 3753: 3752: 3744: 3743: 3735: 3734: 3726: 3725: 3717: 3716: 3708: 3707: 3699: 3698: 3690: 3689: 3666: 3665: 3657: 3656: 3529: 3528: 3501:"Crocodylus" acer 3479: 3478: 3398: 3397: 3389: 3388: 3380: 3379: 3371: 3370: 3362: 3361: 3353: 3352: 3238: 3237: 3229: 3228: 3220: 3219: 3131:Orientalosuchina 3114: 3113: 3105: 3104: 3096: 3095: 2915: 2914: 2906: 2905: 2897: 2896: 2888: 2887: 2879: 2878: 2853: 2852: 2748: 2747: 2739: 2738: 2730: 2729: 2721: 2720: 2712: 2711: 2580: 2579: 2523: 2522: 2514: 2513: 2505: 2504: 2328: 2327: 2321: 2320: 2312: 2311: 2303: 2302: 2294: 2293: 2270: 2269: 2261: 2260: 2252: 2251: 2243: 2242: 2234: 2233: 2225: 2224: 2142: 2141: 2133: 2132: 2124: 2123: 2115: 2114: 2106: 2105: 1908: 1907: 1899: 1898: 1736: 1735: 1727: 1726: 1703: 1702: 1694: 1693: 1685: 1684: 1676: 1675: 1667: 1666: 1658: 1657: 1649: 1648: 1640: 1639: 1530: 1529: 1471: 1470: 1462: 1461: 1453: 1452: 1369: 1368: 1272: 1271: 694:were part of the 652:to be described, 315: 314: 260: 7003: 6961: 6960: 6948: 6947: 6935: 6934: 6922: 6921: 6909: 6908: 6907: 6881: 6880: 6868: 6867: 6855: 6854: 6842: 6841: 6829: 6828: 6816: 6815: 6814: 6801: 6800: 6799: 6769: 6737: 6732: 6694:G. pachyrhynchus 6596:Siquisiquesuchus 6493: 6382: 6354: 6221: 6101: 6012: 5932: 5921: 5908: 5897: 5892: 5859:C. wannlangstoni 5852:C. venezuelensis 5694:Globidentosuchus 5657:Centenariosuchus 5627: 5488:Allognathosuchus 5472: 5463: 5354:Orientalosuchina 5262: 5258: 5211: 5200: 5187: 4982: 4968: 4910: 4883: 4874: 4867: 4866: 4822: 4806: 4799: 4792: 4783: 4776: 4775: 4765: 4755: 4731: 4725: 4724: 4714: 4704: 4678: 4669: 4668: 4658: 4648: 4622: 4616: 4615: 4605: 4573: 4562: 4561: 4551: 4530: 4529: 4489: 4456: 4455: 4418: 4399: 4398: 4382: 4359: 4358: 4330: 4291: 4290: 4274: 4177:madtsoiid snakes 4155:Paleoenvironment 3828:carnassial teeth 3815:Melbourne Museum 3801:Ecology and diet 3621: 3597: 3590: 3566: 3544: 3493: 3443: 3436: 3412: 3300: 3276: 3252: 3184: 3160: 3136: 3128: 3060: 3036: 3012: 3005: 2980: 2973: 2951: 2929: 2921: 2817: 2810: 2787: 2762: 2642: 2618: 2594: 2544: 2537: 2469: 2445: 2421: 2414: 2407: 2400: 2395: 2394: 2376:, but excluding 2374:Orientalosuchina 2216: 2196: 2183: 2169: 2156: 2097: 2077: 2064: 2050: 2037: 2023: 2010: 1996: 1983: 1969: 1956: 1949: 1935: 1922: 1890: 1870: 1857: 1843: 1830: 1823: 1809: 1796: 1786: 1772: 1759: 1749: 1742: 1631: 1611: 1598: 1584: 1571: 1557: 1544: 1521: 1501: 1492: 1485: 1444: 1423: 1410: 1396: 1383: 1360: 1340: 1327: 1320: 1313: 1299: 1286: 1263: 1243: 1230: 1223: 1209: 1196: 1186: 1179: 1174: 1173: 1047:lumbar vertebrae 822: 811: 696:Tingamarra Fauna 658:Rundle Formation 362:Rundle Formation 305: 296: 287: 274: 258: 251: 238: 214:Archosauriformes 201:Archosauromorpha 149: 148: 122: 112: 49: 38:Temporal range: 30: 7011: 7010: 7006: 7005: 7004: 7002: 7001: 7000: 6971: 6970: 6969: 6964: 6956: 6951: 6943: 6938: 6930: 6925: 6917: 6912: 6903: 6902: 6897: 6884: 6876: 6871: 6863: 6858: 6850: 6845: 6837: 6832: 6824: 6819: 6810: 6809: 6804: 6795: 6794: 6789: 6776: 6766: 6761: 6738: 6708: 6670:G. curvirostris 6647:G. bengawanicus 6630: 6625:Toyotamaphimeia 6545:Hesperogavialis 6486: 6474: 6427: 6375: 6367: 6350: 6341: 6293:Megadontosuchus 6217: 6196: 6184:C. palaeindicus 6146: 6090: 6041: 6020:Aldabrachampsus 6008: 5999: 5928: 5915: 5898: 5864: 5838:C. brevirostris 5814: 5785: 5779:Wannaganosuchus 5764:Stangerochampsa 5729:Orthogenysuchus 5672:Chinatichampsus 5612: 5548: 5542:Wannaganosuchus 5461: 5423: 5410: 5398:Orientalosuchus 5348: 5343:Stangerochampsa 5254: 5245: 5207: 5194: 5173: 5160:Prodiplocynodon 5144:Portugalosuchus 5073: 4976: 4946: 4926: 4899: 4868: 4827: 4826: 4816: 4810: 4780: 4779: 4733: 4732: 4728: 4680: 4679: 4672: 4624: 4623: 4619: 4575: 4574: 4565: 4553: 4552: 4533: 4494:Kambara taraina 4491: 4490: 4459: 4420: 4419: 4402: 4384: 4383: 4362: 4332: 4331: 4294: 4276: 4275: 4248: 4243: 4157: 4144:Kambara taraina 4106: 4086: 4035: 4011:Kambara taraina 3975:Kambara taraina 3911: 3887:Kambara taraina 3836:Kambara molnari 3813:species at the 3803: 3798: 3777:Kambara molnari 3763: 3754: 3745: 3736: 3727: 3718: 3709: 3700: 3691: 3667: 3658: 3629:Kambara taraina 3530: 3480: 3399: 3390: 3381: 3372: 3363: 3354: 3342:Mekosuchus spp. 3239: 3230: 3221: 3115: 3106: 3097: 2916: 2907: 2898: 2889: 2880: 2854: 2749: 2740: 2731: 2722: 2713: 2581: 2524: 2515: 2506: 2453:Kambara taraina 2322: 2313: 2304: 2295: 2271: 2262: 2253: 2244: 2235: 2226: 2143: 2134: 2125: 2116: 2107: 1909: 1900: 1865:Kambara taraina 1737: 1728: 1704: 1695: 1686: 1677: 1668: 1659: 1650: 1641: 1531: 1472: 1463: 1454: 1370: 1273: 1238:Kambara taraina 1094: 1059: 1043:cervical series 1023:shoulder blades 969: 843: 842: 841: 840: 825: 824: 823: 814: 813: 812: 801: 747: 732:, the teeth of 617:The fossils of 589: 465: 366:Kambara taraina 358:Kambara molnari 257: 249: 236: 143: 133:Kambara molnari 113: 111: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 44: 43: 36: 26: 19: 12: 11: 5: 7009: 7007: 6999: 6998: 6993: 6988: 6983: 6973: 6972: 6966: 6965: 6963: 6962: 6949: 6936: 6923: 6910: 6894: 6892: 6886: 6885: 6883: 6882: 6869: 6856: 6843: 6830: 6817: 6802: 6786: 6784: 6778: 6777: 6772: 6763: 6762: 6760: 6759: 6754: 6747: 6744: 6743: 6740: 6739: 6727: 6725: 6722: 6721: 6718: 6717: 6714: 6713: 6710: 6709: 6707: 6706: 6698: 6690: 6682: 6674: 6666: 6658: 6650: 6642: 6640: 6632: 6631: 6629: 6628: 6621: 6614: 6607: 6599: 6592: 6585: 6578: 6571: 6563: 6559:Maomingosuchus 6555: 6548: 6541: 6537:Harpacochampsa 6533: 6526: 6519: 6511: 6504: 6496: 6490: 6480: 6479: 6476: 6475: 6473: 6472: 6464: 6460:T. lusitanicum 6456: 6448: 6444:T. calaritanum 6439: 6437: 6429: 6428: 6426: 6425: 6417: 6409: 6402: 6394: 6385: 6379: 6369: 6368: 6366: 6365: 6357: 6351: 6346: 6343: 6342: 6340: 6339: 6332: 6325: 6317: 6310: 6303: 6296: 6289: 6282: 6279:Maomingosuchus 6275: 6271:Leptorrhamphus 6267: 6260: 6253: 6246: 6239: 6232: 6229:Dollosuchoides 6224: 6218: 6213: 6210: 6209: 6206: 6205: 6202: 6201: 6198: 6197: 6195: 6194: 6187: 6180: 6177:C. falconensis 6173: 6166: 6158: 6156: 6148: 6147: 6145: 6144: 6137: 6129: 6121: 6113: 6104: 6098: 6092: 6091: 6089: 6088: 6080: 6073: 6066: 6059: 6051: 6049: 6043: 6042: 6040: 6039: 6031: 6027:Dzungarisuchus 6023: 6015: 6009: 6004: 6001: 6000: 5998: 5997: 5989: 5981: 5974: 5966: 5962:Australosuchus 5958: 5951: 5948:Antecrocodylus 5944: 5935: 5929: 5924: 5917: 5916: 5911: 5904: 5903: 5900: 5899: 5887: 5885: 5882: 5881: 5878: 5877: 5874: 5873: 5870: 5869: 5866: 5865: 5863: 5862: 5855: 5848: 5841: 5834: 5826: 5824: 5816: 5815: 5813: 5812: 5809:M. latrubessei 5805: 5797: 5795: 5787: 5786: 5784: 5783: 5775: 5768: 5760: 5753: 5746: 5739: 5732: 5725: 5718: 5711: 5704: 5697: 5690: 5682: 5675: 5668: 5660: 5653: 5645: 5638: 5630: 5624: 5618: 5617: 5614: 5613: 5611: 5610: 5603: 5596: 5589: 5582: 5575: 5568: 5560: 5558: 5550: 5549: 5547: 5546: 5538: 5534:Procaimanoidea 5530: 5522: 5518:Hassiacosuchus 5514: 5507: 5499: 5491: 5484: 5475: 5469: 5460: 5459: 5451: 5447:Menatalligator 5443: 5435: 5426: 5424: 5419: 5416: 5415: 5412: 5411: 5409: 5408: 5405:Protoalligator 5401: 5394: 5387: 5380: 5373: 5366: 5363:Dongnanosuchus 5358: 5356: 5350: 5349: 5347: 5346: 5339: 5335:Procaimanoidea 5331: 5324: 5316: 5313:Hassiacosuchus 5309: 5302: 5294: 5287: 5280: 5273: 5270:Albertochampsa 5265: 5255: 5250: 5247: 5246: 5244: 5243: 5236: 5229: 5222: 5214: 5208: 5205:Alligatoroidea 5203: 5196: 5195: 5192:Alligatoroidea 5190: 5183: 5182: 5179: 5178: 5175: 5174: 5172: 5171: 5167:Planocraniidae 5163: 5156: 5152:Pristichampsus 5148: 5140: 5133: 5126: 5123:Charactosuchus 5119: 5112: 5104: 5097: 5089: 5081: 5079: 5075: 5074: 5072: 5071: 5064: 5057: 5054:Trilophosuchus 5050: 5042: 5035: 5028: 5021: 5014: 5010:Harpacochampsa 5006: 4999: 4995:Australosuchus 4990: 4988: 4978: 4977: 4971: 4964: 4963: 4960: 4959: 4956: 4955: 4952: 4951: 4948: 4947: 4945: 4944: 4943: 4942: 4936: 4934: 4928: 4927: 4925: 4924: 4923: 4922: 4913: 4907: 4901: 4900: 4898: 4897: 4896: 4895: 4886: 4880: 4870: 4869: 4865: 4864: 4855: 4846: 4840: 4834: 4825: 4818: 4817: 4811: 4809: 4808: 4801: 4794: 4786: 4778: 4777: 4726: 4670: 4617: 4588:(4): 370–415. 4563: 4531: 4504:(2): 473–486. 4457: 4430:(4): 473–486. 4400: 4360: 4341:(3): 179–226. 4292: 4245: 4244: 4242: 4239: 4165:K. implexidens 4161:K. murgonensis 4156: 4153: 4105: 4102: 4085: 4082: 4062:K. implexidens 4047:K. implexidens 4034: 4031: 3931:K. implexidens 3924:Trilophosuchus 3910: 3907: 3802: 3799: 3797: 3794: 3782:K. implexidens 3767: 3766: 3764: 3760: 3759: 3756: 3755: 3751: 3750: 3747: 3746: 3742: 3741: 3738: 3737: 3733: 3732: 3729: 3728: 3724: 3723: 3720: 3719: 3715: 3714: 3711: 3710: 3706: 3705: 3702: 3701: 3697: 3696: 3693: 3692: 3688: 3687: 3684: 3683: 3676: 3673: 3672: 3669: 3668: 3664: 3663: 3660: 3659: 3655: 3654: 3651: 3650: 3641: 3638: 3637: 3634: 3633: 3624: 3619: 3617: 3614: 3613: 3610: 3609: 3600: 3595: 3593: 3588: 3586: 3583: 3582: 3579: 3578: 3569: 3564: 3562: 3559: 3558: 3555: 3554: 3547: 3542: 3540: 3536: 3535: 3532: 3531: 3527: 3526: 3523: 3522: 3513: 3510: 3509: 3506: 3505: 3496: 3491: 3489: 3486: 3485: 3482: 3481: 3477: 3476: 3473: 3472: 3463: 3460: 3459: 3456: 3455: 3446: 3441: 3439: 3434: 3432: 3429: 3428: 3425: 3424: 3415: 3410: 3408: 3405: 3404: 3401: 3400: 3396: 3395: 3392: 3391: 3387: 3386: 3383: 3382: 3378: 3377: 3374: 3373: 3369: 3368: 3365: 3364: 3360: 3359: 3356: 3355: 3351: 3350: 3347: 3346: 3337: 3334: 3333: 3330: 3329: 3320: 3317: 3316: 3313: 3312: 3303: 3298: 3296: 3293: 3292: 3289: 3288: 3279: 3274: 3272: 3269: 3268: 3265: 3264: 3255: 3250: 3248: 3245: 3244: 3241: 3240: 3236: 3235: 3232: 3231: 3227: 3226: 3223: 3222: 3218: 3217: 3214: 3213: 3204: 3201: 3200: 3197: 3196: 3187: 3182: 3180: 3177: 3176: 3173: 3172: 3163: 3158: 3156: 3153: 3152: 3149: 3148: 3139: 3134: 3132: 3126: 3124: 3121: 3120: 3117: 3116: 3112: 3111: 3108: 3107: 3103: 3102: 3099: 3098: 3094: 3093: 3090: 3089: 3085:Paludirex spp. 3080: 3077: 3076: 3073: 3072: 3063: 3058: 3056: 3053: 3052: 3049: 3048: 3039: 3034: 3032: 3029: 3028: 3025: 3024: 3015: 3010: 3008: 3003: 3001: 2997: 2996: 2993: 2992: 2983: 2978: 2976: 2971: 2969: 2966: 2965: 2962: 2961: 2958:Planocraniidae 2954: 2949: 2947: 2944: 2943: 2940: 2939: 2936:Alligatoroidea 2932: 2927: 2925: 2919: 2917: 2913: 2912: 2909: 2908: 2904: 2903: 2900: 2899: 2895: 2894: 2891: 2890: 2886: 2885: 2882: 2881: 2877: 2876: 2873: 2872: 2863: 2860: 2859: 2856: 2855: 2851: 2850: 2847: 2846: 2837: 2834: 2833: 2830: 2829: 2820: 2815: 2813: 2808: 2806: 2802: 2801: 2798: 2797: 2790: 2785: 2783: 2779: 2778: 2775: 2774: 2765: 2760: 2758: 2755: 2754: 2751: 2750: 2746: 2745: 2742: 2741: 2737: 2736: 2733: 2732: 2728: 2727: 2724: 2723: 2719: 2718: 2715: 2714: 2710: 2709: 2706: 2705: 2696: 2693: 2692: 2689: 2688: 2679: 2676: 2675: 2672: 2671: 2662: 2659: 2658: 2655: 2654: 2645: 2640: 2638: 2635: 2634: 2631: 2630: 2621: 2616: 2614: 2611: 2610: 2607: 2606: 2597: 2592: 2590: 2587: 2586: 2583: 2582: 2578: 2577: 2574: 2573: 2564: 2561: 2560: 2557: 2556: 2547: 2542: 2540: 2535: 2533: 2530: 2529: 2526: 2525: 2521: 2520: 2517: 2516: 2512: 2511: 2508: 2507: 2503: 2502: 2499: 2498: 2489: 2486: 2485: 2482: 2481: 2472: 2467: 2465: 2462: 2461: 2458: 2457: 2448: 2443: 2441: 2438: 2437: 2434: 2433: 2424: 2419: 2417: 2412: 2410: 2405: 2403: 2398: 2378:Australosuchus 2366:Australosuchus 2326: 2325: 2323: 2319: 2318: 2315: 2314: 2310: 2309: 2306: 2305: 2301: 2300: 2297: 2296: 2292: 2291: 2288: 2287: 2280: 2277: 2276: 2273: 2272: 2268: 2267: 2264: 2263: 2259: 2258: 2255: 2254: 2250: 2249: 2246: 2245: 2241: 2240: 2237: 2236: 2232: 2231: 2228: 2227: 2223: 2222: 2219: 2218: 2206: 2203: 2202: 2199: 2198: 2186: 2181: 2179: 2176: 2175: 2172: 2171: 2164:Paludirex spp. 2159: 2154: 2152: 2149: 2148: 2145: 2144: 2140: 2139: 2136: 2135: 2131: 2130: 2127: 2126: 2122: 2121: 2118: 2117: 2113: 2112: 2109: 2108: 2104: 2103: 2100: 2099: 2087: 2084: 2083: 2080: 2079: 2067: 2062: 2060: 2057: 2056: 2053: 2052: 2040: 2035: 2033: 2030: 2029: 2026: 2025: 2013: 2008: 2006: 2003: 2002: 1999: 1998: 1986: 1981: 1979: 1976: 1975: 1972: 1971: 1959: 1954: 1952: 1947: 1945: 1942: 1941: 1938: 1937: 1925: 1920: 1918: 1915: 1914: 1911: 1910: 1906: 1905: 1902: 1901: 1897: 1896: 1893: 1892: 1880: 1877: 1876: 1873: 1872: 1860: 1855: 1853: 1850: 1849: 1846: 1845: 1833: 1828: 1826: 1821: 1819: 1816: 1815: 1812: 1811: 1799: 1794: 1792: 1784: 1782: 1779: 1778: 1775: 1774: 1762: 1757: 1755: 1747: 1745: 1740: 1738: 1734: 1733: 1730: 1729: 1725: 1724: 1721: 1720: 1713: 1710: 1709: 1706: 1705: 1701: 1700: 1697: 1696: 1692: 1691: 1688: 1687: 1683: 1682: 1679: 1678: 1674: 1673: 1670: 1669: 1665: 1664: 1661: 1660: 1656: 1655: 1652: 1651: 1647: 1646: 1643: 1642: 1638: 1637: 1634: 1633: 1621: 1618: 1617: 1614: 1613: 1606:Trilophosuchus 1601: 1596: 1594: 1591: 1590: 1587: 1586: 1574: 1569: 1567: 1564: 1563: 1560: 1559: 1547: 1542: 1540: 1537: 1536: 1533: 1532: 1528: 1527: 1524: 1523: 1511: 1508: 1507: 1504: 1503: 1495: 1490: 1488: 1483: 1481: 1478: 1477: 1474: 1473: 1469: 1468: 1465: 1464: 1460: 1459: 1456: 1455: 1451: 1450: 1447: 1446: 1433: 1430: 1429: 1426: 1425: 1413: 1408: 1406: 1403: 1402: 1399: 1398: 1386: 1381: 1379: 1376: 1375: 1372: 1371: 1367: 1366: 1363: 1362: 1350: 1347: 1346: 1343: 1342: 1330: 1325: 1323: 1318: 1316: 1311: 1309: 1306: 1305: 1302: 1301: 1289: 1284: 1282: 1279: 1278: 1275: 1274: 1270: 1269: 1266: 1265: 1253: 1250: 1249: 1246: 1245: 1233: 1228: 1226: 1221: 1219: 1216: 1215: 1212: 1211: 1204:Australosuchus 1199: 1194: 1192: 1184: 1182: 1177: 1151:Australosuchus 1143:Australosuchus 1103:Australosuchus 1093: 1090: 1082:K. murgonensis 1078:K. implexidens 1058: 1055: 968: 965: 961:K. murgonensis 957:K. implexidens 925:K. implexidens 921:K. murgonensis 906:K. implexidens 902:K. implexidens 894:K. murgonensis 889:K. implexidens 885:K. implexidens 870:K. implexidens 868:However, both 859:K. murgonensis 833:K. implexidens 829:K. murgonensis 827: 826: 817: 816: 815: 806: 805: 804: 803: 802: 800: 797: 793:K. implexidens 760:K. implexidens 756:K. murgonensis 746: 743: 742: 741: 730:K. implexidens 717: 716: 708: 707: 704:K. implexidens 692:K. murgonensis 687: 686: 683:K. murgonensis 678: 677: 670:K. implexidens 666:K. murgonensis 645: 644: 636: 635: 632:K. implexidens 627:K. implexidens 623:K. murgonensis 619:K. implexidens 614: 613: 610:K. implexidens 588: 585: 539:K. implexidens 507:K. murgonensis 494:Australosuchus 464: 461: 452:Australosuchus 429:K. implexidens 425:K. murgonensis 389:K. implexidens 381:K. murgonensis 313: 312: 311: 310: 309:Buchanan, 2009 301: 292: 289:K. implexidens 283: 276:K. murgonensis 267: 266: 262: 261: 247: 243: 242: 234: 227: 226: 221: 217: 216: 211: 204: 203: 198: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 137: 136: 124: 123: 115: 114: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 50: 37: 17: 13: 10: 9: 6: 4: 3: 2: 7008: 6997: 6994: 6992: 6989: 6987: 6984: 6982: 6979: 6978: 6976: 6959: 6954: 6950: 6946: 6941: 6937: 6933: 6928: 6924: 6920: 6915: 6911: 6906: 6900: 6896: 6895: 6893: 6891: 6887: 6879: 6874: 6870: 6866: 6861: 6857: 6853: 6848: 6844: 6840: 6835: 6831: 6827: 6822: 6818: 6813: 6807: 6803: 6798: 6792: 6788: 6787: 6785: 6783: 6779: 6775: 6770: 6758: 6757:Gryposuchinae 6755: 6753: 6749: 6748: 6745: 6736: 6731: 6704: 6703: 6699: 6696: 6695: 6691: 6688: 6687: 6683: 6680: 6679: 6675: 6672: 6671: 6667: 6664: 6663: 6659: 6656: 6655: 6651: 6649: 6648: 6644: 6643: 6641: 6639: 6638: 6633: 6627: 6626: 6622: 6620: 6619: 6615: 6613: 6612: 6608: 6605: 6604: 6600: 6598: 6597: 6593: 6591: 6590: 6589:Rhamphosuchus 6586: 6584: 6583: 6582:Piscogavialis 6579: 6577: 6576: 6572: 6569: 6568: 6567:Paratomistoma 6564: 6561: 6560: 6556: 6554: 6553: 6552:Ikanogavialis 6549: 6547: 6546: 6542: 6539: 6538: 6534: 6532: 6531: 6527: 6525: 6524: 6520: 6517: 6516: 6515:Gavialosuchus 6512: 6510: 6509: 6505: 6503: 6502: 6501:Aktiogavialis 6498: 6497: 6494: 6491: 6489: 6485: 6481: 6470: 6469: 6468:T. taiwanicum 6465: 6462: 6461: 6457: 6454: 6453: 6449: 6446: 6445: 6441: 6440: 6438: 6436: 6435: 6430: 6423: 6422: 6418: 6415: 6414: 6413:Paratomistoma 6410: 6408: 6407: 6403: 6400: 6399: 6398:Gavialosuchus 6395: 6392: 6391: 6390:Brasilosuchus 6387: 6386: 6383: 6380: 6378: 6377:sensu stricto 6374: 6370: 6364: 6363: 6359: 6358: 6355: 6352: 6349: 6344: 6338: 6337: 6333: 6331: 6330: 6326: 6323: 6322: 6318: 6316: 6315: 6311: 6309: 6308: 6307:Paratomistoma 6304: 6302: 6301: 6297: 6295: 6294: 6290: 6288: 6287: 6286:Maroccosuchus 6283: 6281: 6280: 6276: 6273: 6272: 6268: 6266: 6265: 6261: 6259: 6258: 6257:Gunggamarandu 6254: 6252: 6251: 6250:Gavialosuchus 6247: 6245: 6244: 6243:Ferganosuchus 6240: 6238: 6237: 6233: 6231: 6230: 6226: 6225: 6222: 6219: 6216: 6211: 6193: 6192: 6188: 6186: 6185: 6181: 6179: 6178: 6174: 6172: 6171: 6167: 6165: 6164: 6160: 6159: 6157: 6155: 6154: 6149: 6143: 6142: 6138: 6135: 6134: 6133:Tzaganosuchus 6130: 6127: 6126: 6122: 6119: 6118: 6114: 6111: 6110: 6106: 6105: 6102: 6099: 6097: 6093: 6086: 6085: 6081: 6079: 6078: 6074: 6072: 6071: 6067: 6065: 6064: 6060: 6058: 6057: 6053: 6052: 6050: 6048: 6047:Osteolaeminae 6044: 6037: 6036: 6035:Oxysdonsaurus 6032: 6029: 6028: 6024: 6022: 6021: 6017: 6016: 6013: 6010: 6007: 6002: 5995: 5994: 5990: 5987: 5986: 5985:Jiangxisuchus 5982: 5980: 5979: 5975: 5972: 5971: 5967: 5964: 5963: 5959: 5957: 5956: 5955:Astorgosuchus 5952: 5950: 5949: 5945: 5942: 5941: 5940:Albertosuchus 5937: 5936: 5933: 5930: 5927: 5926:Crocodyloidea 5922: 5918: 5914: 5909: 5905: 5896: 5891: 5861: 5860: 5856: 5854: 5853: 5849: 5847: 5846: 5845:C. gasparinae 5842: 5840: 5839: 5835: 5833: 5832: 5828: 5827: 5825: 5823: 5822: 5817: 5811: 5810: 5806: 5804: 5803: 5799: 5798: 5796: 5794: 5793: 5788: 5781: 5780: 5776: 5774: 5773: 5769: 5766: 5765: 5761: 5759: 5758: 5754: 5752: 5751: 5747: 5745: 5744: 5740: 5738: 5737: 5733: 5731: 5730: 5726: 5724: 5723: 5719: 5717: 5716: 5712: 5710: 5709: 5708:Kuttanacaiman 5705: 5703: 5702: 5698: 5696: 5695: 5691: 5688: 5687: 5683: 5681: 5680: 5679:Culebrasuchus 5676: 5674: 5673: 5669: 5666: 5665: 5661: 5659: 5658: 5654: 5651: 5650: 5649:Brachychampsa 5646: 5644: 5643: 5639: 5637: 5636: 5632: 5631: 5628: 5625: 5623: 5619: 5609: 5608: 5604: 5602: 5601: 5600:A. prenasalis 5597: 5595: 5594: 5590: 5588: 5587: 5583: 5581: 5580: 5576: 5574: 5573: 5569: 5567: 5566: 5562: 5561: 5559: 5557: 5556: 5551: 5544: 5543: 5539: 5536: 5535: 5531: 5528: 5527: 5523: 5520: 5519: 5515: 5513: 5512: 5511:Chrysochampsa 5508: 5505: 5504: 5500: 5497: 5496: 5492: 5490: 5489: 5485: 5482: 5481: 5480:Akanthosuchus 5477: 5476: 5473: 5470: 5468: 5467:Alligatorinae 5464: 5457: 5456: 5455:Sajkanosuchus 5452: 5449: 5448: 5444: 5441: 5440: 5439:Lianghusuchus 5436: 5433: 5432: 5428: 5427: 5425: 5422: 5421:Alligatoridae 5417: 5407: 5406: 5402: 5400: 5399: 5395: 5393: 5392: 5388: 5386: 5385: 5384:Jiangxisuchus 5381: 5379: 5378: 5374: 5372: 5371: 5367: 5365: 5364: 5360: 5359: 5357: 5355: 5351: 5345: 5344: 5340: 5337: 5336: 5332: 5330: 5329: 5325: 5322: 5321: 5317: 5315: 5314: 5310: 5308: 5307: 5303: 5300: 5299: 5295: 5293: 5292: 5288: 5286: 5285: 5284:Brachychampsa 5281: 5279: 5278: 5274: 5272: 5271: 5267: 5266: 5263: 5259: 5256: 5253: 5248: 5242: 5241: 5237: 5235: 5234: 5230: 5228: 5227: 5223: 5221: 5220: 5216: 5215: 5212: 5209: 5206: 5201: 5197: 5193: 5188: 5184: 5169: 5168: 5164: 5162: 5161: 5157: 5154: 5153: 5149: 5146: 5145: 5141: 5139: 5138: 5134: 5132: 5131: 5127: 5125: 5124: 5120: 5118: 5117: 5113: 5110: 5109: 5108:Borealosuchus 5105: 5103: 5102: 5098: 5095: 5094: 5090: 5088: 5087: 5086:Albertosuchus 5083: 5082: 5080: 5076: 5070: 5069: 5065: 5063: 5062: 5058: 5056: 5055: 5051: 5048: 5047: 5043: 5041: 5040: 5036: 5034: 5033: 5029: 5027: 5026: 5022: 5020: 5019: 5015: 5012: 5011: 5007: 5005: 5004: 5000: 4997: 4996: 4992: 4991: 4989: 4987: 4983: 4979: 4974: 4969: 4965: 4940: 4939: 4938: 4937: 4935: 4933: 4929: 4921: 4917: 4916: 4915: 4914: 4911: 4908: 4906: 4902: 4894: 4890: 4889: 4888: 4887: 4884: 4881: 4879: 4875: 4871: 4863: 4859: 4856: 4854: 4850: 4847: 4845: 4841: 4839: 4835: 4833: 4829: 4828: 4823: 4819: 4815: 4807: 4802: 4800: 4795: 4793: 4788: 4787: 4784: 4773: 4769: 4764: 4759: 4754: 4749: 4745: 4741: 4737: 4730: 4727: 4722: 4718: 4713: 4708: 4703: 4698: 4694: 4690: 4689: 4684: 4677: 4675: 4671: 4666: 4662: 4657: 4652: 4647: 4642: 4638: 4634: 4633: 4628: 4621: 4618: 4613: 4609: 4604: 4599: 4595: 4591: 4587: 4583: 4579: 4572: 4570: 4568: 4564: 4559: 4558: 4550: 4548: 4546: 4544: 4542: 4540: 4538: 4536: 4532: 4527: 4523: 4519: 4515: 4511: 4507: 4503: 4499: 4495: 4488: 4486: 4484: 4482: 4480: 4478: 4476: 4474: 4472: 4470: 4468: 4466: 4464: 4462: 4458: 4453: 4449: 4445: 4441: 4437: 4433: 4429: 4425: 4417: 4415: 4413: 4411: 4409: 4407: 4405: 4401: 4396: 4392: 4388: 4381: 4379: 4377: 4375: 4373: 4371: 4369: 4367: 4365: 4361: 4356: 4352: 4348: 4344: 4340: 4336: 4329: 4327: 4325: 4323: 4321: 4319: 4317: 4315: 4313: 4311: 4309: 4307: 4305: 4303: 4301: 4299: 4297: 4293: 4288: 4284: 4280: 4273: 4271: 4269: 4267: 4265: 4263: 4261: 4259: 4257: 4255: 4253: 4251: 4247: 4240: 4238: 4236: 4232: 4227: 4223: 4218: 4216: 4212: 4209: 4205: 4201: 4196: 4194: 4190: 4186: 4182: 4178: 4174: 4170: 4166: 4162: 4154: 4152: 4150: 4145: 4140: 4135: 4133: 4129: 4125: 4124:osteomyelitis 4120: 4110: 4103: 4101: 4099: 4095: 4091: 4083: 4081: 4077: 4075: 4071: 4067: 4063: 4054: 4050: 4048: 4044: 4040: 4032: 4030: 4028: 4024: 4019: 4014: 4012: 4008: 4004: 4000: 3997: 3993: 3989: 3985: 3980: 3976: 3968: 3963: 3959: 3956: 3952: 3948: 3943: 3938: 3936: 3932: 3927: 3925: 3921: 3916: 3908: 3906: 3904: 3900: 3896: 3892: 3888: 3884: 3883:chelid turtle 3880: 3871: 3867: 3865: 3861: 3857: 3853: 3849: 3843: 3841: 3837: 3832: 3829: 3824: 3816: 3812: 3807: 3800: 3795: 3793: 3791: 3787: 3783: 3778: 3774: 3758: 3757: 3749: 3748: 3740: 3739: 3731: 3730: 3722: 3721: 3713: 3712: 3704: 3703: 3695: 3694: 3686: 3685: 3682: 3681: 3675: 3674: 3671: 3670: 3662: 3661: 3653: 3652: 3649: 3648: 3647: 3640: 3639: 3636: 3635: 3632: 3631: 3630: 3623: 3622: 3616: 3615: 3612: 3611: 3608: 3607: 3606: 3599: 3598: 3592: 3591: 3585: 3584: 3581: 3580: 3577: 3576: 3575: 3568: 3567: 3561: 3560: 3557: 3556: 3553: 3552: 3546: 3545: 3539:Longirostres 3538: 3537: 3534: 3533: 3525: 3524: 3521: 3520: 3519: 3512: 3511: 3508: 3507: 3504: 3503: 3502: 3495: 3494: 3488: 3487: 3484: 3483: 3475: 3474: 3471: 3470: 3469: 3462: 3461: 3458: 3457: 3454: 3453: 3452: 3445: 3444: 3438: 3437: 3431: 3430: 3427: 3426: 3423: 3422: 3421: 3414: 3413: 3407: 3406: 3403: 3402: 3394: 3393: 3385: 3384: 3376: 3375: 3367: 3366: 3358: 3357: 3349: 3348: 3345: 3344: 3343: 3336: 3335: 3332: 3331: 3328: 3327: 3326: 3319: 3318: 3315: 3314: 3311: 3310: 3309: 3302: 3301: 3295: 3294: 3291: 3290: 3287: 3286: 3285: 3284:"Baru" huberi 3278: 3277: 3271: 3270: 3267: 3266: 3263: 3262: 3261: 3254: 3253: 3247: 3246: 3243: 3242: 3234: 3233: 3225: 3224: 3216: 3215: 3212: 3211: 3210: 3203: 3202: 3199: 3198: 3195: 3194: 3193: 3186: 3185: 3179: 3178: 3175: 3174: 3171: 3170: 3169: 3162: 3161: 3155: 3154: 3151: 3150: 3147: 3146: 3145: 3138: 3137: 3130: 3129: 3123: 3122: 3119: 3118: 3110: 3109: 3101: 3100: 3092: 3091: 3088: 3087: 3086: 3079: 3078: 3075: 3074: 3071: 3070: 3069: 3062: 3061: 3055: 3054: 3051: 3050: 3047: 3046: 3045: 3044:Quinkana spp. 3038: 3037: 3031: 3030: 3027: 3026: 3023: 3022: 3021: 3014: 3013: 3007: 3006: 3000:Mekosuchinae 2999: 2998: 2995: 2994: 2991: 2990: 2989: 2982: 2981: 2975: 2974: 2968: 2967: 2964: 2963: 2960: 2959: 2953: 2952: 2946: 2945: 2942: 2941: 2938: 2937: 2931: 2930: 2923: 2922: 2911: 2910: 2902: 2901: 2893: 2892: 2884: 2883: 2875: 2874: 2871: 2870: 2869: 2862: 2861: 2858: 2857: 2849: 2848: 2845: 2844: 2843: 2836: 2835: 2832: 2831: 2828: 2827: 2826: 2819: 2818: 2812: 2811: 2805:Crocodylinae 2804: 2803: 2800: 2799: 2796: 2795: 2794:Osteolaeminae 2789: 2788: 2782:Crocodylidae 2781: 2780: 2777: 2776: 2773: 2772: 2771: 2764: 2763: 2757: 2756: 2753: 2752: 2744: 2743: 2735: 2734: 2726: 2725: 2717: 2716: 2708: 2707: 2704: 2703: 2702: 2695: 2694: 2691: 2690: 2687: 2686: 2685: 2678: 2677: 2674: 2673: 2670: 2669: 2668: 2661: 2660: 2657: 2656: 2653: 2652: 2651: 2644: 2643: 2637: 2636: 2633: 2632: 2629: 2628: 2627: 2620: 2619: 2613: 2612: 2609: 2608: 2605: 2604: 2603: 2602:"Baru" huberi 2596: 2595: 2589: 2588: 2585: 2584: 2576: 2575: 2572: 2571: 2570: 2563: 2562: 2559: 2558: 2555: 2554: 2553: 2546: 2545: 2539: 2538: 2532: 2531: 2528: 2527: 2519: 2518: 2510: 2509: 2501: 2500: 2497: 2496: 2495: 2488: 2487: 2484: 2483: 2480: 2479: 2478: 2471: 2470: 2464: 2463: 2460: 2459: 2456: 2455: 2454: 2447: 2446: 2440: 2439: 2436: 2435: 2432: 2431: 2430: 2423: 2422: 2416: 2415: 2409: 2408: 2402: 2401: 2396: 2393: 2391: 2387: 2383: 2379: 2375: 2371: 2367: 2363: 2359: 2355: 2351: 2350: 2345: 2341: 2337: 2333: 2317: 2316: 2308: 2307: 2299: 2298: 2290: 2289: 2286: 2285: 2279: 2278: 2275: 2274: 2266: 2265: 2257: 2256: 2248: 2247: 2239: 2238: 2230: 2229: 2221: 2220: 2217: 2213: 2212: 2211:Quinkana spp. 2205: 2204: 2201: 2200: 2197: 2193: 2192: 2185: 2184: 2178: 2177: 2174: 2173: 2170: 2166: 2165: 2158: 2157: 2151: 2150: 2147: 2146: 2138: 2137: 2129: 2128: 2120: 2119: 2111: 2110: 2102: 2101: 2098: 2094: 2093: 2086: 2085: 2082: 2081: 2078: 2074: 2073: 2066: 2065: 2059: 2058: 2055: 2054: 2051: 2047: 2046: 2039: 2038: 2032: 2031: 2028: 2027: 2024: 2020: 2019: 2012: 2011: 2005: 2004: 2001: 2000: 1997: 1993: 1992: 1985: 1984: 1978: 1977: 1974: 1973: 1970: 1966: 1965: 1964:"Baru" huberi 1958: 1957: 1951: 1950: 1944: 1943: 1940: 1939: 1936: 1932: 1931: 1924: 1923: 1917: 1916: 1913: 1912: 1904: 1903: 1895: 1894: 1891: 1887: 1886: 1879: 1878: 1875: 1874: 1871: 1867: 1866: 1859: 1858: 1852: 1851: 1848: 1847: 1844: 1840: 1839: 1832: 1831: 1825: 1824: 1818: 1817: 1814: 1813: 1810: 1806: 1805: 1798: 1797: 1791: 1788: 1787: 1781: 1780: 1777: 1776: 1773: 1769: 1768: 1761: 1760: 1754: 1753:Crocodyloidea 1751: 1750: 1744: 1743: 1732: 1731: 1723: 1722: 1719: 1718: 1712: 1711: 1708: 1707: 1699: 1698: 1690: 1689: 1681: 1680: 1672: 1671: 1663: 1662: 1654: 1653: 1645: 1644: 1636: 1635: 1632: 1628: 1627: 1620: 1619: 1616: 1615: 1612: 1608: 1607: 1600: 1599: 1593: 1592: 1589: 1588: 1585: 1581: 1580: 1573: 1572: 1566: 1565: 1562: 1561: 1558: 1554: 1553: 1546: 1545: 1539: 1538: 1535: 1534: 1526: 1525: 1522: 1518: 1517: 1516:"Baru" huberi 1510: 1509: 1506: 1505: 1502: 1494: 1493: 1487: 1486: 1480: 1479: 1476: 1475: 1467: 1466: 1458: 1457: 1449: 1448: 1445: 1440: 1439: 1432: 1431: 1428: 1427: 1424: 1420: 1419: 1412: 1411: 1405: 1404: 1401: 1400: 1397: 1393: 1392: 1385: 1384: 1378: 1377: 1374: 1373: 1365: 1364: 1361: 1357: 1356: 1355:Pallimnarchus 1349: 1348: 1345: 1344: 1341: 1337: 1336: 1329: 1328: 1322: 1321: 1315: 1314: 1308: 1307: 1304: 1303: 1300: 1296: 1295: 1288: 1287: 1281: 1280: 1277: 1276: 1268: 1267: 1264: 1260: 1259: 1252: 1251: 1248: 1247: 1244: 1240: 1239: 1232: 1231: 1225: 1224: 1218: 1217: 1214: 1213: 1210: 1206: 1205: 1198: 1197: 1191: 1188: 1187: 1181: 1180: 1175: 1172: 1170: 1166: 1162: 1158: 1157: 1152: 1148: 1144: 1140: 1136: 1132: 1128: 1124: 1123: 1118: 1117: 1112: 1108: 1107:Pallimnarchus 1104: 1099: 1091: 1089: 1087: 1083: 1079: 1075: 1074: 1069: 1068: 1063: 1056: 1054: 1052: 1048: 1044: 1040: 1036: 1032: 1028: 1024: 1019: 1017: 1013: 1009: 1004: 1002: 998: 993: 989: 985: 978: 973: 966: 964: 962: 958: 955:, while both 954: 950: 944: 942: 941: 936: 935: 934:Borealosuchus 930: 926: 922: 918: 913: 911: 907: 903: 899: 895: 890: 886: 882: 881: 875: 871: 866: 864: 860: 855: 853: 848: 838: 834: 830: 821: 810: 798: 796: 794: 790: 785: 781: 776: 771: 769: 765: 761: 757: 752: 749:The skull of 744: 739: 735: 731: 727: 723: 719: 718: 715: 714: 710: 709: 705: 701: 697: 693: 689: 688: 685: 684: 680: 679: 675: 671: 667: 663: 659: 655: 651: 647: 646: 643: 642: 638: 637: 633: 628: 624: 620: 616: 615: 612: 611: 607: 606: 602: 598: 593: 586: 584: 582: 577: 575: 574:stratigraphic 571: 567: 563: 559: 555: 551: 547: 542: 540: 536: 532: 527: 525: 520: 516: 512: 508: 504: 498: 496: 495: 490: 486: 482: 478: 474: 470: 462: 460: 458: 454: 453: 448: 447: 442: 438: 433: 430: 426: 422: 418: 414: 409: 404: 402: 398: 394: 390: 386: 382: 378: 373: 371: 367: 363: 359: 355: 351: 347: 343: 339: 335: 332: 328: 325: 321: 320: 308: 302: 299: 293: 290: 284: 281: 277: 271: 270: 268: 263: 256: 255: 248: 245: 244: 241: 235: 232: 229: 228: 225: 222: 219: 218: 215: 212: 209: 206: 205: 202: 199: 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 147: 142: 138: 134: 130: 127:The skull of 125: 121: 116: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 48: 45:54.6–29 41: 35: 31: 28: 24: 16: 6981:Mekosuchinae 6889: 6781: 6752:Brevirostres 6702:G. papuensis 6700: 6692: 6684: 6676: 6668: 6660: 6654:G. breviceps 6652: 6645: 6635: 6623: 6616: 6609: 6601: 6594: 6587: 6580: 6575:Penghusuchus 6573: 6565: 6557: 6550: 6543: 6535: 6528: 6521: 6513: 6508:Dadagavialis 6506: 6499: 6487: 6466: 6458: 6450: 6442: 6432: 6421:Thecachampsa 6419: 6411: 6406:Melitosaurus 6404: 6396: 6388: 6376: 6373:Tomistominae 6362:Sacacosuchus 6360: 6334: 6327: 6319: 6314:Thecachampsa 6312: 6305: 6298: 6291: 6284: 6277: 6269: 6262: 6255: 6248: 6241: 6234: 6227: 6189: 6182: 6175: 6170:C. checchiai 6168: 6161: 6151: 6139: 6131: 6123: 6115: 6107: 6096:Crocodylinae 6082: 6075: 6068: 6061: 6054: 6033: 6025: 6018: 6006:Crocodylidae 5993:Mekosuchinae 5991: 5983: 5976: 5968: 5960: 5953: 5946: 5938: 5913:Longirostres 5857: 5850: 5843: 5836: 5831:C. australis 5829: 5819: 5807: 5800: 5792:Melanosuchus 5790: 5777: 5770: 5762: 5755: 5748: 5743:Paranasuchus 5741: 5736:Paranacaiman 5734: 5727: 5720: 5713: 5706: 5699: 5692: 5684: 5677: 5670: 5664:Ceratosuchus 5662: 5655: 5647: 5640: 5633: 5605: 5598: 5591: 5584: 5577: 5570: 5565:A. hailensis 5563: 5553: 5540: 5532: 5526:Navajosuchus 5524: 5516: 5509: 5503:Ceratosuchus 5501: 5493: 5486: 5478: 5453: 5445: 5437: 5429: 5403: 5396: 5389: 5382: 5375: 5368: 5361: 5341: 5333: 5328:Navajosuchus 5326: 5320:Leidyosuchus 5318: 5311: 5304: 5296: 5291:Ceratosuchus 5289: 5282: 5275: 5268: 5238: 5233:Leidyosuchus 5231: 5226:Diplocynodon 5224: 5217: 5165: 5158: 5150: 5142: 5135: 5128: 5121: 5114: 5106: 5101:Asiatosuchus 5099: 5091: 5084: 5066: 5059: 5052: 5044: 5037: 5030: 5024: 5023: 5016: 5008: 5001: 4993: 4986:Mekosuchinae 4975:crocodilians 4893:Pseudosuchia 4878:Pseudosuchia 4857: 4853:Pseudosuchia 4848: 4814:crocodilians 4743: 4739: 4729: 4692: 4686: 4636: 4630: 4620: 4585: 4581: 4556: 4501: 4497: 4493: 4427: 4423: 4394: 4390: 4338: 4334: 4287:Academia.edu 4282: 4230: 4225: 4221: 4219: 4214: 4203: 4199: 4197: 4164: 4160: 4158: 4148: 4143: 4138: 4136: 4115: 4097: 4093: 4089: 4087: 4078: 4073: 4061: 4059: 4046: 4042: 4038: 4036: 4026: 4022: 4017: 4015: 4010: 3999:rauisuchians 3991: 3978: 3974: 3972: 3954: 3950: 3946: 3941: 3939: 3934: 3930: 3928: 3923: 3919: 3914: 3912: 3902: 3894: 3886: 3878: 3876: 3863: 3859: 3855: 3851: 3847: 3846:prey. Among 3844: 3839: 3835: 3833: 3822: 3820: 3810: 3796:Paleobiology 3789: 3785: 3781: 3776: 3772: 3770: 3680:Crocodylidae 3678: 3645: 3644: 3643: 3628: 3627: 3626: 3604: 3603: 3602: 3572: 3571: 3549: 3516: 3515: 3499: 3498: 3466: 3465: 3449: 3448: 3418: 3417: 3340: 3339: 3323: 3322: 3306: 3305: 3282: 3281: 3258: 3257: 3207: 3206: 3190: 3189: 3166: 3165: 3142: 3141: 3083: 3082: 3066: 3065: 3042: 3041: 3018: 3017: 2986: 2985: 2956: 2934: 2866: 2865: 2840: 2839: 2823: 2822: 2792: 2768: 2767: 2699: 2698: 2682: 2681: 2665: 2664: 2648: 2647: 2624: 2623: 2600: 2599: 2569:Baru wickeni 2567: 2566: 2552:Baru darrowi 2550: 2549: 2492: 2491: 2476: 2475: 2474: 2452: 2451: 2450: 2428: 2427: 2426: 2390:Crocodylidae 2385: 2381: 2377: 2369: 2365: 2361: 2358:Asiatosuchus 2357: 2347: 2343: 2339: 2335: 2331: 2329: 2284:Crocodylidae 2282: 2209: 2208: 2189: 2188: 2162: 2161: 2090: 2089: 2070: 2069: 2043: 2042: 2016: 2015: 1989: 1988: 1962: 1961: 1928: 1927: 1884: 1883: 1882: 1864: 1863: 1862: 1837: 1836: 1835: 1802: 1801: 1790:Mekosuchinae 1765: 1764: 1717:Longirostres 1715: 1624: 1623: 1604: 1603: 1577: 1576: 1550: 1549: 1514: 1513: 1497: 1436: 1435: 1418:Baru darrowi 1416: 1415: 1391:Baru wickeni 1389: 1388: 1353: 1352: 1333: 1332: 1293: 1292: 1291: 1257: 1256: 1255: 1237: 1236: 1235: 1202: 1201: 1190:Mekosuchinae 1168: 1164: 1160: 1154: 1150: 1146: 1142: 1138: 1130: 1126: 1120: 1114: 1110: 1106: 1102: 1097: 1095: 1085: 1081: 1077: 1071: 1065: 1061: 1060: 1035:pelvic bones 1020: 1015: 1011: 1005: 1000: 996: 991: 983: 981: 976: 960: 956: 952: 948: 945: 940:Diplocynodon 938: 932: 928: 924: 920: 916: 914: 909: 905: 901: 897: 893: 888: 884: 878: 873: 869: 867: 858: 856: 851: 846: 844: 836: 832: 828: 792: 788: 783: 774: 772: 763: 759: 755: 750: 748: 733: 729: 728:. Like with 725: 721: 712: 711: 703: 691: 682: 681: 673: 669: 665: 661: 653: 649: 640: 639: 631: 626: 622: 618: 609: 608: 600: 596: 580: 578: 569: 561: 557: 545: 543: 538: 534: 530: 528: 523: 518: 511:type species 506: 502: 499: 492: 489:Ralph Molnar 468: 466: 456: 450: 444: 441:Phylogenetic 436: 434: 428: 424: 412: 407: 405: 400: 392: 388: 380: 376: 374: 369: 365: 357: 349: 345: 318: 317: 316: 306: 297: 288: 275: 253: 252: 240:Mekosuchinae 230: 207: 194: 132: 128: 33: 27: 15: 6806:Wikispecies 6678:G. leptodus 6530:Hanyusuchus 6523:Gryposuchus 6452:T. gaudense 6321:Tienosuchus 6300:Ocepesuchus 6264:Kentisuchus 6236:Dollosuchus 6215:Gavialoidea 5757:Purussaurus 5750:Protocaiman 5722:Necrosuchus 5715:Mourasuchus 5701:Gnatusuchus 5642:Bottosaurus 5607:A. thomsoni 5586:A. munensis 5579:A. mefferdi 5495:Arambourgia 5431:Balanerodus 5391:Krabisuchus 5370:Eoalligator 5298:Deinosuchus 5277:Arambourgia 5219:Deinosuchus 5093:Arenysuchus 5061:Ultrastenos 5018:Kalthifrons 4104:Pathologies 3852:K. taraina 3788:, although 3551:Gavialoidea 2924:Crocodilia 1335:Kalthifrons 1161:Kalthifrons 1156:Kalthifrons 1039:spinal cord 1031:finger bone 745:Description 515:Paul Willis 467:Remains of 446:Kalthifrons 375:Species of 334:crocodylian 331:mekosuchine 6975:Categories 6750:See also: 6488:sensu lato 6484:Gavialinae 6348:Gavialidae 6153:Crocodylus 6077:Rimasuchus 6070:Euthecodon 6056:Brochuchus 5802:M. fisheri 5635:Acresuchus 5622:Caimaninae 5572:A. mcgrewi 5252:Globidonta 5032:Mekosuchus 4941:see below↓ 4932:Crocodilia 4844:Sauropsida 4695:: e12094. 4397:: 207–218. 4335:Alcheringa 4241:References 4215:K. taraina 4204:K. taraina 4200:K. molnari 4181:lacustrine 4074:K. taraina 4027:Crocodylus 3988:astragalus 3864:K. taraina 3856:K. taraina 3840:K. molnari 3790:K. molnari 2868:Crocodylus 1579:Mekosuchus 1122:Mekosuchus 1086:K. molnari 1051:PhD thesis 967:Postcrania 953:K. molnari 949:K. taraina 917:K. molnari 910:K. taraina 898:K. taraina 880:Crocodylus 874:K. taraina 837:K. taraina 789:K. molnari 784:K. taraina 734:K. taraina 726:K. molnari 713:K. taraina 674:K. taraina 654:K. molnari 641:K. molnari 570:K. molnari 562:K. taraina 546:K. taraina 535:K. molnari 531:K. molnari 485:Queensland 401:K. molnari 397:crocodiles 393:K. taraina 307:K. taraina 298:K. molnari 224:Crocodilia 6905:Q20718518 6686:G. lewisi 6662:G. browni 6434:Tomistoma 5772:Tsoabichi 5593:A. olseni 5555:Alligator 5039:Paludirex 4920:Neosuchia 4905:Neosuchia 4830:Kingdom: 4746:: e3501. 4612:258878554 4452:129195186 4211:mud flats 4193:billabong 4100:to nest. 4070:taphonomy 4016:Overall, 4003:aetosaurs 3850:species, 3068:Baru spp. 2354:Paleocene 2191:Baru spp. 1092:Phylogeny 1073:Paludirex 1027:coracoids 863:alligator 566:sympatric 473:Paleogene 432:drought. 417:saltwater 385:alligator 360:from the 342:Australia 340:epoch in 164:Kingdom: 158:Eukaryota 6919:42335027 6899:Wikidata 6826:10588969 6797:Q5230226 6791:Wikidata 6637:Gavialis 6125:Quinkana 5686:Eocaiman 5306:Eocaiman 5046:Quinkana 4862:Eusuchia 4838:Chordata 4836:Phylum: 4832:Animalia 4812:Extinct 4772:28674657 4721:34567843 4665:30051855 4639:(1881). 4526:86254159 4208:lagoonal 4189:teleosts 4185:lungfish 4033:Sympatry 3996:Triassic 3986:and the 3920:Quinkana 3899:necrosis 3891:plastron 2825:Quinkana 2362:Quinkana 2332:Kambara' 1626:Quinkana 1135:polytomy 1116:Quinkana 1008:condyles 738:Darumbal 576:layers. 265:Species 188:Reptilia 178:Chordata 174:Phylum: 168:Animalia 154:Domain: 6945:6149935 6932:8610649 6865:3605941 6852:1201546 6839:4822295 6812:Kambara 6782:Kambara 6117:Kinyang 6084:Kinyang 5025:Kambara 4842:Class: 4763:5494174 4712:8428266 4656:6030529 4590:Bibcode 4506:Bibcode 4432:Bibcode 4343:Bibcode 4231:Kambara 4226:Kambara 4222:Kambara 4149:Kambara 4139:Kambara 4098:Kambara 4094:Kambara 4090:Kambara 4084:Nesting 4066:caimans 4023:Kambara 4018:Kambara 3992:Kambara 3979:Kambara 3955:Kambara 3951:Kambara 3947:Kambara 3942:Kambara 3935:Kambara 3915:Kambara 3903:Kambara 3895:Kambara 3879:Kambara 3860:Kambara 3848:Kambara 3823:Kambara 3811:Kambara 3786:Kambara 2386:Kambara 2382:Kambara 2344:Kambara 2340:Kambara 2336:Kambara 1441:Alcoota 1169:Kambara 1165:Kambara 1139:Kambara 1131:Kambara 1098:Kambara 1062:Kambara 1016:Kambara 1012:Kambara 1001:Kambara 997:Kambara 992:Kambara 988:humerus 984:Kambara 929:Kambara 852:Kambara 847:Kambara 775:Kambara 764:Kambara 751:Kambara 722:Kambara 662:Kambara 650:Kambara 587:Species 581:Kambara 554:humerus 519:Kambara 509:as the 505:, with 503:Kambara 469:Kambara 457:Kambara 437:Kambara 413:Kambara 408:Kambara 387:, both 377:Kambara 370:Kambara 324:extinct 319:Kambara 254:Kambara 246:Genus: 220:Order: 184:Class: 34:Kambara 6958:265611 6878:157974 5821:Caiman 5078:Others 4770: 4760: 4719: 4709: 4663: 4653: 4610: 4524: 4450: 4283:Kaupia 4128:fibula 1147:et al. 558:et al. 550:thesis 481:Murgon 354:Murgon 338:Eocene 322:is an 40:Eocene 6847:IRMNG 5068:Volia 4973:Basal 4858:Clade 4849:Clade 4740:PeerJ 4688:PeerJ 4608:S2CID 4522:S2CID 4448:S2CID 4159:Both 3984:ilium 2370:et al 1552:Volia 896:. In 883:. In 568:with 411:that 327:genus 231:Clade 208:Clade 195:Clade 6927:GBIF 6834:GBIF 6141:Voay 5003:Baru 4918:see 4891:see 4768:PMID 4717:PMID 4661:PMID 4202:and 4187:and 4175:and 4163:and 4142:the 4007:knee 4001:and 3922:and 2380:and 2364:and 2352:, a 1438:Baru 1163:and 1113:and 1111:Baru 1070:and 1067:Baru 1057:Size 959:and 937:and 872:and 835:and 672:and 599:and 427:and 419:and 391:and 348:and 280:type 53:PreꞒ 6914:EoL 6821:EoL 4758:PMC 4748:doi 4707:PMC 4697:doi 4651:PMC 4641:doi 4637:285 4598:doi 4514:doi 4440:doi 4351:doi 3965:An 1159:or 877:of 865:. 857:In 449:or 329:of 6977:: 6955:: 6942:: 6929:: 6916:: 6901:: 6875:: 6862:: 6849:: 6836:: 6823:: 6808:: 6793:: 4860:: 4851:: 4766:. 4756:. 4742:. 4738:. 4715:. 4705:. 4691:. 4685:. 4673:^ 4659:. 4649:. 4635:. 4629:. 4606:. 4596:. 4586:47 4584:. 4580:. 4566:^ 4534:^ 4520:. 4512:. 4502:29 4500:. 4460:^ 4446:. 4438:. 4428:36 4426:. 4403:^ 4395:50 4393:. 4389:. 4363:^ 4349:. 4339:20 4337:. 4295:^ 4281:. 4249:^ 4171:, 4151:. 4134:. 4076:. 1153:, 1109:, 1105:, 1033:, 1025:, 1018:. 943:. 770:. 541:. 483:, 399:. 233:: 210:: 197:: 135:. 103:Pg 47:Ma 42:, 6705:? 6697:? 6689:? 6681:? 6673:? 6665:? 6657:? 6606:? 6570:? 6562:? 6540:? 6518:? 6471:? 6463:? 6455:? 6447:? 6424:? 6416:? 6401:? 6393:? 6324:? 6274:? 6136:? 6128:? 6120:? 6112:? 6087:? 6038:? 6030:? 5996:? 5988:? 5973:? 5965:? 5943:? 5782:? 5767:? 5689:? 5667:? 5652:? 5545:? 5537:? 5529:? 5521:? 5506:? 5498:? 5483:? 5458:? 5450:? 5442:? 5434:? 5338:? 5323:? 5301:? 5170:? 5155:? 5147:? 5111:? 5096:? 5049:? 5013:? 4998:? 4805:e 4798:t 4791:v 4774:. 4750:: 4744:5 4723:. 4699:: 4693:9 4667:. 4643:: 4614:. 4600:: 4592:: 4528:. 4516:: 4508:: 4454:. 4442:: 4434:: 4357:. 4353:: 4345:: 4289:. 2215:† 2195:† 2168:† 2096:† 2076:† 2049:† 2022:† 1995:† 1968:† 1934:† 1889:† 1869:† 1842:† 1808:† 1771:† 1630:† 1610:† 1583:† 1556:† 1520:† 1500:† 1443:† 1422:† 1395:† 1359:† 1339:† 1298:† 1262:† 1242:† 1208:† 706:. 304:† 295:† 286:† 282:) 273:† 250:† 237:† 108:N 98:K 93:J 88:T 83:P 78:C 73:D 68:S 63:O 58:Ꞓ 25:.

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