109:
235:
beneath lateral margins, usually a ventrianal shield with 1–5 pairs of setae in addition to circumanal setae, peritreme usually looped around stigma, and movable cheliceral digit usually with a well-developed arthrodial brush at the base. Adult males are similar but with a holoventral shield or
1338:
de
Azevedo, Letícia H.; Emberson, Rowan M.; Esteca, Fernanda de C. N.; de Moraes, Gilberto José (2015), Carrillo, Daniel; de Moraes, Gilberto José; Peña, Jorge E. (eds.),
1698:
1456:"Life history of Macrocheles muscaedomesticae (Parasitiformes: Macrochelidae): new insights on life history and evidence of facultative parasitism on Drosophila"
300:
eggs and larvae). Dung offers high prey availability and shelter from the weather, but it is a temporary resource. Melicharids therefore attach to insects (e.g.
1737:
1380:
288:
Mites of this family are predators. The majority of species are coprophilous, meaning they live in animal dung and feed on the prey available there (
1672:
1711:
1359:
231:, a sternal shield with three pairs of setae, a genital shield with one pair of setae and with a pair of conspicuous accessory
324:
Other macrochelids occupy habitats such as forest litter, decaying plant debris, beach wrack or the nests of various animals.
1716:
1724:
317:
1581:
1558:
1651:
1130:
1742:
1664:
1288:
1060:
1026:
944:
1120:
910:
568:
1806:
814:
624:
558:
333:
1511:
892:
848:
704:
1586:
1801:
1070:
308:. Adult females tend to be the ones that engage in phoresis, though males and nymphs can do it as well.
1778:
108:
1752:
1620:
882:
634:
249:
598:
578:
540:
245:
1549:
1491:
1436:
124:
1760:
1607:
1483:
1475:
1428:
1355:
588:
512:
1703:
227:
Adult females of
Macrochelidae have: an undivided dorsal shield bearing at least 28 pairs of
1467:
1420:
1347:
530:
1765:
1257:
1659:
1646:
1339:
1277:
373:
1455:
1795:
1563:
1495:
1299:
1217:
1157:
795:
645:
481:
413:
301:
59:
1440:
1612:
1408:
1344:
Prospects for
Biological Control of Plant Feeding Mites and Other Harmful Organisms
1237:
1177:
1015:
461:
393:
216:
183:
1729:
1685:
1543:
1351:
995:
987:
975:
955:
775:
755:
433:
289:
277:
34:
1534:
1471:
1424:
1340:"Macrochelid Mites (Mesostigmata: Macrochelidae) as Biological Control Agents"
1197:
501:
312:
79:
44:
1479:
1633:
353:
297:
261:
147:
84:
1487:
1432:
1594:
1573:
1528:
1409:"Reflections on the biology, morphology and ecology of the Macrochelidae"
305:
304:
beetles or flies) to be carried to fresh dung deposits; this is known as
293:
232:
159:
74:
69:
54:
49:
39:
1599:
264:, the production of female offspring from unfertilised eggs. Species of
332:
Because of their predatory nature, some macrochelids have potential as
89:
64:
1690:
135:
1677:
1505:
1638:
1773:
171:
119:. (The legs on the right side have been omitted in this drawing.)
1625:
228:
212:
1509:
336:
agents of pest insects, such as flies and thrips. The species
1346:, Cham: Springer International Publishing, pp. 103–132,
1454:
Durkin, Emily S.; Proctor, Heather; Luong, Lien T. (2019).
280:, the production of male offspring from unfertilised eggs.
348:
Macrochelidae contains the following genera and species:
315:
on their hosts, rather than phoretic. An example is
1518:
311:Some macrochelids are believed to be facultatively
236:separate sternigenital and ventrianal shields.
8:
340:is commercially available for this purpose.
1506:
107:
20:
1325:
7:
1779:3251473D-DE45-FFF7-FF6F-FAEEFC06056A
1753:a87b92af-78fa-449b-8a2c-932333390e8a
1665:9b6ce70b-3d1f-42aa-b98e-3f87a61e58c3
1413:Experimental & Applied Acarology
1402:
1400:
1333:
1331:
1329:
741:van-Driel, Loots & Marais, 1977
1460:Experimental and Applied Acarology
14:
853:Filipponi & Pegazzano, 1967
733:Filipponi & Pegazzano, 1960
650:Filipponi & Pegazzano, 1960
1:
1171:(G.& R. Canestrini, 1882)
835:(van Driel & Loots, 1975)
318:Macrocheles muscaedomesticae
1352:10.1007/978-3-319-15042-0_4
1313:van Driel & Loots, 1975
1304:van Driel & Loots, 1975
1047:Samsinak & Daniel, 1978
672:Glyptholaspis baichengensis
593:Valle & Mazzoleni, 1967
553:Valle & Mazzoleni, 1967
427:van Driel & Loots, 1975
418:van Driel & Loots, 1975
1823:
1472:10.1007/s10493-019-00431-y
1131:Neopodocinum vanderhammeni
1125:Hartini & Takaku, 2003
1065:Hartini & Takaku, 2003
928:Holostaspella similiornata
683:de-Jesus & Rueda, 1990
1289:Trigonholaspis trigonarum
1268:Tricholocelaeno longicoma
1248:Tricholaspis marginipilis
1168:Nothrholaspis tridentatus
1112:Neopodocinum spinirostris
1061:Neopodocinum halimunensis
1027:Neopodocinum caputmedusae
945:Holostaspella tuberilinea
492:Evholocelaeno bursiformis
384:Ancistrocheles bregetovae
125:Scientific classification
123:
115:
106:
23:
1121:Neopodocinum subjaspersi
936:Holostaspella tropicalis
911:Holostaspella scatophila
902:Holostaspella orientalis
866:Holostaspella halawanyii
832:Holostaspella congoensis
746:Glyptholaspis wuhouyongi
722:Glyptholaspis orientalis
696:Glyptholaspis filipponii
569:Geholaspis foroliviensis
1425:10.1023/A:1006097811592
1188:Proholaspina micrarhena
1148:Neopodocinum yunnanense
1140:Neopodocinum wainsteini
1036:Neopodocinum dehongense
1006:Neoholaspis coprophilus
840:Holostaspella crenulata
815:Holostaspella bifoliata
664:Glyptholaspis asperrima
656:Glyptholaspis americana
625:Geholaspis mandibularis
608:Geholaspis longispinosa
559:Geholaspis comelicensis
404:Andhrolaspis trinitatis
364:Aethosoma burchellestes
893:Holostaspella moderata
849:Holostaspella exornata
705:Glyptholaspis fimicola
680:Glyptholaspis cariasoi
452:Calholaspis taiwanicus
424:Bellatocheles variatus
338:Macrocheles robustulus
1407:Krantz, G.W. (1998).
1310:Venatiolaspis pilosus
1208:Synaphasis congoensis
1096:Neopodocinum petrovae
1071:Neopodocinum jaspersi
1052:Neopodocinum gigantum
966:Lordocheles desaegeri
920:Holostaspella sculpta
874:Holostaspella krantzi
824:Holostaspella caelata
806:Holostaspella ateucha
730:Glyptholaspis pontina
688:Glyptholaspis confusa
117:Macrocheles carinatus
1660:Fauna Europaea (new)
1104:Neopodocinum sinicum
1039:Li & Chang, 1979
883:Holostaspella macula
766:Gonatothrix carinata
738:Glyptholaspis thorri
714:Glyptholaspis indica
635:Geholaspis pauperior
444:Calholaspis superbus
1228:Tigonholaspis saiti
1044:Neopodocinum galfyi
599:Geholaspis lagrecai
579:Geholaspis hortorum
550:Geholaspis bianchii
541:Geholaspis berlesei
472:Cophrolaspis glabra
250:reproduce asexually
1088:Neopodocinum maius
1080:Neopodocinum magna
858:Holostaspella foai
769:G. W. Krantz, 1988
760:G. W. Krantz, 1988
616:Geholaspis longula
334:biological control
328:Biological control
248:and some can also
246:reproduce sexually
30:Quaternary–present
1789:
1788:
1761:Open Tree of Life
1512:Taxon identifiers
1361:978-3-319-15041-3
1314:
1305:
1294:
1283:
1272:
1263:
1252:
1243:
1232:
1223:
1212:
1203:
1192:
1183:
1172:
1163:
1152:
1151:Li & Gu, 1987
1144:
1136:
1126:
1116:
1108:
1107:Li & Gu, 1987
1100:
1092:
1084:
1076:
1066:
1056:
1055:Gu & Li, 1987
1048:
1040:
1032:
1021:
1010:
1001:
981:
970:
961:
950:
940:
932:
924:
916:
906:
898:
888:
878:
870:
862:
854:
844:
836:
828:
820:
810:
801:
790:
786:Holocelaeno mitis
781:
770:
761:
750:
742:
734:
726:
718:
710:
700:
692:
684:
676:
668:
660:
651:
640:
630:
620:
612:
604:
594:
589:Geholaspis ilvana
584:
574:
564:
554:
546:
536:
526:
522:Geholaspis alpina
518:
513:Geholaspis aeneus
507:
496:
487:
476:
467:
456:
448:
439:
428:
419:
408:
399:
388:
379:
368:
359:
206:
205:
1814:
1782:
1781:
1769:
1768:
1756:
1755:
1746:
1745:
1733:
1732:
1730:NHMSYS0021127280
1720:
1719:
1707:
1706:
1694:
1693:
1681:
1680:
1668:
1667:
1655:
1654:
1642:
1641:
1629:
1628:
1616:
1615:
1603:
1602:
1590:
1589:
1577:
1576:
1567:
1566:
1554:
1553:
1552:
1539:
1538:
1537:
1507:
1500:
1499:
1466:(3–4): 309–321.
1451:
1445:
1444:
1404:
1395:
1394:
1392:
1391:
1377:
1371:
1370:
1369:
1368:
1335:
1312:
1303:
1293:(Vitzthum, 1930)
1292:
1281:
1270:
1261:
1250:
1241:
1230:
1221:
1210:
1201:
1190:
1181:
1170:
1161:
1150:
1142:
1134:
1124:
1114:
1106:
1098:
1090:
1082:
1074:
1064:
1054:
1046:
1038:
1030:
1019:
1008:
999:
979:
968:
959:
948:
938:
930:
922:
914:
904:
896:
886:
876:
868:
860:
852:
842:
834:
826:
819:(Trägårdh, 1952)
818:
808:
799:
788:
779:
768:
759:
748:
740:
732:
724:
716:
709:(Sellnick, 1931)
708:
698:
690:
682:
674:
666:
658:
649:
638:
628:
618:
610:
602:
592:
582:
573:Lombardini, 1943
572:
563:Lombardini, 1962
562:
552:
544:
534:
531:Geholaspis asper
524:
516:
505:
494:
485:
474:
465:
454:
446:
437:
426:
417:
406:
397:
386:
377:
366:
357:
111:
101:
31:
27:Temporal range:
21:
1822:
1821:
1817:
1816:
1815:
1813:
1812:
1811:
1792:
1791:
1790:
1785:
1777:
1772:
1764:
1759:
1751:
1749:
1741:
1736:
1728:
1723:
1715:
1710:
1702:
1697:
1689:
1684:
1676:
1671:
1663:
1658:
1650:
1645:
1637:
1632:
1624:
1619:
1611:
1606:
1598:
1593:
1585:
1580:
1572:
1570:
1562:
1557:
1548:
1547:
1542:
1533:
1532:
1527:
1514:
1504:
1503:
1453:
1452:
1448:
1406:
1405:
1398:
1389:
1387:
1381:"Macrochelidae"
1379:
1378:
1374:
1366:
1364:
1362:
1337:
1336:
1327:
1322:
1317:
1271:(Berlese, 1910)
1258:Tricholocelaeno
1191:(Berlese, 1916)
1143:Arutunian, 1993
1115:(Berlese, 1910)
1031:(Berlese, 1908)
980:Latreille, 1829
725:Iavorschi, 1980
667:(Berlese, 1905)
659:(Berlese, 1888)
639:(Berlese, 1918)
629:(Berlese, 1904)
619:(Berlese, 1882)
583:(Berlese, 1904)
525:(Berlese, 1887)
495:(Berlese, 1910)
346:
330:
286:
242:
225:
211:is a family of
202:
199:
198:
186:
174:
162:
150:
138:
102:
100:
99:
98:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
16:Family of mites
12:
11:
5:
1820:
1818:
1810:
1809:
1807:Acari families
1804:
1794:
1793:
1787:
1786:
1784:
1783:
1770:
1757:
1747:
1734:
1721:
1708:
1695:
1682:
1669:
1656:
1647:Fauna Europaea
1643:
1630:
1617:
1604:
1591:
1578:
1568:
1555:
1540:
1524:
1522:
1516:
1515:
1510:
1502:
1501:
1446:
1419:(3): 125–137.
1396:
1372:
1360:
1324:
1323:
1321:
1318:
1316:
1315:
1296:
1295:
1282:Vitzthum, 1930
1278:Trigonholaspis
1274:
1273:
1254:
1253:
1234:
1233:
1231:Vitzthum, 1930
1222:Vitzthum, 1930
1214:
1213:
1194:
1193:
1174:
1173:
1154:
1153:
1145:
1137:
1127:
1117:
1109:
1101:
1099:Davydova, 1979
1093:
1085:
1077:
1075:Oudemans, 1902
1067:
1057:
1049:
1041:
1033:
1020:Oudemans, 1902
1012:
1011:
992:
991:
972:
971:
952:
951:
941:
933:
925:
917:
907:
899:
889:
879:
871:
863:
855:
845:
837:
829:
821:
811:
809:Halliday, 1988
792:
791:
772:
771:
752:
751:
743:
735:
727:
719:
711:
701:
693:
685:
677:
669:
661:
642:
641:
631:
621:
613:
611:(Kramer, 1876)
605:
595:
585:
575:
565:
555:
547:
537:
527:
519:
498:
497:
478:
477:
475:(Müller, 1859)
458:
457:
449:
430:
429:
410:
409:
390:
389:
374:Ancistrocheles
370:
369:
350:
345:
342:
329:
326:
285:
282:
244:Macrochelidae
241:
238:
224:
221:
204:
203:
201:Vitzthum, 1930
200:
194:
192:
188:
187:
182:
180:
176:
175:
170:
168:
164:
163:
158:
156:
152:
151:
146:
144:
140:
139:
134:
132:
128:
127:
121:
120:
113:
112:
104:
103:
95:
94:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
1819:
1808:
1805:
1803:
1800:
1799:
1797:
1780:
1775:
1771:
1767:
1762:
1758:
1754:
1748:
1744:
1739:
1735:
1731:
1726:
1722:
1718:
1713:
1709:
1705:
1700:
1696:
1692:
1687:
1683:
1679:
1674:
1670:
1666:
1661:
1657:
1653:
1648:
1644:
1640:
1635:
1631:
1627:
1622:
1618:
1614:
1609:
1605:
1601:
1596:
1592:
1588:
1583:
1579:
1575:
1569:
1565:
1564:Macrochelidae
1560:
1556:
1551:
1550:Macrochelidae
1545:
1541:
1536:
1530:
1526:
1525:
1523:
1521:
1520:Macrochelidae
1517:
1513:
1508:
1497:
1493:
1489:
1485:
1481:
1477:
1473:
1469:
1465:
1461:
1457:
1450:
1447:
1442:
1438:
1434:
1430:
1426:
1422:
1418:
1414:
1410:
1403:
1401:
1397:
1386:
1382:
1376:
1373:
1363:
1357:
1353:
1349:
1345:
1341:
1334:
1332:
1330:
1326:
1319:
1311:
1308:
1307:
1306:
1302:
1301:
1300:Venatiolaspis
1291:
1290:
1286:
1285:
1284:
1280:
1279:
1269:
1266:
1265:
1264:
1262:Berlese, 1918
1260:
1259:
1249:
1246:
1245:
1244:
1240:
1239:
1229:
1226:
1225:
1224:
1220:
1219:
1218:Tigonholaspis
1209:
1206:
1205:
1204:
1200:
1199:
1189:
1186:
1185:
1184:
1182:Berlese, 1918
1180:
1179:
1169:
1166:
1165:
1164:
1162:Berlese, 1918
1160:
1159:
1158:Nothrholaspis
1149:
1146:
1141:
1138:
1133:
1132:
1128:
1123:
1122:
1118:
1113:
1110:
1105:
1102:
1097:
1094:
1091:Berlese, 1911
1089:
1086:
1081:
1078:
1073:
1072:
1068:
1063:
1062:
1058:
1053:
1050:
1045:
1042:
1037:
1034:
1029:
1028:
1024:
1023:
1022:
1018:
1017:
1007:
1004:
1003:
1002:
998:
997:
990:
989:
984:
983:
982:
978:
977:
967:
964:
963:
962:
958:
957:
947:
946:
942:
937:
934:
929:
926:
923:Berlese, 1903
921:
918:
913:
912:
908:
903:
900:
897:Berlese, 1920
895:
894:
890:
885:
884:
880:
875:
872:
869:Ibrahim, 1992
867:
864:
861:Berlese, 1910
859:
856:
851:
850:
846:
841:
838:
833:
830:
827:Berlese, 1910
825:
822:
817:
816:
812:
807:
804:
803:
802:
800:Berlese, 1903
798:
797:
796:Holostaspella
789:Berlese, 1910
787:
784:
783:
782:
780:Berlese, 1910
778:
777:
767:
764:
763:
762:
758:
757:
747:
744:
739:
736:
731:
728:
723:
720:
715:
712:
707:
706:
702:
697:
694:
689:
686:
681:
678:
673:
670:
665:
662:
657:
654:
653:
652:
648:
647:
646:Glyptholaspis
637:
636:
632:
627:
626:
622:
617:
614:
609:
606:
601:
600:
596:
591:
590:
586:
581:
580:
576:
571:
570:
566:
561:
560:
556:
551:
548:
543:
542:
538:
533:
532:
528:
523:
520:
515:
514:
510:
509:
508:
506:Berlese, 1918
504:
503:
493:
490:
489:
488:
486:Berlese, 1918
484:
483:
482:Evholocelaeno
473:
470:
469:
468:
466:Berlese, 1918
464:
463:
453:
450:
447:Berlese, 1918
445:
442:
441:
440:
438:Berlese, 1918
436:
435:
425:
422:
421:
420:
416:
415:
414:Bellatocheles
405:
402:
401:
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365:
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349:
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314:
309:
307:
303:
299:
295:
291:
283:
281:
279:
275:
271:
270:Holostaspella
267:
266:Glyptholaspis
263:
259:
255:
252:. Species of
251:
247:
239:
237:
234:
230:
222:
220:
218:
215:in the order
214:
210:
209:Macrochelidae
197:
196:Macrochelidae
193:
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137:
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91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
24:Macrochelidae
22:
19:
1802:Mesostigmata
1519:
1463:
1459:
1449:
1416:
1412:
1388:. Retrieved
1384:
1375:
1365:, retrieved
1343:
1309:
1298:
1297:
1287:
1276:
1275:
1267:
1256:
1255:
1247:
1238:Tricholaspis
1236:
1235:
1227:
1216:
1215:
1211:Krantz, 1961
1207:
1202:Krantz, 1961
1196:
1195:
1187:
1178:Proholaspina
1176:
1175:
1167:
1156:
1155:
1147:
1139:
1135:Krantz, 1965
1129:
1119:
1111:
1103:
1095:
1087:
1083:Krantz, 1965
1079:
1069:
1059:
1051:
1043:
1035:
1025:
1016:Neopodocinum
1014:
1013:
1005:
994:
993:
986:
974:
973:
969:Krantz, 1961
965:
960:Krantz, 1961
954:
953:
949:(Karg, 1994)
943:
935:
927:
919:
915:Takaku, 1994
909:
901:
891:
881:
873:
865:
857:
847:
843:Krantz, 1967
839:
831:
823:
813:
805:
794:
793:
785:
774:
773:
765:
754:
753:
745:
737:
729:
721:
713:
703:
695:
687:
679:
671:
663:
655:
644:
643:
633:
623:
615:
607:
597:
587:
577:
567:
557:
549:
539:
529:
521:
517:Krauss, 1970
511:
500:
499:
491:
480:
479:
471:
462:Cophrolaspis
460:
459:
451:
443:
432:
431:
423:
412:
411:
403:
394:Andhrolaspis
392:
391:
387:Krantz, 1962
383:
378:Krantz, 1962
372:
371:
367:Krantz, 1962
363:
358:Krantz, 1962
352:
351:
347:
337:
331:
323:
316:
310:
287:
276:can perform
273:
269:
265:
260:can perform
257:
253:
243:
240:Reproduction
226:
217:Mesostigmata
208:
207:
195:
184:Mesostigmata
116:
18:
1686:iNaturalist
1544:Wikispecies
1385:idtools.org
1251:Evans, 1956
1242:Evans, 1956
996:Neoholaspis
988:Macrocheles
976:Macrocheles
956:Lordocheles
776:Holocelaeno
756:Gonatothrix
691:(Foà, 1900)
603:Valle, 1963
545:Valle, 1953
535:Valle, 1953
455:Tseng, 1993
434:Calholaspis
290:oligochaete
278:arrhenotoky
274:Macrocheles
258:Macrocheles
223:Description
1796:Categories
1390:2022-10-14
1367:2022-10-14
1320:References
1198:Synaphasis
1009:Türk, 1948
1000:Türk, 1948
887:Karg, 1979
502:Geholaspis
407:Türk, 1948
398:Türk, 1948
321:on flies.
254:Geholaspis
167:Subclass:
148:Arthropoda
1496:204942853
1480:0168-8162
939:Roy, 1991
931:Roy, 1988
905:Roy, 1988
877:Roy, 1988
717:Roy, 1988
699:Roy, 1988
354:Aethosoma
313:parasitic
298:arthropod
294:nematodes
262:thelytoky
233:sclerites
160:Arachnida
131:Kingdom:
1595:BugGuide
1571:BioLib:
1535:Q3274878
1529:Wikidata
1488:31673886
1441:29284819
749:Ma, 1997
675:Ma, 1997
344:Taxonomy
306:phoresis
191:Family:
143:Phylum:
136:Animalia
1717:1118054
1626:4317924
1433:9519465
292:worms,
284:Ecology
179:Order:
155:Class:
96:↓
1750:NZOR:
1704:108474
1691:172592
1639:1MACHF
1600:265645
1587:452197
1494:
1486:
1478:
1439:
1431:
1358:
302:scarab
1774:Plazi
1766:91760
1743:99224
1699:IRMNG
1652:12465
1613:624KG
1574:19364
1492:S2CID
1437:S2CID
229:setae
213:mites
172:Acari
1738:NCBI
1712:ITIS
1678:2831
1673:GBIF
1634:EPPO
1582:BOLD
1484:PMID
1476:ISSN
1429:PMID
1356:ISBN
985:See
272:and
256:and
35:PreꞒ
1725:NBN
1621:EoL
1608:CoL
1559:AFD
1468:doi
1421:doi
1348:doi
1798::
1776::
1763::
1740::
1727::
1714::
1701::
1688::
1675::
1662::
1649::
1636::
1623::
1610::
1597::
1584::
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1490:.
1482:.
1474:.
1464:79
1462:.
1458:.
1435:.
1427:.
1417:22
1415:.
1411:.
1399:^
1383:.
1354:,
1342:,
1328:^
296:,
268:,
219:.
85:Pg
1498:.
1470::
1443:.
1423::
1393:.
1350::
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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