1780:
146:
132:
1557:
1588:. The afterfeathers are downy plumes that attach directly to the main feathers and were once believed to account for the bird's ability to conserve heat when under water; the plumules are small down feathers that attach directly to the skin, and are much more dense in penguins than other birds; lastly the filoplumes are small (less than 1 cm long) naked shafts that end in a splay of fibers— filoplumes were believed to give flying birds a sense of where their plumage was and whether or not it needed preening, so their presence in penguins may seem inconsistent, but penguins also preen extensively.
1461:, which live in the North Pacific and North Atlantic, developed similar characteristics to survive in the Arctic and sub-Arctic environments. Like the penguins, puffins have a white chest, black back and short stubby wings providing excellent swimming ability in icy water. But, unlike penguins, puffins can fly, as flightless birds would not survive alongside land-based predators such as polar bears and foxes; there are no such predators in the Antarctic. Their similarities indicate that similar environments, although at great distances, can result in similar evolutionary developments, i.e.
162:
627:
1170:, 19 high-coverage genomes that, together with two previously published genomes, encompass all extant penguin species have been sequenced. The origin of the Spheniscinae lies probably in the latest Paleogene and, geographically, it must have been much the same as the general area in which the order evolved: the oceans between the Australia-New Zealand region and the Antarctic. Presumably diverging from other penguins around 40 mya, it seems that the Spheniscinae were for quite some time limited to their ancestral area, as the well-researched deposits of the
1687:
1372:
924:
176:
199:
1483:
1596:
penguins, even those that live in warmer climates, have a counter-current heat exchanger called the humeral plexus. The flippers of penguins have at least three branches of the axillary artery, which allows cold blood to be heated by blood that has already been warmed and limits heat loss from the flippers. This system allows penguins to efficiently use their body heat and explains why such small animals can survive in the extreme cold.
8259:
8283:
1627:
4156:; Mirarab, S.; Aberer, A. J.; Li, B.; Houde, P.; Li, C.; Ho, S. Y. W.; Faircloth, B. C.; Nabholz, B.; Howard, J. T.; Suh, A.; Weber, C. C.; Da Fonseca, R. R.; Li, J.; Zhang, F.; Li, H.; Zhou, L.; Narula, N.; Liu, L.; Ganapathy, G.; Boussau, B.; Bayzid, M. S.; Zavidovych, V.; Subramanian, S.; Gabaldon, T.; Capella-Gutierrez, S.; Huerta-Cepas, J.; Rekepalli, B.; Munch, K.; et al. (2014).
2283:
2305:
1474:
8295:
394:
8271:
1826:
and 16% of the weight of a penguin egg, presumably to reduce the effects of dehydration and to minimize the risk of breakage in an adverse nesting environment. The yolk, too, is large and comprises 22–31% of the egg. Some yolk often remains when a chick is born, and is thought to help sustain the chick if the parents are delayed in returning with food.
1751:
2366:
1534:
usually dive deep; they catch their prey near the surface in dives that normally last only one or two minutes. Larger penguins can dive deep in case of need. Emperor penguins are the world's deepest-diving birds. They can dive to depths of approximately 550 meters (1,800 feet) while searching for food.
2347:, it was suffering from heat exhaustion and had to undergo a number of operations to remove objects like driftwood and sand from its stomach. Happy Feet was a media sensation, with extensive coverage on TV and the web, including a live stream that had thousands of views and a visit from English actor
1829:
When emperor penguin mothers lose a chick, they sometimes attempt to "steal" another mother's chick, usually unsuccessfully as other females in the vicinity assist the defending mother in keeping her chick. In some species, such as emperor and king penguins, the chicks assemble in large groups called
1825:
Penguin eggs are smaller than any other bird species when compared proportionally to the weight of the parent birds; at 52 g (2 oz), the little penguin egg is 4.7% of its mothers' weight, and the 450 g (1 lb) emperor penguin egg is 2.3%. The relatively thick shell forms between 10
1802:
Penguins for the most part breed in large colonies, the exceptions being the yellow-eyed and
Fiordland species; these colonies may range in size from as few as 100 pairs for gentoo penguins to several hundred thousand in the case of king, macaroni and chinstrap penguins. Living in colonies results in
1591:
The emperor penguin has the largest body mass of all penguins, which further reduces relative surface area and heat loss. They also are able to control blood flow to their extremities, reducing the amount of blood that gets cold, but still keeping the extremities from freezing. In the extreme cold of
1187:
appears to be the basalmost divergence among living penguins. They have bright yellow-orange neck, breast, and bill patches; incubate by placing their eggs on their feet, and when they hatch the chicks are almost naked. This genus has a distribution centred on the
Antarctic coasts and barely extends
4239:
Li, C.; Zhang, Y.; Li, J.; Kong, L.; Hu, H.; Pan, H.; Xu, L.; Deng, Y.; Li, Q.; Jin, L.; Yu, H.; Chen, Y.; Liu, B.; Yang, L.; Liu, S.; Zhang, Y.; Lang, Y.; Xia, J.; He, W.; Shi, Q.; Subramanian, S.; Millar, C. D.; Meader, S.; Rands, C. M.; Fujita, M. K.; Greenwold, M. J.; Castoe, T. A.; Pollock, D.
1537:
Penguins either waddle on their feet or slide on their bellies across the snow while using their feet to propel and steer themselves, a movement called "tobogganing", which conserves energy while moving quickly. They also jump with both feet together if they want to move more quickly or cross steep
1698:
where larger-bodied populations live at higher latitudes than smaller-bodied populations. There is some disagreement about this and several other authors have noted that there are fossil penguin species that contradict this hypothesis and that ocean currents and upwellings are likely to have had a
1319:
some 35 million years later. With habitat on the
Antarctic coasts declining, by the Priabonian more hospitable conditions for most penguins existed in the Subantarctic regions rather than in Antarctica itself. Notably, the cold Antarctic Circumpolar Current also started as a continuous circumpolar
1533:
Gentoo penguins are the fastest underwater birds in the world. They are capable of reaching speeds up to 36 km (about 22 miles) per hour while searching for food or escaping from predators. They are also able to dive to depths of 170–200 meters (about 560–660 feet). The small penguins do not
1595:
Calculations of the heat loss and retention ability of marine endotherms suggest that most extant penguins are too small to survive in such cold environments. In 2007, Thomas and
Fordyce wrote about the "heterothermic loophole" that penguins utilize in order to survive in Antarctica. All extant
6222:
Geier, Thom; Jensen, Jeff; Jordan, Tina; Lyons, Margaret; Markovitz, Adam; Nashawaty, Chris; Pastorek, Whitney; Rice, Lynette; Rottenberg, Josh; Schwartz, Missy; Slezak, Michael; Snierson, Dan; Stack, Tim; Stroup, Kate; Tucker, Ken; Vary, Adam B.; Vozick-Levinson, Simon; Ward, Kate,
833:
The birds currently known as penguins were discovered later and were so named by sailors because of their physical resemblance to the great auk. Despite this resemblance, however, they are not auks, and are not closely related to the great auk. They do not belong in the genus
1287:), has its highest diversity in the New Zealand region, and represents a westward dispersal. They are characterized by hairy yellow ornamental head feathers; their bills are at least partly red. These two genera diverged apparently in the Middle Miocene (
1755:
1592:
the
Antarctic winter, the females are at sea fishing for food, leaving the males to brave the weather by themselves. They often huddle together to keep warm and rotate positions to make sure that each penguin gets a turn in the centre of the heat pack.
1788:
1752:
1106:
lineage, or whether gigantism was evolved independently in a restricted
Palaeeudyptinae and the Anthropornithinae – whether they were considered valid, or whether there was a wide size range present in the Palaeeudyptinae as delimited (i.e., including
590:), also known as the fairy penguin, which stands around 30–33 cm (12–13 in) tall and weighs 1.2–1.3 kg (2.6–2.9 lb). Today, larger penguins generally inhabit colder regions, and smaller penguins inhabit regions with temperate or
1012:
were flightless, with short wings adapted for deep diving. They swam on the surface using mainly their feet, but the wings were – as opposed to most other diving birds (both living and extinct) – already adapting to underwater locomotion.
1477:
Penguin wings have the same general bone structure as flighted birds, but the bones are shorter and stouter to allow them to serve as fins. 1). Humerus 2). Sesamoid Bone 3). Radius 4). Ulna 5). Radial Carpal bone 6). Carpometacarpus 7).
1754:
1571:
has a maximum feather density of about nine feathers per square centimeter which is actually much lower than other birds that live in antarctic environments. However, they have been identified as having at least four different
1446:
and consequently would have to be included in that order, or that the plotopterids were not as close to other pelecaniforms as generally assumed, which would necessitate splitting the traditional
Pelecaniformes into three.
910:
Sphenisciformes, i.e., including any flying basal "proto-penguins" to be discovered eventually. Given that neither the relationships of the penguin subfamilies to each other nor the placement of the penguins in the avian
905:
Spheniscidae to what here is referred to as
Spheniscinae. Furthermore, they restrict the phylogenetic taxon Sphenisciformes to flightless taxa, and establish the phylogenetic taxon Pansphenisciformes as equivalent to the
1326:
to South
America and eventually beyond. Despite this, there is no fossil evidence to support the idea of crown radiation from the Antarctic continent in the Paleogene, although DNA study favors such a radiation.
3686:
Slack, Kerryn E.; Jones, Craig M.; Ando, Tatsuro; Harrison, G.L. "Abby"; Fordyce, R. Ewan; Arnason, Ulfur; Penny, David (2006). "Early
Penguin Fossils, plus Mitochondrial Genomes, Calibrate Avian Evolution".
1810:
Penguins form monogamous pairs for a breeding season, though the rate the same pair recouples varies drastically. Most penguins lay two eggs in a clutch, although the two largest species, the emperor and the
2513:
in 1982. Set in Antarctica, the player controls a penguin character who must navigate mazes of ice cubes. The player is rewarded with cut-scenes of animated penguins marching, dancing, saluting and playing
1648:
penguins. Isabellinism is different from albinism. Isabelline penguins tend to live shorter lives than normal penguins, as they are not well-camouflaged against the deep and are often passed over as mates.
1498:
have evolved to become flippers, useless for flight in the air. In the water, however, penguins are astonishingly agile. Penguins' swimming looks very similar to birds' flight in the air. Within the smooth
5386:
Meyer-Rochow V.B. (2015). "Examples of four incompletely resolved aspects of the biology of penguins elated to digestive and reproductive physiology, vision and locomotion". In Jenkins Owen P. (ed.).
2915:
3882:
Pan, Hailin; Cole, Theresa L.; Bi, Xupeng; Fang, Miaoquan; Zhou, Chengran; Yang, Zhengtao; Ksepka, Daniel T.; Hart, Tom; Bouzat, Juan L.; Argilla, Lisa S.; Bertelsen, Mads F. (September 1, 2019).
1196:
contains species with a fairly simple black-and-white head pattern; their distribution is intermediate, centred on Antarctic coasts but extending somewhat northwards from there. In external
2320:
or the nearby offshore islands. They are preyed upon by other birds like skuas, especially in eggs and as fledglings. Other birds like petrels, sheathbills, and gulls also eat the chicks.
6201:
2343:
In June 2011, an emperor penguin came ashore on New Zealand's Peka Peka Beach, 3,200 kilometres (2,000 mi) off course on its journey to Antarctica. Nicknamed Happy Feet, after the
1338:
was already much like today in volume and extent. The emergence of most of today's Subantarctic penguin species almost certainly was caused by this sequence of Neogene climate shifts.
6108:
6078:
1256:
colouration and the former genus has a conspicuous banded head pattern; they are unique among living penguins by nesting in burrows. This group probably radiated eastwards with the
4635:
4128:
6056:
145:
1842:
The majority of living penguin species have declining populations. According to the IUCN Red List, their conservation statuses range from Least Concern through to Endangered.
6138:
1222:
seems to have diverged during the Bartonian, but the range expansion and radiation that led to the present-day diversity probably did not occur until much later; around the
5226:
6168:
5432:
6012:
1753:
5256:
7171:
4971:
4033:
6225:"The 100 Greatest Movies, TV Shows, Albums, Books, Characters, Scenes, Episodes, Songs, Dresses, Music Videos, and Trends That Entertained Us Over the Past 10 Years"
8639:
8478:
5068:
3329:
2734:
Hackett, S. J.; Kimball, R. T.; Reddy, S.; Bowie, R. C. K.; Braun, E. L.; Braun, M. J. (2008). "A Phylogenomic Study of Birds Reveals Their Evolutionary History".
1779:
8517:
1334:, a sharp drop in global average temperature from 14 to 12 mya, and similar abrupt cooling events followed at 8 mya and 4 mya; by the end of the Tortonian, the
1549:. Their eyes are adapted for underwater vision and are their primary means of locating prey and avoiding predators; in air it has been suggested that they are
4831:
2899:
1530:) has difficulty distinguishing between a white penguin belly and the reflective water surface. The dark plumage on their backs camouflages them from above.
1315:. The emergence of the Subantarctic lineage at the end of the Bartonian corresponds with the onset of the slow period of cooling that eventually led to the
1057:
was probably the heaviest, weighing 80 kilograms (180 lb) or more. Both were found on New Zealand, the former also in the Antarctic farther eastwards.
131:
6333:
3174:
958:
6392:
1683:
that flows around these islands. Also, though the climate of the Arctic and Antarctic regions is similar, there are no penguins found in the Arctic.
8644:
8452:
6372:
6016:
2876:
3542:
1556:
8491:
7166:
6376:
1803:
a high level of social interaction between birds, which has led to a large repertoire of visual as well as vocal displays in all penguin species.
3642:
Bertelli, S.; Giannini, N. P. (2005). "A phylogeny of extant penguins (Aves: Sphenisciformes) combining morphology and mitochondrial sequences".
6190:
1098:, Antarctica, for example, around 10 known species of penguins ranging in size from medium to large apparently coexisted some 35 mya during the
1264:(Late Oligocene), starting approximately 28 mya. While the two genera separated during this time, the present-day diversity is the result of a
2316:
Penguins have no special fear of humans and will often approach groups of people. This is probably because penguins have no land predators in
6350:
6288:
5395:
5370:
4890:
4365:
3940:
3485:
3108:
3074:
1658:
582:): on average, adults are about 1.1 m (3 ft 7 in) tall and weigh 35 kg (77 lb). The smallest penguin species is the
1178:
have not yielded Paleogene fossils of the subfamily. Also, the earliest spheniscine lineages are those with the most southern distribution.
6304:
6100:
6070:
2965:
6386:
4242:"Two Antarctic penguin genomes reveal insights into their evolutionary history and molecular changes related to the Antarctic environment"
571:
which they catch with their bills and swallow whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey.
5187:
4912:
4805:
4643:
4528:
Jouventin, P; Aubin, T; Lengagne, T (1999). "Finding a parent in a king penguin colony: The acoustic system of individual recognition".
1503:
a layer of air is preserved, ensuring buoyancy. The air layer also helps insulate the birds in cold waters. On land, penguins use their
4048:
3728:
1567:
Penguins have a thick layer of insulating feathers that keeps them warm in water (heat loss in water is much greater than in air). The
6433:
4302:
4105:
3508:"Description of the earliest fossil penguin from South America and first Paleogene vertebrate locality of Tierra Del Fuego, Argentina"
2938:
1352:
remains unknown and not well-resolved by molecular or morphological analyses. The latter tend to be confounded by the strong adaptive
1075:
if possible, it is becoming accepted that there were at least two major extinct lineages. One or two closely related ones occurred in
8634:
6256:
4965:
4858:
3460:
3422:
3323:
2909:
1331:
6130:
2709:
8576:
6361:
5424:
4106:"Tertiary plotopterids (Aves, Plotopteridae) and a novel hypothesis on the phylogenetic relationships of penguins (Spheniscidae)"
1050:
969:, these areas were at that time less than 1,500 kilometres (930 mi) apart rather than 4,000 kilometres (2,500 mi). The
6227:
6160:
5165:
5095:"Paleogene equatorial penguins challenge the proposed relationship between biogeography, diversity, and Cenozoic climate change"
3842:
1320:
flow only around 30 mya, on the one hand forcing the Antarctic cooling, and on the other facilitating the eastward expansion of
6710:
4435:
3689:
3272:
2360:
4404:
3207:
5248:
2518:. Several remakes and enhanced editions have followed, most recently in 2012. Penguins are also sometimes depicted in music.
2383:, swimming ability and (compared to other birds) lack of fear of humans. Their black-and-white plumage is often likened to a
1822:
Penguins generally only lay one brood; the exception is the little penguin, which can raise two or three broods in a season.
1663:
Although almost all penguin species are native to the Southern Hemisphere, they are not found only in cold climates, such as
8496:
8299:
4955:
4308:
2340:. Typically, penguins do not approach closer than around 9 feet (2.7 meters), at which point they appear to become nervous.
1422:
Inside this group, penguin relationships are far less clear. Depending on the analysis and dataset, a close relationship to
8504:
2628:, is also described as hailing from the Falklands. Opus was a comical, "existentialist" penguin character in the cartoons
2545:
2328:, but dogs have long since been banned from Antarctica. Instead, adult penguins are at risk at sea from predators such as
1442:) may actually be a sister group of the penguins and those penguins may have ultimately shared a common ancestor with the
1257:
1142:, which includes smaller and stout-legged forms, had already arisen in southernmost South America by that time. The early
6030:
1807:
displays are those intended to confront or drive off, or alternately appease and avoid conflict with, other individuals.
5029:
4381:
4334:
3313:
1614:
was originally used for that bird centuries ago. They are only distantly related to the penguins, but are an example of
709:
2846:
626:
198:
8400:
8377:
7156:
6661:
6656:
6380:
4941:
3159:
1977:
1932:
1109:
970:
3131:
650:. When European explorers discovered what are today known as penguins in the Southern Hemisphere, they noticed their
3766:"High flight costs, but low dive costs, in auks support the biomechanical hypothesis for flightlessness in penguins"
3446:
749:, which means 'fat' or 'oil'. Support for this etymology can be found in the alternative Germanic word for penguin,
6938:
4993:
Ashton, K. (2002). "Patterns of within-species body size variation of birds: strong evidence for Bergmann's rule".
4835:
2648:
2439:
1686:
1371:
4157:
3232:
8604:
8522:
6853:
6843:
2596:
1644:
Perhaps one in 50,000 penguins (of most species) are born with brown rather than black plumage. These are called
1054:
703:
3522:
1603:
filters excess salt from the bloodstream. The salt is excreted in a concentrated fluid from the nasal passages.
161:
8629:
7179:
7161:
2992:"State-dependent climate sensitivity in past warm climates and its implications for future climate projections"
2656:
870:
839:
603:
375:
336:
285:
20:
2660:
2464:
1815:, lay only one. With the exception of the emperor penguin, where the male does it all, all penguins share the
923:
574:
They spend about half of their lives on land and the other half in the sea. The largest living species is the
3182:
598:
penguin species were enormous: as tall or heavy as an adult human. There was a great diversity of species in
7184:
6557:
6280:
5990:
5959:
5928:
5897:
5866:
5835:
5804:
5773:
5742:
5711:
5680:
5649:
5618:
5587:
5556:
5525:
5494:
5463:
3452:
2607:
6224:
1166:
and another two more basal genera with more ambiguous relationships. To help resolve the evolution of this
8317:
7151:
6963:
6913:
6426:
5154:"The oldest fossil record of the extant penguin genus Spheniscus – a new species from the Miocene of Peru"
3698:
2592:
2434:
349:
268:
2868:
8624:
8619:
8553:
6988:
6651:
4027:
3507:
2665:
2636:
2458:
2444:
2324:
preyed upon penguins while they were allowed in Antarctica during the age of early human exploration as
2292:
2070:
1316:
1079:, and at least one other—which is or includes the paleeudyptines as recognized today – occurred on most
954:
902:
838:, and are not classified in the same family and order as the great auk. They were classified in 1831 by
323:
310:
8405:
6344:
1450:
A 2014 analysis of whole genomes of 48 representative bird species has concluded that penguins are the
3764:
Elliott, K. H.; Ricklefs, R. E.; Gaston, A. J.; Hatch, S. A.; Speakman, J. R.; Davoren, G. K. (2013).
1307:
The geographical and temporal pattern of spheniscine evolution corresponds closely to two episodes of
8581:
8532:
8426:
8364:
7616:
5106:
5044:
5002:
4674:
4584:
4474:
4172:
3777:
3003:
2743:
2484:
2253:
1819:
duties. These incubation shifts can last days and even weeks as one member of the pair feeds at sea.
1676:
1672:
1615:
1462:
1284:
1211:
1197:
725:, on the basis that the name was originally applied to the great auk, either because it was found on
651:
540:
3703:
950:. Some seminal articles about the evolutionary history of penguins have been published since 2005.
7146:
7038:
6978:
6923:
6703:
6614:
6323:
2588:
2526:
2428:
2093:
1831:
1742:
has collapsed, with barely 200,000 remaining, according to a study published in Antarctic Science.
1645:
1631:
1335:
1171:
655:
532:
362:
8282:
4887:
4762:
Thomas, D.B.; Fordyce, R.E. (2012). "Biological plasticity in penguin heat-retention structures".
2655:
In the mid-2000s, penguins became one of the most publicized species of animals that form lasting
1577:
401:
Breeding range of penguins, all species (aqua); some species have wider seasonal migration ranges
7929:
6993:
6953:
6918:
6579:
5060:
4787:
4698:
4616:
4600:
4553:
4492:
4375:
4079:
3986:"Multiple gene evidence for expansion of extant penguins out of Antarctica due to global cooling"
3659:
3592:"Multiple gene evidence for expansion of extant penguins out of Antarctica due to global cooling"
3534:
2819:
2767:
2642:
2550:
2505:
2469:
2231:
1667:. In fact, only a few species of penguin actually live so far south. Several species live in the
1600:
1147:
1091:
799:
715:
253:
193:
4356:
Buskey, Theresa (March 2001). "The Antarctic Polar Region". In Alan Christopherson, M.S. (ed.).
3414:
3407:
1739:
1695:
737:) in Newfoundland, or because it had white circles around its eyes (though the head was black).
8558:
8355:
8275:
1454:
of Procellariiformes, from which they diverged about 60 million years ago (95% CI, 56.8–62.7).
1146:
saw the emergence of another morphotype in the same area, the similarly sized but more gracile
175:
8614:
8609:
8540:
8413:
8055:
8002:
7952:
7095:
7085:
7028:
6828:
6823:
6641:
6589:
6525:
6520:
6419:
6284:
6252:
6050:
5391:
5366:
5191:
5134:
4961:
4908:
4779:
4690:
4608:
4545:
4510:
4361:
4298:
4273:
4206:
4071:
4015:
3936:
3913:
3805:
3716:
3621:
3481:
3456:
3418:
3384:
3319:
3104:
3070:
3039:
3021:
2905:
2759:
2625:
2600:
2472:
in 1939. Penguins have also appeared in a number of cartoons and television dramas, including
2309:
2287:
2139:
2116:
2000:
1759:
1634:
1491:
1482:
1427:
1376:
974:
915:
is presently resolved, this is confusing, so the established Linnean system is followed here.
726:
151:
8483:
4809:
2482:
and Erika Brueggemann in 1990 and covering more than 100 short episodes. At the end of 2009,
1610:
of the Northern Hemisphere, now extinct, was superficially similar to penguins, and the word
1356:
of the Sphenisciformes; a sometimes perceived fairly close relationship between penguins and
8545:
7921:
7063:
6735:
6676:
6671:
6584:
6456:
5337:
5124:
5114:
5052:
5010:
4771:
4744:
4682:
4592:
4537:
4500:
4482:
4263:
4253:
4196:
4188:
4180:
4120:
4087:
4063:
4005:
3997:
3903:
3895:
3795:
3785:
3750:(2006) mentioned in this section are not as precisely resolved, due to uncertainties of the
3725:
3708:
3651:
3611:
3603:
3526:
3518:
3374:
3364:
3029:
3011:
2809:
2751:
2674:
2621:
2555:
2208:
1955:
1816:
1680:
1626:
1312:
1138:, which competed with them for food and were ultimately more successful. A new lineage, the
1115:
907:
864:
674:
591:
484:
427:
418:
384:
8568:
2942:
2351:. Once he had recovered, Happy Feet was released back into the water south of New Zealand.
8153:
7981:
7451:
6973:
6958:
6896:
6795:
6723:
6631:
6619:
6594:
6499:
6452:
6231:
4894:
3751:
3732:
3267:
2573:
2453:
2297:
2185:
1865:
1568:
1542:
1416:
1167:
1068:
966:
895:
847:
827:
815:
721:
670:
666:
631:
575:
475:
414:
181:
137:
6406:
4866:
5110:
5048:
5006:
4678:
4588:
4478:
4176:
3781:
3007:
2747:
2282:
1360:
is almost certainly an error based on both groups' strong diving adaptations, which are
1060:
Traditionally, most extinct species of penguins, giant or small, had been placed in the
1008:
in New Zealand. While they were not as well-adapted to aquatic life as modern penguins,
8263:
8193:
8145:
8079:
8018:
7973:
7960:
7834:
7797:
7648:
7422:
7259:
7013:
7008:
7003:
6983:
6968:
6863:
6858:
6838:
6833:
6815:
6763:
6696:
6646:
6569:
6551:
6530:
6273:
5129:
5094:
4571:
Sivak, J; Howland, H. C.; McGill-Harelstad, P (1987). "Vision of the Humboldt penguin (
4268:
4241:
4201:
4010:
3985:
3908:
3883:
3800:
3765:
3616:
3591:
3402:
3100:
3094:
3034:
2991:
2694:
2568:
2449:
2162:
2023:
1909:
1791:
1638:
1560:
1546:
1511:
1443:
1308:
1139:
1131:
1095:
878:
811:
741:
730:
687:
583:
548:
167:
4505:
4458:
3979:
3977:
3379:
3352:
8598:
8369:
8201:
8185:
8108:
8100:
7934:
7732:
7582:
7469:
7427:
7227:
7219:
7141:
7105:
7023:
6947:
6666:
5214:
5153:
5056:
5014:
4124:
3857:
3655:
2814:
2797:
2617:
2584:
2479:
2046:
1431:
1423:
1249:
1135:
1030:
982:
77:
8382:
5064:
4791:
4702:
4620:
4557:
4427:
3663:
3538:
3262:
2823:
2488:
put it on its end-of-the-decade "best-of" list, saying, "Whether they were walking (
8287:
8031:
7876:
7692:
7530:
7478:
7350:
7311:
7080:
7068:
7058:
7048:
6745:
6494:
4732:
4665:
Downhower, J.F.; Blumer, L.S. (1988). "Calculating just how small a whale can be".
4396:
4153:
4083:
3199:
3060:
2771:
2670:
2630:
2612:
2577:
2563:
2530:
2397:
2373:
2369:
2337:
1887:
1812:
1734:. Satellite images and photos released in 2018 show the population of 2 million in
1731:
1581:
1527:
1451:
1396:
1365:
1353:
1151:
1103:
1084:
1061:
943:
928:
843:
599:
5342:
5321:
5093:
Clarke, J. A.; Ksepka; Stucchi; Urbina; Giannini; Bertelli; Narváez; Boyd (2007).
4158:"Whole-genome analyses resolve early branches in the tree of life of modern birds"
1130:, around 25 mya. Their decline and disappearance coincided with the spread of the
8418:
6015:. Natural Environment Research Council – British Antarctic Survey. Archived from
4292:
3899:
3240:
3064:
1375:
Humboldt penguins in an aquarium. The penguin is an accomplished swimmer, having
8509:
8465:
8349:
8092:
7939:
7745:
7724:
7713:
7678:
7545:
7535:
7493:
7384:
7251:
7053:
6878:
6873:
6805:
6790:
6758:
6484:
4897:
by Marcel Van Tuinen, Dave Brian Butvill, John A. W. Kirsch and S. Blair Hedges.
4360:. LIFEPAC. 804 N. 2nd Ave. E., Rock Rapids, IA: Alpha Omegan Publications, Inc.
2348:
1795:
1727:
1550:
1361:
1223:
1207:
1183:
646:
first appears in literature at the end of the 16th century as a synonym for the
595:
305:
52:
5099:
Proceedings of the National Academy of Sciences of the United States of America
4467:
Proceedings of the National Academy of Sciences of the United States of America
2966:"Largest penguin ever discovered weighed a whopping 340 pounds, fossils reveal"
2379:
Penguins are widely considered endearing for their unusually upright, waddling
2304:
1545:
for birds; this is used by parents and chicks to locate one another in crowded
8209:
8071:
8010:
7989:
7821:
7805:
7740:
7685:
7657:
7589:
7554:
7432:
7280:
7115:
7090:
6848:
6605:
6510:
6318:
4933:
4330:
3956:
2559:
2500:), these oddly adorable birds took flight at the box office all decade long."
2422:
2410:
2388:
2344:
2317:
1763:
1707:
1664:
1515:
1388:
1322:
1253:
1238:
1200:, these apparently still resemble the common ancestor of the Spheniscinae, as
1192:
1099:
1017:
1005:
946:
history of Sphenisciformes is well-researched, many prehistoric forms are not
874:
611:
411:
370:
357:
344:
115:
97:
62:
3181:(in European Spanish) (22nd ed.). Real Academia Española. Archived from
3025:
2468:
is a children's book written by Richard and Florence Atwater; it was named a
8439:
8169:
8124:
7913:
7706:
7623:
7607:
7596:
7506:
7442:
7401:
7100:
6901:
6888:
6868:
6768:
6541:
5119:
4184:
3790:
3712:
3016:
2838:
2755:
2534:
2522:
2406:
2384:
1715:
1711:
1668:
1607:
1585:
1435:
1400:
1296:
1244:
1175:
1127:
1080:
1076:
1072:
1064:
1046:
1026:
1022:
978:
962:
939:
912:
803:
789:
647:
331:
210:
102:
46:
5138:
4783:
4596:
4549:
4541:
4277:
4258:
4210:
4075:
4019:
4001:
3917:
3809:
3720:
3625:
3607:
3388:
3369:
3154:
3043:
2763:
1679:, but this is only made possible by the cold, rich waters of the Antarctic
393:
4694:
4612:
4514:
4487:
3955:
It is likely that, during the Bartonian, there was a near-synchronous but
3127:
1473:
8340:
8334:
8137:
7867:
7842:
7788:
7699:
7671:
7561:
7525:
7517:
7499:
7394:
7379:
7363:
7110:
7073:
6906:
2515:
2325:
1408:
1288:
1280:
1265:
1261:
1113:). The oldest well-described giant penguin, the 5-foot (1.5 m)-tall
1034:
947:
932:
818:
in the mid-19th century. As confirmed by a 2004 genetic study, the genus
568:
318:
230:
92:
87:
72:
67:
57:
39:
8392:
4214:
1102:(Late Eocene). It is not known whether the palaeeudyptines constitute a
7900:
7779:
7664:
7630:
7568:
7389:
7243:
7235:
6928:
6800:
6785:
6775:
6474:
5357:
Reilly PN, Balmford P (1975). "A breeding study of the little penguin,
4192:
1723:
1500:
1439:
1412:
1384:
1348:
1227:
1215:
1143:
999:
658:
and named them after this bird, although they are not closely related.
607:
552:
544:
536:
107:
82:
6688:
8177:
8116:
7892:
7884:
7813:
7753:
7637:
7575:
7458:
7125:
7120:
6998:
4938:
Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals
4775:
4686:
4604:
4496:
4457:
Wever, E. G.; Herman, P. N.; Simmons, J. A.; Hertzler, D. R. (1969).
3530:
2537:
2414:
2392:
1735:
1703:
1458:
1295:
are the product of a later radiation, stretching from about the late
994:
615:
220:
35:
8470:
8457:
8311:
5826:
International), BirdLife International (BirdLife (August 16, 2018).
5733:
International), BirdLife International (BirdLife (October 1, 2016).
4748:
1576:
of feather: in addition to the traditional feather, the emperor has
8444:
5981:
International), BirdLife International (BirdLife (August 9, 2018).
5950:
International), BirdLife International (BirdLife (August 9, 2018).
5919:
International), BirdLife International (BirdLife (August 9, 2018).
5888:
International), BirdLife International (BirdLife (August 8, 2018).
5857:
International), BirdLife International (BirdLife (August 7, 2018).
5795:
International), BirdLife International (BirdLife (August 7, 2018).
5764:
International), BirdLife International (BirdLife (August 7, 2018).
5702:
International), BirdLife International (BirdLife (August 7, 2018).
5671:
International), BirdLife International (BirdLife (August 7, 2018).
5640:
International), BirdLife International (BirdLife (August 7, 2018).
5609:
International), BirdLife International (BirdLife (August 7, 2018).
5578:
International), BirdLife International (BirdLife (August 7, 2018).
5547:
International), BirdLife International (BirdLife (August 7, 2018).
5516:
International), BirdLife International (BirdLife (August 8, 2018).
5485:
International), BirdLife International (BirdLife (August 7, 2018).
4067:
2798:"The phylogeny of the living and fossil Sphenisciformes (penguins)"
1694:
Several authors have suggested that penguins are a good example of
1248:
contain species with a mostly Subantarctic distribution centred on
8063:
7850:
7413:
6466:
5611:"IUCN Red List of Threatened Species: Northern Rockhopper Penguin"
5549:"IUCN Red List of Threatened Species: Southern Rockhopper Penguin"
3506:
Clarke, Julia A.; Olivero, Eduardo B. & Puerta, Pablo (2003).
2474:
2364:
2329:
2303:
2281:
1749:
1719:
1685:
1625:
1555:
1481:
1472:
1404:
1370:
1357:
1234:
1163:
1071:. More recently, with new taxa being discovered and placed in the
1029:(Middle Eocene), some 39–38 mya, primitive penguins had spread to
922:
891:
625:
564:
556:
295:
5982:
5951:
5920:
5889:
5858:
5827:
5796:
5765:
5734:
5703:
5672:
5641:
5610:
5579:
5548:
5517:
5486:
5455:
3590:
Baker, A. J; Pereira, S. L.; Haddrath, O. P; Edge, K.-A. (2006).
1330:
Later, an interspersed period of slight warming was ended by the
1053:
was the tallest, growing nearly 1.80 meters (5.9 feet) tall. The
8039:
7406:
7369:
7339:
6753:
6719:
5454:
International), BirdLife International (BirdLife (May 1, 2004).
5322:"Prolonged foraging trips and egg desertion in little penguins (
2695:"Antarctic birds (Neornithes) during the Cretaceous–Eocene time"
2510:
2420:
Penguins have been the subject of many books and films, such as
2402:
2380:
2333:
1523:
1522:
with white fronts. A predator looking up from below (such as an
1519:
1504:
1495:
1392:
1120:
560:
240:
8315:
7203:
6692:
6415:
6367:
2606:
Penguins featured regularly in the cartoons of U.K. cartoonist
2583:
Several pro, minor, college and high school sport teams in the
1021:
from northern Peru was dated to 42 mya. An unnamed fossil from
8161:
7018:
6780:
6401:
6356:
3884:"High-coverage genomes to elucidate the evolution of penguins"
2365:
2321:
823:
520:
457:
8431:
2837:
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2023).
1260:
out of the ancestral range of modern penguins throughout the
6364:
on the Internet Bird Collection (archived December 27, 2015)
6339:
5735:"IUCN Red List of Threatened Species: Erect-crested Penguin"
4888:
Convergence and divergence in the evolution of aquatic birds
4735:(2007). "The heterothermic loophole exploited by penguins".
2941:. Penguin Sentinels/University of Washington. Archived from
2646:. He was also the star in the animated Christmas TV special
547:. Highly adapted for life in the ocean water, penguins have
514:
6411:
4113:
Journal of Zoological Systematics and Evolutionary Research
3523:
10.1206/0003-0082(2003)423<0001:dotefp>2.0.co;2
1252:; some, however, range quite far northwards. They all lack
508:
466:
5766:"IUCN Red List of Threatened Species: Yellow-eyed Penguin"
2387:
suit. Some writers and artists have penguins based at the
5952:"IUCN Red List of Threatened Species: Magellanic Penguin"
5425:"If you think penguins are cute and cuddly, you're wrong"
3843:"Eocene penguins of Seymour Island, Antarctica: taxonomy"
3130:. Centre National de Ressources Textuelles et Lexicales.
1699:
greater effect on species diversity than latitude alone.
1150:, as well as the radiation that gave rise to the current
1090:
Size plasticity was significant at this initial stage of
1049:(40–30 mya), some lineages of gigantic penguins existed.
677:
origin (the latter two are attributed to the French word
502:
493:
451:
445:
436:
5983:"IUCN Red List of Threatened Species: Galapagos Penguin"
5828:"IUCN Red List of Threatened Species: Chinstrap Penguin"
5642:"IUCN Red List of Threatened Species: Fiordland Penguin"
5249:"World's biggest king penguin colony shrinks 90 percent"
1507:
and wings to maintain balance for their upright stance.
1368:
datasets do not agree in detail with each other either.
665:
is still debated. The English word is not apparently of
5921:"IUCN Red List of Threatened Species: Humboldt Penguin"
5580:"IUCN Red List of Threatened Species: Macaroni Penguin"
3990:
Proceedings of the Royal Society B: Biological Sciences
3984:
Baker, A., Pereira, SL, Haddrath, OP, Edge, KA (2006).
3596:
Proceedings of the Royal Society B: Biological Sciences
2587:
have named themselves after the species, including the
1553:, although research has not supported this hypothesis.
1383:
What seems clear is that penguins belong to a clade of
5890:"IUCN Red List of Threatened Species: African Penguin"
5456:"IUCN Red List of Threatened Species: Emperor Penguin"
5390:. Hauppauge, N.Y.: Nova Sci Publ Inc. pp. 65–80.
5253:
World's biggest king penguin colony shrinks 90 percent
2990:
Caballero, Rodrigo; Huber, Matthew (August 27, 2013).
5859:"IUCN Red List of Threatened Species: Gentoo Penguin"
5797:"IUCN Red List of Threatened Species: Adelie Penguin"
5673:"IUCN Red List of Threatened Species: Snares Penguin"
5518:"IUCN Red List of Threatened Species: Little Penguin"
3480:. London, England, UK: Helm Publishing. p. 361.
3066:
The Great Auk: The Extinction of the Original Penguin
2796:
Ksepka, D. T.; Bertelli, S.; Giannini, N. P. (2006).
2562:'s reimagining of the story, the character played by
2405:, as the title penguin would interact with Arctic or
1291:, roughly 15–14 mya), although the living species of
523:
511:
505:
499:
490:
487:
448:
442:
433:
430:
6037:. 2011. Archived from the original on April 17, 2021
5704:"IUCN Red List of Threatened Species: Royal Penguin"
4049:"Parallel Radiations in the Primary Clades of Birds"
3351:
Thomas, G. H.; Wills, M. A.; Székely, T. S. (2004).
2558:, he was one of the most popular characters, and in
2544:#58). He became one of the most enduring enemies in
1563:
swimming underwater at the Nagasaki Penguin Aquarium
1126:
Gigantic penguins had disappeared by the end of the
463:
454:
8324:
8136:
8091:
8030:
8001:
7972:
7951:
7912:
7875:
7866:
7833:
7796:
7787:
7778:
7723:
7647:
7606:
7544:
7516:
7492:
7468:
7441:
7362:
7349:
7310:
7279:
7218:
7134:
7037:
6937:
6887:
6814:
6744:
6629:
6603:
6567:
6539:
6508:
6482:
6464:
5487:"IUCN Red List of Threatened Species: King Penguin"
3967:, and the common ancestor of all remaining genera (
517:
496:
439:
6272:
5190:. Southern Indian Education Center. Archived from
3413:. Lincoln: University of Nebraska Press. pp.
3406:
3294:Tui De Roy; Mark Jones; Julie Cornthwaite (2013).
1299:(Late Miocene, 8 mya) to the end of the Pliocene.
2616:newspaper, particularly during and following the
555:and flippers for swimming. Most penguins feed on
6055:: CS1 maint: bot: original URL status unknown (
2839:"Kagu, Sunbittern, tropicbirds, loons, penguins"
2312:encountering a human during the Antarctic summer
1430:has been suggested. Some think the penguin-like
961:in the general area of southern New Zealand and
5361:, in Australia". In Stonehouse, Bernard (ed.).
3770:Proceedings of the National Academy of Sciences
3455:, Australia: CSIRO Publishing. pp. 86–87.
2996:Proceedings of the National Academy of Sciences
6459:: Spheniscidae · subfamily: Spheniscinae)
5215:"Penguin past: The current state of knowledge"
3318:. Oxford University Press (USA). p. 152.
2572:, employed an actual army of penguins (mostly
606:in a region around 2,000 km south of the
6704:
6427:
5320:Numata, M.; Davis, L. S.; Renner, M. (2000).
5288:
5286:
4240:D.; Gu, W.; et al. (December 12, 2014).
3353:"A supertree approach to shorebird phylogeny"
3088:
3086:
2525:introduced the avian-themed character of the
869:The family name of Spheniscidae was given by
842:in several distinct genera within the family
8:
6334:Two new fossil penguin species found in Peru
6013:"Removal of the sledge dogs from Antarctica"
4859:"African Penguins and Penguins of the World"
4577:Proceedings of the Royal Society of London B
4463:, as represented by the cochlear potentials"
4032:: CS1 maint: multiple names: authors list (
3933:Systematics and Taxonomy of Australian Birds
3478:The Helm Dictionary of Scientific Bird Names
1399:" to distinguish them from the more ancient
776:, and a group of penguins in the water is a
750:
686:Some dictionaries suggest a derivation from
678:
6340:Information about penguins at pinguins.info
4957:Book: BSCS Biology, By Arnold Brams Grobman
3448:Australian Bird Names: Origins and Meanings
3069:. Bunker Hill Publishing, Inc. p. 17.
1702:Major populations of penguins are found in
1395:) that comprises what is sometimes called "
882:
756:
755:or 'fat-goose', and the related Dutch word
744:
740:An alternative etymology links the word to
696:
690:
8312:
7872:
7793:
7784:
7775:
7513:
7359:
7346:
7307:
7303:
7276:
7215:
7211:
7200:
7196:
6711:
6697:
6689:
6434:
6420:
6412:
6101:"Happy Feet pulls through third operation"
6071:"Emperor penguin not out of the woods yet"
3935:. Canberra: CSIRO Publishing. p. 97.
3637:
3635:
3307:
3305:
3298:. Princeton University Press. p. 151.
2693:Tambussi, C.; Hospitaleche, C. A. (2007).
1162:Modern penguins constitute two undisputed
1033:and were in the process of expanding into
683:), but first appears in English or Dutch.
392:
27:
6407:Live 24/7 camera inside a penguin habitat
6393:"Lessons in a Land of Wind and Ice" from
6191:"Happy Feet starts the long journey home"
5341:
5303:
5301:
5128:
5118:
4504:
4486:
4331:"Penguin swimming under water, Galapagos"
4267:
4257:
4200:
4009:
3907:
3799:
3789:
3702:
3615:
3585:
3583:
3501:
3499:
3497:
3378:
3368:
3055:
3053:
3033:
3015:
2813:
1158:Origin and systematics of modern penguins
764:Adult male penguins are sometimes called
618:, a climate decidedly warmer than today.
3743:The exact divergence dates according to
3681:
3679:
3677:
3675:
3673:
3581:
3579:
3577:
3575:
3573:
3571:
3569:
3567:
3565:
3563:
3128:"PINGOUIN : Etymologie de PINGOUIN"
3122:
3120:
2791:
2789:
2787:
2785:
2783:
2781:
2391:, but there are no wild penguins in the
1844:
1599:They can drink salt water because their
877:, the name of that genus comes from the
6377:Te Ara: The Encyclopedia of New Zealand
6345:Integrated Taxonomic Information System
5188:"Penguins of Australia and New Zealand"
5074:from the original on September 24, 2015
2845:. International Ornithologists' Union.
2685:
531:). They live almost exclusively in the
8640:Taxa named by Charles Lucien Bonaparte
6141:from the original on December 10, 2013
6111:from the original on December 14, 2011
6081:from the original on December 10, 2013
6048:
5388:Advances in Animal Science and Zoology
5365:. London: Macmillan. pp. 161–87.
5168:from the original on November 13, 2014
4636:"Busting Myths About Penguin Feathers"
4373:
4025:
3227:
3225:
1690:Gálapagos Penguins near Isabela Island
1268:radiation, taking place some 4–2 mya.
1210:are, in most cases, fairly pronounced
957:penguins lived around the time of the
6238:. December 11, 2009 (1079/1080):74-84
6204:from the original on February 2, 2012
6171:from the original on February 1, 2012
4974:from the original on October 18, 2021
4459:"Hearing in the blackfooted penguin,
4438:from the original on January 25, 2021
4407:from the original on January 25, 2021
4134:from the original on October 13, 2006
3332:from the original on October 18, 2021
3275:from the original on October 29, 2013
3134:from the original on October 26, 2008
3099:. New York: Crestwood House. p.
1659:List of Sphenisciformes by population
1518:– that is, they have black backs and
1415:, with the possible exception of the
1045:During the Late Eocene and the Early
959:Cretaceous–Paleogene extinction event
7:
8533:cde60235-d69f-4d81-a72d-1914ff14f319
8270:
6357:Penguin research projects on the web
5423:Gorvett, Zaria (December 23, 2015).
5030:"On the validity of Bergmann's rule"
4915:from the original on August 22, 2016
4720:. New York: Oxford University Press.
1403:. This group contains such birds as
1188:to some Subantarctic islands today.
8294:
6249:Understanding popular music culture
6131:"The truth about Happy Feet's home"
5987:IUCN Red List of Threatened Species
5956:IUCN Red List of Threatened Species
5925:IUCN Red List of Threatened Species
5894:IUCN Red List of Threatened Species
5863:IUCN Red List of Threatened Species
5832:IUCN Red List of Threatened Species
5801:IUCN Red List of Threatened Species
5770:IUCN Red List of Threatened Species
5739:IUCN Red List of Threatened Species
5708:IUCN Red List of Threatened Species
5677:IUCN Red List of Threatened Species
5646:IUCN Red List of Threatened Species
5615:IUCN Red List of Threatened Species
5584:IUCN Red List of Threatened Species
5553:IUCN Red List of Threatened Species
5522:IUCN Red List of Threatened Species
5491:IUCN Red List of Threatened Species
5460:IUCN Red List of Threatened Species
5259:from the original on August 1, 2018
4832:"Humboldt Penguin: Saint Louis Zoo"
3445:Fraser, Ian; Gray, Jeannie (2019).
2849:from the original on April 27, 2020
1119:, existed as far north as northern
826:(Alcidae), within the order of the
772:; a group of penguins on land is a
5993:from the original on March 4, 2020
5962:from the original on March 4, 2020
5931:from the original on March 4, 2020
5900:from the original on March 4, 2020
5869:from the original on March 4, 2020
5838:from the original on March 4, 2020
5807:from the original on March 4, 2020
5776:from the original on March 4, 2020
5745:from the original on March 4, 2020
5714:from the original on March 4, 2020
5683:from the original on March 4, 2020
5652:from the original on March 4, 2020
5621:from the original on March 4, 2020
5590:from the original on March 4, 2020
5559:from the original on March 4, 2020
5528:from the original on March 4, 2020
5497:from the original on March 4, 2020
5466:from the original on March 4, 2020
5435:from the original on March 8, 2020
5229:from the original on March 7, 2022
4311:from the original on July 21, 2020
4297:. Cengage Publishing. p. 30.
3548:from the original on July 24, 2007
2939:"Little Penguin – Penguin Project"
2879:from the original on July 10, 2024
2715:from the original on April 2, 2021
2702:Revista de la Asociación Geológica
1794:watching over a sleeping chick at
1541:Penguins have an average sense of
118:origin according to molecular data
16:Family of aquatic flightless birds
14:
7172:Late Quaternary prehistoric birds
4337:from the original on June 8, 2014
3210:from the original on June 4, 2016
3179:Diccionario de la lengua española
2918:from the original on June 2, 2021
2873:Sea World Parks and Entertainment
1490:Penguins are superbly adapted to
1434:(usually considered relatives of
1342:Relationship to other bird orders
1332:Middle Miocene Climate Transition
1218:conditions. As the former genus,
1134:and other primitive, fish-eating
8293:
8281:
8269:
8258:
8257:
8148:(New World vultures and condors)
5057:10.1046/j.1365-2699.2003.00837.x
5015:10.1046/j.1466-822X.2002.00313.x
4125:10.1111/j.1439-0469.2004.00291.x
3656:10.1111/j.1096-0031.2005.00065.x
2964:Harry Baker (February 8, 2023).
2815:10.1111/j.1096-0031.2006.00116.x
1778:
1283:(though not as far north as the
810:, meaning "plump or fat without
483:
426:
197:
174:
160:
144:
130:
8645:Extant Danian first appearances
6161:"Stephen Fry visits Happy Feet"
4995:Global Ecology and Biogeography
4434:. Britannica Encyclopedia Inc.
4403:. Encyclopedia Britannica Inc.
4333:. Youtube.com. April 14, 2008.
4047:Fain, M. G.; Houde, P. (2004).
3959:split between the ancestors of
3931:Christidis L, Boles WE (2008).
3690:Molecular Biology and Evolution
2462:, a parody of the documentary.
2361:Cultural depictions of penguins
1364:. On the other hand, different
1214:related to that genus' extreme
6351:Penguin information on 70South
5330:New Zealand Journal of Zoology
4954:Grobman, Arnold Brams (1964).
4432:Britannica Online Encyclopedia
4294:Introduction to Marine Biology
927:Penguin tracks in the sand on
901:Some recent sources apply the
385:List of penguins#Fossil genera
1:
6336:. news.nationalgeographic.com
5410:
5343:10.1080/03014223.2000.9518236
5307:
5292:
5277:
5158:Acta Palaeontologica Polonica
4907:Askew, Nick (June 24, 2009).
4737:Australian Journal of Zoology
4634:Ed Young (October 20, 2015).
3409:Diving Birds of North America
2448:, a documentary based on the
1258:Antarctic Circumpolar Current
1051:Nordenskjoeld's giant penguin
822:belongs in the family of the
6560:(or northern little penguin)
6387:Seaworld Penguin Information
6383:(archived September 5, 2008)
6347:(archived February 17, 2006)
6324:Resources in other libraries
5028:Meiri, S; Dayan, T. (2003).
2843:World Bird List Version 13.1
1675:, lives as far north as the
985:boundary, around 70–68 mya.
977:can be roughly dated to the
710:American Heritage Dictionary
383:For prehistoric genera, see
8212:(woodpeckers and relatives)
6662:Northern rockhopper penguin
6657:Southern rockhopper penguin
6389:(archived October 17, 2013)
6381:Encyclopedia of New Zealand
6275:The Penguins – Spheniscidae
4291:Karleskint, George (2012).
3841:Jadwiszczak, Piotr (2006).
3296:Penguins:The Ultimate Guide
3204:Online Etymology Dictionary
3160:Online Etymology Dictionary
3096:Gone Forever: The Great Auk
1978:Northern rockhopper penguin
1933:Southern rockhopper penguin
1110:Anthropornis nordenskjoeldi
971:most recent common ancestor
894:" used for the shape of an
794:Since 1871, the Latin word
768:, females sometimes called
8661:
8074:(cormorants and relatives)
7167:Extinct species since 1500
6271:Williams; Tony D. (1995).
4911:. BirdLife International.
4718:The penguins: Spheniscidae
3968:
3900:10.1093/gigascience/giz117
3856:(#1): 3–62. Archived from
3825:
3744:
3731:December 16, 2009, at the
3476:Jobling, James A. (2010).
2649:A Wish for Wings That Work
2358:
1656:
862:
787:
661:The etymology of the word
604:at least one giant species
18:
8253:
8204:(kingfishers and rollers)
8058:(albatrosses and petrels)
7942:(swifts and hummingbirds)
7924:(nightjars and relatives)
7774:
7748:(pheasants and relatives)
7306:
7275:
7214:
7210:
7199:
7195:
6731:
6455:: Sphenisciformes ·
6449:
6362:Penguin videos and photos
6353:(archived March 15, 2006)
6319:Resources in your library
4909:"List of Penguin Species"
3515:American Museum Novitates
3315:Who Killed the Great Auk?
2937:Grabski, Valerie (2009).
2597:Youngstown State Penguins
1486:Taxidermized penguin skin
1387:(living birds except for
1055:New Zealand giant penguin
883:
854:Systematics and evolution
802:to name the genus of the
800:scientific classification
704:Oxford English Dictionary
701:, 'white', including the
400:
391:
301:
293:
194:Scientific classification
192:
124:
30:
8635:Danian first appearances
8103:(seriemas and relatives)
8082:(pelicans and relatives)
6554:(or little blue penguin)
5213:Jadwiszczak, P. (2009).
4380:: CS1 maint: location (
3357:BMC Evolutionary Biology
3312:Gaskell, Jeremy (2000).
3093:Crofford, Emily (1989).
2869:"Diet and Eating Habits"
2554:TV series, as played by
1653:Distribution and habitat
1346:Penguin ancestry beyond
1279:clade occurs at similar
871:Charles Lucien Bonaparte
840:Charles Lucien Bonaparte
654:to the great auk of the
543:, is found north of the
21:Penguin (disambiguation)
8196:(hornbills and hoopoes)
8111:(falcons and relatives)
6558:White-flippered penguin
6514:(brush-tailed penguins)
6281:Oxford University Press
6230:April 15, 2014, at the
5363:The Biology of Penguins
5152:Gohlich, U. B. (2007).
5120:10.1073/pnas.0611099104
5037:Journal of Biogeography
4716:Williams, T.D. (1995).
4401:Encyclopedia Britannica
4185:10.1126/science.1253451
3791:10.1073/pnas.1304838110
3453:Clayton South, Victoria
3017:10.1073/pnas.1303365110
2756:10.1126/science.1157704
2671:one such penguin family
2624:, from the cartoons of
2546:Batman's rogues gallery
2401:helped perpetuate this
2236:Spheniscus magellanicus
1892:Aptenodytes patagonicus
8180:(trogons and quetzals)
7992:(cranes and relatives)
7262:(emus and cassowaries)
7152:Glossary of bird terms
6964:Confuciusornithiformes
4597:10.1098/rspb.1987.0005
4542:10.1006/anbe.1999.1086
4259:10.1186/2047-217X-3-27
4002:10.1098/rspb.2005.3260
3726:Supplementary Material
3608:10.1098/rspb.2005.3260
3370:10.1186/1471-2148-4-28
3263:"Penguin – Definition"
2593:National Hockey League
2435:Penguins of Madagascar
2376:
2313:
2301:
1766:
1691:
1641:
1564:
1487:
1479:
1469:Anatomy and physiology
1457:The distantly related
1380:
935:
898:'s swimming flippers.
757:
751:
745:
734:
697:
691:
679:
639:
8554:Paleobiology Database
8021:(kagu and sunbittern)
7984:(gulls and relatives)
6989:Songlingornithiformes
6954:Omnivoropterygiformes
6652:Erect-crested penguin
6019:on November 13, 2014.
5219:Polish Polar Research
4893:May 29, 2008, at the
4838:on September 28, 2006
4575:) in air and water".
4488:10.1073/pnas.63.3.676
3850:Polish Polar Research
3713:10.1093/molbev/msj124
2666:And Tango Makes Three
2490:March of the Penguins
2465:Mr. Popper's Penguins
2459:Farce of the Penguins
2445:March of the Penguins
2395:. The cartoon series
2368:
2345:film of the same name
2307:
2285:
2258:Spheniscus mendiculus
2144:Pygoscelis antarctica
2071:Erect-crested penguin
2005:Eudyptes pachyrynchus
1960:Eudyptes chrysolophus
1851:IUCN Red List Status
1758:
1689:
1629:
1559:
1485:
1476:
1374:
1230:, roughly 20–15 mya.
965:, Antarctica. Due to
926:
863:Further information:
629:
34:Temporal range: Late
7843:Phoenicopteriformes
6236:Entertainment Weekly
5194:on February 17, 2012
5105:(#28): 11545–11550.
4857:van der Merwe, H.J.
4806:"Animal Fact Sheets"
4573:Spheniscus humboldti
4220:on February 24, 2015
4171:(#6215): 1320–1331.
3243:on September 5, 2011
2945:on December 16, 2011
2669:, was written about
2503:A video game called
2485:Entertainment Weekly
2372:, the mascot of the
2213:Spheniscus humboldti
2098:Megadyptes antipodes
1870:Aptenodytes forsteri
1616:convergent evolution
1463:convergent evolution
1313:paleoclimatic record
1025:proves that, by the
636:Aptenodytes forsteri
580:Aptenodytes forsteri
19:For other uses, see
7814:Mesitornithiformes
7808:(doves and pigeons)
7142:Families and orders
7004:Ichthyornithiformes
6979:Patagopterygiformes
6615:Yellow-eyed penguin
6247:Shuker, R. (2012).
6200:. August 29, 2011.
6167:. August 11, 2011.
6031:"beautyofbirds.com"
5111:2007PNAS..10411545C
5049:2003JBiog..30..331M
5007:2002GloEB..11..505A
4869:on October 12, 2006
4679:1988Natur.335..675D
4640:National Geographic
4589:1987RSPSB.229..467S
4479:1969PNAS...63..676W
4461:Spheniscus demersus
4177:2014Sci...346.1320J
3782:2013PNAS..110.9380E
3271:. August 31, 2012.
3008:2013PNAS..11014162C
3002:(35): 14162–14167.
2748:2008Sci...320.1763H
2742:(5884): 1763–1768.
2589:Pittsburgh Penguins
2496:), or hanging ten (
2278:Penguins and humans
2190:Spheniscus demersus
2094:Yellow-eyed penguin
1937:Eudyptes chrysocome
1857:Mature Individuals
1838:Conservation status
1622:Isabelline penguins
1336:Antarctic ice sheet
1226:stage of the Early
1172:Antarctic Peninsula
997:penguin species is
656:Northern Hemisphere
584:little blue penguin
567:and other forms of
533:Southern Hemisphere
8156:(eagles and hawks)
8056:Procellariiformes
7961:Opisthocomiformes
7930:Steatornithiformes
7157:List by population
6994:Hongshanornithidae
6919:Evolution of birds
6635:(crested penguins)
6580:Magellanic penguin
6397:Magazine 1/15/2010
6137:. August 8, 2011.
4646:on October 7, 2018
3776:(#23): 9380–9384.
3403:Johnsgard, Paul A.
2657:homosexual couples
2470:Newbery Honor Book
2377:
2355:In popular culture
2314:
2302:
2232:Magellanic penguin
2121:Pygoscelis adeliae
2051:Eudyptes schlegeli
1767:
1760:Chinstrap penguins
1692:
1642:
1601:supraorbital gland
1565:
1538:or rocky terrain.
1488:
1480:
1381:
1311:documented in the
1148:Palaeospheniscinae
936:
903:phylogenetic taxon
808:Pinguinus impennis
716:Century Dictionary
652:similar appearance
640:
614:, during the Late
254:Austrodyptornithes
8592:
8591:
8541:Open Tree of Life
8318:Taxon identifiers
8309:
8308:
8249:
8248:
8245:
8244:
8241:
8240:
8237:
8236:
8233:
8232:
8229:
8228:
8225:
8224:
8221:
8220:
8042:(loons or divers)
8011:Phaethontiformes
8003:Phaethontimorphae
7953:Opisthocomiformes
7922:Caprimulgiformes
7862:
7861:
7851:Podicipediformes
7770:
7769:
7766:
7765:
7762:
7761:
7488:
7487:
7271:
7270:
7228:Struthioniformes
7096:Waterfowl hunting
7029:Gastornithiformes
7024:Aepyornithiformes
7014:Lithornithiformes
6686:
6685:
6642:Fiordland penguin
6590:Galapagos penguin
6573:(banded penguins)
6545:(little penguins)
6526:Chinstrap penguin
6395:National Wildlife
6305:Library resources
6290:978-0-19-854667-2
6107:. June 27, 2011.
6077:. June 25, 2011.
5397:978-1-63483-328-8
5372:978-0-333-16791-5
4764:Anatomical Record
4583:(#1257): 467–72.
4536:(#6): 1175–1183.
4428:"Emperor Penguin"
4367:978-1-58095-156-2
4358:The Polar Regions
4104:Mayr, G. (2005).
4093:on April 7, 2013.
3942:978-0-643-06511-6
3697:(#6): 1144–1155.
3487:978-1-4081-2501-4
3185:on June 9, 2011.
3153:Harper, Douglas.
3110:978-0-89686-459-7
3076:978-1-59373-003-1
2626:Berkeley Breathed
2601:college athletics
2566:in the 1992 film
2533:adversary of the
2409:species, such as
2275:
2274:
2254:Galápagos penguin
2140:Chinstrap penguin
2075:Eudyptes sclateri
2028:Eudyptes robustus
2001:Fiordland penguin
1982:Eudyptes moseleyi
1756:
1677:Galápagos Islands
1673:Galápagos penguin
1510:All penguins are
1428:Procellariiformes
1397:higher waterbirds
1379:instead of wings.
1285:Galápagos penguin
1004:, which lived 62
993:The oldest known
975:Procellariiformes
814:"), which became
798:has been used in
727:White Head Island
592:tropical climates
541:Galápagos penguin
405:
404:
379:
366:
353:
340:
327:
314:
289:
272:
119:
8652:
8605:Flightless birds
8585:
8584:
8572:
8571:
8562:
8561:
8549:
8548:
8536:
8535:
8526:
8525:
8513:
8512:
8510:NBNSYS0000164612
8500:
8499:
8487:
8486:
8474:
8473:
8461:
8460:
8448:
8447:
8435:
8434:
8422:
8421:
8409:
8408:
8396:
8395:
8386:
8385:
8373:
8372:
8360:
8359:
8358:
8345:
8344:
8343:
8313:
8297:
8296:
8285:
8273:
8272:
8261:
8260:
8213:
8205:
8197:
8189:
8186:Leptosomiformes
8181:
8173:
8165:
8157:
8154:Accipitriformes
8149:
8128:
8127:(perching birds)
8120:
8112:
8104:
8083:
8075:
8067:
8059:
8051:
8048:Sphenisciformes
8043:
8022:
8014:
7993:
7985:
7982:Charadriiformes
7964:
7943:
7925:
7904:
7896:
7893:Musophagiformes
7888:
7873:
7854:
7846:
7825:
7817:
7809:
7794:
7785:
7776:
7749:
7736:
7514:
7510:
7503:
7416:
7409:
7373:
7360:
7356:
7347:
7343:
7308:
7304:
7277:
7263:
7255:
7247:
7239:
7231:
7216:
7212:
7201:
7197:
7064:Bird collections
7019:Dinornithiformes
6924:Darwin's finches
6914:Origin of flight
6854:Seabird breeding
6844:Sexual selection
6713:
6706:
6699:
6690:
6672:Macaroni penguin
6585:Humboldt penguin
6488:(great penguins)
6436:
6429:
6422:
6413:
6402:Curious Penguins
6294:
6278:
6259:
6245:
6239:
6220:
6214:
6213:
6211:
6209:
6195:
6187:
6181:
6180:
6178:
6176:
6157:
6151:
6150:
6148:
6146:
6127:
6121:
6120:
6118:
6116:
6097:
6091:
6090:
6088:
6086:
6067:
6061:
6060:
6054:
6046:
6044:
6042:
6027:
6021:
6020:
6009:
6003:
6002:
6000:
5998:
5978:
5972:
5971:
5969:
5967:
5947:
5941:
5940:
5938:
5936:
5916:
5910:
5909:
5907:
5905:
5885:
5879:
5878:
5876:
5874:
5854:
5848:
5847:
5845:
5843:
5823:
5817:
5816:
5814:
5812:
5792:
5786:
5785:
5783:
5781:
5761:
5755:
5754:
5752:
5750:
5730:
5724:
5723:
5721:
5719:
5699:
5693:
5692:
5690:
5688:
5668:
5662:
5661:
5659:
5657:
5637:
5631:
5630:
5628:
5626:
5606:
5600:
5599:
5597:
5595:
5575:
5569:
5568:
5566:
5564:
5544:
5538:
5537:
5535:
5533:
5513:
5507:
5506:
5504:
5502:
5482:
5476:
5475:
5473:
5471:
5451:
5445:
5444:
5442:
5440:
5420:
5414:
5408:
5402:
5401:
5383:
5377:
5376:
5354:
5348:
5347:
5345:
5317:
5311:
5305:
5296:
5290:
5281:
5275:
5269:
5268:
5266:
5264:
5245:
5239:
5238:
5236:
5234:
5210:
5204:
5203:
5201:
5199:
5184:
5178:
5177:
5175:
5173:
5149:
5143:
5142:
5132:
5122:
5090:
5084:
5083:
5081:
5079:
5073:
5034:
5025:
5019:
5018:
4990:
4984:
4983:
4981:
4979:
4960:. Kendall/Hunt.
4951:
4945:
4931:
4925:
4924:
4922:
4920:
4904:
4898:
4885:
4879:
4878:
4876:
4874:
4865:. Archived from
4854:
4848:
4847:
4845:
4843:
4834:. Archived from
4828:
4822:
4821:
4819:
4817:
4812:on July 20, 2006
4808:. Archived from
4802:
4796:
4795:
4776:10.1002/ar.21538
4759:
4753:
4752:
4728:
4722:
4721:
4713:
4707:
4706:
4687:10.1038/335675b0
4662:
4656:
4655:
4653:
4651:
4642:. Archived from
4631:
4625:
4624:
4568:
4562:
4561:
4530:Animal Behaviour
4525:
4519:
4518:
4508:
4490:
4454:
4448:
4447:
4445:
4443:
4426:Rafferty, John.
4423:
4417:
4416:
4414:
4412:
4397:"Gentoo Penguin"
4395:Rafferty, John.
4392:
4386:
4385:
4379:
4371:
4353:
4347:
4346:
4344:
4342:
4327:
4321:
4320:
4318:
4316:
4288:
4282:
4281:
4271:
4261:
4236:
4230:
4229:
4227:
4225:
4219:
4213:. Archived from
4204:
4162:
4150:
4144:
4143:
4141:
4139:
4133:
4110:
4101:
4095:
4094:
4092:
4086:. Archived from
4053:
4044:
4038:
4037:
4031:
4023:
4013:
3981:
3972:
3953:
3947:
3946:
3928:
3922:
3921:
3911:
3879:
3873:
3872:
3870:
3868:
3863:on July 24, 2007
3862:
3847:
3838:
3832:
3820:
3814:
3813:
3803:
3793:
3761:
3755:
3741:
3735:
3724:
3706:
3683:
3668:
3667:
3639:
3630:
3629:
3619:
3587:
3558:
3557:
3555:
3553:
3547:
3512:
3503:
3492:
3491:
3473:
3467:
3466:
3442:
3436:
3435:
3433:
3431:
3412:
3399:
3393:
3392:
3382:
3372:
3348:
3342:
3341:
3339:
3337:
3309:
3300:
3299:
3291:
3285:
3284:
3282:
3280:
3259:
3253:
3252:
3250:
3248:
3239:. Archived from
3229:
3220:
3219:
3217:
3215:
3196:
3190:
3189:
3187:Del fr. pingouin
3171:
3165:
3164:
3150:
3144:
3143:
3141:
3139:
3124:
3115:
3114:
3090:
3081:
3080:
3057:
3048:
3047:
3037:
3019:
2987:
2981:
2980:
2978:
2976:
2961:
2955:
2954:
2952:
2950:
2934:
2928:
2927:
2925:
2923:
2895:
2889:
2888:
2886:
2884:
2865:
2859:
2858:
2856:
2854:
2834:
2828:
2827:
2817:
2793:
2776:
2775:
2731:
2725:
2724:
2722:
2720:
2714:
2699:
2690:
2622:Opus the Penguin
2610:in his strip in
2574:African penguins
2556:Burgess Meredith
2542:Detective Comics
2509:was released by
2478:, co-created by
2300:for consumption.
2240:Near Threatened
2209:Humboldt penguin
2167:Pygoscelis papua
2056:Near Threatened
1956:Macaroni penguin
1874:Near Threatened
1860:Last assessment
1845:
1782:
1757:
1681:Humboldt Current
1206:
1116:Icadyptes salasi
973:of penguins and
886:
885:
865:List of penguins
760:
754:
748:
700:
694:
682:
632:emperor penguins
530:
529:
526:
525:
522:
519:
516:
513:
510:
507:
504:
501:
498:
495:
492:
489:
473:
472:
469:
468:
465:
460:
459:
456:
453:
450:
447:
444:
441:
438:
435:
432:
415:flightless birds
396:
374:
361:
348:
335:
322:
309:
284:
267:
202:
201:
178:
164:
148:
134:
113:
112:
49:
28:
8660:
8659:
8655:
8654:
8653:
8651:
8650:
8649:
8630:Sphenisciformes
8595:
8594:
8593:
8588:
8580:
8575:
8567:
8565:
8557:
8552:
8544:
8539:
8531:
8529:
8521:
8516:
8508:
8503:
8495:
8490:
8482:
8477:
8469:
8464:
8456:
8451:
8443:
8438:
8430:
8425:
8417:
8412:
8404:
8399:
8391:
8389:
8381:
8376:
8368:
8363:
8354:
8353:
8348:
8339:
8338:
8333:
8320:
8310:
8305:
8217:
8211:
8203:
8195:
8194:Bucerotiformes
8188:(cuckoo-roller)
8187:
8179:
8171:
8163:
8155:
8147:
8146:Cathartiformes
8132:
8126:
8118:
8117:Psittaciformes
8110:
8102:
8087:
8081:
8080:Pelecaniformes
8073:
8065:
8057:
8049:
8041:
8026:
8020:
8019:Eurypygiformes
8012:
7997:
7991:
7983:
7968:
7962:
7947:
7941:
7923:
7908:
7902:
7894:
7886:
7858:
7852:
7844:
7829:
7823:
7822:Pterocliformes
7815:
7807:
7758:
7747:
7734:
7719:
7643:
7602:
7540:
7505:
7504:
7497:
7496:
7484:
7464:
7437:
7414:
7407:
7367:
7366:
7354:
7353:
7337:
7336:
7333:
7331:
7329:
7327:
7325:
7323:
7321:
7319:
7317:
7315:
7313:
7298:
7296:
7294:
7292:
7290:
7288:
7286:
7284:
7282:
7267:
7261:
7260:Casuariiformes
7253:
7252:Apterygiformes
7245:
7237:
7229:
7206:
7191:
7162:Lists by region
7130:
7040:
7033:
6974:Chaoyangiformes
6959:Jeholornithidae
6933:
6897:Origin of birds
6883:
6864:Brood parasites
6810:
6740:
6727:
6717:
6687:
6682:
6677:Chatham penguin
6625:
6620:Waitaha penguin
6599:
6595:African penguin
6563:
6535:
6521:Adélie penguin
6504:
6500:Emperor penguin
6478:
6460:
6445:
6440:
6330:
6329:
6328:
6313:
6312:
6308:
6301:
6291:
6270:
6267:
6262:
6246:
6242:
6232:Wayback Machine
6221:
6217:
6207:
6205:
6193:
6189:
6188:
6184:
6174:
6172:
6159:
6158:
6154:
6144:
6142:
6129:
6128:
6124:
6114:
6112:
6099:
6098:
6094:
6084:
6082:
6069:
6068:
6064:
6047:
6040:
6038:
6035:Beauty Of Birds
6029:
6028:
6024:
6011:
6010:
6006:
5996:
5994:
5980:
5979:
5975:
5965:
5963:
5949:
5948:
5944:
5934:
5932:
5918:
5917:
5913:
5903:
5901:
5887:
5886:
5882:
5872:
5870:
5856:
5855:
5851:
5841:
5839:
5825:
5824:
5820:
5810:
5808:
5794:
5793:
5789:
5779:
5777:
5763:
5762:
5758:
5748:
5746:
5732:
5731:
5727:
5717:
5715:
5701:
5700:
5696:
5686:
5684:
5670:
5669:
5665:
5655:
5653:
5639:
5638:
5634:
5624:
5622:
5608:
5607:
5603:
5593:
5591:
5577:
5576:
5572:
5562:
5560:
5546:
5545:
5541:
5531:
5529:
5515:
5514:
5510:
5500:
5498:
5484:
5483:
5479:
5469:
5467:
5453:
5452:
5448:
5438:
5436:
5422:
5421:
5417:
5409:
5405:
5398:
5385:
5384:
5380:
5373:
5359:Eudyptula minor
5356:
5355:
5351:
5336:(#4): 277–289.
5324:Eudyptula minor
5319:
5318:
5314:
5306:
5299:
5291:
5284:
5276:
5272:
5262:
5260:
5247:
5246:
5242:
5232:
5230:
5212:
5211:
5207:
5197:
5195:
5186:
5185:
5181:
5171:
5169:
5151:
5150:
5146:
5092:
5091:
5087:
5077:
5075:
5071:
5043:(#3): 331–351.
5032:
5027:
5026:
5022:
5001:(#6): 505–523.
4992:
4991:
4987:
4977:
4975:
4968:
4953:
4952:
4948:
4942:Greenwood Press
4932:
4928:
4918:
4916:
4906:
4905:
4901:
4895:Wayback Machine
4886:
4882:
4872:
4870:
4856:
4855:
4851:
4841:
4839:
4830:
4829:
4825:
4815:
4813:
4804:
4803:
4799:
4770:(#2): 249–256.
4761:
4760:
4756:
4749:10.1071/zo07053
4743:(#5): 317–321.
4730:
4729:
4725:
4715:
4714:
4710:
4664:
4663:
4659:
4649:
4647:
4633:
4632:
4628:
4570:
4569:
4565:
4527:
4526:
4522:
4456:
4455:
4451:
4441:
4439:
4425:
4424:
4420:
4410:
4408:
4394:
4393:
4389:
4372:
4368:
4355:
4354:
4350:
4340:
4338:
4329:
4328:
4324:
4314:
4312:
4305:
4290:
4289:
4285:
4238:
4237:
4233:
4223:
4221:
4217:
4160:
4152:
4151:
4147:
4137:
4135:
4131:
4108:
4103:
4102:
4098:
4090:
4062:(11): 2558–73.
4051:
4046:
4045:
4041:
4024:
3996:(#1582): 11–7.
3983:
3982:
3975:
3954:
3950:
3943:
3930:
3929:
3925:
3881:
3880:
3876:
3866:
3864:
3860:
3845:
3840:
3839:
3835:
3821:
3817:
3763:
3762:
3758:
3752:molecular clock
3742:
3738:
3733:Wayback Machine
3704:10.1.1.113.4549
3685:
3684:
3671:
3641:
3640:
3633:
3602:(#1582): 11–7.
3589:
3588:
3561:
3551:
3549:
3545:
3510:
3505:
3504:
3495:
3488:
3475:
3474:
3470:
3463:
3444:
3443:
3439:
3429:
3427:
3425:
3401:
3400:
3396:
3350:
3349:
3345:
3335:
3333:
3326:
3311:
3310:
3303:
3293:
3292:
3288:
3278:
3276:
3268:Merriam-Webster
3261:
3260:
3256:
3246:
3244:
3231:
3230:
3223:
3213:
3211:
3198:
3197:
3193:
3173:
3172:
3168:
3152:
3151:
3147:
3137:
3135:
3126:
3125:
3118:
3111:
3092:
3091:
3084:
3077:
3059:
3058:
3051:
2989:
2988:
2984:
2974:
2972:
2970:livescience.com
2963:
2962:
2958:
2948:
2946:
2936:
2935:
2931:
2921:
2919:
2912:
2897:
2896:
2892:
2882:
2880:
2867:
2866:
2862:
2852:
2850:
2836:
2835:
2831:
2808:(#5): 412–441.
2795:
2794:
2779:
2733:
2732:
2728:
2718:
2716:
2712:
2697:
2692:
2691:
2687:
2683:
2661:children's book
2454:emperor penguin
2452:process of the
2363:
2357:
2298:emperor penguin
2280:
2186:African penguin
1914:Eudyptula minor
1866:Emperor penguin
1840:
1800:
1799:
1798:
1790:
1785:
1784:
1783:
1772:
1750:
1748:
1740:Ile aux Cochons
1696:Bergmann's Rule
1671:zone; one, the
1661:
1655:
1624:
1569:emperor penguin
1471:
1417:Charadriiformes
1344:
1305:
1204:
1160:
1069:Palaeeudyptinae
1043:
1041:Palaeeudyptines
991:
967:plate tectonics
948:fully described
921:
896:African penguin
873:from the genus
867:
861:
856:
848:Sphenisciformes
828:Charadriiformes
812:flight feathers
792:
786:
722:Merriam-Webster
638:) in Antarctica
624:
588:Eudyptula minor
576:emperor penguin
551:dark and white
486:
482:
479:Sphenisciformes
462:
429:
425:
410:are a group of
283:
266:
264:Sphenisciformes
196:
188:
187:
186:
185:
184:
179:
171:
170:
165:
156:
155:
154:
149:
141:
140:
135:
120:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
44:
43:
32:
24:
17:
12:
11:
5:
8658:
8656:
8648:
8647:
8642:
8637:
8632:
8627:
8622:
8617:
8612:
8607:
8597:
8596:
8590:
8589:
8587:
8586:
8573:
8563:
8550:
8537:
8527:
8514:
8501:
8488:
8475:
8462:
8449:
8436:
8423:
8410:
8397:
8387:
8374:
8361:
8346:
8330:
8328:
8322:
8321:
8316:
8307:
8306:
8304:
8303:
8291:
8279:
8267:
8254:
8251:
8250:
8247:
8246:
8243:
8242:
8239:
8238:
8235:
8234:
8231:
8230:
8227:
8226:
8223:
8222:
8219:
8218:
8216:
8215:
8207:
8202:Coraciiformes
8199:
8191:
8183:
8178:Trogoniformes
8175:
8167:
8159:
8151:
8142:
8140:
8134:
8133:
8131:
8130:
8125:Passeriformes
8122:
8114:
8109:Falconiformes
8106:
8101:Cariamiformes
8097:
8095:
8089:
8088:
8086:
8085:
8077:
8069:
8064:Ciconiiformes
8061:
8053:
8045:
8036:
8034:
8028:
8027:
8025:
8024:
8016:
8007:
8005:
7999:
7998:
7996:
7995:
7987:
7978:
7976:
7974:Cursorimorphae
7970:
7969:
7967:
7966:
7957:
7955:
7949:
7948:
7946:
7945:
7937:
7932:
7927:
7918:
7916:
7910:
7909:
7907:
7906:
7898:
7890:
7881:
7879:
7870:
7864:
7863:
7860:
7859:
7857:
7856:
7848:
7839:
7837:
7835:Mirandornithes
7831:
7830:
7828:
7827:
7819:
7811:
7806:Columbiformes
7802:
7800:
7798:Columbimorphae
7791:
7782:
7772:
7771:
7768:
7767:
7764:
7763:
7760:
7759:
7757:
7756:
7751:
7743:
7738:
7733:Meleagridinae
7729:
7727:
7721:
7720:
7718:
7717:
7710:
7703:
7696:
7689:
7682:
7675:
7668:
7661:
7653:
7651:
7649:Odontophoridae
7645:
7644:
7642:
7641:
7634:
7627:
7620:
7612:
7610:
7604:
7603:
7601:
7600:
7593:
7586:
7579:
7572:
7565:
7558:
7550:
7548:
7542:
7541:
7539:
7538:
7533:
7528:
7522:
7520:
7511:
7490:
7489:
7486:
7485:
7483:
7482:
7474:
7472:
7466:
7465:
7463:
7462:
7455:
7447:
7445:
7439:
7438:
7436:
7435:
7430:
7428:Stictonettinae
7425:
7423:Dendrocygninae
7420:
7419:
7418:
7411:
7399:
7398:
7397:
7392:
7387:
7376:
7374:
7357:
7344:
7301:
7273:
7272:
7269:
7268:
7266:
7265:
7257:
7249:
7241:
7233:
7224:
7222:
7208:
7207:
7204:
7193:
7192:
7190:
7189:
7188:
7187:
7182:
7176:Notable birds
7174:
7169:
7164:
7159:
7154:
7149:
7144:
7138:
7136:
7132:
7131:
7129:
7128:
7126:Egg collecting
7123:
7118:
7113:
7108:
7103:
7098:
7093:
7088:
7083:
7078:
7077:
7076:
7066:
7061:
7056:
7051:
7045:
7043:
7035:
7034:
7032:
7031:
7026:
7021:
7016:
7011:
7009:Hesperornithes
7006:
7001:
6996:
6991:
6986:
6984:Ambiortiformes
6981:
6976:
6971:
6969:Enantiornithes
6966:
6961:
6956:
6951:
6943:
6941:
6935:
6934:
6932:
6931:
6926:
6921:
6916:
6911:
6910:
6909:
6904:
6893:
6891:
6885:
6884:
6882:
6881:
6876:
6871:
6866:
6861:
6856:
6851:
6846:
6841:
6836:
6831:
6826:
6820:
6818:
6812:
6811:
6809:
6808:
6803:
6798:
6793:
6788:
6783:
6778:
6773:
6772:
6771:
6761:
6756:
6750:
6748:
6742:
6741:
6739:
6738:
6732:
6729:
6728:
6718:
6716:
6715:
6708:
6701:
6693:
6684:
6683:
6681:
6680:
6674:
6669:
6664:
6659:
6654:
6649:
6647:Snares penguin
6644:
6638:
6636:
6627:
6626:
6624:
6623:
6617:
6611:
6609:
6601:
6600:
6598:
6597:
6592:
6587:
6582:
6576:
6574:
6565:
6564:
6562:
6561:
6555:
6552:Little penguin
6548:
6546:
6537:
6536:
6534:
6533:
6531:Gentoo penguin
6528:
6523:
6517:
6515:
6506:
6505:
6503:
6502:
6497:
6491:
6489:
6480:
6479:
6472:
6470:
6462:
6461:
6450:
6447:
6446:
6441:
6439:
6438:
6431:
6424:
6416:
6410:
6409:
6404:
6399:
6390:
6384:
6370:
6365:
6359:
6354:
6348:
6342:
6337:
6327:
6326:
6321:
6315:
6314:
6303:
6302:
6300:
6299:External links
6297:
6296:
6295:
6289:
6266:
6263:
6261:
6260:
6240:
6215:
6182:
6152:
6122:
6092:
6062:
6022:
6004:
5973:
5942:
5911:
5880:
5849:
5818:
5787:
5756:
5725:
5694:
5663:
5632:
5601:
5570:
5539:
5508:
5477:
5446:
5415:
5403:
5396:
5378:
5371:
5349:
5312:
5297:
5282:
5270:
5255:(in Turkish).
5240:
5205:
5179:
5144:
5085:
5020:
4985:
4966:
4946:
4926:
4899:
4880:
4849:
4823:
4797:
4754:
4731:Thomas, D.B.;
4723:
4708:
4673:(#6192): 675.
4657:
4626:
4563:
4520:
4473:(#3): 676–80.
4449:
4418:
4387:
4366:
4348:
4322:
4304:978-1285402222
4303:
4283:
4231:
4145:
4096:
4068:10.1554/04-235
4039:
3973:
3948:
3941:
3923:
3874:
3833:
3815:
3756:
3736:
3669:
3650:(3): 209–239.
3631:
3559:
3517:(3423): 1–18.
3493:
3486:
3468:
3461:
3437:
3423:
3394:
3343:
3324:
3301:
3286:
3254:
3221:
3191:
3166:
3145:
3116:
3109:
3082:
3075:
3049:
2982:
2956:
2929:
2910:
2890:
2860:
2829:
2777:
2726:
2708:(4): 604–617.
2684:
2682:
2679:
2569:Batman Returns
2359:Main article:
2356:
2353:
2310:Adélie penguin
2279:
2276:
2273:
2272:
2269:
2266:
2263:
2260:
2250:
2249:
2246:
2244:
2241:
2238:
2228:
2227:
2224:
2221:
2218:
2215:
2205:
2204:
2201:
2198:
2195:
2192:
2182:
2181:
2178:
2175:
2172:
2171:Least Concern
2169:
2163:Gentoo penguin
2159:
2158:
2155:
2152:
2149:
2148:Least Concern
2146:
2136:
2135:
2132:
2129:
2126:
2125:Least Concern
2123:
2117:Adélie penguin
2113:
2112:
2109:
2106:
2103:
2100:
2090:
2089:
2086:
2083:
2080:
2077:
2067:
2066:
2063:
2060:
2057:
2054:
2043:
2042:
2039:
2036:
2033:
2030:
2024:Snares penguin
2020:
2019:
2016:
2013:
2010:
2007:
1997:
1996:
1993:
1990:
1987:
1984:
1974:
1973:
1970:
1968:
1965:
1962:
1952:
1951:
1948:
1945:
1942:
1939:
1929:
1928:
1925:
1922:
1919:
1918:Least Concern
1916:
1910:Little penguin
1906:
1905:
1902:
1900:
1897:
1896:Least Concern
1894:
1884:
1883:
1880:
1878:
1875:
1872:
1862:
1861:
1858:
1855:
1852:
1849:
1839:
1836:
1792:Gentoo penguin
1787:
1786:
1777:
1776:
1775:
1774:
1773:
1771:
1768:
1747:
1744:
1654:
1651:
1639:Gourdin Island
1635:Adélie penguin
1623:
1620:
1561:Gentoo penguin
1470:
1467:
1444:Pelecaniformes
1354:autapomorphies
1343:
1340:
1309:global cooling
1304:
1301:
1208:autapomorphies
1159:
1156:
1140:Paraptenodytes
1136:toothed whales
1132:Squalodontidae
1123:about 36 mya.
1096:Seymour Island
1042:
1039:
990:
987:
920:
917:
860:
857:
855:
852:
788:Main article:
785:
782:
623:
620:
403:
402:
398:
397:
389:
388:
381:
380:
367:
354:
341:
328:
315:
299:
298:
291:
290:
278:
274:
273:
261:
257:
256:
251:
244:
243:
238:
234:
233:
228:
224:
223:
218:
214:
213:
208:
204:
203:
190:
189:
180:
173:
172:
166:
159:
158:
157:
150:
143:
142:
136:
129:
128:
127:
126:
125:
122:
121:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
50:
33:
15:
13:
10:
9:
6:
4:
3:
2:
8657:
8646:
8643:
8641:
8638:
8636:
8633:
8631:
8628:
8626:
8623:
8621:
8618:
8616:
8613:
8611:
8608:
8606:
8603:
8602:
8600:
8583:
8578:
8574:
8570:
8564:
8560:
8555:
8551:
8547:
8542:
8538:
8534:
8528:
8524:
8519:
8515:
8511:
8506:
8502:
8498:
8493:
8489:
8485:
8480:
8476:
8472:
8467:
8463:
8459:
8454:
8450:
8446:
8441:
8437:
8433:
8428:
8424:
8420:
8415:
8411:
8407:
8402:
8398:
8394:
8388:
8384:
8379:
8375:
8371:
8366:
8362:
8357:
8351:
8347:
8342:
8336:
8332:
8331:
8329:
8327:
8323:
8319:
8314:
8302:
8301:
8292:
8290:
8289:
8284:
8280:
8278:
8277:
8268:
8266:
8265:
8256:
8255:
8252:
8214:
8208:
8206:
8200:
8198:
8192:
8190:
8184:
8182:
8176:
8174:
8168:
8166:
8162:Strigiformes
8160:
8158:
8152:
8150:
8144:
8143:
8141:
8139:
8135:
8129:
8123:
8121:
8115:
8113:
8107:
8105:
8099:
8098:
8096:
8094:
8090:
8084:
8078:
8076:
8070:
8068:
8062:
8060:
8054:
8052:
8046:
8044:
8038:
8037:
8035:
8033:
8029:
8023:
8017:
8015:
8013:(tropicbirds)
8009:
8008:
8006:
8004:
8000:
7994:
7988:
7986:
7980:
7979:
7977:
7975:
7971:
7965:
7959:
7958:
7956:
7954:
7950:
7944:
7938:
7936:
7935:Podargiformes
7933:
7931:
7928:
7926:
7920:
7919:
7917:
7915:
7911:
7905:
7899:
7897:
7891:
7889:
7885:Cuculiformes
7883:
7882:
7880:
7878:
7874:
7871:
7869:
7865:
7855:
7849:
7847:
7841:
7840:
7838:
7836:
7832:
7826:
7820:
7818:
7812:
7810:
7804:
7803:
7801:
7799:
7795:
7792:
7790:
7786:
7783:
7781:
7777:
7773:
7755:
7752:
7750:
7744:
7742:
7739:
7737:
7731:
7730:
7728:
7726:
7722:
7716:
7715:
7711:
7709:
7708:
7704:
7702:
7701:
7697:
7695:
7694:
7690:
7688:
7687:
7683:
7681:
7680:
7676:
7674:
7673:
7669:
7667:
7666:
7662:
7660:
7659:
7655:
7654:
7652:
7650:
7646:
7640:
7639:
7635:
7633:
7632:
7628:
7626:
7625:
7621:
7619:
7618:
7614:
7613:
7611:
7609:
7605:
7599:
7598:
7594:
7592:
7591:
7587:
7585:
7584:
7583:Macrocephalon
7580:
7578:
7577:
7573:
7571:
7570:
7566:
7564:
7563:
7559:
7557:
7556:
7552:
7551:
7549:
7547:
7543:
7537:
7534:
7532:
7529:
7527:
7524:
7523:
7521:
7519:
7515:
7512:
7508:
7501:
7495:
7491:
7481:
7480:
7476:
7475:
7473:
7471:
7470:Anseranatidae
7467:
7461:
7460:
7456:
7454:
7453:
7449:
7448:
7446:
7444:
7440:
7434:
7431:
7429:
7426:
7424:
7421:
7417:
7412:
7410:
7405:
7404:
7403:
7400:
7396:
7393:
7391:
7388:
7386:
7383:
7382:
7381:
7378:
7377:
7375:
7371:
7365:
7361:
7358:
7352:
7348:
7345:
7341:
7335:
7309:
7305:
7302:
7300:
7278:
7274:
7264:
7258:
7256:
7250:
7248:
7244:Tinamiformes
7242:
7240:
7234:
7232:
7226:
7225:
7223:
7221:
7220:Palaeognathae
7217:
7213:
7209:
7202:
7198:
7194:
7186:
7183:
7181:
7178:
7177:
7175:
7173:
7170:
7168:
7165:
7163:
7160:
7158:
7155:
7153:
7150:
7148:
7145:
7143:
7140:
7139:
7137:
7133:
7127:
7124:
7122:
7119:
7117:
7114:
7112:
7109:
7107:
7106:Pigeon racing
7104:
7102:
7099:
7097:
7094:
7092:
7089:
7087:
7084:
7082:
7079:
7075:
7072:
7071:
7070:
7067:
7065:
7062:
7060:
7057:
7055:
7052:
7050:
7047:
7046:
7044:
7042:
7036:
7030:
7027:
7025:
7022:
7020:
7017:
7015:
7012:
7010:
7007:
7005:
7002:
7000:
6997:
6995:
6992:
6990:
6987:
6985:
6982:
6980:
6977:
6975:
6972:
6970:
6967:
6965:
6962:
6960:
6957:
6955:
6952:
6950:
6949:
6948:Archaeopteryx
6945:
6944:
6942:
6940:
6936:
6930:
6927:
6925:
6922:
6920:
6917:
6915:
6912:
6908:
6905:
6903:
6900:
6899:
6898:
6895:
6894:
6892:
6890:
6886:
6880:
6877:
6875:
6872:
6870:
6867:
6865:
6862:
6860:
6857:
6855:
6852:
6850:
6847:
6845:
6842:
6840:
6837:
6835:
6832:
6830:
6827:
6825:
6822:
6821:
6819:
6817:
6813:
6807:
6804:
6802:
6799:
6797:
6794:
6792:
6789:
6787:
6784:
6782:
6779:
6777:
6774:
6770:
6767:
6766:
6765:
6762:
6760:
6757:
6755:
6752:
6751:
6749:
6747:
6743:
6737:
6734:
6733:
6730:
6725:
6721:
6714:
6709:
6707:
6702:
6700:
6695:
6694:
6691:
6678:
6675:
6673:
6670:
6668:
6667:Royal penguin
6665:
6663:
6660:
6658:
6655:
6653:
6650:
6648:
6645:
6643:
6640:
6639:
6637:
6634:
6633:
6628:
6621:
6618:
6616:
6613:
6612:
6610:
6608:
6607:
6602:
6596:
6593:
6591:
6588:
6586:
6583:
6581:
6578:
6577:
6575:
6572:
6571:
6566:
6559:
6556:
6553:
6550:
6549:
6547:
6544:
6543:
6538:
6532:
6529:
6527:
6524:
6522:
6519:
6518:
6516:
6513:
6512:
6507:
6501:
6498:
6496:
6493:
6492:
6490:
6487:
6486:
6481:
6477:
6476:
6471:
6469:
6468:
6463:
6458:
6454:
6448:
6444:
6437:
6432:
6430:
6425:
6423:
6418:
6417:
6414:
6408:
6405:
6403:
6400:
6398:
6396:
6391:
6388:
6385:
6382:
6378:
6374:
6371:
6369:
6368:Penguin World
6366:
6363:
6360:
6358:
6355:
6352:
6349:
6346:
6343:
6341:
6338:
6335:
6332:
6331:
6325:
6322:
6320:
6317:
6316:
6311:
6306:
6298:
6292:
6286:
6282:
6277:
6276:
6269:
6268:
6264:
6258:
6257:9781317440895
6254:
6250:
6244:
6241:
6237:
6233:
6229:
6226:
6219:
6216:
6203:
6199:
6192:
6186:
6183:
6170:
6166:
6162:
6156:
6153:
6140:
6136:
6132:
6126:
6123:
6110:
6106:
6102:
6096:
6093:
6080:
6076:
6072:
6066:
6063:
6058:
6052:
6036:
6032:
6026:
6023:
6018:
6014:
6008:
6005:
5992:
5988:
5984:
5977:
5974:
5961:
5957:
5953:
5946:
5943:
5930:
5926:
5922:
5915:
5912:
5899:
5895:
5891:
5884:
5881:
5868:
5864:
5860:
5853:
5850:
5837:
5833:
5829:
5822:
5819:
5806:
5802:
5798:
5791:
5788:
5775:
5771:
5767:
5760:
5757:
5744:
5740:
5736:
5729:
5726:
5713:
5709:
5705:
5698:
5695:
5682:
5678:
5674:
5667:
5664:
5651:
5647:
5643:
5636:
5633:
5620:
5616:
5612:
5605:
5602:
5589:
5585:
5581:
5574:
5571:
5558:
5554:
5550:
5543:
5540:
5527:
5523:
5519:
5512:
5509:
5496:
5492:
5488:
5481:
5478:
5465:
5461:
5457:
5450:
5447:
5434:
5430:
5426:
5419:
5416:
5412:
5407:
5404:
5399:
5393:
5389:
5382:
5379:
5374:
5368:
5364:
5360:
5353:
5350:
5344:
5339:
5335:
5331:
5327:
5325:
5316:
5313:
5309:
5304:
5302:
5298:
5294:
5289:
5287:
5283:
5279:
5274:
5271:
5258:
5254:
5250:
5244:
5241:
5228:
5224:
5220:
5216:
5209:
5206:
5193:
5189:
5183:
5180:
5167:
5163:
5159:
5155:
5148:
5145:
5140:
5136:
5131:
5126:
5121:
5116:
5112:
5108:
5104:
5100:
5096:
5089:
5086:
5070:
5066:
5062:
5058:
5054:
5050:
5046:
5042:
5038:
5031:
5024:
5021:
5016:
5012:
5008:
5004:
5000:
4996:
4989:
4986:
4973:
4969:
4967:9780787290085
4963:
4959:
4958:
4950:
4947:
4943:
4939:
4935:
4930:
4927:
4914:
4910:
4903:
4900:
4896:
4892:
4889:
4884:
4881:
4868:
4864:
4860:
4853:
4850:
4837:
4833:
4827:
4824:
4811:
4807:
4801:
4798:
4793:
4789:
4785:
4781:
4777:
4773:
4769:
4765:
4758:
4755:
4750:
4746:
4742:
4738:
4734:
4733:Fordyce, R.E.
4727:
4724:
4719:
4712:
4709:
4704:
4700:
4696:
4692:
4688:
4684:
4680:
4676:
4672:
4668:
4661:
4658:
4645:
4641:
4637:
4630:
4627:
4622:
4618:
4614:
4610:
4606:
4602:
4598:
4594:
4590:
4586:
4582:
4578:
4574:
4567:
4564:
4559:
4555:
4551:
4547:
4543:
4539:
4535:
4531:
4524:
4521:
4516:
4512:
4507:
4502:
4498:
4494:
4489:
4484:
4480:
4476:
4472:
4468:
4464:
4462:
4453:
4450:
4437:
4433:
4429:
4422:
4419:
4406:
4402:
4398:
4391:
4388:
4383:
4377:
4369:
4363:
4359:
4352:
4349:
4336:
4332:
4326:
4323:
4310:
4306:
4300:
4296:
4295:
4287:
4284:
4279:
4275:
4270:
4265:
4260:
4255:
4251:
4247:
4243:
4235:
4232:
4216:
4212:
4208:
4203:
4198:
4194:
4190:
4186:
4182:
4178:
4174:
4170:
4166:
4159:
4155:
4154:Jarvis, E. D.
4149:
4146:
4130:
4126:
4122:
4118:
4114:
4107:
4100:
4097:
4089:
4085:
4081:
4077:
4073:
4069:
4065:
4061:
4057:
4050:
4043:
4040:
4035:
4029:
4021:
4017:
4012:
4007:
4003:
3999:
3995:
3991:
3987:
3980:
3978:
3974:
3970:
3966:
3962:
3958:
3952:
3949:
3944:
3938:
3934:
3927:
3924:
3919:
3915:
3910:
3905:
3901:
3897:
3893:
3889:
3885:
3878:
3875:
3859:
3855:
3851:
3844:
3837:
3834:
3830:
3827:
3824:
3819:
3816:
3811:
3807:
3802:
3797:
3792:
3787:
3783:
3779:
3775:
3771:
3767:
3760:
3757:
3753:
3749:
3746:
3740:
3737:
3734:
3730:
3727:
3722:
3718:
3714:
3710:
3705:
3700:
3696:
3692:
3691:
3682:
3680:
3678:
3676:
3674:
3670:
3665:
3661:
3657:
3653:
3649:
3645:
3638:
3636:
3632:
3627:
3623:
3618:
3613:
3609:
3605:
3601:
3597:
3593:
3586:
3584:
3582:
3580:
3578:
3576:
3574:
3572:
3570:
3568:
3566:
3564:
3560:
3544:
3540:
3536:
3532:
3528:
3524:
3520:
3516:
3509:
3502:
3500:
3498:
3494:
3489:
3483:
3479:
3472:
3469:
3464:
3462:9781486311637
3458:
3454:
3450:
3449:
3441:
3438:
3426:
3424:0-8032-2566-0
3420:
3416:
3411:
3410:
3404:
3398:
3395:
3390:
3386:
3381:
3376:
3371:
3366:
3362:
3358:
3354:
3347:
3344:
3331:
3327:
3325:0-19-856478-3
3321:
3317:
3316:
3308:
3306:
3302:
3297:
3290:
3287:
3274:
3270:
3269:
3264:
3258:
3255:
3242:
3238:
3234:
3228:
3226:
3222:
3209:
3205:
3201:
3195:
3192:
3188:
3184:
3180:
3176:
3170:
3167:
3162:
3161:
3156:
3149:
3146:
3133:
3129:
3123:
3121:
3117:
3112:
3106:
3102:
3098:
3097:
3089:
3087:
3083:
3078:
3072:
3068:
3067:
3062:
3061:Fuller, Errol
3056:
3054:
3050:
3045:
3041:
3036:
3031:
3027:
3023:
3018:
3013:
3009:
3005:
3001:
2997:
2993:
2986:
2983:
2971:
2967:
2960:
2957:
2949:September 24,
2944:
2940:
2933:
2930:
2917:
2913:
2911:9781465459008
2907:
2903:
2902:
2894:
2891:
2878:
2874:
2870:
2864:
2861:
2848:
2844:
2840:
2833:
2830:
2825:
2821:
2816:
2811:
2807:
2803:
2799:
2792:
2790:
2788:
2786:
2784:
2782:
2778:
2773:
2769:
2765:
2761:
2757:
2753:
2749:
2745:
2741:
2737:
2730:
2727:
2711:
2707:
2703:
2696:
2689:
2686:
2680:
2678:
2676:
2672:
2668:
2667:
2662:
2658:
2653:
2651:
2650:
2645:
2644:
2639:
2638:
2633:
2632:
2627:
2623:
2619:
2618:Falklands War
2615:
2614:
2609:
2604:
2602:
2598:
2594:
2590:
2586:
2585:United States
2581:
2579:
2578:king penguins
2575:
2571:
2570:
2565:
2561:
2557:
2553:
2552:
2548:. In the 60s
2547:
2543:
2539:
2536:
2532:
2528:
2524:
2519:
2517:
2512:
2508:
2507:
2501:
2499:
2495:
2491:
2487:
2486:
2481:
2480:Otmar Gutmann
2477:
2476:
2471:
2467:
2466:
2461:
2460:
2455:
2451:
2447:
2446:
2441:
2437:
2436:
2431:
2430:
2425:
2424:
2418:
2416:
2412:
2408:
2404:
2400:
2399:
2394:
2390:
2386:
2382:
2375:
2371:
2367:
2362:
2354:
2352:
2350:
2346:
2341:
2339:
2338:leopard seals
2335:
2331:
2327:
2323:
2319:
2311:
2306:
2299:
2296:preparing an
2295:
2294:
2289:
2284:
2277:
2270:
2267:
2264:
2261:
2259:
2255:
2252:
2251:
2247:
2245:
2242:
2239:
2237:
2233:
2230:
2229:
2225:
2222:
2219:
2216:
2214:
2210:
2207:
2206:
2202:
2199:
2196:
2193:
2191:
2187:
2184:
2183:
2179:
2176:
2173:
2170:
2168:
2164:
2161:
2160:
2156:
2153:
2150:
2147:
2145:
2141:
2138:
2137:
2133:
2130:
2127:
2124:
2122:
2118:
2115:
2114:
2110:
2107:
2104:
2101:
2099:
2095:
2092:
2091:
2087:
2084:
2081:
2078:
2076:
2072:
2069:
2068:
2064:
2061:
2058:
2055:
2052:
2048:
2047:Royal penguin
2045:
2044:
2040:
2037:
2034:
2031:
2029:
2025:
2022:
2021:
2017:
2014:
2011:
2008:
2006:
2002:
1999:
1998:
1994:
1991:
1988:
1985:
1983:
1979:
1976:
1975:
1971:
1969:
1966:
1963:
1961:
1957:
1954:
1953:
1949:
1946:
1943:
1940:
1938:
1934:
1931:
1930:
1926:
1923:
1920:
1917:
1915:
1911:
1908:
1907:
1903:
1901:
1898:
1895:
1893:
1889:
1886:
1885:
1881:
1879:
1876:
1873:
1871:
1867:
1864:
1863:
1859:
1856:
1853:
1850:
1847:
1846:
1843:
1837:
1835:
1833:
1827:
1823:
1820:
1818:
1814:
1813:king penguins
1808:
1806:
1797:
1793:
1789:
1781:
1769:
1765:
1761:
1745:
1743:
1741:
1737:
1733:
1729:
1725:
1721:
1717:
1713:
1709:
1705:
1700:
1697:
1688:
1684:
1682:
1678:
1674:
1670:
1666:
1660:
1652:
1650:
1647:
1640:
1636:
1633:
1628:
1621:
1619:
1617:
1613:
1609:
1604:
1602:
1597:
1593:
1589:
1587:
1583:
1579:
1578:afterfeathers
1575:
1570:
1562:
1558:
1554:
1552:
1548:
1544:
1539:
1535:
1531:
1529:
1525:
1521:
1517:
1513:
1512:countershaded
1508:
1506:
1502:
1497:
1493:
1484:
1475:
1468:
1466:
1464:
1460:
1455:
1453:
1448:
1445:
1441:
1437:
1433:
1429:
1425:
1424:Ciconiiformes
1420:
1418:
1414:
1410:
1406:
1402:
1398:
1394:
1390:
1386:
1378:
1373:
1369:
1367:
1363:
1359:
1355:
1351:
1350:
1341:
1339:
1337:
1333:
1328:
1325:
1324:
1318:
1314:
1310:
1302:
1300:
1298:
1294:
1290:
1286:
1282:
1278:
1274:
1269:
1267:
1263:
1259:
1255:
1251:
1250:South America
1247:
1246:
1241:
1240:
1236:
1231:
1229:
1225:
1221:
1217:
1213:
1209:
1203:
1199:
1195:
1194:
1189:
1186:
1185:
1179:
1177:
1173:
1169:
1165:
1157:
1155:
1154:of penguins.
1153:
1149:
1145:
1141:
1137:
1133:
1129:
1124:
1122:
1118:
1117:
1112:
1111:
1105:
1101:
1097:
1093:
1088:
1086:
1082:
1078:
1074:
1070:
1066:
1063:
1058:
1056:
1052:
1048:
1040:
1038:
1036:
1032:
1031:South America
1028:
1024:
1020:
1019:
1014:
1011:
1007:
1003:
1001:
996:
989:Basal fossils
988:
986:
984:
983:Maastrichtian
980:
976:
972:
968:
964:
960:
956:
951:
949:
945:
944:biogeographic
941:
938:Although the
934:
930:
925:
918:
916:
914:
909:
908:Linnean taxon
904:
899:
897:
893:
889:
880:
876:
872:
866:
858:
853:
851:
849:
845:
841:
837:
831:
829:
825:
821:
817:
813:
809:
805:
801:
797:
791:
783:
781:
779:
775:
771:
767:
762:
759:
753:
747:
743:
738:
736:
732:
728:
724:
723:
718:
717:
712:
711:
706:
705:
699:
695:, 'head' and
693:
689:
684:
681:
676:
672:
668:
664:
659:
657:
653:
649:
645:
637:
633:
628:
621:
619:
617:
613:
609:
605:
602:regions, and
601:
597:
593:
589:
585:
581:
577:
572:
570:
566:
562:
558:
554:
550:
549:countershaded
546:
542:
538:
534:
528:
480:
477:
471:
423:
420:
416:
413:
409:
399:
395:
390:
387:
386:
377:
373:
372:
368:
364:
360:
359:
355:
351:
350:Milne-Edwards
347:
346:
342:
338:
334:
333:
329:
325:
321:
320:
316:
312:
311:Miller, J. F.
308:
307:
303:
302:
300:
297:
292:
287:
282:
279:
276:
275:
270:
265:
262:
259:
258:
255:
252:
249:
246:
245:
242:
239:
236:
235:
232:
229:
226:
225:
222:
219:
216:
215:
212:
209:
206:
205:
200:
195:
191:
183:
177:
169:
163:
153:
147:
139:
133:
123:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
48:
41:
37:
29:
26:
22:
8625:Spheniscinae
8620:Spheniscidae
8383:Spheniscidae
8370:Spheniscidae
8356:Spheniscidae
8326:Spheniscidae
8325:
8298:
8286:
8274:
8262:
8172:(mousebirds)
8170:Coliiformes
8047:
8040:Gaviiformes
8032:Aequornithes
7940:Apodiformes
7901:Otidiformes
7877:Otidimorphae
7824:(sandgrouse)
7746:Phasianinae
7712:
7705:
7698:
7693:Odontophorus
7691:
7684:
7677:
7670:
7663:
7656:
7636:
7629:
7622:
7615:
7595:
7588:
7581:
7574:
7567:
7560:
7553:
7531:Oreophasinae
7477:
7457:
7450:
7355:(waterfowls)
7351:Anseriformes
7101:Cockfighting
7086:Conservation
7081:Bird feeding
7069:Birdwatching
7059:Ornithomancy
6999:Gansuiformes
6946:
6939:Fossil birds
6829:Intelligence
6630:
6604:
6568:
6540:
6509:
6495:King penguin
6483:
6473:
6465:
6442:
6394:
6309:
6274:
6265:Bibliography
6251:. Routledge.
6248:
6243:
6235:
6218:
6208:November 21,
6206:. Retrieved
6197:
6185:
6175:November 21,
6173:. Retrieved
6164:
6155:
6145:November 21,
6143:. Retrieved
6134:
6125:
6115:November 21,
6113:. Retrieved
6104:
6095:
6085:November 21,
6083:. Retrieved
6074:
6065:
6039:. Retrieved
6034:
6025:
6017:the original
6007:
5995:. Retrieved
5986:
5976:
5964:. Retrieved
5955:
5945:
5933:. Retrieved
5924:
5914:
5902:. Retrieved
5893:
5883:
5871:. Retrieved
5862:
5852:
5840:. Retrieved
5831:
5821:
5809:. Retrieved
5800:
5790:
5778:. Retrieved
5769:
5759:
5747:. Retrieved
5738:
5728:
5716:. Retrieved
5707:
5697:
5685:. Retrieved
5676:
5666:
5654:. Retrieved
5645:
5635:
5623:. Retrieved
5614:
5604:
5592:. Retrieved
5583:
5573:
5561:. Retrieved
5552:
5542:
5530:. Retrieved
5521:
5511:
5499:. Retrieved
5490:
5480:
5468:. Retrieved
5459:
5449:
5437:. Retrieved
5428:
5418:
5406:
5387:
5381:
5362:
5358:
5352:
5333:
5329:
5323:
5315:
5273:
5261:. Retrieved
5252:
5243:
5233:November 13,
5231:. Retrieved
5222:
5218:
5208:
5198:September 8,
5196:. Retrieved
5192:the original
5182:
5172:November 13,
5170:. Retrieved
5161:
5157:
5147:
5102:
5098:
5088:
5078:November 13,
5076:. Retrieved
5040:
5036:
5023:
4998:
4994:
4988:
4978:November 18,
4976:. Retrieved
4956:
4949:
4937:
4929:
4917:. Retrieved
4902:
4883:
4871:. Retrieved
4867:the original
4862:
4852:
4840:. Retrieved
4836:the original
4826:
4814:. Retrieved
4810:the original
4800:
4767:
4763:
4757:
4740:
4736:
4726:
4717:
4711:
4670:
4666:
4660:
4648:. Retrieved
4644:the original
4639:
4629:
4580:
4576:
4572:
4566:
4533:
4529:
4523:
4470:
4466:
4460:
4452:
4440:. Retrieved
4431:
4421:
4409:. Retrieved
4400:
4390:
4357:
4351:
4341:September 8,
4339:. Retrieved
4325:
4313:. Retrieved
4293:
4286:
4249:
4245:
4234:
4222:. Retrieved
4215:the original
4168:
4164:
4148:
4136:. Retrieved
4116:
4112:
4099:
4088:the original
4059:
4055:
4042:
4028:cite journal
3993:
3989:
3964:
3960:
3951:
3932:
3926:
3891:
3887:
3877:
3865:. Retrieved
3858:the original
3853:
3849:
3836:
3828:
3822:
3818:
3773:
3769:
3759:
3747:
3739:
3694:
3688:
3647:
3643:
3599:
3595:
3550:. Retrieved
3514:
3477:
3471:
3447:
3440:
3428:. Retrieved
3408:
3397:
3360:
3356:
3346:
3336:November 18,
3334:. Retrieved
3314:
3295:
3289:
3279:September 8,
3277:. Retrieved
3266:
3257:
3245:. Retrieved
3241:the original
3236:
3212:. Retrieved
3203:
3194:
3186:
3183:the original
3178:
3169:
3158:
3148:
3136:. Retrieved
3095:
3065:
2999:
2995:
2985:
2973:. Retrieved
2969:
2959:
2947:. Retrieved
2943:the original
2932:
2922:November 18,
2920:. Retrieved
2900:
2893:
2881:. Retrieved
2872:
2863:
2851:. Retrieved
2842:
2832:
2805:
2801:
2739:
2735:
2729:
2717:. Retrieved
2705:
2701:
2688:
2675:New York Zoo
2664:
2654:
2647:
2641:
2635:
2631:Bloom County
2629:
2613:The Guardian
2611:
2605:
2591:team in the
2582:
2567:
2564:Danny Devito
2549:
2541:
2531:supervillain
2520:
2504:
2502:
2497:
2493:
2492:), dancing (
2489:
2483:
2473:
2463:
2457:
2443:
2433:
2427:
2421:
2419:
2398:Chilly Willy
2396:
2378:
2374:Linux kernel
2342:
2315:
2291:
2257:
2235:
2212:
2189:
2166:
2143:
2120:
2108:2,528–3,480
2097:
2074:
2050:
2027:
2015:2,500–9,999
2004:
1981:
1959:
1936:
1913:
1891:
1888:King penguin
1869:
1841:
1828:
1824:
1821:
1809:
1804:
1801:
1732:South Africa
1701:
1693:
1662:
1643:
1611:
1605:
1598:
1594:
1590:
1573:
1566:
1540:
1536:
1532:
1528:leopard seal
1509:
1494:life. Their
1489:
1456:
1452:sister group
1449:
1432:plotopterids
1421:
1382:
1366:DNA sequence
1347:
1345:
1329:
1321:
1306:
1292:
1276:
1272:
1270:
1243:
1237:
1232:
1219:
1201:
1191:
1190:
1182:
1180:
1161:
1152:biodiversity
1125:
1114:
1108:
1104:monophyletic
1089:
1085:Subantarctic
1062:paraphyletic
1059:
1044:
1016:
1015:
1009:
998:
992:
952:
940:evolutionary
937:
929:Bruny Island
900:
887:
868:
844:Spheniscidae
835:
832:
819:
807:
795:
793:
777:
773:
769:
765:
763:
739:
720:
714:
708:
702:
685:
662:
660:
643:
641:
635:
600:subantarctic
587:
579:
573:
478:
422:Spheniscidae
421:
407:
406:
382:
369:
356:
343:
330:
317:
304:
281:Spheniscidae
280:
263:
247:
25:
8466:iNaturalist
8350:Wikispecies
8300:WikiProject
8210:Piciformes
8093:Australaves
8072:Suliformes
7990:Gruiformes
7845:(flamingos)
7754:Tetraoninae
7725:Phasianidae
7714:Rhynchortyx
7679:Dactylortyx
7546:Megapodidae
7536:Penelopinae
7494:Galliformes
7236:Rheiformes
7230:(ostriches)
7180:individuals
7054:Ornithology
7041:interaction
6796:Preen gland
6485:Aptenodytes
5164:: 285–298.
4934:Piper, Ross
4863:iafrica.com
4442:January 20,
4411:January 20,
4246:GigaScience
4193:10072/67425
3961:Aptenodytes
3888:GigaScience
3867:February 2,
3138:January 25,
2904:. Penguin.
2898:DK (2016).
2411:polar bears
2349:Stephen Fry
2265:Decreasing
2262:Endangered
2243:Decreasing
2220:Decreasing
2217:Vulnerable
2197:Decreasing
2194:Endangered
2151:Decreasing
2128:Increasing
2105:Decreasing
2102:Endangered
2082:Decreasing
2079:Endangered
2053:(disputed)
2032:Vulnerable
2012:Decreasing
2009:Vulnerable
1989:Decreasing
1986:Endangered
1967:Decreasing
1964:Vulnerable
1944:Decreasing
1941:Vulnerable
1899:Increasing
1796:Brown Bluff
1728:New Zealand
1551:nearsighted
1389:paleognaths
1362:homoplasies
1224:Burdigalian
1212:adaptations
1202:Aptenodytes
1184:Aptenodytes
630:A group of
596:prehistoric
535:: only one
306:Aptenodytes
8599:Categories
8050:(penguins)
7903:(bustards)
7741:Perdicinae
7686:Dendrortyx
7658:Callipepla
7590:Megapodius
7555:Aepypodius
7433:Tadorninae
7415:true geese
7246:(tinamous)
7205:Neornithes
7116:Pheasantry
7091:Aviculture
6859:Incubation
6849:Lek mating
6606:Megadyptes
6570:Spheniscus
6511:Pygoscelis
6279:. Oxford:
4650:October 7,
4315:August 15,
4252:(#1): 27.
4224:August 28,
3965:Pygoscelis
3957:allopatric
3644:Cladistics
3552:January 4,
3247:October 7,
3175:"Pingüino"
2802:Cladistics
2681:References
2608:Steve Bell
2560:Tim Burton
2494:Happy Feet
2423:Happy Feet
2407:sub-Arctic
2389:North Pole
2318:Antarctica
2154:8,000,000
2131:7,580,000
2062:1,700,000
1947:2,500,000
1817:incubation
1764:Antarctica
1738:'s remote
1708:Antarctica
1665:Antarctica
1657:See also:
1646:isabelline
1632:isabelline
1586:filoplumes
1516:camouflage
1436:cormorants
1411:, and the
1323:Spheniscus
1273:Megadyptes
1254:carotenoid
1239:Spheniscus
1220:Pygoscelis
1198:morphology
1193:Pygoscelis
1181:The genus
1100:Priabonian
1018:Perudyptes
1002:manneringi
875:Spheniscus
846:and order
371:Spheniscus
358:Pygoscelis
345:Megadyptes
116:Cretaceous
45:62–0
8119:(parrots)
7963:(hoatzin)
7914:Strisores
7895:(turacos)
7887:(cuckoos)
7816:(mesites)
7735:(turkeys)
7707:Philortyx
7624:Agelastes
7617:Acryllium
7608:Numididae
7597:Talegalla
7507:gamebirds
7500:landfowls
7479:Anseranas
7443:Anhimidae
7402:Anserinae
7185:fictional
6907:dinosaurs
6902:Theropoda
6889:Evolution
6834:Migration
6816:Behaviour
6679:(extinct)
6622:(extinct)
6542:Eudyptula
6198:3 News NZ
6165:3 News NZ
6135:3 News NZ
6105:3 News NZ
6075:3 News NZ
6041:April 17,
5429:BBC Earth
5263:August 1,
4919:August 8,
4376:cite book
4119:: 61–71.
4056:Evolution
3699:CiteSeerX
3531:2246/2788
3233:"penguin"
3214:April 29,
3200:"penguin"
3155:"penguin"
3026:0027-8424
2975:August 4,
2853:April 16,
2719:March 21,
2535:superhero
2523:DC Comics
2521:In 1941,
2498:Surf's Up
2450:migration
2429:Surf's Up
2385:white tie
2326:sled dogs
2293:Endurance
1805:Agonistic
1746:Behaviour
1716:Australia
1712:Argentina
1669:temperate
1608:great auk
1478:Phalanges
1401:waterfowl
1303:Geography
1297:Tortonian
1281:latitudes
1245:Eudyptula
1176:Patagonia
1128:Paleogene
1092:radiation
1081:Antarctic
1077:Patagonia
1073:phylogeny
1065:subfamily
1047:Oligocene
1027:Bartonian
1023:Argentina
979:Campanian
963:Byrd Land
919:Evolution
913:phylogeny
836:Pinguinus
820:Pinguinus
804:great auk
796:Pinguinus
790:Great auk
784:Pinguinus
648:great auk
642:The word
622:Etymology
474:) of the
417:from the
337:Bonaparte
332:Eudyptula
286:Bonaparte
217:Kingdom:
211:Eukaryota
152:Chinstrap
114:Possible
8615:Seabirds
8610:Penguins
8390:BioLib:
8335:Wikidata
8264:Category
8138:Afroaves
8066:(storks)
7868:Passerea
7853:(grebes)
7789:Columbea
7700:Oreortyx
7672:Cyrtonyx
7562:Alectura
7526:Cracinae
7518:Cracidae
7395:Oxyurini
7385:Aythyini
7380:Anatinae
7364:Anatidae
7111:Falconry
7074:big year
6929:Seabirds
6839:Foraging
6786:Feathers
6632:Eudyptes
6443:Penguins
6373:Penguins
6228:Archived
6202:Archived
6169:Archived
6139:Archived
6109:Archived
6079:Archived
6051:cite web
5991:Archived
5960:Archived
5929:Archived
5898:Archived
5867:Archived
5836:Archived
5805:Archived
5774:Archived
5743:Archived
5712:Archived
5681:Archived
5650:Archived
5619:Archived
5588:Archived
5557:Archived
5526:Archived
5495:Archived
5464:Archived
5439:March 9,
5433:Archived
5411:Williams
5308:Williams
5293:Williams
5278:Williams
5257:Archived
5227:Archived
5225:: 3–28.
5166:Archived
5139:17601778
5069:Archived
5065:11954818
4972:Archived
4936:(2007),
4913:Archived
4891:Archived
4873:July 21,
4842:July 21,
4816:July 21,
4792:23175291
4784:22213564
4703:47562824
4621:32860474
4558:45578269
4550:10373249
4436:Archived
4405:Archived
4335:Archived
4309:Archived
4278:25671092
4211:25504713
4129:Archived
4076:15612298
4020:16519228
3918:31531675
3810:23690614
3729:Archived
3721:16533822
3664:85071808
3626:16519228
3543:Archived
3539:19041678
3405:(1987).
3389:15329156
3330:Archived
3273:Archived
3208:Archived
3132:Archived
3063:(2003).
3044:23918397
2916:Archived
2883:July 19,
2877:Archived
2847:Archived
2824:85673628
2764:18583609
2710:Archived
2595:and the
2516:peekaboo
2415:walruses
2177:774,000
2085:150,000
1992:480,600
1924:469,760
1877:Unknown
1848:Species
1770:Breeding
1582:plumules
1547:colonies
1440:anhingas
1413:seabirds
1377:flippers
1317:ice ages
1293:Eudyptes
1289:Langhian
1277:Eudyptes
1266:Pliocene
1262:Chattian
1087:coasts.
1037:waters.
1035:Atlantic
933:Tasmania
859:Taxonomy
752:fettgans
735:Pen Gwyn
680:pingouin
610:35
569:sea life
408:Penguins
324:Vieillot
319:Eudyptes
277:Family:
231:Chordata
227:Phylum:
221:Animalia
207:Domain:
31:Penguins
8276:Commons
7780:Neoaves
7665:Colinus
7631:Guttera
7569:Eulipoa
7390:Mergini
7254:(kiwis)
7238:(rheas)
7049:Ringing
6874:Hybrids
6869:Nesting
6824:Singing
6801:Plumage
6776:Dactyly
6746:Anatomy
6736:Outline
6726:: Aves)
6475:Species
6310:Penguin
6194:(Video)
5997:June 2,
5966:June 2,
5935:June 2,
5904:June 2,
5873:June 2,
5842:June 2,
5811:June 2,
5780:June 2,
5749:June 2,
5718:June 2,
5687:June 2,
5656:June 2,
5625:June 2,
5594:June 2,
5563:June 2,
5532:June 2,
5501:June 2,
5470:June 2,
5413:, p. 24
5310:, p. 23
5295:, p. 57
5280:, p. 17
5130:1913862
5107:Bibcode
5045:Bibcode
5003:Bibcode
4695:3173490
4675:Bibcode
4613:2881308
4585:Bibcode
4515:5259756
4475:Bibcode
4269:4322438
4202:4405904
4173:Bibcode
4165:Science
4138:July 8,
4084:1296408
4011:1560011
3909:6904868
3831:(2006).
3801:3677478
3778:Bibcode
3617:1560011
3430:May 11,
3415:265–266
3237:Wordnik
3035:3761583
3004:Bibcode
2901:Animal!
2772:6472805
2744:Bibcode
2736:Science
2673:in the
2637:Outland
2527:Penguin
2442:films;
2290:on the
2223:32,000
2200:50,000
2174:Stable
2059:Stable
2038:63,000
2035:Stable
1921:Stable
1832:crèches
1724:Namibia
1612:penguin
1543:hearing
1501:plumage
1492:aquatic
1459:Puffins
1385:Neoaves
1349:Waimanu
1228:Miocene
1216:habitat
1144:Neogene
1067:called
1010:Waimanu
1000:Waimanu
816:extinct
758:vetgans
746:pinguis
675:Spanish
663:penguin
644:penguin
608:equator
594:. Some
553:plumage
545:Equator
537:species
412:aquatic
376:Brisson
294:Modern
260:Order:
237:Class:
182:African
138:Emperor
8582:196034
8566:uBio:
8546:494367
8530:NZOR:
8497:174443
8484:100371
8445:1ZPHNF
8288:Portal
8164:(owls)
7638:Numida
7576:Leipoa
7459:Chauna
7452:Anhima
7147:Genera
7121:Imping
7039:Human
6879:Colony
6806:Vision
6791:Flight
6457:family
6307:about
6287:
6255:
5394:
5369:
5137:
5127:
5063:
4964:
4790:
4782:
4701:
4693:
4667:Nature
4619:
4611:
4603:
4556:
4548:
4513:
4506:223504
4503:
4495:
4364:
4301:
4276:
4266:
4209:
4199:
4082:
4074:
4018:
4008:
3939:
3916:
3906:
3829:et al.
3823:Contra
3808:
3798:
3748:et al.
3719:
3701:
3662:
3624:
3614:
3537:
3484:
3459:
3421:
3387:
3380:515296
3377:
3363:: 28.
3322:
3107:
3073:
3042:
3032:
3024:
2908:
2822:
2770:
2762:
2551:Batman
2538:Batman
2456:; and
2438:, all
2393:Arctic
2336:, and
2330:sharks
2268:1,200
1854:Trend
1736:France
1730:, and
1704:Angola
1584:, and
1426:or to
1405:storks
1358:grebes
1235:genera
1164:clades
995:fossil
774:waddle
713:, the
707:, the
671:Breton
667:French
616:Eocene
539:, the
419:family
363:Wagler
296:genera
288:, 1831
271:, 1891
269:Sharpe
168:Snares
40:Recent
36:Danian
8577:WoRMS
8569:21597
8559:39710
8479:IRMNG
8341:Q9147
7408:swans
7370:ducks
7340:fowls
7135:Lists
6759:Brain
6724:class
6720:Birds
6467:Genus
6453:order
5072:(PDF)
5061:S2CID
5033:(PDF)
4788:S2CID
4699:S2CID
4617:S2CID
4605:36191
4601:JSTOR
4554:S2CID
4497:59401
4493:JSTOR
4218:(PDF)
4161:(PDF)
4132:(PDF)
4109:(PDF)
4091:(PDF)
4080:S2CID
4052:(PDF)
3969:Baker
3894:(9).
3861:(PDF)
3846:(PDF)
3826:Baker
3754:used.
3745:Baker
3660:S2CID
3546:(PDF)
3535:S2CID
3511:(PDF)
2820:S2CID
2768:S2CID
2713:(PDF)
2698:(PDF)
2529:as a
2506:Pengo
2475:Pingu
2334:orcas
2271:2018
2248:2018
2226:2018
2203:2018
2180:2018
2157:2018
2134:2018
2111:2018
2088:2016
2065:2018
2041:2018
2018:2018
1995:2018
1972:2018
1950:2018
1927:2018
1904:2018
1882:2018
1720:Chile
1574:types
1526:or a
1520:wings
1505:tails
1496:wings
1409:rails
1205:'
1168:order
1094:: on
955:basal
892:wedge
888:sphēn
881:word
879:Greek
766:cocks
742:Latin
731:Welsh
688:Welsh
565:squid
557:krill
476:order
248:Clade
8523:9231
8518:NCBI
8492:ITIS
8471:3806
8458:5284
8453:GBIF
8440:EPPO
8432:7986
8406:1484
8401:BOLD
8393:8345
6781:Eggs
6769:milk
6764:Crop
6754:Beak
6379:the
6285:ISBN
6253:ISBN
6210:2012
6177:2012
6147:2012
6117:2012
6087:2012
6057:link
6043:2021
5999:2020
5968:2020
5937:2020
5906:2020
5875:2020
5844:2020
5813:2020
5782:2020
5751:2020
5720:2020
5689:2020
5658:2020
5627:2020
5596:2020
5565:2020
5534:2020
5503:2020
5472:2020
5441:2020
5392:ISBN
5367:ISBN
5265:2018
5235:2014
5200:2013
5174:2014
5135:PMID
5080:2014
4980:2020
4962:ISBN
4921:2016
4875:2006
4844:2006
4818:2006
4780:PMID
4691:PMID
4652:2018
4609:PMID
4546:PMID
4511:PMID
4444:2021
4413:2021
4382:link
4362:ISBN
4343:2013
4317:2018
4299:ISBN
4274:PMID
4226:2015
4207:PMID
4140:2006
4072:PMID
4034:link
4016:PMID
3937:ISBN
3914:PMID
3869:2007
3806:PMID
3717:PMID
3622:PMID
3554:2007
3482:ISBN
3457:ISBN
3432:2010
3419:ISBN
3385:PMID
3338:2020
3320:ISBN
3281:2013
3249:2014
3216:2016
3140:2010
3105:ISBN
3071:ISBN
3040:PMID
3022:ISSN
2977:2024
2951:2022
2924:2020
2906:ISBN
2885:2024
2855:2023
2760:PMID
2721:2021
2659:. A
2643:Opus
2640:and
2576:and
2511:Sega
2432:and
2413:and
2403:myth
2381:gait
2322:Dogs
2288:cook
2286:The
1606:The
1524:orca
1514:for
1438:and
1393:fowl
1391:and
1271:The
1242:and
1233:The
1174:and
1121:Peru
1083:and
953:The
942:and
884:σφήν
824:auks
778:raft
770:hens
719:and
698:gwyn
561:fish
378:1760
365:1832
352:1880
339:1856
326:1816
313:1778
241:Aves
53:PreꞒ
8505:NBN
8427:EoL
8419:GHC
8414:CoL
8378:AFD
8365:ADW
6375:in
5338:doi
5125:PMC
5115:doi
5103:104
5053:doi
5011:doi
4772:doi
4768:295
4745:doi
4683:doi
4671:335
4593:doi
4581:229
4538:doi
4501:PMC
4483:doi
4264:PMC
4254:doi
4197:PMC
4189:hdl
4181:doi
4169:346
4121:doi
4064:doi
4006:PMC
3998:doi
3994:273
3904:PMC
3896:doi
3796:PMC
3786:doi
3774:110
3709:doi
3652:doi
3612:PMC
3604:doi
3600:273
3527:hdl
3519:doi
3375:PMC
3365:doi
3030:PMC
3012:doi
3000:110
2810:doi
2752:doi
2740:320
2599:in
2580:).
2440:CGI
2370:Tux
2308:An
1762:in
1637:on
1630:An
1006:mya
692:pen
673:or
612:mya
515:ɔːr
8601::
8579::
8556::
8543::
8520::
8507::
8494::
8481::
8468::
8455::
8442::
8429::
8416::
8403::
8380::
8367::
8352::
8337::
6283:.
6234:.
6196:.
6163:.
6133:.
6103:.
6073:.
6053:}}
6049:{{
6033:.
5989:.
5985:.
5958:.
5954:.
5927:.
5923:.
5896:.
5892:.
5865:.
5861:.
5834:.
5830:.
5803:.
5799:.
5772:.
5768:.
5741:.
5737:.
5710:.
5706:.
5679:.
5675:.
5648:.
5644:.
5617:.
5613:.
5586:.
5582:.
5555:.
5551:.
5524:.
5520:.
5493:.
5489:.
5462:.
5458:.
5431:.
5427:.
5334:27
5332:.
5328:.
5326:)"
5300:^
5285:^
5251:.
5223:30
5221:.
5217:.
5162:52
5160:.
5156:.
5133:.
5123:.
5113:.
5101:.
5097:.
5067:.
5059:.
5051:.
5041:30
5039:.
5035:.
5009:.
4999:11
4997:.
4970:.
4940:,
4861:.
4786:.
4778:.
4766:.
4741:55
4739:.
4697:.
4689:.
4681:.
4669:.
4638:.
4615:.
4607:.
4599:.
4591:.
4579:.
4552:.
4544:.
4534:57
4532:.
4509:.
4499:.
4491:.
4481:.
4471:63
4469:.
4465:.
4430:.
4399:.
4378:}}
4374:{{
4307:.
4272:.
4262:.
4248:.
4244:.
4205:.
4195:.
4187:.
4179:.
4167:.
4163:.
4127:.
4117:43
4115:.
4111:.
4078:.
4070:.
4060:58
4058:.
4054:.
4030:}}
4026:{{
4014:.
4004:.
3992:.
3988:.
3976:^
3971:).
3963:,
3912:.
3902:.
3890:.
3886:.
3854:27
3852:.
3848:.
3804:.
3794:.
3784:.
3772:.
3768:.
3715:.
3707:.
3695:23
3693:.
3672:^
3658:.
3648:21
3646:.
3634:^
3620:.
3610:.
3598:.
3594:.
3562:^
3541:.
3533:.
3525:.
3513:.
3496:^
3451:.
3417:.
3383:.
3373:.
3359:.
3355:.
3328:.
3304:^
3265:.
3235:.
3224:^
3206:.
3202:.
3177:.
3157:.
3119:^
3103:.
3101:10
3085:^
3052:^
3038:.
3028:.
3020:.
3010:.
2998:.
2994:.
2968:.
2914:.
2875:.
2871:.
2841:.
2818:.
2806:22
2804:.
2800:.
2780:^
2766:.
2758:.
2750:.
2738:.
2706:62
2704:.
2700:.
2677:.
2663:,
2652:.
2634:,
2620:.
2603:.
2426:,
2417:.
2332:,
2256:,
2234:,
2211:,
2188:,
2165:,
2142:,
2119:,
2096:,
2073:,
2049:,
2026:,
2003:,
1980:,
1958:,
1935:,
1912:,
1890:,
1868:,
1834:.
1726:,
1722:,
1718:,
1714:,
1710:,
1706:,
1618:.
1580:,
1465:.
1419:.
1407:,
931:,
850:.
830:.
780:.
761:.
733::
669:,
563:,
559:,
521:iː
467:aɪ
461:,-
458:iː
250::
103:Pg
47:Ma
42:,
7509:)
7502:-
7498:(
7372:)
7368:(
7342:)
7338:(
7334:e
7332:a
7330:r
7328:e
7326:s
7324:n
7322:a
7320:o
7318:l
7316:l
7314:a
7312:G
7299:e
7297:a
7295:h
7293:t
7291:a
7289:n
7287:g
7285:o
7283:e
7281:N
6722:(
6712:e
6705:t
6698:v
6451:(
6435:e
6428:t
6421:v
6293:.
6212:.
6179:.
6149:.
6119:.
6089:.
6059:)
6045:.
6001:.
5970:.
5939:.
5908:.
5877:.
5846:.
5815:.
5784:.
5753:.
5722:.
5691:.
5660:.
5629:.
5598:.
5567:.
5536:.
5505:.
5474:.
5443:.
5400:.
5375:.
5346:.
5340::
5267:.
5237:.
5202:.
5176:.
5141:.
5117::
5109::
5082:.
5055::
5047::
5017:.
5013::
5005::
4982:.
4944:.
4923:.
4877:.
4846:.
4820:.
4794:.
4774::
4751:.
4747::
4705:.
4685::
4677::
4654:.
4623:.
4595::
4587::
4560:.
4540::
4517:.
4485::
4477::
4446:.
4415:.
4384:)
4370:.
4345:.
4319:.
4280:.
4256::
4250:3
4228:.
4191::
4183::
4175::
4142:.
4123::
4066::
4036:)
4022:.
4000::
3945:.
3920:.
3898::
3892:8
3871:.
3812:.
3788::
3780::
3723:.
3711::
3666:.
3654::
3628:.
3606::
3556:.
3529::
3521::
3490:.
3465:.
3434:.
3391:.
3367::
3361:4
3340:.
3283:.
3251:.
3218:.
3163:.
3142:.
3113:.
3079:.
3046:.
3014::
3006::
2979:.
2953:.
2926:.
2887:.
2857:.
2826:.
2812::
2774:.
2754::
2746::
2723:.
2540:(
1275:–
981:–
890:"
806:(
729:(
634:(
586:(
578:(
527:/
524:z
518:m
512:f
509:ə
506:s
503:ɪ
500:n
497:ˈ
494:ɪ
491:f
488:s
485:/
481:(
470:/
464:d
455:d
452:ɪ
449:s
446:ɪ
443:n
440:ˈ
437:ɪ
434:f
431:s
428:/
424:(
108:N
98:K
93:J
88:T
83:P
78:C
73:D
68:S
63:O
58:Ꞓ
38:-
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.