565:(sometimes also listed as belonging to the tribes Mylodontini and Lestodontini. The subdivision of the terminal group of mylodonts into the Lestodontinae and Mylodontinae found confirmation in one of the most comprehensive studies of the phylogeny of sloths based on cranial features in 2004, which subsequently found multiple support. However, a later analysis from 2019 cast doubt on it again. A higher-resolution phylogenetic study of the mylodonts published in the same year again supports the branching of terminal forms. According to this, the Mylodontinae and Lestodontinae can be distinguished on the basis of the canine anterior teeth. In the latter, these are large and separated from the posterior teeth by a long diastema; the former, on the other hand, have only small or partially reduced caniniform teeth, which are usually more closely apposed to the molar-like teeth. Numerous other subfamilies have been established in the past, including, for example, the Nematheriinae for representatives from the Lower Miocene or the
139:
168:
2398:
3154:
573:, were established only in 2004. Their basal position consists of the late Miocene representatives of northern South America. In principle, a revision for the entire family is urged, since many of the higher taxonomic units lack a formal diagnosis.
1179:
Boscaini, Alberto; Pujos, François; Gaudin, Timothy J. (November 2019). "A reappraisal of the phylogeny of
Mylodontidae (Mammalia, Xenarthra) and the divergence of mylodontine and lestodontine sloths".
1458:"A new genus and species of Mylodontidae (Mammalia: Xenarthra) from the late Miocene of southern Uruguay, with comments on the systematics of the Mylodontinae"
1627:
3238:
541:
The classification of the
Mylodontidae is complex and often under discussion. The most widely accepted subfamilies are the Mylodontinae with
3243:
1411:"Phylogeny, Macroevolutionary Trends and Historical Biogeography of Sloths: Insights From a Bayesian Morphological Clock Analysis"
3263:
138:
1356:"Archaeomylodon sampedrinensis, gen. et sp. nov., a new mylodontine from the middle Pleistocene of Pampean Region, Argentina"
151:
56:
44:
3258:
3248:
3253:
1620:
1513:"Urumacotheriinae, nova subfamília de Mylodontidae (Mammalia, Tardigrada) do Mioceno superior-Plioceno, América do Sul"
1309:"Baraguatherium takumara, Gen. et Sp. Nov., the Earliest Mylodontoid Sloth (Early Miocene) from Northern South America"
167:
3233:
1552:
Rincón, Ascanio D.; McDonald, H. Gregory; Solórzano, Andrés; Flores, Mónica Núñez; Ruiz-Ramoni, Damián (2015).
1613:
3173:
2758:
3077:
2744:
2118:
683:
533:
is an extinct subfamily of ground sloths that lived from the Early
Miocene to the Early Holocene epochs.
3205:
2608:
3112:
3084:
576:
Below is a phylogenetic tree of the
Mylodontinae, based on the work of Boscaini and colleagues (2019).
2643:
2171:
2008:
1260:"Phylogenetic relationships among sloths (Mammalia, Xenarthra, Tardigrada): the craniodental evidence"
570:
566:
288:
3091:
3014:
2994:
2296:
2090:
2015:
1565:
1469:
1367:
944:
569:
for all basal forms. Their recognition varies mostly depending on the author. Another subfamily, the
457:
2737:
2523:
2492:
2331:
2029:
912:
507:
3049:
3028:
2795:
2688:
2500:
3056:
2622:
2397:
2269:
859:
427:
2847:
2819:
2629:
2594:
2538:
2476:
2310:
2221:
2125:
1493:
1391:
1336:
1289:
1197:
1046:
606:
477:
369:
339:
162:
2207:
2178:
1307:
Rincón, Ascanio D.; Solórzano, Andrés; McDonald, H. Gregory; Flores, Mónica Núñez (2017-06-01).
349:
298:
3210:
3153:
3126:
2924:
2650:
2553:
2214:
359:
2636:
2587:
1880:
1591:
1534:
1485:
1438:
1430:
1383:
1328:
1281:
1240:
3070:
2973:
2917:
2446:
2097:
1949:
1581:
1573:
1524:
1477:
1422:
1375:
1320:
1271:
1232:
1189:
665:
3098:
2838:
2806:
2560:
2262:
1832:
1765:
1409:
Varela, Luciano; Tambusso, P. Sebastián; McDonald, H. Gregory; Fariña, Richard A. (2018).
417:
1569:
1473:
1371:
3105:
3042:
3021:
2903:
2730:
2723:
2661:
2546:
2514:
2428:
2415:
2365:
2241:
2156:
2132:
2076:
2069:
2022:
1987:
1935:
1774:
1754:
1744:
1586:
1553:
764:
624:
387:
267:
28:
1456:
Rinderknecht, Andrés; T., Enrique
Bostelmann; Perea, Daniel; Lecuona, Gustavo (2010).
3227:
3005:
2939:
2910:
2882:
2861:
2854:
2578:
2303:
2276:
1994:
1958:
1917:
1340:
1276:
1201:
996:
816:
561:
546:
467:
437:
96:
40:
1497:
1395:
1293:
3035:
2987:
2980:
2952:
2889:
2826:
2466:
2232:
2185:
2163:
2055:
1972:
1850:
1735:
1724:
717:
583:
308:
277:
232:
1379:
3196:
3119:
3063:
2964:
2868:
2751:
2716:
2681:
2615:
2567:
2338:
2324:
2287:
2248:
2199:
2192:
2104:
2083:
1979:
1794:
1784:
1714:
1355:
1021:
962:
841:
555:
517:
497:
397:
328:
318:
71:
48:
36:
2875:
2785:
2702:
2674:
2601:
2484:
2001:
1873:
1859:
1481:
1324:
116:
81:
1538:
1489:
1434:
1387:
1332:
1285:
1244:
2896:
2695:
2111:
1866:
1667:
1554:"A new enigmatic Late Miocene mylodontoid sloth from northern South America"
1426:
1308:
179:
121:
65:
1595:
1442:
1529:
3190:
2709:
2530:
2456:
2255:
2036:
1901:
1801:
1698:
1655:
782:
407:
199:
111:
106:
91:
86:
76:
52:
32:
1577:
2317:
1965:
1894:
1887:
1064:
487:
126:
101:
1193:
1682:
1661:
1649:
1636:
219:
209:
189:
155:
3167:
1512:
1410:
1259:
1216:
1457:
1236:
1821:
1605:
3171:
2413:
1680:
1609:
1354:
Brambilla, Luciano; Ibarra, Damián
Alberto (2018-11-02).
2782:
2511:
2472:
2462:
2452:
2052:
1946:
1847:
1790:
1780:
1750:
1720:
1710:
513:
503:
493:
483:
473:
463:
453:
445:
433:
423:
413:
403:
393:
383:
375:
365:
355:
345:
335:
324:
314:
304:
294:
284:
273:
263:
3180:
3004:
2963:
2938:
2837:
2805:
2780:
2660:
2577:
2510:
2426:
2364:
2286:
2231:
2142:
2050:
1945:
1915:
1846:
1819:
1764:
1734:
1697:
1219:
Classification of
Mammals: Above the Species Level.
1174:
1172:
1170:
1621:
8:
3168:
2944:
2811:
2802:
2666:
2574:
2438:
2434:
2423:
2410:
2361:
2148:
2061:
1927:
1923:
1843:
1839:
1703:
1694:
1690:
1677:
1628:
1614:
1606:
137:
20:
1585:
1528:
1275:
1264:Zoological Journal of the Linnean Society
1166:
1215:Theodor, Jessica M. (December 1998).
7:
1517:Revista Brasileira de Paleontologia
1221:Malcolm C. McKenna , Susan K. Bell"
1037:
1012:
987:
935:
928:
903:
832:
807:
755:
748:
737:
708:
656:
649:
597:
590:
579:
1462:Journal of Vertebrate Paleontology
1360:Journal of Vertebrate Paleontology
14:
3152:
2396:
1277:10.1111/j.1096-3642.2003.00100.x
553:, which sometimes also includes
166:
1225:The Quarterly Review of Biology
3239:Prehistoric animal subfamilies
1313:Journal of Mammalian Evolution
1:
1380:10.1080/02724634.2018.1542308
16:Extinct subfamily of mammals
3244:Messinian first appearances
1258:Gaudin, Timothy J. (2004).
3280:
1558:Royal Society Open Science
545:as the type genus and the
3150:
2947:
2814:
2669:
2441:
2437:
2422:
2409:
2394:
2360:
2151:
2064:
1930:
1926:
1842:
1706:
1693:
1689:
1676:
1644:
1511:Negri, Francisco (2004).
1482:10.1080/02724631003757997
1325:10.1007/s10914-016-9328-y
1060:
1042:
1035:
1017:
1010:
992:
985:
958:
940:
933:
926:
908:
901:
855:
837:
830:
812:
805:
778:
760:
753:
746:
735:
713:
706:
679:
661:
654:
647:
620:
602:
595:
588:
260:
255:
163:Scientific classification
161:
145:
136:
23:
3264:Pleistocene xenarthrans
549:, whose type genus is
152:Natural History Museum
148:Glossotherium robustum
3206:Paleobiology Database
1530:10.4072/rbp.2004.2.23
1427:10.1093/sysbio/syy058
3259:Pliocene xenarthrans
3249:Holocene extinctions
3254:Miocene xenarthrans
3078:Perezfontanatherium
2119:Paroctodontotherium
1578:10.1098/rsos.150138
1570:2015RSOS....250138R
1474:2010JVPal..30..899R
1372:2018JVPal..38E2308B
684:Paroctodontotherium
3234:Prehistoric sloths
2609:Megalonychotherium
2126:Pseudoprepotherium
1415:Systematic Biology
899:Mylodontini
607:Pseudoprepotherium
3221:
3220:
3174:Taxon identifiers
3165:
3164:
3161:
3160:
3148:
3147:
3144:
3143:
3140:
3139:
3136:
3135:
3113:Pyramiodontherium
3085:Plesiomegatherium
2934:
2933:
2776:
2775:
2772:
2771:
2768:
2767:
2637:Proschismotherium
2405:
2404:
2392:
2391:
2388:
2387:
2384:
2383:
2356:
2355:
2352:
2351:
2348:
2347:
2046:
2045:
1911:
1910:
1815:
1814:
1811:
1810:
1194:10.1111/zsc.12376
1182:Zoologica Scripta
1157:
1156:
1148:
1147:
1139:
1138:
1130:
1129:
1121:
1120:
1112:
1111:
1103:
1102:
1094:
1093:
1085:
1084:
1076:
1075:
974:
973:
889:
888:
880:
879:
871:
870:
794:
793:
695:
694:
636:
635:
528:
527:
251:
3271:
3214:
3213:
3201:
3200:
3199:
3169:
3156:
2945:
2918:Pronothrotherium
2812:
2803:
2784:
2667:
2644:Pseudortotherium
2575:
2513:
2474:
2464:
2454:
2439:
2435:
2424:
2411:
2400:
2362:
2172:Glossotheriopsis
2149:
2098:Octodontotherium
2062:
2054:
2009:Scelidotheridium
1950:Scelidotheriidae
1948:
1928:
1924:
1849:
1844:
1840:
1792:
1782:
1752:
1722:
1712:
1704:
1695:
1691:
1678:
1630:
1623:
1616:
1607:
1600:
1599:
1589:
1549:
1543:
1542:
1532:
1508:
1502:
1501:
1453:
1447:
1446:
1406:
1400:
1399:
1351:
1345:
1344:
1304:
1298:
1297:
1279:
1255:
1249:
1248:
1212:
1206:
1205:
1176:
1038:
1013:
988:
936:
929:
904:
833:
808:
756:
749:
738:
709:
666:Octodontotherium
657:
650:
598:
591:
580:
571:Urumacotheriinae
567:Octomylodontinae
515:
505:
495:
485:
475:
465:
455:
447:
435:
425:
415:
405:
395:
385:
377:
367:
357:
347:
337:
326:
316:
306:
296:
289:Glossotheriopsis
286:
275:
265:
249:
244:
231:
171:
170:
141:
131:
68:
64:15.5–0.010
27:Temporal range:
21:
3279:
3278:
3274:
3273:
3272:
3270:
3269:
3268:
3224:
3223:
3222:
3217:
3209:
3204:
3195:
3194:
3189:
3176:
3166:
3157:
3132:
3099:Proeremotherium
3092:Pliomegatherium
3015:Anisodontherium
3000:
2995:Proprepotherium
2959:
2930:
2839:Nothrotheriinae
2833:
2807:Nothrotheriidae
2801:
2788:
2764:
2656:
2573:
2561:Proplatyarthrus
2506:
2431:
2418:
2401:
2380:
2344:
2297:Glossotheridium
2282:
2263:Magdalenabradys
2227:
2138:
2091:Octodontobradys
2058:
2042:
2016:Scelidotheriops
1941:
1920:
1907:
1838:
1833:Pseudoglyptodon
1824:
1807:
1766:Myrmecophagidae
1760:
1730:
1685:
1672:
1640:
1634:
1604:
1603:
1551:
1550:
1546:
1510:
1509:
1505:
1455:
1454:
1450:
1408:
1407:
1403:
1366:(6): e1542308.
1353:
1352:
1348:
1306:
1305:
1301:
1257:
1256:
1252:
1214:
1213:
1209:
1178:
1177:
1168:
1163:
1158:
1149:
1140:
1131:
1122:
1113:
1104:
1095:
1086:
1077:
975:
945:Glossotheridium
890:
881:
872:
795:
696:
637:
539:
458:Glossotheridium
418:Magdalenabradys
248:
242:
229:
165:
132:
130:
129:
124:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
63:
62:
60:
25:
17:
12:
11:
5:
3277:
3275:
3267:
3266:
3261:
3256:
3251:
3246:
3241:
3236:
3226:
3225:
3219:
3218:
3216:
3215:
3202:
3186:
3184:
3178:
3177:
3172:
3163:
3162:
3159:
3158:
3151:
3149:
3146:
3145:
3142:
3141:
3138:
3137:
3134:
3133:
3131:
3130:
3123:
3116:
3109:
3106:Promegatherium
3102:
3095:
3088:
3081:
3074:
3067:
3060:
3053:
3046:
3043:Megathericulus
3039:
3032:
3025:
3022:Diabolotherium
3018:
3010:
3008:
3002:
3001:
2999:
2998:
2991:
2984:
2977:
2969:
2967:
2961:
2960:
2958:
2957:
2948:
2942:
2936:
2935:
2932:
2931:
2929:
2928:
2921:
2914:
2907:
2904:Nothrotheriops
2900:
2893:
2886:
2879:
2872:
2865:
2858:
2851:
2843:
2841:
2835:
2834:
2832:
2831:
2823:
2815:
2809:
2800:
2799:
2791:
2789:
2781:
2778:
2777:
2774:
2773:
2770:
2769:
2766:
2765:
2763:
2762:
2755:
2748:
2741:
2738:Protomegalonyx
2734:
2731:Pliometanastes
2727:
2724:Pattersonocnus
2720:
2713:
2706:
2699:
2692:
2685:
2678:
2670:
2664:
2662:Megalonychinae
2658:
2657:
2655:
2654:
2647:
2640:
2633:
2626:
2619:
2612:
2605:
2598:
2591:
2583:
2581:
2572:
2571:
2564:
2557:
2550:
2547:Mesopotamocnus
2543:
2535:
2527:
2524:Deseadognathus
2519:
2517:
2515:Megalonychidae
2508:
2507:
2505:
2504:
2496:
2493:Schismotherium
2488:
2480:
2470:
2460:
2450:
2442:
2432:
2429:Megatherioidea
2427:
2420:
2419:
2416:Megatherioidea
2414:
2407:
2406:
2403:
2402:
2395:
2393:
2390:
2389:
2386:
2385:
2382:
2381:
2379:
2378:
2377:
2376:
2370:
2368:
2366:Megatherioidea
2358:
2357:
2354:
2353:
2350:
2349:
2346:
2345:
2343:
2342:
2335:
2332:Pleurolestodon
2328:
2321:
2314:
2307:
2300:
2292:
2290:
2284:
2283:
2281:
2280:
2273:
2266:
2259:
2252:
2245:
2242:Bolivartherium
2237:
2235:
2229:
2228:
2226:
2225:
2218:
2211:
2204:
2196:
2189:
2182:
2175:
2168:
2160:
2157:Archaeomylodon
2152:
2146:
2140:
2139:
2137:
2136:
2133:Urumacotherium
2129:
2122:
2115:
2108:
2101:
2094:
2087:
2080:
2077:Eionaletherium
2073:
2070:Baraguatherium
2065:
2059:
2051:
2048:
2047:
2044:
2043:
2041:
2040:
2033:
2030:Sibyllotherium
2026:
2023:Scelidotherium
2019:
2012:
2005:
1998:
1991:
1988:Neonematherium
1984:
1976:
1969:
1962:
1954:
1952:
1943:
1942:
1940:
1939:
1931:
1921:
1916:
1913:
1912:
1909:
1908:
1906:
1905:
1898:
1891:
1884:
1877:
1870:
1863:
1855:
1853:
1837:
1836:
1827:
1825:
1820:
1817:
1816:
1813:
1812:
1809:
1808:
1806:
1805:
1798:
1788:
1778:
1770:
1768:
1762:
1761:
1759:
1758:
1755:Palaeomyrmidon
1748:
1740:
1738:
1732:
1731:
1729:
1728:
1718:
1707:
1701:
1687:
1686:
1681:
1674:
1673:
1671:
1670:
1664:
1658:
1652:
1645:
1642:
1641:
1635:
1633:
1632:
1625:
1618:
1610:
1602:
1601:
1544:
1523:(2): 281–288.
1503:
1468:(3): 899–910.
1448:
1421:(2): 204–218.
1401:
1346:
1319:(2): 179–191.
1299:
1270:(2): 255–305.
1250:
1237:10.1086/420478
1231:(4): 518–519.
1207:
1188:(6): 691–710.
1165:
1164:
1162:
1159:
1155:
1154:
1151:
1150:
1146:
1145:
1142:
1141:
1137:
1136:
1133:
1132:
1128:
1127:
1124:
1123:
1119:
1118:
1115:
1114:
1110:
1109:
1106:
1105:
1101:
1100:
1097:
1096:
1092:
1091:
1088:
1087:
1083:
1082:
1079:
1078:
1074:
1073:
1070:
1069:
1059:
1056:
1055:
1052:
1051:
1041:
1036:
1034:
1031:
1030:
1027:
1026:
1016:
1011:
1009:
1006:
1005:
1002:
1001:
991:
986:
984:
981:
980:
977:
976:
972:
971:
968:
967:
957:
954:
953:
950:
949:
939:
934:
932:
927:
925:
922:
921:
918:
917:
913:Pleurolestodon
907:
902:
900:
896:
895:
892:
891:
887:
886:
883:
882:
878:
877:
874:
873:
869:
868:
865:
864:
854:
851:
850:
847:
846:
836:
831:
829:
826:
825:
822:
821:
811:
806:
804:
801:
800:
797:
796:
792:
791:
788:
787:
777:
774:
773:
770:
769:
765:Bolivartherium
759:
754:
752:
747:
745:
736:
734:
727:
726:
723:
722:
712:
707:
705:
702:
701:
698:
697:
693:
692:
689:
688:
678:
675:
674:
671:
670:
660:
655:
653:
648:
646:
643:
642:
639:
638:
634:
633:
630:
629:
625:Urumacotherium
619:
616:
615:
612:
611:
601:
596:
594:
589:
587:
578:
538:
537:Classification
535:
526:
525:
524:
523:
522:
521:
511:
508:Pleurolestodon
501:
491:
481:
471:
461:
443:
442:
441:
431:
421:
411:
401:
391:
388:Bolivartherium
373:
363:
353:
343:
333:
322:
312:
302:
292:
282:
271:
268:Archaeomylodon
258:
257:
253:
252:
240:
236:
235:
227:
223:
222:
217:
213:
212:
207:
203:
202:
197:
193:
192:
187:
183:
182:
177:
173:
172:
159:
158:
143:
142:
134:
133:
125:
120:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
69:
33:Early Holocene
29:Middle Miocene
26:
15:
13:
10:
9:
6:
4:
3:
2:
3276:
3265:
3262:
3260:
3257:
3255:
3252:
3250:
3247:
3245:
3242:
3240:
3237:
3235:
3232:
3231:
3229:
3212:
3207:
3203:
3198:
3192:
3188:
3187:
3185:
3183:
3179:
3175:
3170:
3155:
3129:
3128:
3124:
3122:
3121:
3117:
3115:
3114:
3110:
3108:
3107:
3103:
3101:
3100:
3096:
3094:
3093:
3089:
3087:
3086:
3082:
3080:
3079:
3075:
3073:
3072:
3068:
3066:
3065:
3061:
3059:
3058:
3054:
3052:
3051:
3050:Megatheridium
3047:
3045:
3044:
3040:
3038:
3037:
3033:
3031:
3030:
3029:Eomegatherium
3026:
3024:
3023:
3019:
3017:
3016:
3012:
3011:
3009:
3007:
3006:Megatheriinae
3003:
2997:
2996:
2992:
2990:
2989:
2985:
2983:
2982:
2978:
2976:
2975:
2971:
2970:
2968:
2966:
2962:
2955:
2954:
2950:
2949:
2946:
2943:
2941:
2940:Megatheriidae
2937:
2927:
2926:
2922:
2920:
2919:
2915:
2913:
2912:
2911:Nothrotherium
2908:
2906:
2905:
2901:
2899:
2898:
2894:
2892:
2891:
2887:
2885:
2884:
2883:Mcdonaldocnus
2880:
2878:
2877:
2873:
2871:
2870:
2866:
2864:
2863:
2862:Chasicobradys
2859:
2857:
2856:
2855:Aymaratherium
2852:
2850:
2849:
2845:
2844:
2842:
2840:
2836:
2829:
2828:
2824:
2822:
2821:
2817:
2816:
2813:
2810:
2808:
2804:
2798:
2797:
2796:Analcimorphus
2793:
2792:
2790:
2787:
2779:
2761:
2760:
2756:
2754:
2753:
2749:
2747:
2746:
2742:
2740:
2739:
2735:
2733:
2732:
2728:
2726:
2725:
2721:
2719:
2718:
2714:
2712:
2711:
2707:
2705:
2704:
2700:
2698:
2697:
2693:
2691:
2690:
2689:Megalonychops
2686:
2684:
2683:
2679:
2677:
2676:
2672:
2671:
2668:
2665:
2663:
2659:
2653:
2652:
2648:
2646:
2645:
2641:
2639:
2638:
2634:
2632:
2631:
2627:
2625:
2624:
2620:
2618:
2617:
2613:
2611:
2610:
2606:
2604:
2603:
2599:
2597:
2596:
2592:
2590:
2589:
2585:
2584:
2582:
2580:
2579:Ortotheriinae
2576:
2570:
2569:
2565:
2563:
2562:
2558:
2556:
2555:
2551:
2549:
2548:
2544:
2541:
2540:
2536:
2533:
2532:
2528:
2526:
2525:
2521:
2520:
2518:
2516:
2509:
2503:
2502:
2501:Similhapalops
2497:
2495:
2494:
2489:
2487:
2486:
2481:
2479:
2478:
2471:
2469:
2468:
2461:
2459:
2458:
2451:
2449:
2448:
2444:
2443:
2440:
2436:
2433:
2430:
2425:
2421:
2417:
2412:
2408:
2399:
2374:
2373:
2372:
2371:
2369:
2367:
2363:
2359:
2341:
2340:
2336:
2334:
2333:
2329:
2327:
2326:
2322:
2320:
2319:
2315:
2313:
2312:
2308:
2306:
2305:
2304:Glossotherium
2301:
2299:
2298:
2294:
2293:
2291:
2289:
2285:
2279:
2278:
2277:Thinobadistes
2274:
2272:
2271:
2267:
2265:
2264:
2260:
2258:
2257:
2253:
2251:
2250:
2246:
2244:
2243:
2239:
2238:
2236:
2234:
2230:
2224:
2223:
2219:
2217:
2216:
2212:
2210:
2209:
2205:
2202:
2201:
2197:
2195:
2194:
2190:
2188:
2187:
2183:
2181:
2180:
2176:
2174:
2173:
2169:
2166:
2165:
2161:
2159:
2158:
2154:
2153:
2150:
2147:
2145:
2141:
2135:
2134:
2130:
2128:
2127:
2123:
2121:
2120:
2116:
2114:
2113:
2109:
2107:
2106:
2102:
2100:
2099:
2095:
2093:
2092:
2088:
2086:
2085:
2081:
2079:
2078:
2074:
2072:
2071:
2067:
2066:
2063:
2060:
2057:
2049:
2039:
2038:
2034:
2032:
2031:
2027:
2025:
2024:
2020:
2018:
2017:
2013:
2011:
2010:
2006:
2004:
2003:
1999:
1997:
1996:
1995:Proscelidodon
1992:
1990:
1989:
1985:
1982:
1981:
1977:
1975:
1974:
1970:
1968:
1967:
1963:
1961:
1960:
1959:Analcitherium
1956:
1955:
1953:
1951:
1944:
1938:
1937:
1933:
1932:
1929:
1925:
1922:
1919:
1918:Mylodontoidea
1914:
1904:
1903:
1899:
1897:
1896:
1892:
1890:
1889:
1885:
1883:
1882:
1878:
1876:
1875:
1871:
1869:
1868:
1864:
1862:
1861:
1857:
1856:
1854:
1852:
1845:
1841:
1835:
1834:
1829:
1828:
1826:
1823:
1818:
1804:
1803:
1799:
1797:
1796:
1789:
1787:
1786:
1779:
1777:
1776:
1772:
1771:
1769:
1767:
1763:
1757:
1756:
1749:
1747:
1746:
1742:
1741:
1739:
1737:
1733:
1727:
1726:
1719:
1717:
1716:
1709:
1708:
1705:
1702:
1700:
1696:
1692:
1688:
1684:
1679:
1675:
1669:
1665:
1663:
1659:
1657:
1653:
1651:
1647:
1646:
1643:
1638:
1631:
1626:
1624:
1619:
1617:
1612:
1611:
1608:
1597:
1593:
1588:
1583:
1579:
1575:
1571:
1567:
1564:(4): 150138.
1563:
1559:
1555:
1548:
1545:
1540:
1536:
1531:
1526:
1522:
1518:
1514:
1507:
1504:
1499:
1495:
1491:
1487:
1483:
1479:
1475:
1471:
1467:
1463:
1459:
1452:
1449:
1444:
1440:
1436:
1432:
1428:
1424:
1420:
1416:
1412:
1405:
1402:
1397:
1393:
1389:
1385:
1381:
1377:
1373:
1369:
1365:
1361:
1357:
1350:
1347:
1342:
1338:
1334:
1330:
1326:
1322:
1318:
1314:
1310:
1303:
1300:
1295:
1291:
1287:
1283:
1278:
1273:
1269:
1265:
1261:
1254:
1251:
1246:
1242:
1238:
1234:
1230:
1226:
1222:
1220:
1211:
1208:
1203:
1199:
1195:
1191:
1187:
1183:
1175:
1173:
1171:
1167:
1160:
1153:
1152:
1144:
1143:
1135:
1134:
1126:
1125:
1117:
1116:
1108:
1107:
1099:
1098:
1090:
1089:
1081:
1080:
1072:
1071:
1068:
1067:
1066:
1058:
1057:
1054:
1053:
1050:
1049:
1048:
1040:
1039:
1033:
1032:
1029:
1028:
1025:
1024:
1023:
1015:
1014:
1008:
1007:
1004:
1003:
1000:
999:
998:
997:Glossotherium
990:
989:
983:
982:
979:
978:
970:
969:
966:
965:
964:
956:
955:
952:
951:
948:
947:
946:
938:
937:
931:
930:
924:
923:
920:
919:
916:
915:
914:
906:
905:
898:
897:
894:
893:
885:
884:
876:
875:
867:
866:
863:
862:
861:
853:
852:
849:
848:
845:
844:
843:
835:
834:
828:
827:
824:
823:
820:
819:
818:
817:Thinobadistes
810:
809:
803:
802:
799:
798:
790:
789:
786:
785:
784:
776:
775:
772:
771:
768:
767:
766:
758:
757:
751:
750:
743:
740:
739:
732:
729:
728:
725:
724:
721:
720:
719:
711:
710:
704:
703:
700:
699:
691:
690:
687:
686:
685:
677:
676:
673:
672:
669:
668:
667:
659:
658:
652:
651:
645:
644:
641:
640:
632:
631:
628:
627:
626:
618:
617:
614:
613:
610:
609:
608:
600:
599:
593:
592:
585:
582:
581:
577:
574:
572:
568:
564:
563:
562:Glossotherium
558:
557:
552:
548:
547:Lestodontinae
544:
536:
534:
532:
520:
519:
512:
510:
509:
502:
500:
499:
492:
490:
489:
482:
480:
479:
472:
470:
469:
468:Glossotherium
462:
460:
459:
452:
451:
450:
444:
440:
439:
438:Thinobadistes
432:
430:
429:
422:
420:
419:
412:
410:
409:
402:
400:
399:
392:
390:
389:
382:
381:
380:
374:
372:
371:
364:
362:
361:
354:
352:
351:
344:
342:
341:
334:
331:
330:
323:
321:
320:
313:
311:
310:
303:
301:
300:
293:
291:
290:
283:
280:
279:
272:
270:
269:
262:
261:
259:
254:
247:
241:
238:
237:
234:
228:
225:
224:
221:
218:
215:
214:
211:
208:
205:
204:
201:
198:
195:
194:
191:
188:
185:
184:
181:
178:
175:
174:
169:
164:
160:
157:
153:
149:
144:
140:
135:
128:
123:
118:
113:
108:
103:
98:
93:
88:
83:
78:
73:
67:
58:
54:
50:
46:
42:
41:Rancholabrean
38:
34:
30:
22:
19:
3182:Mylodontinae
3181:
3125:
3118:
3111:
3104:
3097:
3090:
3083:
3076:
3069:
3062:
3057:Megatheriops
3055:
3048:
3041:
3036:Eremotherium
3034:
3027:
3020:
3013:
2993:
2988:Prepoplanops
2986:
2981:Prepotherium
2979:
2972:
2953:Thalassocnus
2951:
2923:
2916:
2909:
2902:
2895:
2890:Mionothropus
2888:
2881:
2874:
2867:
2860:
2853:
2846:
2827:Thalassocnus
2825:
2818:
2794:
2757:
2750:
2743:
2736:
2729:
2722:
2715:
2708:
2701:
2694:
2687:
2680:
2673:
2649:
2642:
2635:
2628:
2623:Paranabradys
2621:
2614:
2607:
2600:
2593:
2586:
2566:
2559:
2552:
2545:
2537:
2529:
2522:
2499:
2491:
2483:
2475:
2467:Hiskatherium
2465:
2455:
2445:
2337:
2330:
2323:
2316:
2309:
2302:
2295:
2275:
2270:Sphenotherus
2268:
2261:
2254:
2247:
2240:
2233:Lestodontini
2220:
2213:
2206:
2198:
2191:
2186:Mylodonopsis
2184:
2177:
2170:
2164:Brievabradys
2162:
2155:
2144:Mylodontinae
2143:
2131:
2124:
2117:
2110:
2103:
2096:
2089:
2082:
2075:
2068:
2056:Mylodontidae
2035:
2028:
2021:
2014:
2007:
2000:
1993:
1986:
1978:
1973:Chubutherium
1971:
1964:
1957:
1934:
1900:
1893:
1886:
1879:
1872:
1865:
1858:
1851:Megalocnidae
1831:
1800:
1793:
1783:
1775:Myrmecophaga
1773:
1753:
1743:
1736:Cyclopedidae
1725:Orthoarthrus
1723:
1713:
1666:Superorder:
1561:
1557:
1547:
1520:
1516:
1506:
1465:
1461:
1451:
1418:
1414:
1404:
1363:
1359:
1349:
1316:
1312:
1302:
1267:
1263:
1253:
1228:
1224:
1218:
1210:
1185:
1181:
1063:
1061:
1045:
1043:
1020:
1018:
995:
993:
961:
959:
943:
941:
911:
909:
860:Sphenotherus
858:
856:
840:
838:
815:
813:
781:
779:
763:
761:
742:Lestodontini
741:
731:Mylodontinae
730:
718:Brievabradys
716:
714:
682:
680:
664:
662:
623:
621:
605:
603:
584:Mylodontidae
575:
560:
554:
550:
542:
540:
531:Mylodontinae
530:
529:
516:
506:
496:
486:
476:
466:
456:
448:
436:
428:Sphenotherus
426:
416:
406:
396:
386:
379:Lestodontini
378:
368:
358:
348:
338:
327:
317:
309:Mylodonopsis
307:
297:
287:
278:Brievabradys
276:
266:
246:Mylodontinae
245:
233:Mylodontidae
147:
146:Skeleton of
24:Mylodontinae
18:
3120:Sibotherium
3064:Megatherium
2965:Planopsinae
2869:Huilabradys
2848:Amphibradys
2820:Neohapalops
2759:Zacatzontli
2752:Xibalbaonyx
2717:Nohochichak
2682:Australonyx
2630:Pliomorphus
2616:Ortotherium
2595:Eucholoeops
2568:Urumacocnus
2539:Hyperleptus
2477:Hyperleptus
2339:Simomylodon
2325:Paramylodon
2311:Kiyumylodon
2288:Mylodontini
2249:Lestobradys
2222:Strabosodon
2200:Oreomylodon
2193:Ocnotherium
2105:Octomylodon
2084:Nematherium
1980:Nematherium
1795:Protamandua
1785:Neotamandua
1715:Argyromanis
1699:Vermilingua
1047:Kiyumylodon
1022:Paramylodon
963:Simomylodon
842:Lestobradys
556:Paramylodon
518:Simomylodon
498:Paramylodon
478:Kiyumylodon
449:Mylodontini
398:Lestobradys
370:Strabosodon
340:Prolestodon
329:Oreomylodon
319:Ocnotherium
239:Subfamily:
49:Colloncuran
37:Hemphillian
3228:Categories
2876:Lakukullus
2786:Megatheria
2745:Sinclairia
2703:Megistonyx
2675:Ahytherium
2602:Hapaloides
2485:Pelecyodon
2375:see below↓
2208:Promylodon
2179:Megabradys
2002:Scelidodon
1874:Megalocnus
1860:Acratocnus
1161:References
350:Promylodon
299:Megabradys
256:Subgroups
250:Gill, 1872
3197:Q21291109
3127:Urumaquia
2925:Xyophorus
2897:Nothropus
2696:Megalonyx
2651:Torcellia
2554:Paulocnus
2215:Ranculcus
2112:Orophodon
1936:Choloepus
1867:Imagocnus
1668:Xenarthra
1648:Kingdom:
1539:1519-7530
1490:0272-4634
1435:1063-5157
1388:0272-4634
1341:254703922
1333:1573-7055
1286:1096-3642
1245:0033-5770
1202:201194980
360:Ranculcus
186:Kingdom:
180:Eukaryota
3191:Wikidata
2710:Meizonyx
2588:Diodomus
2531:Hapalops
2457:Hapalops
2447:Bradypus
2256:Lestodon
2037:Valgipes
1902:Parocnus
1881:Mesocnus
1822:Folivora
1802:Tamandua
1745:Cyclopes
1662:Mammalia
1656:Chordata
1654:Phylum:
1650:Animalia
1596:26065399
1498:86604163
1443:30239971
1396:91874640
1294:38722942
783:Lestodon
551:Lestodon
408:Lestodon
226:Family:
210:Mammalia
200:Chordata
196:Phylum:
190:Animalia
176:Domain:
53:Lujanian
47:) &
3071:Ocnopus
2974:Planops
2318:Mylodon
1966:Catonyx
1895:Neocnus
1888:Miocnus
1660:Class:
1637:Pilosan
1587:4448861
1566:Bibcode
1470:Bibcode
1368:Bibcode
1065:Mylodon
744:
733:
586:
543:Mylodon
488:Mylodon
216:Order:
206:Class:
150:at the
1683:Pilosa
1639:genera
1594:
1584:
1537:
1496:
1488:
1441:
1433:
1394:
1386:
1339:
1331:
1292:
1284:
1243:
1200:
1062:
1044:
1019:
994:
960:
942:
910:
857:
839:
814:
780:
762:
715:
681:
663:
622:
604:
220:Pilosa
156:London
3211:83049
1494:S2CID
1392:S2CID
1337:S2CID
1290:S2CID
1198:S2CID
57:SALMA
45:NALMA
1592:PMID
1535:ISSN
1486:ISSN
1439:PMID
1431:ISSN
1384:ISSN
1329:ISSN
1282:ISSN
1241:ISSN
559:and
72:PreꞒ
1582:PMC
1574:doi
1525:doi
1478:doi
1423:doi
1376:doi
1321:doi
1272:doi
1268:140
1233:doi
1190:doi
31:to
3230::
3208::
3193::
1590:.
1580:.
1572:.
1560:.
1556:.
1533:.
1519:.
1515:.
1492:.
1484:.
1476:.
1466:30
1464:.
1460:.
1437:.
1429:.
1419:68
1417:.
1413:.
1390:.
1382:.
1374:.
1364:38
1362:.
1358:.
1335:.
1327:.
1317:24
1315:.
1311:.
1288:.
1280:.
1266:.
1262:.
1239:.
1229:73
1227:.
1223:.
1196:.
1186:48
1184:.
1169:^
154:,
122:Pg
66:Ma
2956:?
2830:?
2783:†
2542:?
2534:?
2512:†
2498:†
2490:†
2482:†
2473:†
2463:†
2453:†
2203:?
2167:?
2053:†
1983:?
1947:†
1848:†
1830:†
1791:†
1781:†
1751:†
1721:†
1711:†
1629:e
1622:t
1615:v
1598:.
1576::
1568::
1562:2
1541:.
1527::
1521:7
1500:.
1480::
1472::
1445:.
1425::
1398:.
1378::
1370::
1343:.
1323::
1296:.
1274::
1247:.
1235::
1217:"
1204:.
1192::
514:†
504:†
494:†
484:†
474:†
464:†
454:†
446:†
434:†
424:†
414:†
404:†
394:†
384:†
376:†
366:†
356:†
346:†
336:†
332:?
325:†
315:†
305:†
295:†
285:†
281:?
274:†
264:†
243:†
230:†
127:N
117:K
112:J
107:T
102:P
97:C
92:D
87:S
82:O
77:Ꞓ
61:~
59:)
55:(
51:-
43:(
39:-
35:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
