113:
822:(Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
134:
720:(Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
595:(Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian
527:– Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma.
336:
as subfamilies
Permocupedinae and Taldycupedinae respectively. However, other studies have recovered Talycupedidae as more closely related to crown-group beetles than to Permocupedidae.
879:– Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma; considered indeterminate to genus by Kirejtshuk (2020)
1031:
Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.).
953:[Permian fossil insects from the Arkhangelsk District. Part II. Neuroptera, Megaloptera and Coleoptera with the description of two new beetles from Tikhie Gory]
1357:
649:(Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma.
892:– Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma.
1344:
950:Пермские ископаемые насекомые Архангельского края. Часть II. Сетчатокрылые, вислокрылые и жуки, с приложением описания двух новых жуков из Тихих Гор
467:– Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma).
565:– Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
445:– Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
423:– Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
1426:
1436:
1092:"Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea"
918:
Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
1188:
Ponomarenko, A. G.; Yan, E. V.; Huang, Di-ying (2014). "New beetles (Coleoptera) from the terminal Middle
Permian of China".
278:
stem group beetles. They first appeared during the Early
Permian, and were one of the dominant groups of beetles during the
112:
591:(Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma;
133:
978:"First Triassic record of the beetle family Permocupedidae (Insecta: Coleoptera): a peculiar example of a Lazarus taxon"
392:(Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma
229:
1139:
Boudinot, Brendon Elias; Yan, Evgeny
Viktorovich; Prokop, Jakub; Luo, Xiao-Zhu; Beutel, Rolf Georg (2022-07-28).
1141:"Permian parallelisms: Reanalysis of †Tshekardocoleidae sheds light on the earliest evolution of the Coleoptera"
948:
1431:
1416:
1411:
683:), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
1421:
1261:
915:
1383:
782:, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
596:
1331:
1197:
1103:
989:
1299:
797:
558:
353:
580:
520:
490:
416:
292:
1170:
1013:
885:
717:
250:
128:
1388:
907:
702:
664:
371:
638:
438:
341:
1370:
1318:
1162:
1121:
1072:
1054:
1005:
775:
767:
1375:
1362:
460:
1239:
1205:
1152:
1111:
1062:
1044:
997:
502:
305:
260:
779:
592:
498:
389:
297:
275:
211:
198:
1201:
1107:
993:
1067:
1032:
819:
676:
379:
333:
1405:
1174:
1017:
977:
680:
328:
70:
1323:
588:
349:
1284:
1293:
1116:
1091:
309:
279:
45:
1209:
1001:
322:
90:
55:
17:
1244:
1227:
1166:
1125:
1058:
1009:
283:
165:
145:
95:
1076:
1310:
1278:
312:(wood eating), which is presumed to be the ancestral ecology of beetles.
287:
85:
80:
65:
60:
50:
1049:
1349:
1157:
1140:
1033:"Early evolution of beetles regulated by the end-Permian deforestation"
301:
100:
75:
185:
175:
155:
1255:
646:
1336:
1228:"Permian Insects from Paraná Basin, South Brazil, IV Coleoptera"
320:
Kirejtshuk (2020) included the following genera in an expanded (
1259:
976:
Ponomarenko, Alexandr G.; Bashkuev, Alexei S. (December 2018).
356:, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma.
410:– considered indeterminate to genus by Kirejtshuk (2020)
959:
505:
Middle
Triassic, Olenekian/Anisian, −247.2–242.0 Ma.
694:
Shcherbakov, Makarkin, Aristov & Vasilenko, 2009
673:
Shcherbakov, Makarkin, Aristov & Vasilenko, 2009
1268:
778:(Road Cut on Road BR 290 near Minas do Leao),
8:
1221:
1219:
308:of Germany. They are thought to have been
1256:
111:
31:
1243:
1156:
1115:
1066:
1048:
1090:Kirejtshuk, Alexander G. (2020-02-17).
939:
286:, with only one species known from the
7:
363:Geertsema & van den Heever, 1996
346:Geertsema & van den Heever, 1996
656:Ponomarenko, Yan & Huang, 2014
643:Ponomarenko, Yan & Huang, 2014
25:
512:Ponomarenko & Bashkuev, 2018
495:Ponomarenko & Bashkuev, 2018
326:) family, with the traditional (
132:
120:Fossil and life restoration of
1:
858:Protocupoides plavilstshikovi
332:) Permocupedidae and related
1427:Prehistoric animals of China
497:– Germany: Gambach am Main (
1437:Prehistoric insect families
1117:10.3390/geosciences10020073
874:"Protocupoides" rohdendorfi
531:Ichthyocupes kuznetskiensis
1453:
961:(in Russian and English).
282:. They became rare in the
1210:10.1134/S0031030114010109
1002:10.1007/s12542-018-0426-4
256:
249:
242:
237:
129:Scientific classification
127:
119:
110:
34:
27:Extinct family of beetles
1245:10.22456/1807-9806.21678
949:
619:Kaltanicupes kitjakensis
611:Kaltanicupes kargalensis
1190:Paleontological Journal
947:Martynov, A.V. (1933).
842:Protocupoides elongatus
834:Protocupoides carinatus
569:Kaltanocoleus pospelovi
405:"Archicupes" reichardti
896:Tatarocupes granulatus
599:of the Lower Permian.
1384:Paleobiology Database
1145:Systematic Entomology
866:Protocupoides sharovi
786:Pintolla ponomarenkoi
757:Permocupes sojanensis
724:Permocupes distinctus
687:Maricoleus valentinae
653:Linicupes yinpinensis
627:Kaltanicupes richteri
547:Ichthyocupes tyzhnovi
479:Eocupes lukjanovitshi
427:Cytocupoides longatus
122:Permocupes sojanensis
1226:Pinto, I.D. (1987).
509:Frankencupes ultimus
396:Archicupes jacobsoni
293:Frankencupes ultimus
1202:2014PalJ...48..191P
1108:2020Geosc..10...73K
1050:10.7554/eLife.72692
994:2018PalZ...92..587P
925:(Ponomarenko, 1963)
916:Belebeevo Formation
850:Protocupoides esini
748:Permocupes semenovi
718:Arkhangel’sk Region
679:(Cape, Novosilsky,
603:Kaltanicupes acutus
354:Whitehill Formation
1158:10.1111/syen.12562
869:(Rohdendorf, 1961)
837:(Rohdendorf, 1961)
829:(Rohdendorf, 1961)
826:Protocupoides acer
732:Permocupes grandis
542:(Rohdendorf, 1961)
539:Ichthyocupes skoki
471:Eocupes cellulosus
449:Cytocupes angustus
41:Cisuralian–Anisian
1399:
1398:
1371:Open Tree of Life
1262:Taxon identifiers
912:Ponomarenko, 1969
899:Ponomarenko, 2004
890:Ponomarenko, 2004
853:Ponomarenko, 2013
845:Ponomarenko, 2013
776:Rio Grande do Sul
760:Ponomarenko, 1963
743:Ponomarenko, 2013
735:Ponomarenko, 1963
622:Ponomarenko, 1963
614:Ponomarenko, 1963
606:Ponomarenko, 1963
597:Barakar Formation
534:Ponomarenko, 1969
474:Ponomarenko, 1969
430:Ponomarenko, 1969
421:Ponomarenko, 1969
352:(Kenmoore Farm),
269:
268:
233:
16:(Redirected from
1444:
1392:
1391:
1379:
1378:
1366:
1365:
1353:
1352:
1340:
1339:
1327:
1326:
1314:
1313:
1304:
1303:
1302:
1289:
1288:
1287:
1257:
1250:
1249:
1247:
1223:
1214:
1213:
1185:
1179:
1178:
1160:
1136:
1130:
1129:
1119:
1087:
1081:
1080:
1070:
1052:
1028:
1022:
1021:
973:
967:
966:
956:
944:
926:
922:Uralocupes major
913:
900:
891:
878:
870:
862:
861:Rohdendorf, 1961
854:
846:
838:
830:
817:
816:Rohdendorf, 1961
810:
809:Rohdendorf, 1961
803:
802:Rohdendorf, 1961
790:
773:
772:Kirejtshuk, 2020
761:
752:
744:
740:Permocupes latus
736:
728:
727:(Martynov, 1933)
715:
708:
695:
674:
657:
644:
631:
630:Rohdendorf, 1961
623:
615:
607:
586:
585:Rohdendorf, 1961
573:
572:Rohdendorf, 1961
564:
563:Rohdendorf, 1961
551:
550:Rohdendorf, 1961
543:
535:
526:
525:Rohdendorf, 1961
513:
496:
483:
482:Rohdendorf, 1961
475:
466:
465:Rohdendorf, 1961
453:
452:Rohdendorf, 1961
444:
443:Rohdendorf, 1961
431:
422:
409:
408:Rohdendorf, 1961
400:
399:Rohdendorf, 1961
387:
386:Rohdendorf, 1961
377:
376:Rohdendorf, 1961
364:
360:Afrocupes firmae
348:– South Africa:
347:
276:Protocoleopteran
264:
258:Kaltanocoleidae
228:
223:
210:
197:
137:
136:
115:
105:
42:
38:Temporal range:
32:
21:
1452:
1451:
1447:
1446:
1445:
1443:
1442:
1441:
1432:Beetle families
1417:Permian insects
1412:Permocupedoidea
1402:
1401:
1400:
1395:
1387:
1382:
1374:
1369:
1361:
1356:
1348:
1343:
1335:
1330:
1322:
1317:
1309:
1307:
1298:
1297:
1292:
1283:
1282:
1277:
1264:
1254:
1253:
1225:
1224:
1217:
1187:
1186:
1182:
1138:
1137:
1133:
1089:
1088:
1084:
1030:
1029:
1025:
975:
974:
970:
954:
951:
946:
945:
941:
936:
924:
911:
898:
889:
876:
868:
860:
852:
844:
836:
828:
815:
808:
801:
788:
780:Irati Formation
771:
759:
750:
742:
734:
726:
713:
706:
689:
668:
655:
642:
629:
621:
613:
605:
593:Orenburg Region
584:
571:
562:
549:
541:
533:
524:
511:
499:Lower Franconia
494:
481:
473:
464:
451:
442:
429:
420:
407:
398:
390:Kemerovo Region
385:
375:
362:
345:
318:
298:Middle Triassic
274:is a family of
259:
227:
221:
212:Permocupedoidea
208:
199:Protocoleoptera
195:
131:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
1450:
1448:
1440:
1439:
1434:
1429:
1424:
1419:
1414:
1404:
1403:
1397:
1396:
1394:
1393:
1380:
1367:
1354:
1341:
1328:
1315:
1305:
1300:Permocupedidae
1290:
1274:
1272:
1270:Permocupedidae
1266:
1265:
1260:
1252:
1251:
1215:
1196:(2): 191–200.
1180:
1131:
1082:
1023:
988:(4): 587–591.
968:
938:
937:
935:
932:
931:
930:
929:
928:
927:(Type species)
904:
903:
902:
901:(Type species)
882:
881:
880:
871:
863:
855:
847:
839:
831:
820:Vologda Region
794:
793:
792:
791:(Type species)
764:
763:
762:
754:
753:(Type species)
751:Martynov, 1933
745:
737:
729:
714:Martynov, 1933
707:Martynov, 1933
699:
698:
697:
696:(Type species)
677:Primorsky Krai
661:
660:
659:
658:(Type species)
635:
634:
633:
632:(Type species)
624:
616:
608:
577:
576:
575:
574:(Type species)
555:
554:
553:
552:(Type species)
544:
536:
517:
516:
515:
514:(Type species)
487:
486:
485:
484:(Type species)
476:
457:
456:
455:
454:(Type species)
435:
434:
433:
432:(Type species)
413:
412:
411:
402:
401:(Type species)
368:
367:
366:
365:(Type species)
334:Taldycupedidae
317:
314:
280:Middle Permian
272:Permocupedidae
267:
266:
254:
253:
247:
246:
240:
239:
235:
234:
225:Permocupedidae
219:
215:
214:
206:
202:
201:
193:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
125:
124:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
37:
35:Permocupedidae
26:
24:
18:Permocupedinae
14:
13:
10:
9:
6:
4:
3:
2:
1449:
1438:
1435:
1433:
1430:
1428:
1425:
1423:
1422:Permian China
1420:
1418:
1415:
1413:
1410:
1409:
1407:
1390:
1385:
1381:
1377:
1372:
1368:
1364:
1359:
1355:
1351:
1346:
1342:
1338:
1333:
1329:
1325:
1320:
1316:
1312:
1306:
1301:
1295:
1291:
1286:
1280:
1276:
1275:
1273:
1271:
1267:
1263:
1258:
1246:
1241:
1237:
1233:
1229:
1222:
1220:
1216:
1211:
1207:
1203:
1199:
1195:
1191:
1184:
1181:
1176:
1172:
1168:
1164:
1159:
1154:
1150:
1146:
1142:
1135:
1132:
1127:
1123:
1118:
1113:
1109:
1105:
1101:
1097:
1093:
1086:
1083:
1078:
1074:
1069:
1064:
1060:
1056:
1051:
1046:
1042:
1038:
1034:
1027:
1024:
1019:
1015:
1011:
1007:
1003:
999:
995:
991:
987:
983:
979:
972:
969:
964:
960:
952:
943:
940:
933:
923:
920:
919:
917:
910:
909:
905:
897:
894:
893:
888:
887:
883:
875:
872:
867:
864:
859:
856:
851:
848:
843:
840:
835:
832:
827:
824:
823:
821:
814:
807:
800:
799:
798:Protocupoides
795:
789:(Pinto, 1987)
787:
784:
783:
781:
777:
770:
769:
765:
758:
755:
749:
746:
741:
738:
733:
730:
725:
722:
721:
719:
712:
711:Permocupoides
705:
704:
700:
693:
688:
685:
684:
682:
681:Russky Island
678:
672:
667:
666:
662:
654:
651:
650:
648:
641:
640:
636:
628:
625:
620:
617:
612:
609:
604:
601:
600:
598:
594:
590:
583:
582:
578:
570:
567:
566:
561:
560:
559:Kaltanocoleus
556:
548:
545:
540:
537:
532:
529:
528:
523:
522:
518:
510:
507:
506:
504:
503:Röt Formation
500:
493:
492:
488:
480:
477:
472:
469:
468:
463:
462:
458:
450:
447:
446:
441:
440:
436:
428:
425:
424:
419:
418:
414:
406:
403:
397:
394:
393:
391:
384:
381:
374:
373:
369:
361:
358:
357:
355:
351:
344:
343:
339:
338:
337:
335:
331:
330:
329:sensu stricto
325:
324:
315:
313:
311:
307:
306:Röt Formation
303:
299:
295:
294:
289:
285:
281:
277:
273:
265:
262:
255:
252:
248:
245:
241:
236:
231:
226:
220:
217:
216:
213:
207:
205:Superfamily:
204:
203:
200:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
135:
130:
126:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
33:
30:
19:
1269:
1238:(19): 5–12.
1235:
1231:
1193:
1189:
1183:
1148:
1144:
1134:
1099:
1095:
1085:
1040:
1036:
1026:
985:
981:
971:
962:
958:
942:
921:
906:
895:
884:
873:
865:
857:
849:
841:
833:
825:
818:) – Russia:
812:
805:
796:
785:
766:
756:
747:
739:
731:
723:
716:) – Russia:
710:
701:
691:
690:Shcherbakov
686:
670:
669:Shcherbakov
663:
652:
637:
626:
618:
610:
602:
589:Kirov Region
581:Kaltanicupes
579:
568:
557:
546:
538:
530:
521:Ichthyocupes
519:
508:
491:Frankencupes
489:
478:
470:
459:
448:
437:
426:
417:Cytocupoides
415:
404:
395:
388:) – Russia,
382:
370:
359:
350:Western Cape
340:
327:
321:
319:
291:
284:Late Permian
271:
270:
257:
243:
224:
121:
29:
1294:Wikispecies
1096:Geosciences
886:Tatarocupes
877:Pinto, 1987
383:Palaeocupes
310:xylophagous
1406:Categories
1043:: e72692.
934:References
914:– Russia:
908:Uralocupes
806:Tomiocupes
774:– Brazil:
703:Permocupes
675:– Russia,
665:Maricoleus
587:– Russia:
372:Archicupes
323:sensu lato
261:Rohdendorf
192:Suborder:
186:Coleoptera
166:Arthropoda
1285:Q19385267
1232:Pesquisas
1175:251171914
1167:0307-6970
1151:: 69–96.
1126:2076-3263
1102:(2): 73.
1059:2050-084X
1018:133665417
1010:0031-0220
645:– China:
639:Linicupes
439:Cytocupes
342:Afrocupes
296:from the
244:See text
152:Kingdom:
146:Eukaryota
1308:BioLib:
1279:Wikidata
1077:34747694
965:: 63–96.
813:Tricupes
768:Pintolla
316:Taxonomy
288:Triassic
251:Synonyms
230:Martynov
218:Family:
162:Phylum:
156:Animalia
142:Domain:
1376:4664797
1350:4769965
1337:8975518
1198:Bibcode
1104:Bibcode
1068:8585485
990:Bibcode
461:Eocupes
302:Anisian
238:Genera
182:Order:
176:Insecta
172:Class:
1389:153424
1363:103066
1311:854606
1173:
1165:
1124:
1075:
1065:
1057:
1016:
1008:
955:(DjVu)
263:, 1961
232:, 1933
1358:IRMNG
1324:62679
1171:S2CID
1037:eLife
1014:S2CID
804:(syn
709:(syn
647:Anhui
1345:GBIF
1163:ISSN
1122:ISSN
1073:PMID
1055:ISSN
1006:ISSN
982:PalZ
46:PreꞒ
1332:EoL
1319:CoL
1240:doi
1206:doi
1153:doi
1112:doi
1063:PMC
1045:doi
998:doi
501:),
380:syn
1408::
1386::
1373::
1360::
1347::
1334::
1321::
1296::
1281::
1236:19
1234:.
1230:.
1218:^
1204:.
1194:48
1192:.
1169:.
1161:.
1149:48
1147:.
1143:.
1120:.
1110:.
1100:10
1098:.
1094:.
1071:.
1061:.
1053:.
1041:10
1039:.
1035:.
1012:.
1004:.
996:.
986:92
984:.
980:.
957:.
811:,
692:in
671:in
304:)
290:,
96:Pg
1248:.
1242::
1212:.
1208::
1200::
1177:.
1155::
1128:.
1114::
1106::
1079:.
1047::
1020:.
1000::
992::
963:2
378:(
300:(
222:†
209:†
196:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
20:)
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