3234:
5366:
3227:
532:
1731:
2556:
2542:
866:
4818:
4022:
4804:
2535:
5514:
4811:
708:) would be a functional counterpart of the cutting edge of a beak. Thus, increasing the number of teeth would be an adaptation for cutting and ripping, as long as the space between adjacent teeth was preserved...while it would have the effect of working as a beak if spaces were filled with more teeth. The adaption to a cut-and-rip function therefore becomes an exaptation with a slicing effect, eventually leading to the cutting edge seen in most ornithomimosaurs."
1737:
3241:
139:
2549:
118:
5379:
4040:
4028:
4047:
5373:
4798:
497:
589:
4035:
528:, showing two different basic forms. The teeth in the front of the upper jaw were broad and D-shaped in cross-section, while those further back were blade-like, and on the whole the teeth in the upper jaw were larger than those in the lower. All of its teeth were unserrated, and had a constricted "waist" between the crown and the root. Interdental plates were lacking.
5526:
688:, "The phylogenetic hypothesis...supports an unexpected approach, involving exaptation, which might explain the evolutionary process towards the toothless condition in Ornithomimosauria. Until now, a progressive reduction in the number of teeth has been considered as the most likely explanation: the primitive
616:
might have been much like a modern-day crane, wading out in lakes or ponds using its claws and teeth to capture fish and then storing them in its skin flap. Some parts of the impressions revealed wrinkled skin, interpreted as lacking scales or feathers. Filament-like structures were also preserved;
511:
was a small ornithomimosaur, at about 1.9–2.5 m (6.2–8.2 ft) and 17–30 kg (37–66 lb). Its skull was unusually long and narrow, with a maximum length of about 4.5 times its maximum height. It was highly unusual among ornithomimosaurs in its large number of teeth: it had about 220
696:
hypothesis suggests an alternative evolutionary process based on a functional analysis of increasing numbers of teeth. A high number of teeth with enough interdental space and properly placed
604:
Soft-tissue remains preserved by the exceptional preservational environment of the La Hoyas lagerstätte revealed the presence of a small skin or keratin crest on the back of the head, and a
1103:
D. E. G. Briggs, P. R. Wilby, B. Pérez Pérez-Moreno, J. L. Sanz, M. Fregenal-Martinez, (1997). "The mineralization of dinosaur soft tissue in the Lower
Cretaceous of Las Hoyas, Spain."
480:. The only known specimen consists of the articulated front half of a skeleton and includes the skull, lower jaws, all the neck vertebrae and most of the back vertebrae, ribs,
488:, a complete right forelimb and most of the left forelimb. Remains of the soft parts are visible at the back of the skull, around the neck and around the front limbs.
5556:
5458:
585:
bones in the lower arm tightly adhered to each other. The hand was hook-like and had fingers of equal length equipped with rather straight claws.
3233:
5576:
3226:
1140:
Makovicky, P., Kobayashi, Y. and Currie, P. J. (2004). "Ornithomimosauria." pp. 137-150 in D. B. Weishampel, P. Dodson, et H. Osmólska (eds.),
5365:
1063:
672:. A study by Kobayashi and Lü in 2003 indicated that these two species formed a basal arrangement of steps leading towards the more advanced
962:
473:
1193:
5571:
2555:
704:) would be an adaptation for cutting and ripping. On the other hand, an excessive number of teeth with no interdental space (as in
641:
plates and uncinate processes, which imply flight musculature. These adaptations have been noted years later by the paleontologist
4021:
2541:
4803:
692:
theropods have up to 80 teeth with tall blade-like crowns, and the primitive ornithomimosaurs have only a few small teeth. The
1730:
1156:
Kobayashi, Y., and Lü, J.-C. (2003). "A new ornithomimid dinosaur with gregarious habits from the Late
Cretaceous of China."
2534:
4817:
1736:
5244:
4039:
5551:
3240:
138:
1129:
4810:
2548:
531:
5504:
5378:
383:
4046:
5566:
5561:
5546:
553:, which had far fewer (eleven total, and only in the lower jaw). The presence of such a large number of teeth in
1281:
697:
4027:
995:
Pérez-Moreno, B. P.; Sanz, J. L.; Buscalioni, A. D.; Moratalla, J. J.; Ortega, F.; Rasskin-Gutman, D. (1994).
5372:
1186:
418:
617:
first interpreted as an integument, some of these were later seen as representing preserved muscle fibers.
5398:
399:
375:
5484:
3752:
3664:
865:
421:
is a reference to the large number of teeth possessed by this theropod and is derived from Greek πολύς (
336:
289:
391:
4797:
5445:
5330:
3441:
3409:
1011:
642:
449:
266:
395:
5518:
3765:
3552:
876:
The Las Hoyas lagerstätte has produced numerous other exquisitely preserved species, including the
4034:
5436:
5237:
3741:
3601:
3457:
3397:
2832:
2742:
2625:
1489:
1179:
133:
5489:
5471:
5119:
5036:
5010:
4849:
4836:
4714:
4255:
4207:
3039:
2389:
1468:
1069:
1059:
722:
465:
387:
359:
241:
225:
5476:
573:
leading to the toothless cutting edge found in later ornithomimosaurs. The arms and hands of
5530:
5223:
4980:
4890:
4679:
4595:
4457:
4376:
4348:
4308:
4189:
4159:
3677:
3430:
3294:
3151:
3051:
2900:
2807:
2765:
2332:
1642:
1019:
656:
was by the describers assigned to the
Ornithomimosauria, in the basalmost position. A later
347:
117:
30:
5286:
5024:
4987:
4919:
4774:
4767:
4579:
4521:
4494:
4484:
4443:
4278:
4137:
4121:
3939:
3911:
3889:
3776:
3621:
3328:
3102:
2985:
2963:
2884:
2713:
2690:
2683:
2661:
2647:
2640:
2601:
2452:
2299:
2226:
2215:
2151:
2039:
1926:
1503:
1412:
1266:
1241:
950:
626:
485:
340:
1015:
5251:
5214:
5195:
5157:
5146:
4943:
4656:
4615:
4606:
4588:
4568:
4464:
4178:
3923:
3785:
3612:
3594:
3422:
3389:
3375:
3336:
3110:
2971:
2928:
2921:
2843:
2819:
2772:
2731:
2668:
2654:
2382:
2367:
2194:
2028:
1892:
1860:
1849:
1826:
1788:
1649:
1619:
1543:
1482:
1381:
877:
809:
469:
5540:
5293:
5100:
5079:
4958:
4951:
4878:
4672:
4645:
4503:
4419:
4397:
4390:
4383:
4326:
4243:
4106:
4078:
4065:
3989:
3866:
3706:
3540:
3471:
3357:
3344:
3089:
3067:
3060:
3024:
2997:
2949:
2873:
2609:
2400:
2374:
2290:
2271:
2264:
2201:
2180:
2159:
2144:
2130:
2089:
2055:
2048:
1967:
1819:
1706:
1699:
1626:
1596:
1585:
1475:
1446:
1439:
1365:
1120:
Dinosaurs of the Air: The
Evolution and Loss of Flight in Dinosaurs and Birds, pg 237
882:
637:
or be a recent descendant from a flying animal. This is due to the presence of large
582:
453:
68:
5265:
5203:
5173:
5093:
5086:
5064:
5056:
4936:
4928:
4908:
4623:
4534:
4472:
4362:
4333:
4285:
4114:
4099:
3874:
3806:
3734:
3713:
3692:
3508:
3464:
3272:
3259:
3189:
3142:
3017:
2907:
2859:
2852:
2789:
2758:
2751:
2632:
2459:
2311:
2208:
2110:
2103:
1998:
1982:
1918:
1869:
1801:
1578:
1461:
1454:
1425:
1388:
1357:
997:"A unique multitoothed ornithomimosaur dinosaur from the Lower Cretaceous of Spain"
907:
693:
673:
596:. The small crest in back of the head (colored in light grey) was probably made by
540:
496:
5430:
5319:
5300:
5230:
5180:
5166:
5108:
5049:
4994:
4871:
4665:
4541:
4510:
4426:
4412:
4144:
4092:
3960:
3882:
3845:
3727:
3699:
3652:
3572:
3450:
3307:
3158:
3131:
3031:
2978:
2914:
2893:
2866:
2617:
2430:
2423:
2409:
2235:
2082:
2073:
1975:
1946:
1876:
1841:
1834:
1634:
1612:
1605:
1564:
1556:
1532:
1517:
1496:
1404:
1396:
935:
785:
605:
588:
43:
915:
bones. Coarse sediments of the La Hoyas lagerstätte have produced bones of the
5258:
5187:
5132:
4753:
4552:
4369:
4355:
4340:
4292:
4226:
4152:
3974:
3967:
3953:
3859:
3792:
3720:
3684:
3637:
3579:
3565:
3532:
3524:
3517:
3498:
3321:
3286:
3074:
2675:
2466:
2416:
2353:
2324:
2279:
2257:
2250:
2242:
2166:
2062:
2021:
2014:
2006:
1990:
1910:
1781:
1657:
1510:
1373:
1232:
945:
940:
930:
916:
894:
761:
701:
570:
562:
549:
525:
513:
196:
88:
53:
5421:
1073:
5279:
5272:
4972:
4965:
4638:
4630:
4561:
4214:
3932:
3478:
3382:
3117:
2702:
2587:
2574:
2501:
2437:
2360:
2339:
2096:
1959:
1767:
1754:
1672:
1524:
1343:
1330:
1293:
1202:
921:
888:
711:
689:
681:
657:
565:
for cutting and ripping, a "functional counterpart of the cutting edge of a
557:, coupled with a lack of interdental space, was interpreted by Pérez-Moreno
477:
209:
150:
93:
37:
1039:
Récords y curiosidades de los dinosaurios Terópodos y otros dinosauromorfos
996:
668:
5415:
4863:
4271:
4264:
4236:
4129:
3629:
3010:
2346:
2173:
1934:
1571:
1250:
1226:
912:
902:
441:
363:
343:
183:
170:
83:
78:
63:
58:
48:
608:
similar to the much larger pouches found in modern pelicans, from which
5463:
4404:
4166:
3896:
3799:
3124:
2445:
638:
597:
521:
517:
481:
329:
325:
98:
73:
5072:
4760:
2187:
1220:
1053:
1023:
926:
634:
160:
5392:
3903:
3644:
3586:
864:
622:
587:
530:
495:
457:
407:
355:
351:
332:
5450:
1090:
Barsbold and Perle 1984 (Dinosauria; Theropoda) of
Mongolia." in
680:
has played an important and surprising role in understanding the
5336:
3946:
899:
578:
566:
417:, "mimic", in reference to the long snout and throat pouch. The
5396:
4834:
4063:
3257:
2572:
1752:
1328:
1213:
1175:
621:
was also the first ornithomimosaur discovered with a preserved
431:
423:
1171:
448:, part of the Las Hoyas Collection presently housed at the
547:
Only one other ornithomimosaur is known to possess teeth,
1094:
ed. Carpenter, K. 2005. Indiana
University Press:97-126.
1152:
1150:
1130:
990:
988:
986:
984:
982:
980:
978:
5502:
1086:
Kobayashi, Y., and
Barsbold, R. (2005). "Anatomy of
925:. The lagerstätte beds have also yielded remains of
5405:
5318:
5213:
5156:
5118:
5035:
5008:
4918:
4888:
4847:
4745:
4713:
4655:
4605:
4578:
4551:
4520:
4493:
4482:
4441:
4306:
4254:
4225:
4187:
4176:
4076:
3988:
3922:
3844:
3775:
3751:
3663:
3611:
3551:
3507:
3496:
3440:
3408:
3355:
3305:
3270:
3188:
3141:
3088:
3050:
2996:
2947:
2883:
2842:
2818:
2787:
2741:
2711:
2700:
2585:
2500:
2399:
2310:
2289:
2225:
2128:
2072:
2038:
1945:
1890:
1859:
1799:
1765:
1691:
1671:
1595:
1542:
1423:
1341:
1292:
1265:
1144:. 2nd ed. University of California Press, Berkeley.
625:apparatus (specialized tongue bones in the neck).
684:of the Ornithomimosauria. To quote Pérez-Moreno
577:were more typical of ornithomimosaurs, with the
382:site. In 1994 this was named and described by
1187:
8:
1105:Journal of the Geological Society of London
1058:. Princeton University Press. p. 128.
5393:
5124:
5041:
5016:
4900:
4896:
4855:
4844:
4831:
4742:
4710:
4526:
4490:
4449:
4318:
4314:
4199:
4195:
4184:
4084:
4073:
4060:
3851:
3841:
3757:
3669:
3557:
3504:
3414:
3367:
3363:
3313:
3278:
3267:
3254:
3185:
3094:
3002:
2955:
2824:
2799:
2795:
2723:
2719:
2708:
2593:
2582:
2569:
2497:
2316:
2307:
2136:
1951:
1902:
1898:
1811:
1807:
1773:
1762:
1749:
1548:
1431:
1349:
1338:
1325:
1271:
1262:
1210:
1194:
1180:
1172:
116:
20:
468:locality in Cuenca Province, Spain, from
933:, as well as of the unique early mammal
676:(see cladogram below). The discovery of
5509:
974:
872:(foreground) among contemporary animals
1055:The Princeton Field Guide to Dinosaurs
1037:Molina-Pérez & Larramendi (2016).
512:very small teeth in total, with seven
378:discovered a theropod skeleton at the
7:
5557:Early Cretaceous dinosaurs of Europe
963:Timeline of ornithomimosaur research
354:. It is notable for possessing more
801:
777:
753:
729:
717:
406:. The generic name is derived from
14:
474:Calizas de La Huérguina Formation
366:), most of which were toothless.
5524:
5512:
5377:
5371:
5364:
4816:
4809:
4802:
4796:
4045:
4038:
4033:
4026:
4020:
3239:
3232:
3225:
2554:
2547:
2540:
2533:
1735:
1729:
137:
1041:. Spain: Larousse. p. 268.
911:remains, and a few fragmentary
592:Illustration of the skull from
714:after Kobayashi and Lü, 2003:
1:
5577:Fossil taxa described in 1994
1158:Acta Palaeontologica Polonica
384:Bernardino Pérez Pérez-Moreno
358:than any other member of the
432:
424:
633:might have been capable of
5593:
520:, and seventy-five in the
16:Extinct genus of dinosaurs
5361:
5127:
5044:
5019:
4903:
4899:
4858:
4843:
4830:
4794:
4741:
4709:
4529:
4452:
4321:
4317:
4202:
4198:
4087:
4072:
4059:
4018:
3854:
3840:
3760:
3672:
3560:
3417:
3370:
3366:
3316:
3281:
3266:
3253:
3222:
3184:
3097:
3005:
2958:
2827:
2802:
2798:
2726:
2722:
2596:
2581:
2568:
2530:
2496:
2319:
2139:
1954:
1905:
1901:
1814:
1810:
1776:
1761:
1748:
1726:
1551:
1434:
1352:
1337:
1324:
1274:
1261:
1209:
1092:The Carnivorous Dinosaurs
1052:Paul, Gregory S. (2016).
806:
799:
782:
775:
758:
751:
734:
727:
295:
288:
134:Scientific classification
132:
124:
115:
23:
398:, Francisco Ortega and
5572:La Huérguina Formation
1088:Harpymimus okladnikovi
873:
664:(2005) confirmed that
660:analysis by Makovicky
601:
544:
505:
504:, compared to a human.
464:was recovered at the
404:Pelecanimimus polyodon
302:Pelecanimimus polyodon
5485:Paleobiology Database
3753:Carcharodontosaurinae
3665:Carcharodontosauridae
868:
591:
534:
499:
429:), "many" and ὀδούς (
4746:Dubious coelurosaurs
3442:Metriacanthosaurinae
3410:Metriacanthosauridae
1692:Dubious neotheropods
629:has speculated that
516:teeth, about thirty
476:dating to the Lower
400:Diego Rasskin-Gutman
376:Armando Díaz Romeral
370:Discovery and naming
5552:Feathered dinosaurs
3766:Carcharodontosaurus
3553:Carcharodontosauria
1016:1994Natur.370..363P
5238:Archaeornithomimus
3742:Veterupristisaurus
3602:Veterupristisaurus
3458:Metriacanthosaurus
3398:Piatnitzkysauridae
2833:Eustreptospondylus
2743:Piatnitzkysauridae
2626:Chuandongocoelurus
1490:Notatesseraeraptor
874:
602:
545:
506:
500:Estimated size of
402:as a new species:
341:ornithomimosaurian
5500:
5499:
5472:Open Tree of Life
5399:Taxon identifiers
5390:
5389:
5386:
5385:
5359:
5358:
5355:
5354:
5351:
5350:
5347:
5346:
5314:
5313:
5310:
5309:
5120:Macrocheiriformes
5037:Ornithomimosauria
5011:Maniraptoriformes
5004:
5003:
4850:Maniraptoromorpha
4837:Maniraptoromorpha
4826:
4825:
4792:
4791:
4788:
4787:
4784:
4783:
4737:
4736:
4733:
4732:
4715:Maniraptoromorpha
4705:
4704:
4701:
4700:
4697:
4696:
4693:
4692:
4689:
4688:
4437:
4436:
4302:
4301:
4256:Proceratosauridae
4208:Chingkankousaurus
4055:
4054:
4016:
4015:
4012:
4011:
4008:
4007:
3984:
3983:
3836:
3835:
3832:
3831:
3828:
3827:
3824:
3823:
3820:
3819:
3816:
3815:
3492:
3491:
3488:
3487:
3249:
3248:
3220:
3219:
3216:
3215:
3212:
3211:
3208:
3207:
3180:
3179:
3176:
3175:
3172:
3171:
3168:
3167:
3084:
3083:
3040:Vallibonavenatrix
2943:
2942:
2939:
2938:
2783:
2782:
2564:
2563:
2528:
2527:
2524:
2523:
2520:
2519:
2492:
2491:
2488:
2487:
2484:
2483:
2480:
2479:
2476:
2475:
2124:
2123:
2120:
2119:
1886:
1885:
1744:
1743:
1724:
1723:
1720:
1719:
1716:
1715:
1667:
1666:
1320:
1319:
1316:
1315:
1312:
1311:
1065:978-1-78684-190-2
1010:(6488): 363−367.
898:, along with non-
857:
856:
848:
847:
839:
838:
830:
829:
821:
820:
723:Ornithomimosauria
569:," as well as an
524:. The teeth were
413:, "pelican", and
392:Angela Buscalioni
360:Ornithomimosauria
317:
316:
269:
242:Macrocheiriformes
226:Ornithomimosauria
5584:
5567:Fossils of Spain
5562:Cretaceous Spain
5547:Ornithomimosaurs
5529:
5528:
5527:
5517:
5516:
5515:
5508:
5493:
5492:
5480:
5479:
5467:
5466:
5454:
5453:
5441:
5440:
5439:
5426:
5425:
5424:
5394:
5381:
5375:
5368:
5224:Aepyornithomimus
5125:
5042:
5017:
4981:Sinocalliopteryx
4901:
4897:
4891:Neocoelurosauria
4856:
4845:
4832:
4820:
4813:
4806:
4800:
4743:
4711:
4680:Zhuchengtyrannus
4596:Thanatotheristes
4527:
4491:
4458:Appalachiosaurus
4450:
4319:
4315:
4309:Pantyrannosauria
4200:
4196:
4190:Tyrannosauroidea
4185:
4160:Xinjiangovenator
4085:
4074:
4061:
4049:
4042:
4037:
4030:
4024:
3852:
3842:
3758:
3678:Acrocanthosaurus
3670:
3558:
3505:
3431:Yangchuanosaurus
3415:
3368:
3364:
3314:
3295:Lourinhanosaurus
3279:
3268:
3255:
3243:
3236:
3229:
3186:
3152:Sigilmassasaurus
3095:
3052:Ceratosuchopsini
3003:
2956:
2901:Dubreuillosaurus
2825:
2808:Streptospondylus
2800:
2796:
2766:Piatnitzkysaurus
2724:
2720:
2709:
2594:
2583:
2570:
2558:
2551:
2544:
2537:
2498:
2333:Ekrixinatosaurus
2317:
2308:
2137:
1952:
1903:
1899:
1812:
1808:
1774:
1763:
1750:
1739:
1733:
1643:Procompsognathus
1549:
1432:
1350:
1339:
1326:
1272:
1263:
1256:
1255:
1211:
1196:
1189:
1182:
1173:
1161:
1154:
1145:
1138:
1132:
1127:
1121:
1118:
1112:
1101:
1095:
1084:
1078:
1077:
1049:
1043:
1042:
1034:
1028:
1027:
1024:10.1038/370363a0
1001:
992:
953:are also known.
802:
778:
754:
730:
718:
538:being chased by
472:beds within the
435:
427:
380:Las Hoyas Unit 3
348:Early Cretaceous
304:
300:
281:P. polyodon
260:
253:
240:
224:
142:
141:
120:
110:
40:
31:Early Cretaceous
29:Temporal range:
21:
5592:
5591:
5587:
5586:
5585:
5583:
5582:
5581:
5537:
5536:
5535:
5525:
5523:
5513:
5511:
5503:
5501:
5496:
5488:
5483:
5475:
5470:
5462:
5457:
5449:
5444:
5435:
5434:
5429:
5420:
5419:
5414:
5401:
5391:
5382:
5369:
5343:
5306:
5287:Sinornithomimus
5209:
5152:
5114:
5031:
5025:Compsognathidae
5013:
5000:
4988:Sinosauropteryx
4920:Compsognathidae
4914:
4893:
4884:
4852:
4839:
4822:
4821:
4807:
4780:
4775:Shanyangosaurus
4768:Phaedrolosaurus
4729:
4685:
4651:
4601:
4580:Daspletosaurini
4574:
4547:
4522:Tyrannosaurinae
4516:
4495:Albertosaurinae
4487:
4485:Tyrannosauridae
4478:
4446:
4444:Eutyrannosauria
4433:
4311:
4298:
4279:Proceratosaurus
4250:
4221:
4192:
4181:
4172:
4138:Richardoestesia
4122:Chilantaisaurus
4081:
4068:
4051:
4050:
4031:
4004:
3980:
3940:Australovenator
3918:
3912:Chilantaisaurus
3890:Phuwiangvenator
3812:
3777:Giganotosaurini
3771:
3747:
3659:
3622:Chilantaisaurus
3607:
3547:
3501:
3484:
3436:
3404:
3360:
3351:
3329:Monolophosaurus
3310:
3301:
3275:
3262:
3245:
3244:
3230:
3204:
3164:
3137:
3103:Camarillasaurus
3080:
3046:
2992:
2986:Ostafrikasaurus
2964:Camarillasaurus
2952:
2935:
2885:Afrovenatorinae
2879:
2838:
2814:
2792:
2779:
2737:
2716:
2714:Megalosauroidea
2705:
2696:
2691:Vectaerovenator
2684:Szechuanosaurus
2662:Monolophosaurus
2648:Kaijiangosaurus
2641:Cryolophosaurus
2602:Calamospondylus
2590:
2577:
2560:
2559:
2552:
2545:
2538:
2516:
2472:
2453:Pycnonemosaurus
2395:
2306:
2300:Majungasaurinae
2285:
2227:Majungasaurinae
2221:
2216:Xenotarsosaurus
2152:Dryptosauroides
2133:
2116:
2068:
2040:Elaphrosaurinae
2034:
1941:
1927:Ornithomimoides
1895:
1882:
1855:
1804:
1795:
1770:
1757:
1740:
1734:
1712:
1687:
1663:
1591:
1538:
1504:Shuangbaisaurus
1428:
1419:
1346:
1333:
1308:
1288:
1282:Avemetatarsalia
1267:Avemetatarsalia
1257:
1242:Avemetatarsalia
1216:
1215:
1205:
1200:
1170:
1165:
1164:
1155:
1148:
1139:
1135:
1128:
1124:
1119:
1115:
1102:
1098:
1085:
1081:
1066:
1051:
1050:
1046:
1036:
1035:
1031:
999:
994:
993:
976:
971:
959:
951:crocodylomorphs
878:enantiornithine
863:
858:
849:
840:
831:
822:
651:
643:Mickey Mortimer
627:Gregory S. Paul
612:took its name.
494:
486:pectoral girdle
450:Museo de Cuenca
372:
313:
306:
298:
297:
284:
259:
251:
238:
222:
136:
125:Restoration of
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
5590:
5588:
5580:
5579:
5574:
5569:
5564:
5559:
5554:
5549:
5539:
5538:
5534:
5533:
5521:
5498:
5497:
5495:
5494:
5481:
5468:
5455:
5442:
5427:
5411:
5409:
5403:
5402:
5397:
5388:
5387:
5384:
5383:
5363:
5362:
5360:
5357:
5356:
5353:
5352:
5349:
5348:
5345:
5344:
5342:
5341:
5340:
5339:
5333:
5324:
5322:
5316:
5315:
5312:
5311:
5308:
5307:
5305:
5304:
5297:
5290:
5283:
5276:
5269:
5262:
5255:
5252:Dromiceiomimus
5248:
5241:
5234:
5227:
5219:
5217:
5215:Ornithomimidae
5211:
5210:
5208:
5207:
5200:
5196:Paraxenisaurus
5192:
5184:
5177:
5170:
5162:
5160:
5158:Deinocheiridae
5154:
5153:
5151:
5150:
5147:Shenzhousaurus
5143:
5136:
5128:
5122:
5116:
5115:
5113:
5112:
5105:
5097:
5090:
5083:
5076:
5069:
5061:
5053:
5045:
5039:
5033:
5032:
5030:
5029:
5020:
5014:
5009:
5006:
5005:
5002:
5001:
4999:
4998:
4991:
4984:
4977:
4969:
4962:
4955:
4948:
4944:Beipiaognathus
4940:
4933:
4924:
4922:
4916:
4915:
4913:
4912:
4904:
4894:
4889:
4886:
4885:
4883:
4882:
4875:
4868:
4859:
4853:
4848:
4841:
4840:
4835:
4828:
4827:
4824:
4823:
4815:
4801:
4795:
4793:
4790:
4789:
4786:
4785:
4782:
4781:
4779:
4778:
4771:
4764:
4757:
4749:
4747:
4739:
4738:
4735:
4734:
4731:
4730:
4728:
4727:
4726:
4725:
4719:
4717:
4707:
4706:
4703:
4702:
4699:
4698:
4695:
4694:
4691:
4690:
4687:
4686:
4684:
4683:
4676:
4669:
4661:
4659:
4657:Tyrannosaurini
4653:
4652:
4650:
4649:
4642:
4635:
4627:
4620:
4616:Bistahieversor
4611:
4609:
4607:Teratophoneini
4603:
4602:
4600:
4599:
4592:
4589:Daspletosaurus
4584:
4582:
4576:
4575:
4573:
4572:
4569:Qianzhousaurus
4565:
4557:
4555:
4549:
4548:
4546:
4545:
4538:
4530:
4524:
4518:
4517:
4515:
4514:
4507:
4499:
4497:
4488:
4483:
4480:
4479:
4477:
4476:
4469:
4465:Bistahieversor
4461:
4453:
4447:
4442:
4439:
4438:
4435:
4434:
4432:
4431:
4423:
4416:
4409:
4401:
4394:
4387:
4380:
4373:
4366:
4359:
4352:
4345:
4337:
4330:
4322:
4312:
4307:
4304:
4303:
4300:
4299:
4297:
4296:
4289:
4282:
4275:
4268:
4260:
4258:
4252:
4251:
4249:
4248:
4240:
4232:
4230:
4223:
4222:
4220:
4219:
4211:
4203:
4193:
4188:
4182:
4179:Tyrannoraptora
4177:
4174:
4173:
4171:
4170:
4163:
4156:
4149:
4141:
4134:
4126:
4118:
4111:
4103:
4096:
4088:
4082:
4077:
4070:
4069:
4064:
4057:
4056:
4053:
4052:
4044:
4025:
4019:
4017:
4014:
4013:
4010:
4009:
4006:
4005:
4003:
4002:
4001:
4000:
3994:
3992:
3986:
3985:
3982:
3981:
3979:
3978:
3971:
3964:
3957:
3950:
3943:
3936:
3928:
3926:
3924:Megaraptoridae
3920:
3919:
3917:
3916:
3908:
3900:
3893:
3886:
3879:
3871:
3863:
3855:
3849:
3838:
3837:
3834:
3833:
3830:
3829:
3826:
3825:
3822:
3821:
3818:
3817:
3814:
3813:
3811:
3810:
3803:
3796:
3789:
3786:Giganotosaurus
3781:
3779:
3773:
3772:
3770:
3769:
3761:
3755:
3749:
3748:
3746:
3745:
3738:
3731:
3724:
3717:
3710:
3703:
3696:
3689:
3681:
3673:
3667:
3661:
3660:
3658:
3657:
3649:
3641:
3634:
3626:
3617:
3615:
3613:Neovenatoridae
3609:
3608:
3606:
3605:
3598:
3595:Ulughbegsaurus
3591:
3583:
3576:
3569:
3561:
3555:
3549:
3548:
3546:
3545:
3537:
3529:
3521:
3513:
3511:
3502:
3497:
3494:
3493:
3490:
3489:
3486:
3485:
3483:
3482:
3475:
3468:
3461:
3454:
3446:
3444:
3438:
3437:
3435:
3434:
3427:
3423:Xuanhanosaurus
3418:
3412:
3406:
3405:
3403:
3402:
3394:
3390:Xuanhanosaurus
3386:
3379:
3376:Asfaltovenator
3371:
3361:
3356:
3353:
3352:
3350:
3349:
3341:
3337:Megalosauridae
3333:
3325:
3317:
3311:
3306:
3303:
3302:
3300:
3299:
3291:
3282:
3276:
3271:
3264:
3263:
3258:
3251:
3250:
3247:
3246:
3238:
3224:
3223:
3221:
3218:
3217:
3214:
3213:
3210:
3209:
3206:
3205:
3203:
3202:
3201:
3200:
3194:
3192:
3182:
3181:
3178:
3177:
3174:
3173:
3170:
3169:
3166:
3165:
3163:
3162:
3155:
3147:
3145:
3139:
3138:
3136:
3135:
3128:
3121:
3114:
3111:Ichthyovenator
3107:
3098:
3092:
3086:
3085:
3082:
3081:
3079:
3078:
3071:
3064:
3056:
3054:
3048:
3047:
3045:
3044:
3036:
3028:
3021:
3014:
3006:
3000:
2994:
2993:
2991:
2990:
2982:
2975:
2972:Cristatusaurus
2968:
2959:
2953:
2948:
2945:
2944:
2941:
2940:
2937:
2936:
2934:
2933:
2929:Poekilopleuron
2925:
2922:Piveteausaurus
2918:
2911:
2904:
2897:
2889:
2887:
2881:
2880:
2878:
2877:
2870:
2863:
2856:
2848:
2846:
2844:Megalosaurinae
2840:
2839:
2837:
2836:
2828:
2822:
2820:Megalosauridae
2816:
2815:
2813:
2812:
2803:
2793:
2788:
2785:
2784:
2781:
2780:
2778:
2777:
2773:Xuanhanosaurus
2769:
2762:
2755:
2747:
2745:
2739:
2738:
2736:
2735:
2732:Yunyangosaurus
2727:
2717:
2712:
2706:
2701:
2698:
2697:
2695:
2694:
2687:
2680:
2672:
2669:Pandoravenator
2665:
2658:
2655:Kayentavenator
2651:
2644:
2637:
2629:
2622:
2614:
2606:
2597:
2591:
2586:
2579:
2578:
2573:
2566:
2565:
2562:
2561:
2553:
2546:
2539:
2532:
2531:
2529:
2526:
2525:
2522:
2521:
2518:
2517:
2515:
2514:
2513:
2512:
2506:
2504:
2494:
2493:
2490:
2489:
2486:
2485:
2482:
2481:
2478:
2477:
2474:
2473:
2471:
2470:
2463:
2456:
2449:
2442:
2434:
2427:
2420:
2413:
2405:
2403:
2397:
2396:
2394:
2393:
2386:
2383:Skorpiovenator
2379:
2371:
2364:
2357:
2350:
2343:
2336:
2329:
2320:
2314:
2305:
2304:
2295:
2293:
2287:
2286:
2284:
2283:
2276:
2268:
2261:
2254:
2247:
2239:
2231:
2229:
2223:
2222:
2220:
2219:
2212:
2205:
2198:
2195:Spectrovenator
2191:
2184:
2177:
2170:
2163:
2156:
2148:
2140:
2134:
2129:
2126:
2125:
2122:
2121:
2118:
2117:
2115:
2114:
2107:
2100:
2093:
2086:
2078:
2076:
2070:
2069:
2067:
2066:
2059:
2052:
2044:
2042:
2036:
2035:
2033:
2032:
2029:Spinostropheus
2025:
2018:
2011:
2003:
1995:
1987:
1979:
1972:
1964:
1955:
1949:
1943:
1942:
1940:
1939:
1931:
1923:
1915:
1906:
1896:
1893:Abelisauroidea
1891:
1888:
1887:
1884:
1883:
1881:
1880:
1873:
1865:
1863:
1861:Ceratosauridae
1857:
1856:
1854:
1853:
1850:Saltriovenator
1846:
1838:
1831:
1827:Fosterovenator
1823:
1815:
1805:
1800:
1797:
1796:
1794:
1793:
1789:Lophostropheus
1785:
1777:
1771:
1766:
1759:
1758:
1753:
1746:
1745:
1742:
1741:
1728:
1727:
1725:
1722:
1721:
1718:
1717:
1714:
1713:
1711:
1710:
1703:
1695:
1693:
1689:
1688:
1686:
1685:
1684:
1683:
1677:
1675:
1669:
1668:
1665:
1664:
1662:
1661:
1654:
1650:Pterospondylus
1646:
1639:
1631:
1623:
1620:Lucianovenator
1616:
1609:
1601:
1599:
1593:
1592:
1590:
1589:
1582:
1575:
1568:
1561:
1552:
1546:
1544:Coelophysoidea
1540:
1539:
1537:
1536:
1529:
1521:
1514:
1507:
1500:
1493:
1486:
1483:Lophostropheus
1479:
1472:
1465:
1458:
1451:
1443:
1435:
1429:
1424:
1421:
1420:
1418:
1417:
1409:
1401:
1393:
1385:
1382:Erythrovenator
1378:
1370:
1362:
1353:
1347:
1342:
1335:
1334:
1329:
1322:
1321:
1318:
1317:
1314:
1313:
1310:
1309:
1307:
1306:
1305:
1304:
1298:
1296:
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1287:
1286:
1285:
1284:
1275:
1269:
1259:
1258:
1254:
1253:
1244:
1235:
1229:
1223:
1214:
1207:
1206:
1201:
1199:
1198:
1191:
1184:
1176:
1169:
1168:External links
1166:
1163:
1162:
1146:
1142:The Dinosauria
1133:
1122:
1113:
1096:
1079:
1064:
1044:
1029:
973:
972:
970:
967:
966:
965:
958:
955:
862:
859:
855:
854:
851:
850:
846:
845:
842:
841:
837:
836:
833:
832:
828:
827:
824:
823:
819:
818:
815:
814:
810:Ornithomimidae
805:
800:
798:
795:
794:
791:
790:
781:
776:
774:
771:
770:
767:
766:
757:
752:
750:
747:
746:
743:
742:
733:
728:
726:
716:
650:
649:Classification
647:
493:
490:
396:José Moratalla
388:José Luis Sanz
371:
368:
362:(or any other
339:("primitive")
328:mimic") is an
315:
314:
307:
293:
292:
286:
285:
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6:
4:
3:
2:
5589:
5578:
5575:
5573:
5570:
5568:
5565:
5563:
5560:
5558:
5555:
5553:
5550:
5548:
5545:
5544:
5542:
5532:
5522:
5520:
5510:
5506:
5491:
5486:
5482:
5478:
5473:
5469:
5465:
5460:
5456:
5452:
5447:
5443:
5438:
5437:Pelecanimimus
5432:
5428:
5423:
5417:
5413:
5412:
5410:
5408:
5407:Pelecanimimus
5404:
5400:
5395:
5380:
5376:
5374:
5367:
5338:
5334:
5332:
5328:
5327:
5326:
5325:
5323:
5321:
5317:
5303:
5302:
5298:
5296:
5295:
5294:Struthiomimus
5291:
5289:
5288:
5284:
5282:
5281:
5277:
5275:
5274:
5270:
5268:
5267:
5263:
5261:
5260:
5256:
5254:
5253:
5249:
5247:
5246:
5245:"Coelosaurus"
5242:
5240:
5239:
5235:
5233:
5232:
5228:
5226:
5225:
5221:
5220:
5218:
5216:
5212:
5206:
5205:
5201:
5198:
5197:
5193:
5190:
5189:
5185:
5183:
5182:
5178:
5176:
5175:
5171:
5169:
5168:
5164:
5163:
5161:
5159:
5155:
5149:
5148:
5144:
5142:
5141:
5140:Pelecanimimus
5137:
5135:
5134:
5130:
5129:
5126:
5123:
5121:
5117:
5111:
5110:
5106:
5103:
5102:
5101:Thecocoelurus
5098:
5096:
5095:
5091:
5089:
5088:
5084:
5082:
5081:
5080:Kinnareemimus
5077:
5075:
5074:
5070:
5067:
5066:
5062:
5059:
5058:
5054:
5052:
5051:
5047:
5046:
5043:
5040:
5038:
5034:
5027:
5026:
5022:
5021:
5018:
5015:
5012:
5007:
4997:
4996:
4992:
4990:
4989:
4985:
4983:
4982:
4978:
4975:
4974:
4970:
4968:
4967:
4963:
4961:
4960:
4959:Huaxiagnathus
4956:
4954:
4953:
4952:Compsognathus
4949:
4946:
4945:
4941:
4939:
4938:
4934:
4931:
4930:
4926:
4925:
4923:
4921:
4917:
4911:
4910:
4906:
4905:
4902:
4898:
4895:
4892:
4887:
4881:
4880:
4879:Ornitholestes
4876:
4874:
4873:
4869:
4866:
4865:
4861:
4860:
4857:
4854:
4851:
4846:
4842:
4838:
4833:
4829:
4819:
4814:
4812:
4805:
4799:
4777:
4776:
4772:
4770:
4769:
4765:
4763:
4762:
4758:
4756:
4755:
4751:
4750:
4748:
4744:
4740:
4723:
4722:
4721:
4720:
4718:
4716:
4712:
4708:
4682:
4681:
4677:
4675:
4674:
4673:Tyrannosaurus
4670:
4668:
4667:
4663:
4662:
4660:
4658:
4654:
4648:
4647:
4646:Teratophoneus
4643:
4641:
4640:
4636:
4633:
4632:
4628:
4626:
4625:
4621:
4618:
4617:
4613:
4612:
4610:
4608:
4604:
4598:
4597:
4593:
4591:
4590:
4586:
4585:
4583:
4581:
4577:
4571:
4570:
4566:
4564:
4563:
4559:
4558:
4556:
4554:
4550:
4544:
4543:
4539:
4537:
4536:
4532:
4531:
4528:
4525:
4523:
4519:
4513:
4512:
4508:
4506:
4505:
4504:Albertosaurus
4501:
4500:
4498:
4496:
4492:
4489:
4486:
4481:
4475:
4474:
4470:
4467:
4466:
4462:
4460:
4459:
4455:
4454:
4451:
4448:
4445:
4440:
4429:
4428:
4424:
4422:
4421:
4420:Xiongguanlong
4417:
4415:
4414:
4410:
4407:
4406:
4402:
4400:
4399:
4398:Suskityrannus
4395:
4393:
4392:
4391:Stokesosaurus
4388:
4386:
4385:
4384:Santanaraptor
4381:
4379:
4378:
4374:
4372:
4371:
4367:
4365:
4364:
4360:
4358:
4357:
4353:
4351:
4350:
4346:
4343:
4342:
4338:
4336:
4335:
4331:
4329:
4328:
4327:Alectrosaurus
4324:
4323:
4320:
4316:
4313:
4310:
4305:
4295:
4294:
4290:
4288:
4287:
4283:
4281:
4280:
4276:
4274:
4273:
4269:
4267:
4266:
4262:
4261:
4259:
4257:
4253:
4246:
4245:
4244:Tanycolagreus
4241:
4239:
4238:
4234:
4233:
4231:
4228:
4224:
4217:
4216:
4212:
4210:
4209:
4205:
4204:
4201:
4197:
4194:
4191:
4186:
4183:
4180:
4175:
4169:
4168:
4164:
4162:
4161:
4157:
4155:
4154:
4150:
4147:
4146:
4142:
4140:
4139:
4135:
4132:
4131:
4127:
4124:
4123:
4119:
4117:
4116:
4112:
4109:
4108:
4107:Bahariasaurus
4104:
4102:
4101:
4097:
4095:
4094:
4090:
4089:
4086:
4083:
4080:
4079:Coelurosauria
4075:
4071:
4067:
4066:Coelurosauria
4062:
4058:
4048:
4043:
4041:
4036:
4029:
4023:
3998:
3997:
3996:
3995:
3993:
3991:
3990:Coelurosauria
3987:
3977:
3976:
3972:
3970:
3969:
3965:
3963:
3962:
3958:
3956:
3955:
3951:
3949:
3948:
3944:
3942:
3941:
3937:
3935:
3934:
3930:
3929:
3927:
3925:
3921:
3915:
3913:
3909:
3907:
3905:
3901:
3899:
3898:
3894:
3892:
3891:
3887:
3885:
3884:
3880:
3877:
3876:
3872:
3869:
3868:
3867:Bahariasaurus
3864:
3862:
3861:
3857:
3856:
3853:
3850:
3847:
3843:
3839:
3809:
3808:
3804:
3802:
3801:
3797:
3795:
3794:
3790:
3788:
3787:
3783:
3782:
3780:
3778:
3774:
3768:
3767:
3763:
3762:
3759:
3756:
3754:
3750:
3744:
3743:
3739:
3737:
3736:
3732:
3730:
3729:
3725:
3723:
3722:
3718:
3716:
3715:
3711:
3709:
3708:
3707:Kelmayisaurus
3704:
3702:
3701:
3697:
3695:
3694:
3690:
3687:
3686:
3682:
3680:
3679:
3675:
3674:
3671:
3668:
3666:
3662:
3655:
3654:
3650:
3647:
3646:
3642:
3640:
3639:
3635:
3632:
3631:
3627:
3624:
3623:
3619:
3618:
3616:
3614:
3610:
3604:
3603:
3599:
3597:
3596:
3592:
3589:
3588:
3584:
3582:
3581:
3577:
3575:
3574:
3570:
3568:
3567:
3563:
3562:
3559:
3556:
3554:
3550:
3543:
3542:
3541:Saurophaganax
3538:
3535:
3534:
3530:
3527:
3526:
3522:
3520:
3519:
3515:
3514:
3512:
3510:
3506:
3503:
3500:
3495:
3481:
3480:
3476:
3474:
3473:
3472:Siamotyrannus
3469:
3467:
3466:
3462:
3460:
3459:
3455:
3453:
3452:
3448:
3447:
3445:
3443:
3439:
3433:
3432:
3428:
3425:
3424:
3420:
3419:
3416:
3413:
3411:
3407:
3400:
3399:
3395:
3392:
3391:
3387:
3385:
3384:
3380:
3378:
3377:
3373:
3372:
3369:
3365:
3362:
3359:
3358:Allosauroidea
3354:
3347:
3346:
3345:Spinosauridae
3342:
3339:
3338:
3334:
3331:
3330:
3326:
3324:
3323:
3319:
3318:
3315:
3312:
3309:
3304:
3297:
3296:
3292:
3289:
3288:
3284:
3283:
3280:
3277:
3274:
3269:
3265:
3261:
3256:
3252:
3242:
3237:
3235:
3228:
3198:
3197:
3196:
3195:
3193:
3191:
3187:
3183:
3161:
3160:
3156:
3154:
3153:
3149:
3148:
3146:
3144:
3140:
3134:
3133:
3129:
3127:
3126:
3122:
3120:
3119:
3115:
3113:
3112:
3108:
3105:
3104:
3100:
3099:
3096:
3093:
3091:
3090:Spinosaurinae
3087:
3077:
3076:
3072:
3070:
3069:
3068:Riparovenator
3065:
3063:
3062:
3061:Ceratosuchops
3058:
3057:
3055:
3053:
3049:
3042:
3041:
3037:
3034:
3033:
3029:
3027:
3026:
3025:Riojavenatrix
3022:
3020:
3019:
3015:
3013:
3012:
3008:
3007:
3004:
3001:
2999:
2998:Baryonychinae
2995:
2988:
2987:
2983:
2981:
2980:
2976:
2974:
2973:
2969:
2966:
2965:
2961:
2960:
2957:
2954:
2951:
2950:Spinosauridae
2946:
2931:
2930:
2926:
2924:
2923:
2919:
2917:
2916:
2912:
2910:
2909:
2905:
2903:
2902:
2898:
2896:
2895:
2891:
2890:
2888:
2886:
2882:
2876:
2875:
2874:Wiehenvenator
2871:
2869:
2868:
2864:
2862:
2861:
2857:
2855:
2854:
2850:
2849:
2847:
2845:
2841:
2835:
2834:
2830:
2829:
2826:
2823:
2821:
2817:
2810:
2809:
2805:
2804:
2801:
2797:
2794:
2791:
2786:
2775:
2774:
2770:
2768:
2767:
2763:
2761:
2760:
2756:
2754:
2753:
2749:
2748:
2746:
2744:
2740:
2734:
2733:
2729:
2728:
2725:
2721:
2718:
2715:
2710:
2707:
2704:
2699:
2693:
2692:
2688:
2686:
2685:
2681:
2678:
2677:
2673:
2671:
2670:
2666:
2664:
2663:
2659:
2657:
2656:
2652:
2650:
2649:
2645:
2643:
2642:
2638:
2635:
2634:
2630:
2628:
2627:
2623:
2620:
2619:
2615:
2612:
2611:
2610:Chienkosaurus
2607:
2604:
2603:
2599:
2598:
2595:
2592:
2589:
2584:
2580:
2576:
2571:
2567:
2557:
2550:
2543:
2536:
2510:
2509:
2508:
2507:
2505:
2503:
2499:
2495:
2469:
2468:
2464:
2462:
2461:
2457:
2455:
2454:
2450:
2448:
2447:
2443:
2440:
2439:
2435:
2433:
2432:
2428:
2426:
2425:
2421:
2419:
2418:
2414:
2412:
2411:
2407:
2406:
2404:
2402:
2401:Furileusauria
2398:
2392:
2391:
2387:
2385:
2384:
2380:
2377:
2376:
2375:Rahiolisaurus
2372:
2370:
2369:
2365:
2363:
2362:
2358:
2356:
2355:
2351:
2349:
2348:
2344:
2342:
2341:
2337:
2335:
2334:
2330:
2327:
2326:
2322:
2321:
2318:
2315:
2313:
2309:
2302:
2301:
2297:
2296:
2294:
2292:
2291:Carnotaurinae
2288:
2282:
2281:
2277:
2274:
2273:
2272:Rahiolisaurus
2269:
2267:
2266:
2265:Majungasaurus
2262:
2260:
2259:
2255:
2253:
2252:
2248:
2245:
2244:
2240:
2238:
2237:
2233:
2232:
2230:
2228:
2224:
2218:
2217:
2213:
2211:
2210:
2206:
2204:
2203:
2202:Tarascosaurus
2199:
2197:
2196:
2192:
2190:
2189:
2185:
2183:
2182:
2178:
2176:
2175:
2171:
2169:
2168:
2164:
2162:
2161:
2160:Eoabelisaurus
2157:
2154:
2153:
2149:
2147:
2146:
2145:Chenanisaurus
2142:
2141:
2138:
2135:
2132:
2131:Abelisauridae
2127:
2113:
2112:
2108:
2106:
2105:
2101:
2099:
2098:
2094:
2092:
2091:
2090:Masiakasaurus
2087:
2085:
2084:
2080:
2079:
2077:
2075:
2071:
2065:
2064:
2060:
2058:
2057:
2056:Huinculsaurus
2053:
2051:
2050:
2049:Elaphrosaurus
2046:
2045:
2043:
2041:
2037:
2031:
2030:
2026:
2024:
2023:
2019:
2017:
2016:
2012:
2009:
2008:
2004:
2001:
2000:
1996:
1993:
1992:
1988:
1985:
1984:
1980:
1978:
1977:
1973:
1970:
1969:
1968:Austrocheirus
1965:
1962:
1961:
1957:
1956:
1953:
1950:
1948:
1944:
1937:
1936:
1932:
1929:
1928:
1924:
1921:
1920:
1916:
1913:
1912:
1908:
1907:
1904:
1900:
1897:
1894:
1889:
1879:
1878:
1874:
1872:
1871:
1867:
1866:
1864:
1862:
1858:
1852:
1851:
1847:
1844:
1843:
1839:
1837:
1836:
1832:
1829:
1828:
1824:
1822:
1821:
1820:Berberosaurus
1817:
1816:
1813:
1809:
1806:
1803:
1798:
1791:
1790:
1786:
1784:
1783:
1779:
1778:
1775:
1772:
1769:
1764:
1760:
1756:
1751:
1747:
1738:
1732:
1709:
1708:
1707:Halticosaurus
1704:
1702:
1701:
1700:Teinurosaurus
1697:
1696:
1694:
1690:
1681:
1680:
1679:
1678:
1676:
1674:
1670:
1660:
1659:
1655:
1652:
1651:
1647:
1645:
1644:
1640:
1637:
1636:
1632:
1629:
1628:
1627:Megapnosaurus
1624:
1622:
1621:
1617:
1615:
1614:
1610:
1608:
1607:
1603:
1602:
1600:
1598:
1597:Coelophysidae
1594:
1588:
1587:
1586:Powellvenator
1583:
1581:
1580:
1576:
1574:
1573:
1569:
1567:
1566:
1562:
1559:
1558:
1554:
1553:
1550:
1547:
1545:
1541:
1535:
1534:
1530:
1527:
1526:
1522:
1520:
1519:
1515:
1513:
1512:
1508:
1506:
1505:
1501:
1499:
1498:
1494:
1492:
1491:
1487:
1485:
1484:
1480:
1478:
1477:
1476:Liliensternus
1473:
1471:
1470:
1466:
1464:
1463:
1459:
1457:
1456:
1452:
1449:
1448:
1447:Dolichosuchus
1444:
1442:
1441:
1440:Dilophosaurus
1437:
1436:
1433:
1430:
1427:
1422:
1415:
1414:
1413:Herrerasauria
1410:
1407:
1406:
1402:
1399:
1398:
1394:
1391:
1390:
1386:
1384:
1383:
1379:
1376:
1375:
1371:
1368:
1367:
1366:Daemonosaurus
1363:
1360:
1359:
1355:
1354:
1351:
1348:
1345:
1340:
1336:
1332:
1327:
1323:
1302:
1301:
1300:
1299:
1297:
1295:
1291:
1283:
1279:
1278:
1277:
1276:
1273:
1270:
1268:
1264:
1260:
1252:
1248:
1245:
1243:
1239:
1236:
1234:
1230:
1228:
1224:
1222:
1218:
1217:
1212:
1208:
1204:
1197:
1192:
1190:
1185:
1183:
1178:
1177:
1174:
1167:
1159:
1153:
1151:
1147:
1143:
1137:
1134:
1131:
1126:
1123:
1117:
1114:
1110:
1106:
1100:
1097:
1093:
1089:
1083:
1080:
1075:
1071:
1067:
1061:
1057:
1056:
1048:
1045:
1040:
1033:
1030:
1025:
1021:
1017:
1013:
1009:
1005:
998:
991:
989:
987:
985:
983:
981:
979:
975:
968:
964:
961:
960:
956:
954:
952:
948:
947:
942:
938:
937:
932:
928:
924:
923:
918:
914:
910:
909:
904:
901:
897:
896:
891:
890:
885:
884:
883:Iberomesornis
879:
871:
870:Pelecanimimus
867:
860:
853:
852:
844:
843:
835:
834:
826:
825:
817:
816:
813:
812:
811:
804:
803:
797:
796:
793:
792:
789:
788:
787:
780:
779:
773:
772:
769:
768:
765:
764:
763:
756:
755:
749:
748:
745:
744:
741:
740:
739:
738:Pelecanimimus
732:
731:
725:
724:
720:
719:
715:
713:
709:
707:
706:Pelecanimimus
703:
699:
695:
691:
687:
683:
679:
678:Pelecanimimus
675:
674:ornithomimids
671:
667:
666:Pelecanimimus
663:
659:
655:
654:Pelecanimimus
648:
646:
644:
640:
636:
632:
631:Pelecanimimus
628:
624:
620:
619:Pelecanimimus
615:
614:Pelecanimimus
611:
610:Pelecanimimus
607:
599:
595:
594:Pelecanimimus
590:
586:
584:
580:
576:
575:Pelecanimimus
572:
568:
564:
560:
556:
555:Pelecanimimus
552:
551:
543:
542:
537:
536:Pelecanimimus
533:
529:
527:
523:
519:
515:
510:
509:Pelecanimimus
503:
502:Pelecanimimus
498:
491:
489:
487:
483:
479:
475:
471:
467:
463:
462:Pelecanimimus
459:
455:
451:
447:
443:
438:
436:
434:
428:
426:
420:
419:specific name
416:
412:
409:
405:
401:
397:
393:
389:
385:
381:
377:
374:In July 1993
369:
367:
365:
361:
357:
353:
349:
345:
342:
338:
334:
331:
327:
323:
322:
321:Pelecanimimus
311:
308:Perez-Moreno
305:
303:
294:
291:
290:Binomial name
287:
283:
282:
276:
273:
272:
268:
264:
261:Perez-Moreno
258:
257:
256:Pelecanimimus
250:
247:
246:
243:
237:
234:
231:
230:
227:
221:
218:
215:
214:
211:
208:
205:
202:
201:
198:
195:
192:
189:
188:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Pelecanimimus
22:
19:
5519:Paleontology
5406:
5370:
5299:
5292:
5285:
5278:
5271:
5266:Ornithomimus
5264:
5257:
5250:
5243:
5236:
5229:
5222:
5204:Tyrannomimus
5202:
5194:
5186:
5179:
5174:Deinocheirus
5172:
5165:
5145:
5139:
5138:
5131:
5107:
5099:
5094:Nqwebasaurus
5092:
5087:Nedcolbertia
5085:
5078:
5071:
5065:Calamosaurus
5063:
5057:Aviatyrannis
5055:
5048:
5023:
4993:
4986:
4979:
4971:
4964:
4957:
4950:
4942:
4937:Aristosuchus
4935:
4929:Aniksosaurus
4927:
4909:Aniksosaurus
4907:
4877:
4870:
4862:
4808:
4773:
4766:
4759:
4752:
4678:
4671:
4664:
4644:
4637:
4629:
4624:Dynamoterror
4622:
4614:
4594:
4587:
4567:
4560:
4540:
4535:Asiatyrannus
4533:
4509:
4502:
4473:Dryptosaurus
4471:
4463:
4456:
4425:
4418:
4411:
4403:
4396:
4389:
4382:
4375:
4368:
4363:Jinbeisaurus
4361:
4354:
4347:
4339:
4334:Aviatyrannis
4332:
4325:
4291:
4286:Sinotyrannus
4284:
4277:
4270:
4263:
4242:
4235:
4213:
4206:
4165:
4158:
4151:
4143:
4136:
4128:
4120:
4115:Bicentenaria
4113:
4105:
4100:Asiamericana
4098:
4091:
4032:
3973:
3966:
3959:
3952:
3945:
3938:
3931:
3910:
3902:
3895:
3888:
3881:
3875:Deltadromeus
3873:
3865:
3858:
3807:Tyrannotitan
3805:
3798:
3791:
3784:
3764:
3740:
3735:Taurovenator
3733:
3726:
3719:
3714:Lajasvenator
3712:
3705:
3698:
3693:Concavenator
3691:
3683:
3676:
3651:
3643:
3636:
3628:
3620:
3600:
3593:
3585:
3578:
3571:
3564:
3539:
3531:
3523:
3516:
3509:Allosauridae
3477:
3470:
3465:Shidaisaurus
3463:
3456:
3449:
3429:
3421:
3396:
3388:
3381:
3374:
3343:
3335:
3327:
3320:
3293:
3285:
3273:Avetheropoda
3260:Avetheropoda
3231:
3190:Avetheropoda
3157:
3150:
3143:Spinosaurini
3130:
3123:
3116:
3109:
3101:
3073:
3066:
3059:
3038:
3030:
3023:
3018:Protathlitis
3016:
3009:
2984:
2977:
2970:
2962:
2927:
2920:
2913:
2908:Leshansaurus
2906:
2899:
2892:
2872:
2865:
2860:Megalosaurus
2858:
2853:Duriavenator
2851:
2831:
2806:
2790:Megalosauria
2771:
2764:
2759:Marshosaurus
2757:
2752:Condorraptor
2750:
2730:
2689:
2682:
2674:
2667:
2660:
2653:
2646:
2639:
2633:Cruxicheiros
2631:
2624:
2616:
2608:
2600:
2465:
2460:Quilmesaurus
2458:
2451:
2444:
2436:
2429:
2422:
2415:
2408:
2388:
2381:
2373:
2366:
2359:
2352:
2345:
2338:
2331:
2323:
2312:Brachyrostra
2298:
2278:
2270:
2263:
2256:
2249:
2241:
2234:
2214:
2209:Tralkasaurus
2207:
2200:
2193:
2186:
2179:
2172:
2165:
2158:
2150:
2143:
2111:Vespersaurus
2109:
2104:Velocisaurus
2102:
2095:
2088:
2081:
2061:
2054:
2047:
2027:
2020:
2013:
2005:
1999:Deltadromeus
1997:
1989:
1983:Compsosuchus
1981:
1974:
1966:
1958:
1933:
1925:
1919:Lametasaurus
1917:
1909:
1875:
1870:Ceratosaurus
1868:
1848:
1840:
1833:
1825:
1818:
1802:Ceratosauria
1787:
1780:
1705:
1698:
1656:
1648:
1641:
1633:
1625:
1618:
1611:
1604:
1584:
1579:Podokesaurus
1577:
1570:
1563:
1555:
1531:
1523:
1516:
1509:
1502:
1495:
1488:
1481:
1474:
1467:
1462:Gojirasaurus
1460:
1455:Dracovenator
1453:
1445:
1438:
1426:Neotheropoda
1411:
1403:
1395:
1389:Guaibasaurus
1387:
1380:
1372:
1364:
1358:Chindesaurus
1356:
1246:
1237:
1157:
1141:
1136:
1125:
1116:
1108:
1104:
1099:
1091:
1087:
1082:
1054:
1047:
1038:
1032:
1007:
1003:
944:
934:
920:
908:Concavenator
906:
893:
887:
881:
875:
869:
861:Paleoecology
808:
807:
784:
783:
760:
759:
737:
736:
735:
721:
710:
705:
694:phylogenetic
685:
677:
669:
665:
661:
653:
652:
630:
618:
613:
609:
603:
593:
574:
558:
554:
548:
546:
541:Concavenator
539:
535:
514:premaxillary
508:
507:
501:
461:
446:LH 7777
445:
439:
430:
422:
414:
410:
403:
379:
373:
320:
319:
318:
309:
301:
296:
280:
279:
262:
255:
254:
232:
216:
203:
190:
177:
126:
24:
18:
5431:Wikispecies
5331:Maniraptora
5320:Maniraptora
5301:Tototlmimus
5231:Anserimimus
5181:Garudimimus
5167:Beishanlong
5109:Valdoraptor
5050:Arkansaurus
4995:Xunmenglong
4872:Juravenator
4666:Tarbosaurus
4542:Nanuqsaurus
4511:Gorgosaurus
4427:Megaraptora
4413:Timurlengia
4145:Sciurumimus
4093:Aratasaurus
3961:Murusraptor
3883:Fukuiraptor
3846:Megaraptora
3728:Shaochilong
3700:Eocarcharia
3653:Megaraptora
3573:Lusovenator
3451:Alpkarakush
3308:Carnosauria
3159:Spinosaurus
3132:Siamosaurus
3032:Suchosaurus
2979:Iberospinus
2915:Magnosaurus
2894:Afrovenator
2867:Torvosaurus
2618:Chilesaurus
2431:Carnotaurus
2424:Caletodraco
2410:Abelisaurus
2236:Arcovenator
2083:Laevisuchus
2074:Noasaurinae
1976:Berthasaura
1947:Noasauridae
1877:Genyodectes
1842:Lukousaurus
1835:Jubbulpuria
1635:Panguraptor
1613:Coelophysis
1606:Camposaurus
1565:Panguraptor
1557:Dracoraptor
1533:Zupaysaurus
1518:Tachiraptor
1497:Sarcosaurus
1397:Nhandumirim
1160:48:235-259.
936:Spinolestes
931:salamanders
786:Garudimimus
702:troodontids
606:gular pouch
492:Description
470:lagerstätte
437:) "tooth".
127:P. polyodon
5541:Categories
5259:Gallimimus
5188:Harpymimus
5133:Harpymimus
4754:Iliosuchus
4724:see below↓
4553:Alioramini
4370:Juratyrant
4356:Eotyrannus
4341:Bagaraatan
4293:Yutyrannus
4227:Coeluridae
4153:Vayuraptor
3999:see below↓
3975:Tratayenia
3968:Orkoraptor
3954:Megaraptor
3860:Aoniraptor
3793:Mapusaurus
3721:Sauroniops
3685:Altispinax
3638:Neovenator
3580:Siamraptor
3566:Datanglong
3533:Epanterias
3525:Antrodemus
3518:Allosaurus
3499:Allosauria
3322:Altispinax
3287:Gasosaurus
3199:see below↓
3075:Suchomimus
2676:Sinosaurus
2511:see below↓
2467:Viavenator
2417:Aucasaurus
2354:Ilokelesia
2325:Dahalokely
2280:Rajasaurus
2258:Indosaurus
2251:Genusaurus
2243:Dahalokely
2167:Indosuchus
2063:Limusaurus
2022:Ligabueino
2015:Kiyacursor
2007:Genusaurus
1991:Dahalokely
1911:Betasuchus
1782:Dornraptor
1682:see below↓
1658:Segisaurus
1511:Sinosaurus
1374:Eodromaeus
1303:see below↓
1251:Dinosauria
1233:Sauropsida
1111:: 587-588.
969:References
946:Europejara
941:pterosaurs
939:. Several
917:ornithopod
895:Eoalulavis
762:Harpymimus
670:Harpymimus
571:exaptation
563:adaptation
550:Harpymimus
526:heterodont
444:specimen,
324:(meaning "
197:Saurischia
184:Dinosauria
5531:Dinosaurs
5335:includes
5280:Rativates
5273:Qiupalong
4973:Scipionyx
4966:Mirischia
4639:Lythronax
4631:Labocania
4562:Alioramus
4215:Labocania
3933:Aerosteon
3479:Sinraptor
3383:Erectopus
3118:Irritator
2703:Orionides
2588:Tetanurae
2575:Tetanurae
2502:Tetanurae
2438:Elemgasem
2361:Llukalkan
2340:Elemgasem
2097:Noasaurus
1960:Afromimus
1768:Averostra
1755:Averostra
1673:Averostra
1525:Velocipes
1344:Theropoda
1331:Theropoda
1294:Theropoda
1219:Kingdom:
1203:Theropoda
1074:985402380
922:Iguanodon
919:dinosaur
889:Concornis
712:Cladogram
698:denticles
690:tetanuran
682:evolution
658:cladistic
518:maxillary
478:Barremian
411:pelecanus
346:from the
274:Species:
210:Theropoda
157:Kingdom:
151:Eukaryota
36:130
5451:47458714
5416:Wikidata
4864:Coelurus
4272:Kileskus
4265:Guanlong
4237:Coelurus
4130:Gualicho
3630:Gualicho
3011:Baryonyx
2347:Guemesia
2174:Kryptops
1935:Ozraptor
1572:Pendraig
1227:Chordata
1225:Phylum:
1221:Animalia
957:See also
913:sauropod
903:theropod
466:La Hoyas
442:holotype
364:theropod
344:dinosaur
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
5505:Portals
5477:4127289
5464:4822643
5422:Q134151
4405:Timimus
4167:Zuolong
3897:Rapator
3800:Meraxes
3125:Oxalaia
2446:Koleken
1469:Lepidus
1231:Class:
1012:Bibcode
927:lizards
905:teeth,
700:(as in
639:sternal
598:keratin
522:dentary
482:sternum
330:extinct
326:pelican
299:†
278:†
248:Genus:
105:↓
5073:Hexing
4761:Kakuru
4349:Dilong
2390:Thanos
2368:Niebla
2188:Rugops
2181:Kurupi
1072:
1062:
1004:Nature
892:, and
880:birds
686:et al.
662:et al.
635:flight
583:radius
561:as an
559:et al.
484:, the
454:Cuenca
312:, 1994
310:et al.
263:et al.
5490:64117
5337:birds
4377:Moros
3904:Siats
3645:Siats
3587:Siats
1247:Clade
1238:Clade
1000:(PDF)
943:like
900:avian
623:hyoid
460:, of
458:Spain
433:odous
425:polys
415:mimus
408:Latin
356:teeth
352:Spain
337:basal
333:genus
233:Clade
217:Clade
204:Clade
191:Clade
178:Clade
5459:GBIF
5329:see
3947:Maip
1405:Tawa
1280:see
1070:OCLC
1060:ISBN
949:and
929:and
581:and
579:ulna
567:beak
440:The
267:1994
44:PreꞒ
5446:EoL
1109:154
1020:doi
1008:370
350:of
335:of
5543::
5487::
5474::
5461::
5448::
5433::
5418::
1249::
1240::
1149:^
1107:,
1068:.
1018:.
1006:.
1002:.
977:^
886:,
645:.
456:,
452:,
394:,
390:,
386:,
265:,
235::
219::
206::
193::
180::
94:Pg
38:Ma
33:,
5507::
5199:?
5191:?
5104:?
5068:?
5060:?
5028:?
4976:?
4947:?
4932:?
4867:?
4634:?
4619:?
4468:?
4430:?
4408:?
4344:?
4247:?
4229:?
4218:?
4148:?
4133:?
4125:?
4110:?
3914:?
3906:?
3878:?
3870:?
3848:?
3688:?
3656:?
3648:?
3633:?
3625:?
3590:?
3544:?
3536:?
3528:?
3426:?
3401:?
3393:?
3348:?
3340:?
3332:?
3298:?
3290:?
3106:?
3043:?
3035:?
2989:?
2967:?
2932:?
2811:?
2776:?
2679:?
2636:?
2621:?
2613:?
2605:?
2441:?
2378:?
2328:?
2303:?
2275:?
2246:?
2155:?
2010:?
2002:?
1994:?
1986:?
1971:?
1963:?
1938:?
1930:?
1922:?
1914:?
1845:?
1830:?
1792:?
1653:?
1638:?
1630:?
1560:?
1528:?
1450:?
1416:?
1408:?
1400:?
1392:?
1377:?
1369:?
1361:?
1195:e
1188:t
1181:v
1076:.
1026:.
1022::
1014::
600:.
252:†
239:†
223:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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