124:
2480:
1615:
326:. They range in size from 0.2 to 1.4 millimetres (0.008 to 0.055 in). There are currently 12,000 species that have been identified, but researchers estimate that there may be anywhere from 60,000 to 120,000 total species. Oribatid mites are by far the most prevalent of all arthropods in forest soils, and are essential for breaking down organic detritus and distributing fungi.
102:
384:
Many species have a mineralized exoskeleton as adults. In some, this includes a pair of pteromorphae: wing-like flaps that overhang the legs on either side. Some oribatids can also tuck in their legs underneath their protective armor, an ability known as ptychoidy, for more defence against predation.
409:
Many species of oribatid mites require extremely specific habitats, resulting in large diversity within the order due to the many niches they evolve to. Some species are especially suited to dry conditions, or on bare lichen covered rocks, but that largest section of
Oribatida prefers the moist
433:, though once considered a separate group, are now considered part of Oribatida. They are quite different from other oribatids (e.g. many astigmatans are soft-bodied and some are parasitic), resulting in them often being treated separately.
421:
oribatid mite lineages have been hypothesized to be adapted to occupy narrow specialized ecological niches. However, it was recently shown that parthenogenetic oribatid mite species actually possess a widely adapted general-purpose
410:
forest floor and its accompanying litter. There are a small number of species who have evolved to live on aquatic plants, often spending the majority of their life submersed underwater.
1816:
Norton, Roy A.; Behan-Pelletier, Valerie M. (June 1991). "Calcium carbonate and calcium oxalate as cuticular hardening agents in oribatid mites (Acari: Oribatida)".
2763:
2802:
417:
lineages are short lived, four species-rich parthenogenetic clusters of the order
Oribatida are very ancient and likely arose 400-300 million years ago.
2079:
Maraun, Mark; Bischof, Paul S. P.; Klemp, Finn L.; Pollack, Jule; Raab, Linnea; Schmerbach, Jan; Schaefer, Ina; Scheu, Stefan; Caruso, Tancredi (2022).
2737:
2776:
2165:[Systematic and biogeographic list, with synonymies, of the oribatid mites (Acariformes: Oribatida) of the world (excluding fossils)]
2323:
2296:
2269:
2215:
1800:
1727:
1692:
1952:
2162:
1930:
Denegri, G.M. Review of oribatid mites as intermediate hosts of tapeworms of the
Anoplocephalidae. Exp Appl Acarol 17, 567â580 (1993).
2145:
2242:
1684:
2854:
1790:
2354:
2163:"Listado sistemĂĄtico, sinonĂmico y biogeogrĂĄfico de los ĂĄcaros oribĂĄtidos (Acariformes: Oribatida) del mundo (Excepto fĂłsiles)"
1874:
Saporito, Ralph A.; Donnelly, Maureen A.; Norton, Roy A.; Garraffo, H. Martin; Spande, Thomas F.; Daly, John W. (2007-05-22).
2781:
2789:
1750:"Listado sistemĂĄtico, sinonĂmico y biogeogrĂĄfico de los ĂĄcaros oribĂĄtidos (Acariformes, Oribatida) del mundo (1758-2002)"
357:
instars and the adult. All these stages after the prelarva feed on a wide variety of material including living and dead
2659:
334:
2716:
1189:
123:
2807:
2729:
1629:
2887:
2207:
2612:
200:
533:
2828:
2257:
1636:
834:
712:
562:
546:
1337:
1161:
1048:
846:
777:
759:
728:
688:
667:
638:
622:
554:
2846:
1573:
1528:
1481:
1355:
1292:
1212:
1076:
1016:
938:
866:
790:
751:
743:
704:
696:
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2859:
2698:
1519:
1472:
1454:
1373:
1319:
1310:
1096:
1028:
950:
906:
886:
614:
590:
2315:
1564:
1555:
1546:
1400:
918:
489:
460:
2419:
2347:
1537:
1463:
1436:
1364:
1240:
970:
517:
502:
250:
2414:
1714:, Developments in Hydrobiology, vol. 198, Dordrecht: Springer Netherlands, pp. 323â328,
1445:
1409:
1382:
1301:
1280:
1146:
521:
447:
238:
2288:
2009:
1591:
1498:
1427:
1391:
818:
402:
species, and by increasing the breakdown of organic material in the soil, in a similar manner to
277:
118:
2833:
2409:
2404:
1346:
1328:
1133:
1000:
805:
233:
228:
1676:
1418:
2815:
2685:
2591:
2583:
2319:
2292:
2265:
2238:
2211:
2203:
2141:
2118:
2100:
2058:
2001:
1993:
1956:
1913:
1895:
1796:
1771:
1723:
1688:
1620:
651:
2820:
2170:
2768:
2568:
2261:
2234:
2108:
2092:
2048:
2040:
2027:
Pachl, Patrick; Uusitalo, Matti; Scheu, Stefan; Schaefer, Ina; Maraun, Mark (January 2021).
1983:
1903:
1887:
1825:
1761:
1715:
1150:
1068:
680:
392:
342:
2479:
1850:
337:
with adults living a relatively long time; for example, estimates of development time from
1766:
1749:
1672:
1510:
1224:
1169:
606:
594:
418:
414:
391:
are produced by some oribatids, presumably as another defence against predation. In turn,
354:
308:
2650:
2225:
Halliday, R. B.; D. E. Walter; H. C. Proctor; R. A. Norton; M. J. Colloff, eds. (2001).
2080:
2724:
2711:
2488:
2465:
2452:
2429:
2340:
2311:
2113:
2053:
2028:
1908:
1643:
1116:
1108:
1060:
878:
323:
110:
2876:
2563:
2437:
2284:
2013:
1944:
473:
338:
59:
1875:
2690:
2573:
2555:
2545:
2510:
2505:
2399:
2394:
1614:
1267:
1252:
982:
577:
259:
223:
218:
1709:
2664:
1719:
381:
and feeding habits may differ between immatures and adults of the same species.
243:
2794:
2750:
2644:
2515:
2497:
2460:
2442:
2377:
1582:
1196:
430:
316:
185:
165:
34:
2635:
1971:
1988:
1610:
1185:
378:
330:
79:
44:
2104:
1997:
1899:
1775:
2535:
2230:
1891:
1177:
403:
155:
135:
84:
2122:
2062:
2005:
1917:
2332:
1972:"Catalogue of oribatid mites (Acari: Oribatida) from Vorarlberg (Austria)"
101:
2672:
2629:
2540:
2254:
Mites in Soil, An interactive key to mites and other soil microarthropods
2029:"Repeated convergent evolution of parthenogenesis in Acariformes (Acari)"
423:
399:
388:
374:
209:
175:
74:
69:
54:
49:
39:
1876:"Oribatid mites as a major dietary source for alkaloids in poison frogs"
426:, and thus each such lineage might be viewed as a âjack-of-all-tradesâ.
395:
that prey on oribatids sequester these alkaloids for their own defence.
2742:
2677:
370:
89:
64:
2755:
2096:
2044:
1931:
366:
362:
346:
145:
2606:
2136:
Krantz, G.W.; Walter, D.E., eds. (2009). "Astigmatina. Chapter 16".
1829:
2841:
358:
350:
320:
2703:
2526:
2368:
2364:
312:
2610:
2336:
1792:
Mites: Ecology, Evolution & Behaviour: Life at a
Microscale
398:
The
Oribatida are of economic importance as hosts of various
1711:
Global diversity of oribatids (Oribatida: Acari: Arachnida)
2227:
441:
The order
Oribatida is divided into the following taxa:
1789:
Walter, David Evans; Proctor, Heather C. (2013-10-08).
333:
rates, slow development and low fecundity. Species are
2308:
Phthiracaroidea (Acari, Oribatida): Systematic
Studies
2619:
2582:
2554:
2524:
2496:
2487:
2451:
2428:
2385:
2376:
1708:Schatz, Heinrich; Behan-Pelletier, Valerie (2008),
341:to adult vary from several months to two years in
1880:Proceedings of the National Academy of Sciences
2348:
2074:
2072:
8:
1943:Edward W. Baker & G. W. Wharton (1952).
413:In contrast to the commonly held view that
2607:
2493:
2382:
2355:
2341:
2333:
2252:Walter, D. E. & H. C. Proctor (2001).
100:
20:
2112:
2081:""Jackâofâallâtrades" is parthenogenetic"
2052:
1987:
1907:
1795:. Springer Science & Business Media.
1765:
295:(formerly Cryptostigmata), also known as
1655:
345:soils. Oribatid mites have six active
272:families, 1,200 genera, 6,600 species
1767:10.3989/graellsia.2004.v60.iextra.218
1667:
1665:
1663:
1661:
1659:
7:
2847:2B2B5066-FFBA-C47F-FF3D-FBB3FE3991EE
2730:45cca84e-2408-46a8-b5fb-96230986901d
1845:
1843:
1841:
1839:
1681:Encyclopedia of Entomology, Volume 1
2281:Acarology: Mites and Human Welfare
1932:https://doi.org/10.1007/BF00053486
329:Oribatid mites generally have low
14:
2138:A Manual of Acarology 3rd Edition
2478:
2140:. Texas Tech. University Press.
1613:
122:
1970:SCHATZ, HEINRICH (2020-05-27).
2256:. ABRS Identification Series.
1748:SubĂas, Luis S. (2004-12-31).
1:
1679:. In John L. Capinera (ed.).
266:
2169:(in Spanish). Archived from
1949:An Introduction to Acarology
1720:10.1007/978-1-4020-8259-7_35
2306:Niedbala, Wojciech (1992).
1818:Canadian Journal of Zoology
2904:
2279:Woolley, Tyler A. (1988).
1851:"All about oribatid mites"
822:Balogh & Mahunka, 1979
2476:
1989:10.11646/zootaxa.4783.1.1
1630:Archegozetes longisetosus
626:Balogh & Balogh, 1992
283:
276:
265:
258:
215:
208:
119:Scientific classification
117:
108:
99:
23:
2208:Oregon State University
2161:Luis S. SubĂas (2007).
1982:(1): zootaxa.4783.1.1.
1892:10.1073/pnas.0702851104
850:Norton & Metz, 1980
838:Norton & Metz, 1980
2198:Krantz, G. W. (1978).
1945:"Oribatei Dugès, 1833"
1687:. pp. 3463â3466.
2829:Paleobiology Database
2258:Collingwood, Victoria
2200:A Manual of Acarology
2085:Ecology and Evolution
2033:Ecology and Evolution
1637:Conoppia palmicinctum
2725:Fauna Europaea (new)
1523:Petrunkevitch, 1955
534:Parhypochthonioidea
524:& Mahunka, 1979
109:Unidentified mite (
2289:Wiley Interscience
2237:. pp. 1â960.
1855:A Chaos of Delight
835:Nehypochthonioidea
713:Sphaerochthoniidae
563:Elliptochthoniidae
547:Parhypochthoniidae
315:, in the "chewing
2870:
2869:
2816:Open Tree of Life
2613:Taxon identifiers
2604:
2603:
2600:
2599:
2584:Opilioacariformes
2474:
2473:
2325:978-8-301-09740-0
2298:978-0-471-04168-9
2271:978-0-643-06790-5
2217:978-0-88246-064-2
2097:10.1002/ece3.9036
2045:10.1002/ece3.7047
1886:(21): 8885â8890.
1802:978-94-007-7164-2
1729:978-1-4020-8258-0
1694:978-1-4020-6242-1
1621:Arthropods portal
1596:
1587:
1578:
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1524:
1515:
1503:
1486:
1477:
1468:
1459:
1450:
1441:
1432:
1423:
1414:
1405:
1396:
1387:
1378:
1369:
1360:
1351:
1342:
1338:Polypterozetoidea
1333:
1324:
1315:
1306:
1297:
1285:
1272:
1257:
1245:
1229:
1217:
1201:
1193:
1182:
1174:
1166:
1162:Thrypochthoniidae
1154:
1138:
1121:
1113:
1101:
1081:
1073:
1065:
1053:
1049:Euphthiracaroidea
1033:
1021:
1005:
987:
975:
955:
943:
923:
911:
891:
883:
871:
851:
847:Nehypochthoniidae
839:
823:
810:
795:
782:
778:Protoplophoroidea
764:
760:Phyllochthoniidae
756:
748:
733:
729:Atopochthonioidea
717:
709:
708:Lavoipierre, 1946
701:
693:
689:Heterochthoniidae
685:
672:
668:Cosmochthonioidea
656:
652:Brachychthoniidae
643:
639:Brachychthonoidea
627:
623:Arborichthoniidae
619:
611:
598:
582:
567:
559:
555:Gehypochthoniidae
551:
538:
525:
507:
494:
478:
465:
452:
393:poison dart frogs
290:
289:
204:
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2823:
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2795:NHMSYS0000865202
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2784:
2772:
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2758:
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2482:
2383:
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2343:
2334:
2329:
2302:
2275:
2262:CSIRO Publishing
2248:
2235:CSIRO Publishing
2221:
2202:(2nd ed.).
2186:
2185:
2183:
2181:
2175:
2168:
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2152:
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2018:
2017:
1991:
1967:
1961:
1960:
1940:
1934:
1928:
1922:
1921:
1911:
1871:
1865:
1864:
1862:
1861:
1847:
1834:
1833:
1824:(6): 1504â1511.
1813:
1807:
1806:
1786:
1780:
1779:
1769:
1760:(Extra): 3â305.
1745:
1739:
1738:
1737:
1736:
1705:
1699:
1698:
1683:(2nd ed.).
1669:
1623:
1618:
1617:
1594:
1585:
1576:
1574:Zetomotrichoidea
1567:
1558:
1549:
1540:
1531:
1529:Unduloribatoidea
1522:
1513:
1501:
1484:
1482:Cymbaeremaeoidea
1475:
1466:
1457:
1448:
1439:
1430:
1421:
1412:
1403:
1394:
1385:
1376:
1367:
1358:
1356:Charassobatoidea
1349:
1340:
1331:
1322:
1313:
1304:
1295:
1293:Hermannielloidea
1283:
1270:
1255:
1243:
1227:
1215:
1213:Nanhermannioidea
1199:
1188:
1180:
1172:
1164:
1149:
1136:
1119:
1111:
1099:
1079:
1077:Synichotritiidae
1071:
1069:Euphthiracaridae
1063:
1051:
1031:
1019:
1017:Mesoplophoroidea
1003:
985:
973:
953:
941:
939:Epilohmannioidea
921:
909:
889:
881:
869:
867:Perlohmannioidea
849:
837:
821:
808:
793:
791:Protoplophoridae
780:
762:
754:
752:Pterochthoniidae
746:
744:Atopochthoniidae
731:
715:
707:
705:Pediculochelidae
699:
697:Haplochthoniidae
691:
683:
681:Cosmochthoniidae
670:
654:
641:
625:
617:
609:
593:
580:
565:
557:
549:
536:
520:
505:
492:
476:
463:
450:
343:temperate forest
271:
268:
247:
199:
127:
126:
104:
94:
31:
30:Devonianâpresent
27:Temporal range:
21:
2903:
2902:
2898:
2897:
2896:
2894:
2893:
2892:
2888:Arachnid orders
2873:
2872:
2871:
2866:
2858:
2853:
2845:
2840:
2832:
2827:
2819:
2814:
2806:
2801:
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2762:
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2684:
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2671:
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2658:
2649:
2648:
2643:
2634:
2633:
2628:
2615:
2605:
2596:
2578:
2550:
2546:Nuttalliellidae
2520:
2483:
2470:
2447:
2424:
2372:
2361:
2326:
2305:
2299:
2278:
2272:
2251:
2245:
2224:
2218:
2197:
2194:
2192:Further reading
2189:
2179:
2177:
2176:on July 6, 2012
2173:
2166:
2160:
2159:
2155:
2148:
2135:
2134:
2130:
2078:
2077:
2070:
2026:
2025:
2021:
1969:
1968:
1964:
1942:
1941:
1937:
1929:
1925:
1873:
1872:
1868:
1859:
1857:
1849:
1848:
1837:
1830:10.1139/z91-210
1815:
1814:
1810:
1803:
1788:
1787:
1783:
1747:
1746:
1742:
1734:
1732:
1730:
1707:
1706:
1702:
1695:
1673:Marjorie A. Hoy
1671:
1670:
1657:
1653:
1619:
1612:
1609:
1604:
1577:Grandjean, 1934
1520:Phenopelopoidea
1514:Grandjean, 1931
1511:Licneremaeoidea
1502:Grandjean, 1954
1473:Ameronothroidea
1467:Grandjean, 1954
1458:Grandjean, 1954
1455:Tectocepheoidea
1404:Grandjean, 1954
1374:Zetorchestoidea
1368:Grandjean, 1936
1359:Grandjean, 1958
1341:Grandjean, 1959
1323:Grandjean, 1954
1320:Gymnodamaeoidea
1311:Plateremaeoidea
1296:Grandjean, 1934
1284:Grandjean, 1954
1225:Nanhermanniidae
1170:Malaconothridae
1137:Grandjean, 1969
1097:Phthiracaroidea
1064:Grandjean, 1954
1029:Mesoplophoridae
1004:Grandjean, 1967
951:Epilohmanniidae
922:Grandjean, 1931
910:Grandjean, 1931
907:Eulohmannioidea
890:Grandjean, 1958
887:Collohmanniidae
882:Grandjean, 1954
870:Grandjean, 1954
809:Grandjean, 1969
755:Grandjean, 1950
747:Grandjean, 1949
732:Grandjean, 1949
716:Grandjean, 1947
692:Grandjean, 1954
684:Grandjean, 1947
671:Grandjean, 1947
618:Grandjean, 1947
615:Eniochthoniidae
607:Hypochthoniidae
591:Hypochthonoidea
581:Grandjean, 1947
550:Grandjean, 1969
537:Grandjean, 1969
506:Grandjean, 1932
493:Grandjean, 1932
477:Grandjean, 1932
464:Grandjean, 1932
451:Grandjean, 1969
439:
419:Parthenogenetic
415:parthenogenetic
285:Cryptostigmata
269:
241:
198:
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
2901:
2899:
2891:
2890:
2885:
2875:
2874:
2868:
2867:
2865:
2864:
2851:
2838:
2825:
2812:
2799:
2786:
2773:
2760:
2747:
2734:
2721:
2712:Fauna Europaea
2708:
2695:
2682:
2669:
2656:
2641:
2625:
2623:
2617:
2616:
2611:
2602:
2601:
2598:
2597:
2595:
2594:
2592:Opilioacaridae
2588:
2586:
2580:
2579:
2577:
2576:
2571:
2566:
2560:
2558:
2552:
2551:
2549:
2548:
2543:
2538:
2532:
2530:
2522:
2521:
2519:
2518:
2513:
2508:
2502:
2500:
2491:
2489:Parasitiformes
2485:
2484:
2477:
2475:
2472:
2471:
2469:
2468:
2466:Sphaerolichida
2463:
2457:
2455:
2453:Trombidiformes
2449:
2448:
2446:
2445:
2440:
2434:
2432:
2430:Sarcoptiformes
2426:
2425:
2423:
2422:
2417:
2412:
2407:
2402:
2397:
2391:
2389:
2380:
2374:
2373:
2362:
2360:
2359:
2352:
2345:
2337:
2331:
2330:
2324:
2316:PWN + Elsevier
2312:Warsaw, Poland
2303:
2297:
2276:
2270:
2249:
2243:
2222:
2216:
2193:
2190:
2188:
2187:
2153:
2147:978-0896726208
2146:
2128:
2068:
2039:(1): 321â337.
2019:
1962:
1935:
1923:
1866:
1835:
1808:
1801:
1781:
1740:
1728:
1700:
1693:
1654:
1652:
1649:
1648:
1647:
1644:Neotrichozetes
1640:
1633:
1625:
1624:
1608:
1605:
1603:
1602:
1601:
1600:
1599:
1598:
1589:
1580:
1571:
1565:Ceratozetoidea
1562:
1556:Oribatelloidea
1553:
1547:Achipterioidea
1544:
1535:
1526:
1517:
1505:
1504:
1493:
1492:
1491:
1490:
1489:
1488:
1485:Sellnick, 1928
1479:
1476:Willmann, 1931
1470:
1461:
1452:
1443:
1434:
1425:
1422:Sellnick, 1937
1416:
1407:
1401:Amerobelboidea
1398:
1395:Oudemans, 1900
1389:
1386:Oudemans, 1900
1380:
1371:
1362:
1353:
1344:
1335:
1326:
1317:
1314:Trägürdh, 1926
1308:
1305:Sellnick, 1928
1299:
1287:
1286:
1275:
1274:
1263:
1262:
1261:
1260:
1259:
1258:
1256:Sellnick, 1928
1247:
1246:
1244:Sellnick, 1928
1235:
1234:
1233:
1232:
1231:
1230:
1228:Sellnick, 1928
1219:
1218:
1216:Sellnick, 1928
1207:
1206:
1205:
1204:
1203:
1202:
1194:
1183:
1175:
1167:
1165:Willmann, 1931
1156:
1155:
1141:
1140:
1129:
1128:
1127:
1126:
1125:
1124:
1123:
1122:
1120:NiedbaĹa, 1986
1117:Steganacaridae
1114:
1109:Phthiracaridae
1103:
1102:
1089:
1088:
1087:
1086:
1085:
1084:
1083:
1082:
1074:
1066:
1061:Oribotritiidae
1055:
1054:
1041:
1040:
1039:
1038:
1037:
1036:
1035:
1034:
1023:
1022:
1009:
1008:
1007:
1006:
995:
994:
993:
992:
991:
990:
989:
988:
977:
976:
963:
962:
961:
960:
959:
958:
957:
956:
954:Oudemans, 1923
945:
944:
942:Oudemans, 1923
931:
930:
929:
928:
927:
926:
925:
924:
919:Eulohmanniidae
913:
912:
899:
898:
897:
896:
895:
894:
893:
892:
884:
879:Perlohmaniidae
873:
872:
859:
858:
857:
856:
855:
854:
853:
852:
841:
840:
827:
826:
825:
824:
813:
812:
801:
800:
799:
798:
797:
796:
785:
784:
772:
771:
770:
769:
768:
767:
766:
765:
757:
749:
736:
735:
723:
722:
721:
720:
719:
718:
710:
702:
694:
686:
675:
674:
673:(c. 14 genera)
662:
661:
660:
659:
658:
657:
646:
645:
644:(c. 11 genera)
633:
632:
631:
630:
629:
628:
620:
612:
601:
600:
585:
584:
573:
572:
571:
570:
569:
568:
560:
558:Strenzke, 1963
552:
541:
540:
528:
527:
513:
512:
511:
510:
509:
508:
497:
496:
490:Palaeacaroidea
484:
483:
482:
481:
480:
479:
468:
467:
461:Acaronychoidea
455:
454:
443:
438:
435:
377:, but none is
324:Sarcoptiformes
297:oribatid mites
288:
287:
281:
280:
274:
273:
263:
262:
256:
255:
254:
253:
248:
236:
231:
226:
221:
213:
212:
206:
205:
193:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
115:
114:
111:Phthiracaridae
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
16:Order of mites
15:
13:
10:
9:
6:
4:
3:
2:
2900:
2889:
2886:
2884:
2881:
2880:
2878:
2861:
2856:
2852:
2848:
2843:
2839:
2835:
2830:
2826:
2822:
2817:
2813:
2809:
2804:
2800:
2796:
2791:
2787:
2783:
2778:
2774:
2770:
2765:
2761:
2757:
2752:
2748:
2744:
2739:
2735:
2731:
2726:
2722:
2718:
2713:
2709:
2705:
2700:
2696:
2692:
2687:
2683:
2679:
2674:
2670:
2666:
2661:
2657:
2652:
2646:
2642:
2637:
2631:
2627:
2626:
2624:
2622:
2618:
2614:
2609:
2593:
2590:
2589:
2587:
2585:
2581:
2575:
2572:
2570:
2567:
2565:
2564:Monogynaspida
2562:
2561:
2559:
2557:
2553:
2547:
2544:
2542:
2539:
2537:
2534:
2533:
2531:
2528:
2523:
2517:
2514:
2512:
2509:
2507:
2504:
2503:
2501:
2499:
2495:
2492:
2490:
2486:
2481:
2467:
2464:
2462:
2459:
2458:
2456:
2454:
2450:
2444:
2441:
2439:
2438:Endeostigmata
2436:
2435:
2433:
2431:
2427:
2421:
2420:Parhyposomata
2418:
2416:
2413:
2411:
2408:
2406:
2403:
2401:
2398:
2396:
2393:
2392:
2390:
2388:
2384:
2381:
2379:
2375:
2370:
2366:
2358:
2353:
2351:
2346:
2344:
2339:
2338:
2335:
2327:
2321:
2317:
2313:
2309:
2304:
2300:
2294:
2290:
2286:
2282:
2277:
2273:
2267:
2263:
2259:
2255:
2250:
2246:
2244:0-643-06658-6
2240:
2236:
2232:
2228:
2223:
2219:
2213:
2209:
2205:
2204:Corvallis, OR
2201:
2196:
2195:
2191:
2172:
2164:
2157:
2154:
2149:
2143:
2139:
2132:
2129:
2124:
2120:
2115:
2110:
2106:
2102:
2098:
2094:
2090:
2086:
2082:
2075:
2073:
2069:
2064:
2060:
2055:
2050:
2046:
2042:
2038:
2034:
2030:
2023:
2020:
2015:
2011:
2007:
2003:
1999:
1995:
1990:
1985:
1981:
1977:
1973:
1966:
1963:
1958:
1954:
1950:
1946:
1939:
1936:
1933:
1927:
1924:
1919:
1915:
1910:
1905:
1901:
1897:
1893:
1889:
1885:
1881:
1877:
1870:
1867:
1856:
1852:
1846:
1844:
1842:
1840:
1836:
1831:
1827:
1823:
1819:
1812:
1809:
1804:
1798:
1794:
1793:
1785:
1782:
1777:
1773:
1768:
1763:
1759:
1755:
1751:
1744:
1741:
1731:
1725:
1721:
1717:
1713:
1712:
1704:
1701:
1696:
1690:
1686:
1682:
1678:
1674:
1668:
1666:
1664:
1662:
1660:
1656:
1650:
1646:
1645:
1641:
1639:
1638:
1634:
1632:
1631:
1627:
1626:
1622:
1616:
1611:
1606:
1593:
1590:
1588:(19 families)
1584:
1581:
1575:
1572:
1566:
1563:
1557:
1554:
1548:
1545:
1539:
1538:Limnozetoidea
1536:
1530:
1527:
1521:
1518:
1512:
1509:
1508:
1507:
1506:
1500:
1497:
1496:
1495:
1494:
1483:
1480:
1474:
1471:
1465:
1464:Hydrozetoidea
1462:
1456:
1453:
1447:
1444:
1438:
1437:Otocepheoidea
1435:
1429:
1426:
1424:(12 families)
1420:
1417:
1411:
1408:
1406:(10 families)
1402:
1399:
1393:
1390:
1384:
1381:
1377:Michael, 1898
1375:
1372:
1366:
1365:Microzetoidea
1363:
1357:
1354:
1350:Berlese, 1896
1348:
1345:
1339:
1336:
1332:Berlese, 1896
1330:
1327:
1321:
1318:
1312:
1309:
1303:
1300:
1294:
1291:
1290:
1289:
1288:
1282:
1279:
1278:
1277:
1276:
1273:
1269:
1265:
1264:
1254:
1251:
1250:
1249:
1248:
1242:
1241:Hermannioidea
1239:
1238:
1237:
1236:
1226:
1223:
1222:
1221:
1220:
1214:
1211:
1210:
1209:
1208:
1200:Thorell, 1876
1198:
1195:
1191:
1187:
1184:
1181:Berlese, 1896
1179:
1176:
1173:Berlese, 1916
1171:
1168:
1163:
1160:
1159:
1158:
1157:
1152:
1148:
1145:
1144:
1143:
1142:
1139:
1135:
1131:
1130:
1118:
1115:
1110:
1107:
1106:
1105:
1104:
1098:
1095:
1094:
1093:
1092:
1091:
1090:
1078:
1075:
1070:
1067:
1062:
1059:
1058:
1057:
1056:
1050:
1047:
1046:
1045:
1044:
1043:
1042:
1030:
1027:
1026:
1025:
1024:
1018:
1015:
1014:
1013:
1012:
1011:
1010:
1002:
999:
998:
997:
996:
986:Berlese, 1916
984:
981:
980:
979:
978:
974:Berlese, 1916
972:
971:Lohmannioidea
969:
968:
967:
966:
965:
964:
952:
949:
948:
947:
946:
940:
937:
936:
935:
934:
933:
932:
920:
917:
916:
915:
914:
908:
905:
904:
903:
902:
901:
900:
888:
885:
880:
877:
876:
875:
874:
868:
865:
864:
863:
862:
861:
860:
848:
845:
844:
843:
842:
836:
833:
832:
831:
830:
829:
828:
820:
817:
816:
815:
814:
811:
807:
803:
802:
792:
789:
788:
787:
786:
783:(c. 7 genera)
779:
776:
775:
774:
773:
761:
758:
753:
750:
745:
742:
741:
740:
739:
738:
737:
730:
727:
726:
725:
724:
714:
711:
706:
703:
698:
695:
690:
687:
682:
679:
678:
677:
676:
669:
666:
665:
664:
663:
653:
650:
649:
648:
647:
640:
637:
636:
635:
634:
624:
621:
616:
613:
610:Berlese, 1910
608:
605:
604:
603:
602:
599:(c. 8 genera)
596:
592:
589:
588:
587:
586:
583:
579:
575:
574:
564:
561:
556:
553:
548:
545:
544:
543:
542:
535:
532:
531:
530:
529:
526:
523:
519:
518:Parhyposomata
515:
514:
504:
503:Palaeacaridae
501:
500:
499:
498:
491:
488:
487:
486:
485:
475:
474:Acaronychidae
472:
471:
470:
469:
462:
459:
458:
457:
456:
453:
449:
445:
444:
442:
436:
434:
432:
427:
425:
420:
416:
411:
407:
405:
401:
396:
394:
390:
386:
382:
380:
376:
372:
368:
364:
360:
356:
352:
348:
344:
340:
336:
332:
327:
325:
322:
318:
314:
310:
306:
302:
298:
294:
286:
282:
279:
275:
264:
261:
257:
252:
251:Parhyposomata
249:
245:
240:
237:
235:
232:
230:
227:
225:
222:
220:
217:
216:
214:
211:
207:
202:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
2620:
2574:Trigynaspida
2556:Mesostigmata
2511:Holothyridae
2506:Allothyridae
2415:Palaeosomata
2400:Enarthronota
2395:Brachypylina
2386:
2307:
2285:New York, NY
2280:
2253:
2226:
2199:
2178:. Retrieved
2171:the original
2156:
2137:
2131:
2088:
2084:
2036:
2032:
2022:
1979:
1975:
1965:
1951:. New York:
1948:
1938:
1926:
1883:
1879:
1869:
1858:. Retrieved
1854:
1821:
1817:
1811:
1791:
1784:
1757:
1753:
1743:
1733:, retrieved
1710:
1703:
1680:
1677:"Soil mites"
1642:
1635:
1628:
1597:(3 families)
1570:(5 families)
1561:(3 families)
1552:(2 families)
1543:(2 families)
1534:(3 families)
1516:(6 families)
1487:(3 families)
1478:(3 families)
1460:(2 families)
1451:(3 families)
1446:Carabodoidea
1442:(4 families)
1440:Balogh, 1961
1433:(6 families)
1415:(7 families)
1413:Balogh, 1961
1410:Eremelloidea
1397:(4 families)
1388:(8 families)
1383:Gustavioidea
1361:(3 families)
1352:(7 families)
1343:(2 families)
1325:(2 families)
1316:(4 families)
1302:Neoliodoidea
1298:(2 families)
1281:Pycnonoticae
1268:Brachypylina
1266:
1253:Hermanniidae
1147:Crotonioidea
1132:
1080:Walker, 1965
983:Lohmanniidae
804:
700:Hammen, 1959
578:Enarthronota
576:
566:Norton, 1975
516:
448:Palaeosomata
446:
440:
428:
412:
408:
397:
387:
383:
349:: prelarva,
328:
305:beetle mites
304:
300:
296:
292:
291:
284:
239:Palaeosomata
224:Enarthronota
219:Brachypylina
195:
182:Superorder:
18:
2883:Acariformes
2751:iNaturalist
2645:Wikispecies
2516:Neothyridae
2498:Holothyrida
2461:Prostigmata
2443:Psoroptidia
2378:Acariformes
1955:. pp.
1595:Jacot, 1925
1592:Galumnoidea
1586:Jacot, 1925
1583:Oripodoidea
1568:Jacot, 1925
1559:Jacot, 1925
1532:Kunst, 1971
1499:Poronoticae
1431:Ewing, 1917
1428:Trizetoidea
1392:Eremaeoidea
1197:Crotoniidae
1112:Perty, 1841
1100:Perty, 1841
1072:Jacot, 1930
1052:Jacot, 1930
1032:Ewing, 1917
1020:Ewing, 1917
819:Dichosomata
794:Ewing, 1917
781:Ewing, 1917
763:TravĂŠ, 1967
437:Systematics
431:Astigmatina
373:; some are
335:iteroparous
317:Acariformes
242: [
186:Acariformes
166:Chelicerata
162:Subphylum:
2877:Categories
2410:Mixonomata
2405:Holosomata
2180:January 5,
1860:2023-07-10
1735:2020-12-01
1651:References
1579:(1 family)
1550:Thor, 1929
1541:Thor, 1937
1525:(1 family)
1469:(1 family)
1449:Koch, 1837
1379:(1 family)
1370:(1 family)
1347:Cepheoidea
1334:(1 family)
1329:Damaeoidea
1307:(1 family)
1271:Hull, 1918
1186:Camisiidae
1134:Holosomata
1001:Euptyctima
806:Mixonomata
734:(3 genera)
655:Thor, 1934
642:Thor, 1934
539:(3 genera)
495:(8 genera)
466:(6 genera)
404:earthworms
365:material,
301:moss mites
270: 200
234:Mixonomata
229:Holosomata
156:Arthropoda
2665:Oribatida
2651:Oribatida
2621:Oribatida
2536:Argasidae
2525:Ixodida (
2387:Oribatida
2231:Melbourne
2105:2045-7758
2014:219739139
1998:1175-5334
1953:Macmillan
1900:0027-8424
1776:1989-953X
1754:Graellsia
1419:Oppioidea
1178:Nothridae
389:Alkaloids
379:parasitic
375:predatory
331:metabolic
307:, are an
293:Oribatida
260:Diversity
210:Suborders
196:Oribatida
176:Arachnida
142:Kingdom:
136:Eukaryota
24:Oribatida
2673:BugGuide
2636:Q1387882
2630:Wikidata
2541:Ixodidae
2123:35784052
2063:33437432
2006:33056509
1918:17502597
1685:Springer
1675:(2008).
1607:See also
1190:Oudemans
424:genotype
400:tapeworm
353:, three
278:Synonyms
152:Phylum:
146:Animalia
132:Domain:
2821:4683995
2743:4653212
2704:2587416
2363:Acari (
2114:9219104
2054:7790623
1976:Zootaxa
1909:1885597
1151:Thorell
595:Berlese
371:carrion
367:lichens
355:nymphal
347:instars
192:Order:
172:Class:
2860:390786
2834:155663
2782:733326
2756:332183
2569:Sejina
2322:
2295:
2268:
2241:
2214:
2144:
2121:
2111:
2103:
2061:
2051:
2012:
2004:
1996:
1916:
1906:
1898:
1799:
1774:
1726:
1691:
1192:, 1900
1153:, 1876
597:, 1910
522:Balogh
363:fungal
203:, 1833
2855:WoRMS
2842:Plazi
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