1835:
3863:
500:
3852:
3001:
3858:
150:
3869:
128:
119:
3892:
618:
3013:
2989:
632:
that possessed enlarged curved claws on the second toe of each foot for predation; these claws were 6.6 cm (2.6 in) long for
Pyroraptor. As in other dromaeosaurids, these claws might have been used as weapons or as climbing aids. Its two known teeth are flattened and curved backwards, with
1757:
were compared to those of South
American Unenlagiines found that although not significantly, some similarities in cross section were noticeable. But Unenlagiine teeth do not have any serrations on their edges and are usually fluted, a trait that is in direct contradiction with what is seen in
1791:
originated from North
America or Euro-America, and then dispersed into Asia. On the basis of newly discovered Early Cretaceous material from China, this dispersal model has since lost support, so Allain and Taquet postulated that perhaps European dromaeosaurids such as
676:, in its initial description it was noted that due to the paucity of remains, not enough diagnostic features can be gathered to create a good comparison with other dromaeosaurids. This concern has been echoed by others, most notably in a 2012 phylogeny of
688:
is known to create issues in phylogenies which are only fixed when it is pruned from the analysis. Hence, this taxon has rarely been used since in phylogenies, which causes difficulty in pinning its precise classification.
1734:, lending support to the idea that European raptors were closely related to Southern forms from Africa and South America. Another subsequent 2022 study that describes a newly discovered dromaeosaurid sickle claw from the same
555:
very clearly distinguished the holotype from any referred material and argue that some of the material very clearly belongs to the same individual while also elucidating that the situation with
1849:
of what is now
Southern France. During the Late Cretaceous, it was one part of the island landmass known as Ibero-Armorica, formed from what is today Southern France and Northern Spain in the
2679:
Santos
Brilhante, Natan; de França, Tainá Constância; Castro, Fabiano; Sanches da Costa, Leandro; Currie, Philip J.; Kugland de Azevedo, Sergio Alex; Delcourt, Rafael (2022-03-08).
2311:
Manning, Phil L., Payne, David., Pennicott, John., Barrett, Paul M., Ennos, Roland A. (2005) "Dinosaur killer claws or climbing crampons?" Biology
Letters (2006) 2; pg. 110-112
2064:
Pereda-Suberbiol, X.; Corral, J.C.; Astibia, H.; Badiola, A.; Bardet, N.; Berreteaga, A.; Buffetaut, E.; Buscalioni, A.D.; Cappetta, H.; Cavin, L.; Díez Díaz, V. (2015-05-08).
2124:"Upper Cretaceous European theropod palaeobiodiversity, palaeobiogeography and the intra‐Maastrichtian faunal turnover: new contributions from the Iberian fossil site of Laño"
510:
Finds of dromaeosaurid dinosaur remains are rare in Europe and typically provide little taxonomic information. The first dromaeosaurid fossils found in France were those of
1712:
lineages, but scant material from
European forms prevented this idea from being tested at the time. A 2012 phylogeny study noted that despite being an unstable taxa,
2896:"Island life in the Cretaceous-faunal composition, biostratigraphy, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago"
4018:
2634:"A new dromaeosaurid dinosaur from the Wessex Formation (Lower Cretaceous, Barremian) of the Isle of Wight, and implications for European palaeobiogeography"
609:
is raised but it is concluded that due to the current lack of overlap in material, this possible synonymy cannot yet be tested and more remains are needed.
598:"unique" traits are widespread features of dromaeosauridae while also ruling out some of the other "unique traits" as results of preservation deformation.
2728:"The Teeth of the Unenlagiine TheropodBuitreraptorfrom the Cretaceous of Patagonia, Argentina, and the Unusual Dentition of the Gondwanan Dromaeosaurids"
1846:
1735:
582:
specimen, some of which are stated to undeniably belong to the holotype individual, thus adding new diagnostic traits and establishing the validity of
3045:
3972:
4063:
4058:
2187:
1796:
were the remnants of Late
Jurassic or Early Cretaceous fauna that evolved separately during the late Cretaceous isolation of Southern Europe.
2542:
2066:"Late cretaceous continental and marine vertebrate assemblages from the Laño quarry (Basque-Cantabrian Region, Iberian Peninsula): an update"
528:, and that the material did not have enough diagnostic traits to warrant the naming of a new species. However, one of the authors that named
586:. The authors also bring light to the fact that Allain and Taquet did not provide enough diagnosable traits to establish the holotype of
2844:
Brum, Arthur S.; Pêgas, Rodrigo V.; Bandeira, Kamila L. N.; Souza, Lucy G.; Campos, Diogenes A.; Kellner, Alexander W. A. (2021-08-05).
3885:
1999:
551:, Phornphen Chanthasit and Eric Buffetaut addressed the claims of Allain & Taquet by mentioning that the original description of
3862:
2968:
563:
remains, it's unknown if they belong to the same individual. This suspicion is affirmed by a 2012 study that states the known
480:
may have also inhabited Spain; however, a 2022 reevaluation of these teeth states that they cannot be confidently assigned to
2034:
Allain, R.; Taquet, P. (2000). "A new genus of
Dromaeosauridae (Dinosauria, Theropoda) from the Upper Cretaceous of France".
3851:
3038:
2459:
Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019-07-10).
3868:
3857:
4028:
2336:
1834:
149:
4038:
4033:
2979:
570:
The other criticism of Allain and Taquet, that there are not enough diagnostic features to support the validity of
4048:
4043:
2352:"A New Specimen of Microraptor (Theropoda: Dromaeosauridae) from the Lower Cretaceous of Western Liaoning, China"
2047:
4053:
3142:
3031:
2122:
Isasmendi, Erik; Torices, Angelica; Canudo, José Ignacio; Currie, Philip J.; Pereda‐Suberbiola, Xabier (2022).
684:
was found to wildly fluctuate in its position within
Dromaeosauridae. Due to its status as a "wildcard taxa",
644:
likely had well-developed forelimbs with curved claws, and probably balanced the body with a long, thin tail.
430:
is derived from Mont Olympe, a mountain in Provence at the foot of which the animal's remains were unearthed.
2956:
2461:"A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight"
4023:
427:
378:, or P. olympius, were discovered in southeastern France, at the La Boucharde locality of the Arc Basin in
3912:
1939:
1700:
This is also consistent with some previous studies, such as a 2005 study where it was suggested that some
524:
on the basis that fossil material of the species was collected from localities different from that of the
3985:
3895:
1933:
1927:
1915:
343:, approximately 72 million years ago. It is known from a single partial specimen that was found in
274:
499:
2633:
3959:
3196:
2680:
2579:
2406:
2188:"New data on the Dromaeosauridae (Dinosauria: Theropoda) from the Late Cretaceous of southern France"
1776:
is an unenlagiid, then this would mean that not all unenlagiids would have fluted, unserrated teeth.
391:
358:
340:
3613:
3005:
1979:
1874:
1383:
759:
4013:
3350:
3169:
2873:
2767:
2708:
2661:
2611:
2438:
2375:
2265:
2161:
426:
for "thief"), due to its remains being discovered after a forest fire that occurred in 1992. The
144:
3990:
3810:
3701:
3637:
3533:
3326:
2963:
2923:
2865:
2826:
2818:
2759:
2700:
2653:
2603:
2595:
2538:
2500:
2482:
2430:
2422:
2367:
2331:
2283:
2257:
2207:
2153:
2097:
1975:
1945:
1868:
1278:
1123:
469:
434:
1962:
3734:
3680:
3540:
3518:
2993:
2913:
2903:
2857:
2808:
2798:
2749:
2739:
2692:
2645:
2587:
2530:
2490:
2472:
2414:
2359:
2312:
2249:
2199:
2143:
2135:
2087:
2077:
2043:
859:
656:
458:
442:
3950:
2395:"New specimen of Archaeopteryx provides insights into the evolution of pennaceous feathers"
2065:
1772:
as being an unenlagiid due to this dissimilarity. However, a 2022 study points out that if
127:
3794:
3648:
3551:
3264:
3242:
3208:
3127:
3054:
2951:
1886:
1819:
1815:
1788:
1784:
1731:
1665:
1294:
and geographically consistent with Cretaceous Europe showing a trend of hosting typically
673:
532:, Jean Le Lœuff, points out in a conversation that the same reasons given for considering
485:
454:
387:
383:
354:
325:
317:
233:
30:
2895:
2583:
2410:
601:
Since the two existed at the same time and place, the possibility of a synonymy between
3452:
2918:
2522:
2495:
2460:
2082:
1987:
1956:
1949:
1907:
1764:
1400:
2894:
Csiki-Sava, Z.; Buffetaut, E.; Ősi, A.; Pereda-Suberbiola, X.; Brusatte, S.L. (2015).
118:
4007:
3773:
3687:
3579:
3564:
3503:
3496:
3489:
3479:
3414:
3407:
3336:
3017:
2877:
2712:
2665:
2327:
2165:
1902:
1880:
1337:
1198:
991:
910:
876:
715:
590:
as a valid species, noting that most features stated for diagnosis are not unique to
462:
415:
337:
69:
2846:"A new unenlagiine (Theropoda, Dromaeosauridae) from the Upper Cretaceous of Brazil"
2803:
2786:
2771:
2615:
2379:
2269:
1803:
along with other European raptors, likely originated from the Southern continent of
3832:
3743:
3664:
3657:
3622:
3601:
3586:
3525:
3392:
3385:
3378:
3368:
3315:
3300:
2442:
1983:
1921:
1891:
1850:
1811:
1650:
1555:
1466:
1442:
1291:
1287:
1164:
1042:
1008:
951:
893:
791:
736:
521:
395:
321:
2696:
2649:
2350:
Pei, Rui; Li, Quanguo; Meng, Qingjin; Gao, Ke-Qin; Norell, Mark A. (2014-12-22).
1762:
teeth which are serrated, as is typical of dromaeosaurs, the 2021 description of
3944:
3787:
3765:
3716:
3672:
3629:
3594:
3572:
3511:
3422:
3357:
3181:
2534:
2203:
1971:
1967:
1896:
1721:
1507:
1147:
1075:
842:
697:
650:
594:
but are simply widespread traits of dromaeosaurs. A 2012 study also agrees that
44:
2351:
3780:
3757:
3709:
3460:
3307:
3093:
3084:
2394:
2237:
1025:
617:
516:, described by Jean Le Lœuff and Eric Buffetaut. Allain and Taquet considered
512:
207:
89:
54:
3935:
2908:
2869:
2822:
2763:
2704:
2657:
2599:
2567:
2486:
2426:
2371:
2261:
2211:
2157:
2101:
437:, MNHN BO001, consists of the second toe claw of the left foot. The assigned
3818:
3750:
3445:
3437:
3343:
3154:
3102:
2726:
Gianechini, Federico A.; Makovicky, Peter J.; Apesteguía, Sebastián (2011).
2287:
1572:
1531:
1181:
934:
808:
634:
629:
333:
220:
161:
94:
38:
2927:
2830:
2607:
2504:
2434:
2316:
2744:
2727:
1753:
Two separate studies from 2011 regarding unenlagiines, where the teeth of
1750:
and other unenlagiines were found, this time in the anatomy of the claws.
3929:
3802:
3723:
3694:
3468:
3278:
3078:
2632:
Longrich, Nicholas R.; Martill, David M.; Jacobs, Megan L. (2022-06-01).
2566:
Makovicky, Peter J.; Apesteguía, Sebastián; Agnolín, Federico L. (2005).
1862:
1823:
1808:
1804:
1717:
1709:
1701:
1295:
710:
706:
525:
438:
379:
344:
313:
194:
181:
84:
79:
64:
59:
49:
2813:
2754:
2681:"A dromaeosaurid-like claw from the Upper Cretaceous of southern France"
2591:
2418:
2092:
449:(long forearm bone); and two teeth. Additional material was referred to
3977:
3825:
3111:
2944:
2477:
2148:
1099:
677:
310:
99:
74:
2861:
2139:
3400:
3292:
3072:
825:
329:
171:
3906:
2845:
2787:"Unenlagiinae revisited: dromaeosaurid theropods from South América"
2236:
Turner, Alan H.; Makovicky, Peter J.; Norell, Mark A. (2012-08-17).
2123:
2393:
Foth, Christian; Tischlinger, Helmut; Rauhut, Oliver W. M. (2014).
2363:
3023:
2253:
616:
498:
423:
403:
306:
3964:
446:
3910:
3065:
3027:
559:
is quite similar. However, due to the disarticulated nature of
547:
In a 2009 description of new dromaeosaurid remains assigned to
472:
of North Eastern Spain have been compared to those referred to
441:
include the equivalent claw of the right foot; the left second
2238:"A Review of Dromaeosaurid Systematics and Paravian Phylogeny"
574:, is also addressed by the description of new and overlapping
648:
was also covered in feathers, as many of its relatives, like
1860:
would have coexisted alongside a variety of species such as
2889:
2887:
2048:
10.1671/0272-4634(2000)020[0404:ANGODD]2.0.CO;2
139:
restored as a generalized dromaeosaurid and an unenlagiine
567:
material to belong to at least two different individuals.
2568:"The earliest dromaeosaurid theropod from South America"
1746:
and other dromaeosaurids, and more similarities between
484:
and may belong to a whole different variety of European
2785:
Gianechini, Federico A.; Apesteguia, Sebastian (2011).
2029:
2027:
2025:
2023:
2021:
2019:
2017:
2015:
2288:"Evidence of predatory behavior by theropod dinosaurs"
2977:
2186:
Chanthasit, Phornphen; Buffetaut, Eric (2009-03-01).
3919:
3733:
3647:
3612:
3550:
3478:
3367:
3325:
3262:
3207:
3180:
3153:
3126:
2529:, Dordrecht: Springer Netherlands, pp. 37–47,
347:in 1992, after a forest fire. The animal was named
2242:Bulletin of the American Museum of Natural History
1730:had more similarities with unenlagiines than with
628:was a dromaeosaurid, a small, bird-like predatory
445:; another, more complete second toe claw; a right
2521:Agnolín, Federico L.; Novas, Fernando E. (2013),
328:Ibero-Armorican island, of what is now southern
578:remains from the same locality as the holotype
3039:
1807:. as its anatomy is more consistent with the
8:
1720:dromaeosaur groups. The 2022 description of
544:'s remains present the exact same problems.
2192:Bulletin de la Société Géologique de France
1931:, and non-dinosaurs such as the pterosaurs
3907:
3556:
3270:
3259:
3186:
3159:
3132:
3123:
3062:
3046:
3032:
3024:
1742:, compared the sickle claws with those of
465:, a dorsal vertebra, and a tail vertebra.
126:
117:
20:
2917:
2907:
2812:
2802:
2753:
2743:
2494:
2476:
2147:
2091:
2081:
2791:Anais da Academia Brasileira de Ciências
1833:
2984:
2974:by Meteor Studios for Discovery Channel
2011:
382:. It was described and named by French
2627:
2625:
2561:
2559:
2516:
2514:
2454:
2452:
1986:and/or anguid squamates, and derived
709:as seen in the cladogram below. This
305:(meaning "fire thief") is an extinct
7:
2231:
2229:
2227:
2225:
2223:
2221:
2181:
2179:
2177:
2175:
2117:
2115:
2113:
2111:
2059:
2057:
1847:Argiles et Grès à Reptiles Formation
1822:European fauna, which are typically
713:was also repeated in a phylogeny of
695:was included in a 2014 phylogeny of
536:to be a nomen dubium would apply to
4019:Late Cretaceous dinosaurs of Europe
1643:
1547:
1523:
1482:
1458:
1434:
1375:
1329:
1322:
1315:
1308:
1301:
1139:
1115:
1091:
1067:
983:
976:
926:
751:
729:
722:
2083:10.5209/rev_JIGE.2015.v41.n1.48658
2036:Journal of Vertebrate Paleontology
2000:Timeline of dromaeosaurid research
1799:However, recent studies show that
1779:During the initial description of
1726:noted that the foot morphology of
14:
2523:"Uncertain Averaptoran Theropods"
457:, one manual digit, a piece of a
3891:
3890:
3867:
3861:
3856:
3850:
3011:
2999:
2987:
633:their rear margins having finer
621:Estimated size compared to human
148:
2804:10.1590/s0001-37652011000100009
1818:. This is congruent with other
1290:. The study noted that this is
701:, as with most trees including
398:and so far the only species is
1:
4064:Taxa named by Philippe Taquet
4059:Fossil taxa described in 2000
2732:Acta Palaeontologica Polonica
2697:10.1080/08912963.2021.2007243
2650:10.1016/j.cretres.2021.105123
1838:Ibero-Armorica labelled as IA
1716:never fell within any of the
1946:Musturzabalsuchus buffetauti
1887:Atsinganosaurus velauciensis
1787:evolution proposed that the
1708:could be considered part of
2535:10.1007/978-94-007-5637-3_4
2334:'s 2003 documentary series
2204:10.2113/gssgfbull.180.2.145
1276:In the 2019 description of
332:. It lived during the late
4080:
2070:Journal of Iberian Geology
16:Extinct genus of dinosaurs
3880:
3848:
3559:
3273:
3258:
3239:
3189:
3162:
3135:
3122:
3061:
2972:computer animation stills
2356:American Museum Novitates
2340:, ep. 2: "Pod's Travels".
1783:, the suggested model of
1663:
1648:
1641:
1569:
1552:
1545:
1528:
1521:
1504:
1487:
1480:
1463:
1456:
1439:
1432:
1397:
1380:
1373:
1351:
1334:
1327:
1320:
1313:
1306:
1282:, a phylogeny tree found
1195:
1178:
1161:
1144:
1137:
1120:
1113:
1096:
1089:
1072:
1065:
1039:
1022:
1005:
988:
981:
974:
948:
931:
924:
907:
890:
873:
856:
839:
822:
805:
788:
773:
756:
749:
734:
727:
293:Allain & Taquet, 2000
280:
273:
253:Allain & Taquet, 2000
145:Scientific classification
143:
134:
125:
116:
23:
2962:, by Meteor Studios for
2909:10.3897/zookeys.469.8439
1903:Tarascosaurus salluvicus
2850:Papers in Palaeontology
2128:Papers in Palaeontology
719:from the same year.
672:has been shown to be a
453:, including five pedal
2317:10.1098/rsbl.2005.0395
1943:, the crocodylomorphs
1940:Hatzegopteryx thambema
1922:Gargantuavis philoinos
1881:Lirainosaurus astibiae
1839:
622:
507:
3986:Paleobiology Database
2745:10.4202/app.2009.0127
1934:Azhdarcho lancicollis
1928:Martinaves cruzyensis
1916:Variraptor mechanorum
1837:
620:
584:Variraptor mechinorum
553:Variraptor mechinorum
549:Variraptor mechinorum
534:Variraptor mechinorum
518:Variraptor mechinorum
513:Variraptor mechinorum
502:
374:The first remains of
1292:biostratigraphically
660:, also had plumage.
640:As a dromaeosaurid,
503:Life restoration of
468:Some teeth from the
365:Discovery and naming
135:Figured material of
3614:Saurornitholestinae
2638:Cretaceous Research
2592:10.1038/nature03996
2584:2005Natur.437.1007M
2578:(7061): 1007–1011.
2419:10.1038/nature13467
2411:2014Natur.511...79F
1897:Arcovenator escotae
1890:, some undescribed
1875:Ampelosaurus atacis
1843:Pyroraptor olympius
1794:Pyroraptor olympius
1781:Pyroraptor olympius
1728:Pyroraptor olympius
1704:dromaeoaurids like
670:Pyroraptor olympius
637:than at the front.
626:Pyroraptor olympius
538:Pyroraptor olympius
505:Pyroraptor olympius
400:Pyroraptor olympius
376:Pyroraptor olympius
370:Naming and Material
350:Pyroraptor olympius
287:Pyroraptor olympius
4029:Maastrichtian life
2960:computer animation
2685:Historical Biology
2478:10.7717/peerj.7247
2284:Carpenter, Kenneth
1845:is known from the
1840:
1736:geologic formation
1723:Vectiraptor greeni
623:
508:
476:, suggesting that
4039:Fossils of France
4034:Cretaceous France
4001:
4000:
3913:Taxon identifiers
3904:
3903:
3876:
3875:
3846:
3845:
3842:
3841:
3811:Saurornitholestes
3702:Saurornitholestes
3638:Saurornitholestes
3327:Halszkaraptorinae
3254:
3253:
3250:
3249:
3235:
3234:
3231:
3230:
3227:
3226:
2964:Discovery Channel
2862:10.1002/spp2.1375
2691:(11): 2195–2204.
2544:978-94-007-5636-6
2332:Discovery Channel
2140:10.1002/spp2.1419
1980:batrachosauroidid
1909:Struthiosaurus sp
1869:Rhabdodon priscus
1694:
1693:
1685:
1684:
1676:
1675:
1630:
1629:
1621:
1620:
1612:
1611:
1603:
1602:
1594:
1593:
1585:
1584:
1421:
1420:
1412:
1411:
1362:
1361:
1279:Hesperornithoides
1273:
1272:
1264:
1263:
1255:
1254:
1246:
1245:
1237:
1236:
1228:
1227:
1219:
1218:
1210:
1209:
1124:Saurornitholestes
1054:
1053:
963:
962:
470:Iberian peninsula
435:holotype specimen
386:Ronan Allain and
298:
297:
254:
4071:
4049:Fossils of Spain
4044:Cretaceous Spain
3994:
3993:
3981:
3980:
3968:
3967:
3955:
3954:
3953:
3940:
3939:
3938:
3908:
3894:
3893:
3871:
3865:
3860:
3854:
3735:Velociraptorinae
3681:Dromaeosauroides
3557:
3541:Zhongjianosaurus
3519:Sinornithosaurus
3271:
3260:
3187:
3160:
3133:
3124:
3117:
3116:
3063:
3048:
3041:
3034:
3025:
3016:
3015:
3014:
3004:
3003:
3002:
2992:
2991:
2990:
2983:
2932:
2931:
2921:
2911:
2891:
2882:
2881:
2856:(4): 2075–2099.
2841:
2835:
2834:
2816:
2806:
2782:
2776:
2775:
2757:
2747:
2723:
2717:
2716:
2676:
2670:
2669:
2629:
2620:
2619:
2563:
2554:
2553:
2552:
2551:
2518:
2509:
2508:
2498:
2480:
2456:
2447:
2446:
2390:
2384:
2383:
2347:
2341:
2325:
2319:
2309:
2303:
2302:
2292:
2280:
2274:
2273:
2233:
2216:
2215:
2183:
2170:
2169:
2151:
2119:
2106:
2105:
2095:
2085:
2061:
2052:
2051:
2031:
1644:
1548:
1524:
1483:
1459:
1435:
1376:
1330:
1323:
1316:
1309:
1302:
1140:
1116:
1092:
1068:
984:
977:
927:
860:Sinornithosaurus
752:
730:
723:
705:, this caused a
657:Sinornithosaurus
418:for "fire") and
289:
285:
266:P. olympius
252:
245:
232:
153:
152:
130:
121:
111:
41:
29:Temporal range:
21:
4079:
4078:
4074:
4073:
4072:
4070:
4069:
4068:
4054:Tremp Formation
4004:
4003:
4002:
3997:
3989:
3984:
3976:
3971:
3963:
3958:
3949:
3948:
3943:
3934:
3933:
3928:
3915:
3905:
3900:
3872:
3838:
3795:Luanchuanraptor
3729:
3649:Dromaeosaurinae
3643:
3608:
3552:Eudromaeosauria
3546:
3474:
3363:
3321:
3267:
3265:Dromaeosauridae
3246:
3243:Dromaeosauridae
3223:
3209:Dromaeosauridae
3203:
3176:
3149:
3143:Avemetatarsalia
3128:Avemetatarsalia
3118:
3068:
3067:
3057:
3055:Dromaeosauridae
3052:
3022:
3012:
3010:
3000:
2998:
2988:
2986:
2978:
2941:
2936:
2935:
2893:
2892:
2885:
2843:
2842:
2838:
2784:
2783:
2779:
2725:
2724:
2720:
2678:
2677:
2673:
2631:
2630:
2623:
2565:
2564:
2557:
2549:
2547:
2545:
2527:Avian Ancestors
2520:
2519:
2512:
2458:
2457:
2450:
2405:(7507): 79–82.
2392:
2391:
2387:
2349:
2348:
2344:
2337:Dinosaur Planet
2326:
2322:
2310:
2306:
2290:
2282:
2281:
2277:
2235:
2234:
2219:
2185:
2184:
2173:
2121:
2120:
2109:
2063:
2062:
2055:
2033:
2032:
2013:
2008:
1996:
1988:alethinophidian
1976:palaeobatrachid
1832:
1820:Late Cretaceous
1697:
1695:
1686:
1677:
1666:Dromaeosauridae
1631:
1622:
1613:
1604:
1595:
1586:
1422:
1413:
1363:
1274:
1265:
1256:
1247:
1238:
1229:
1220:
1211:
1055:
964:
666:
615:
540:as well, since
497:
388:Philippe Taquet
384:paleontologists
372:
367:
326:Late Cretaceous
294:
291:
283:
282:
269:
251:
243:
234:Dromaeosauridae
230:
147:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
31:Late Cretaceous
27:
17:
12:
11:
5:
4077:
4075:
4067:
4066:
4061:
4056:
4051:
4046:
4041:
4036:
4031:
4026:
4024:Campanian life
4021:
4016:
4006:
4005:
3999:
3998:
3996:
3995:
3982:
3969:
3956:
3941:
3925:
3923:
3917:
3916:
3911:
3902:
3901:
3899:
3898:
3888:
3881:
3878:
3877:
3874:
3873:
3849:
3847:
3844:
3843:
3840:
3839:
3837:
3836:
3829:
3822:
3815:
3807:
3799:
3791:
3784:
3777:
3770:
3762:
3754:
3747:
3739:
3737:
3731:
3730:
3728:
3727:
3720:
3713:
3706:
3698:
3691:
3684:
3677:
3669:
3661:
3653:
3651:
3645:
3644:
3642:
3641:
3634:
3626:
3618:
3616:
3610:
3609:
3607:
3606:
3598:
3591:
3583:
3576:
3569:
3560:
3554:
3548:
3547:
3545:
3544:
3537:
3530:
3522:
3515:
3508:
3500:
3493:
3485:
3483:
3476:
3475:
3473:
3472:
3465:
3457:
3453:Unquillosaurus
3449:
3442:
3434:
3426:
3419:
3411:
3404:
3397:
3389:
3382:
3374:
3372:
3365:
3364:
3362:
3361:
3354:
3347:
3340:
3332:
3330:
3323:
3322:
3320:
3319:
3312:
3304:
3297:
3289:
3282:
3274:
3268:
3263:
3256:
3255:
3252:
3251:
3248:
3247:
3240:
3237:
3236:
3233:
3232:
3229:
3228:
3225:
3224:
3222:
3221:
3220:
3219:
3213:
3211:
3205:
3204:
3202:
3201:
3200:
3199:
3190:
3184:
3178:
3177:
3175:
3174:
3173:
3172:
3163:
3157:
3151:
3150:
3148:
3147:
3146:
3145:
3136:
3130:
3120:
3119:
3115:
3114:
3105:
3096:
3087:
3081:
3075:
3066:
3059:
3058:
3053:
3051:
3050:
3043:
3036:
3028:
3021:
3020:
3008:
2996:
2976:
2975:
2966:
2954:
2940:
2939:External links
2937:
2934:
2933:
2902:(469): 1–161.
2883:
2836:
2797:(1): 163–195.
2777:
2738:(2): 279–290.
2718:
2671:
2621:
2555:
2543:
2510:
2448:
2385:
2364:10.1206/3821.1
2358:(3821): 1–28.
2342:
2320:
2304:
2275:
2217:
2198:(2): 145–154.
2171:
2107:
2076:(1): 101–124.
2053:
2042:(2): 404–407.
2010:
2009:
2007:
2004:
2003:
2002:
1995:
1992:
1960:, the turtles
1957:Allodaposuchus
1950:Massaliasuchus
1856:At that time,
1831:
1828:
1816:eudromaeosaurs
1765:Ypupiara lopai
1732:eudromaeosaurs
1692:
1691:
1688:
1687:
1683:
1682:
1679:
1678:
1674:
1673:
1670:
1669:
1662:
1659:
1658:
1655:
1654:
1647:
1642:
1640:
1637:
1636:
1633:
1632:
1628:
1627:
1624:
1623:
1619:
1618:
1615:
1614:
1610:
1609:
1606:
1605:
1601:
1600:
1597:
1596:
1592:
1591:
1588:
1587:
1583:
1582:
1579:
1578:
1568:
1565:
1564:
1561:
1560:
1551:
1546:
1544:
1541:
1540:
1537:
1536:
1527:
1522:
1520:
1517:
1516:
1513:
1512:
1503:
1500:
1499:
1496:
1495:
1486:
1481:
1479:
1476:
1475:
1472:
1471:
1462:
1457:
1455:
1452:
1451:
1448:
1447:
1438:
1433:
1431:
1428:
1427:
1424:
1423:
1419:
1418:
1415:
1414:
1410:
1409:
1406:
1405:
1401:Ningyuansaurus
1396:
1393:
1392:
1389:
1388:
1379:
1374:
1372:
1369:
1368:
1365:
1364:
1360:
1359:
1356:
1355:
1350:
1347:
1346:
1343:
1342:
1333:
1328:
1326:
1321:
1319:
1314:
1312:
1307:
1305:
1300:
1271:
1270:
1267:
1266:
1262:
1261:
1258:
1257:
1253:
1252:
1249:
1248:
1244:
1243:
1240:
1239:
1235:
1234:
1231:
1230:
1226:
1225:
1222:
1221:
1217:
1216:
1213:
1212:
1208:
1207:
1204:
1203:
1194:
1191:
1190:
1187:
1186:
1177:
1174:
1173:
1170:
1169:
1160:
1157:
1156:
1153:
1152:
1143:
1138:
1136:
1133:
1132:
1129:
1128:
1119:
1114:
1112:
1109:
1108:
1105:
1104:
1095:
1090:
1088:
1085:
1084:
1081:
1080:
1071:
1066:
1064:
1061:
1060:
1057:
1056:
1052:
1051:
1048:
1047:
1038:
1035:
1034:
1031:
1030:
1021:
1018:
1017:
1014:
1013:
1004:
1001:
1000:
997:
996:
987:
982:
980:
975:
973:
970:
969:
966:
965:
961:
960:
957:
956:
947:
944:
943:
940:
939:
930:
925:
923:
920:
919:
916:
915:
906:
903:
902:
899:
898:
889:
886:
885:
882:
881:
872:
869:
868:
865:
864:
855:
852:
851:
848:
847:
838:
835:
834:
831:
830:
821:
818:
817:
814:
813:
804:
801:
800:
797:
796:
787:
784:
783:
780:
779:
772:
769:
768:
765:
764:
755:
750:
748:
745:
744:
741:
740:
733:
728:
726:
721:
665:
664:Classification
662:
614:
611:
496:
490:
486:dromaeosaurids
371:
368:
366:
363:
353:by Allain and
296:
295:
292:
278:
277:
271:
270:
262:
260:
256:
255:
241:
237:
236:
228:
224:
223:
218:
211:
210:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
132:
131:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4076:
4065:
4062:
4060:
4057:
4055:
4052:
4050:
4047:
4045:
4042:
4040:
4037:
4035:
4032:
4030:
4027:
4025:
4022:
4020:
4017:
4015:
4012:
4011:
4009:
3992:
3987:
3983:
3979:
3974:
3970:
3966:
3961:
3957:
3952:
3946:
3942:
3937:
3931:
3927:
3926:
3924:
3922:
3918:
3914:
3909:
3897:
3889:
3887:
3883:
3882:
3879:
3870:
3866:
3864:
3859:
3853:
3835:
3834:
3830:
3828:
3827:
3823:
3821:
3820:
3816:
3813:
3812:
3808:
3805:
3804:
3800:
3797:
3796:
3792:
3790:
3789:
3785:
3783:
3782:
3778:
3776:
3775:
3774:Kansaignathus
3771:
3768:
3767:
3763:
3760:
3759:
3755:
3753:
3752:
3748:
3746:
3745:
3741:
3740:
3738:
3736:
3732:
3726:
3725:
3721:
3719:
3718:
3714:
3712:
3711:
3707:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3688:Dromaeosaurus
3685:
3683:
3682:
3678:
3675:
3674:
3670:
3667:
3666:
3662:
3660:
3659:
3655:
3654:
3652:
3650:
3646:
3640:
3639:
3635:
3632:
3631:
3627:
3625:
3624:
3620:
3619:
3617:
3615:
3611:
3604:
3603:
3599:
3597:
3596:
3592:
3589:
3588:
3584:
3582:
3581:
3580:Dineobellator
3577:
3575:
3574:
3570:
3567:
3566:
3562:
3561:
3558:
3555:
3553:
3549:
3543:
3542:
3538:
3536:
3535:
3531:
3528:
3527:
3523:
3521:
3520:
3516:
3514:
3513:
3509:
3506:
3505:
3504:Hesperonychus
3501:
3499:
3498:
3497:Graciliraptor
3494:
3492:
3491:
3490:Changyuraptor
3487:
3486:
3484:
3481:
3480:Microraptoria
3477:
3471:
3470:
3466:
3463:
3462:
3458:
3455:
3454:
3450:
3448:
3447:
3443:
3440:
3439:
3435:
3432:
3431:
3427:
3425:
3424:
3420:
3417:
3416:
3415:Ornithodesmus
3412:
3410:
3409:
3408:Neuquenraptor
3405:
3403:
3402:
3398:
3395:
3394:
3390:
3388:
3387:
3383:
3381:
3380:
3376:
3375:
3373:
3370:
3366:
3360:
3359:
3355:
3353:
3352:
3348:
3346:
3345:
3341:
3339:
3338:
3337:Halszkaraptor
3334:
3333:
3331:
3328:
3324:
3318:
3317:
3313:
3310:
3309:
3305:
3303:
3302:
3298:
3295:
3294:
3290:
3288:
3287:
3283:
3281:
3280:
3276:
3275:
3272:
3269:
3266:
3261:
3257:
3245:
3244:
3238:
3217:
3216:
3215:
3214:
3212:
3210:
3206:
3198:
3194:
3193:
3192:
3191:
3188:
3185:
3183:
3179:
3171:
3167:
3166:
3165:
3164:
3161:
3158:
3156:
3152:
3144:
3140:
3139:
3138:
3137:
3134:
3131:
3129:
3125:
3121:
3113:
3109:
3106:
3104:
3100:
3097:
3095:
3091:
3088:
3086:
3082:
3080:
3076:
3074:
3070:
3069:
3064:
3060:
3056:
3049:
3044:
3042:
3037:
3035:
3030:
3029:
3026:
3019:
3009:
3007:
2997:
2995:
2985:
2981:
2973:
2971:
2967:
2965:
2961:
2959:
2955:
2953:
2949:
2948:official site
2947:
2943:
2942:
2938:
2929:
2925:
2920:
2915:
2910:
2905:
2901:
2897:
2890:
2888:
2884:
2879:
2875:
2871:
2867:
2863:
2859:
2855:
2851:
2847:
2840:
2837:
2832:
2828:
2824:
2820:
2815:
2810:
2805:
2800:
2796:
2792:
2788:
2781:
2778:
2773:
2769:
2765:
2761:
2756:
2751:
2746:
2741:
2737:
2733:
2729:
2722:
2719:
2714:
2710:
2706:
2702:
2698:
2694:
2690:
2686:
2682:
2675:
2672:
2667:
2663:
2659:
2655:
2651:
2647:
2643:
2639:
2635:
2628:
2626:
2622:
2617:
2613:
2609:
2605:
2601:
2597:
2593:
2589:
2585:
2581:
2577:
2573:
2569:
2562:
2560:
2556:
2546:
2540:
2536:
2532:
2528:
2524:
2517:
2515:
2511:
2506:
2502:
2497:
2492:
2488:
2484:
2479:
2474:
2470:
2466:
2462:
2455:
2453:
2449:
2444:
2440:
2436:
2432:
2428:
2424:
2420:
2416:
2412:
2408:
2404:
2400:
2396:
2389:
2386:
2381:
2377:
2373:
2369:
2365:
2361:
2357:
2353:
2346:
2343:
2339:
2338:
2333:
2329:
2328:Scott Sampson
2324:
2321:
2318:
2314:
2308:
2305:
2300:
2296:
2289:
2285:
2279:
2276:
2271:
2267:
2263:
2259:
2255:
2254:10.1206/748.1
2251:
2247:
2243:
2239:
2232:
2230:
2228:
2226:
2224:
2222:
2218:
2213:
2209:
2205:
2201:
2197:
2193:
2189:
2182:
2180:
2178:
2176:
2172:
2167:
2163:
2159:
2155:
2150:
2145:
2141:
2137:
2133:
2129:
2125:
2118:
2116:
2114:
2112:
2108:
2103:
2099:
2094:
2089:
2084:
2079:
2075:
2071:
2067:
2060:
2058:
2054:
2049:
2045:
2041:
2037:
2030:
2028:
2026:
2024:
2022:
2020:
2018:
2016:
2012:
2005:
2001:
1998:
1997:
1993:
1991:
1989:
1985:
1984:amphisbaenian
1981:
1977:
1973:
1969:
1965:
1964:
1959:
1958:
1953:
1951:
1947:
1942:
1941:
1936:
1935:
1930:
1929:
1925:
1923:
1918:
1917:
1912:
1910:
1905:
1904:
1899:
1898:
1893:
1892:titanosaurids
1889:
1888:
1883:
1882:
1877:
1876:
1871:
1870:
1865:
1864:
1859:
1854:
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1848:
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1825:
1821:
1817:
1813:
1810:
1806:
1802:
1797:
1795:
1790:
1786:
1785:dromaeosaurid
1782:
1777:
1775:
1771:
1767:
1766:
1761:
1756:
1751:
1749:
1745:
1741:
1737:
1733:
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1608:
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1599:
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1574:
1567:
1566:
1563:
1562:
1559:
1558:
1557:
1550:
1549:
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1535:
1534:
1533:
1526:
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1519:
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1498:
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1470:
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1461:
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1454:
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1445:
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1437:
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1430:
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1426:
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1417:
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1408:
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1403:
1402:
1395:
1394:
1391:
1390:
1387:
1386:
1385:
1378:
1377:
1371:
1370:
1367:
1366:
1358:
1357:
1354:
1349:
1348:
1345:
1344:
1341:
1340:
1339:
1338:Halszkaraptor
1332:
1331:
1325:
1324:
1318:
1317:
1311:
1310:
1304:
1303:
1299:
1297:
1293:
1289:
1285:
1281:
1280:
1269:
1268:
1260:
1259:
1251:
1250:
1242:
1241:
1233:
1232:
1224:
1223:
1215:
1214:
1206:
1205:
1202:
1201:
1200:
1193:
1192:
1189:
1188:
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1184:
1183:
1176:
1175:
1172:
1171:
1168:
1167:
1166:
1159:
1158:
1155:
1154:
1151:
1150:
1149:
1142:
1141:
1135:
1134:
1131:
1130:
1127:
1126:
1125:
1118:
1117:
1111:
1110:
1107:
1106:
1103:
1102:
1101:
1094:
1093:
1087:
1086:
1083:
1082:
1079:
1078:
1077:
1070:
1069:
1063:
1062:
1059:
1058:
1050:
1049:
1046:
1045:
1044:
1037:
1036:
1033:
1032:
1029:
1028:
1027:
1020:
1019:
1016:
1015:
1012:
1011:
1010:
1003:
1002:
999:
998:
995:
994:
993:
992:Dromaeosaurus
986:
985:
979:
978:
972:
971:
968:
967:
959:
958:
955:
954:
953:
946:
945:
942:
941:
938:
937:
936:
929:
928:
922:
921:
918:
917:
914:
913:
912:
911:Hesperonychus
905:
904:
901:
900:
897:
896:
895:
888:
887:
884:
883:
880:
879:
878:
877:Graciliraptor
871:
870:
867:
866:
863:
862:
861:
854:
853:
850:
849:
846:
845:
844:
837:
836:
833:
832:
829:
828:
827:
820:
819:
816:
815:
812:
811:
810:
803:
802:
799:
798:
795:
794:
793:
786:
785:
782:
781:
778:
777:
771:
770:
767:
766:
763:
762:
761:
754:
753:
747:
746:
743:
742:
739:
738:
732:
731:
725:
724:
720:
718:
717:
716:Archaeopteryx
712:
708:
704:
700:
699:
694:
690:
687:
683:
679:
675:
674:dromaeosaurid
671:
663:
661:
659:
658:
653:
652:
647:
643:
638:
636:
631:
627:
619:
612:
610:
608:
604:
599:
597:
593:
589:
585:
581:
577:
573:
568:
566:
562:
558:
554:
550:
545:
543:
539:
535:
531:
527:
523:
519:
515:
514:
506:
501:
495:
491:
489:
487:
483:
479:
475:
471:
466:
464:
460:
456:
452:
448:
444:
440:
436:
431:
429:
428:specific name
425:
421:
417:
413:
409:
405:
401:
397:
393:
389:
385:
381:
377:
369:
364:
362:
360:
356:
352:
351:
346:
342:
339:
338:Maastrichtian
335:
331:
327:
323:
319:
318:dromaeosaurid
316:, probably a
315:
312:
308:
304:
303:
290:
288:
279:
276:
275:Binomial name
272:
268:
267:
261:
258:
257:
250:
249:
242:
239:
238:
235:
229:
226:
225:
222:
219:
216:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
138:
133:
129:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
26:
22:
19:
3920:
3855:
3833:Velociraptor
3831:
3824:
3817:
3809:
3801:
3793:
3786:
3779:
3772:
3764:
3756:
3749:
3744:Acheroraptor
3742:
3722:
3715:
3708:
3700:
3693:
3686:
3679:
3671:
3665:Dakotaraptor
3663:
3658:Achillobator
3656:
3636:
3628:
3623:Atrociraptor
3621:
3602:Zhenyuanlong
3600:
3593:
3587:Tianyuraptor
3585:
3578:
3571:
3563:
3539:
3532:
3526:Tianyuraptor
3524:
3517:
3510:
3502:
3495:
3488:
3467:
3459:
3451:
3444:
3436:
3429:
3428:
3421:
3413:
3406:
3399:
3393:Dakotaraptor
3391:
3386:Buitreraptor
3384:
3379:Austroraptor
3377:
3369:Unenlagiinae
3356:
3349:
3342:
3335:
3316:Zhenyuanlong
3314:
3306:
3301:Tianyuraptor
3299:
3291:
3285:
3284:
3277:
3241:
3107:
3098:
3089:
3006:Paleontology
2969:
2957:
2945:
2899:
2853:
2849:
2839:
2814:11336/191999
2794:
2790:
2780:
2755:11336/192712
2735:
2731:
2721:
2688:
2684:
2674:
2641:
2637:
2575:
2571:
2548:, retrieved
2526:
2468:
2464:
2402:
2398:
2388:
2355:
2345:
2335:
2323:
2307:
2298:
2294:
2278:
2245:
2241:
2195:
2191:
2131:
2127:
2093:10486/679211
2073:
2069:
2039:
2035:
1974:eutherians,
1961:
1955:
1944:
1938:
1932:
1926:
1920:
1914:
1908:
1901:
1895:
1885:
1879:
1873:
1867:
1861:
1857:
1855:
1851:Tethys Ocean
1842:
1841:
1830:Paleoecology
1812:Unenlagiines
1800:
1798:
1793:
1789:dromaeosaurs
1780:
1778:
1773:
1769:
1763:
1759:
1754:
1752:
1747:
1743:
1739:
1727:
1722:
1713:
1705:
1699:
1696:
1664:
1651:Troodontidae
1649:
1571:
1570:
1556:Dakotaraptor
1554:
1553:
1530:
1529:
1506:
1505:
1490:
1489:
1488:
1467:Buitreraptor
1465:
1464:
1443:Austroraptor
1441:
1440:
1399:
1398:
1382:
1381:
1352:
1336:
1335:
1283:
1277:
1275:
1197:
1196:
1180:
1179:
1165:Velociraptor
1163:
1162:
1146:
1145:
1122:
1121:
1098:
1097:
1074:
1073:
1043:Achillobator
1041:
1040:
1024:
1023:
1009:Atrociraptor
1007:
1006:
990:
989:
952:Buitreraptor
950:
949:
933:
932:
909:
908:
894:Tianyuraptor
892:
891:
875:
874:
858:
857:
841:
840:
824:
823:
807:
806:
792:Austroraptor
790:
789:
775:
774:
758:
757:
737:Troodontidae
735:
714:
702:
696:
692:
691:
685:
681:
669:
667:
655:
649:
645:
641:
639:
625:
624:
606:
602:
600:
596:Pyroraptor's
595:
591:
587:
583:
579:
575:
571:
569:
564:
560:
556:
552:
548:
546:
541:
537:
533:
529:
522:nomen dubium
517:
511:
509:
504:
493:
492:Relation to
481:
477:
473:
467:
450:
432:
419:
411:
407:
404:generic name
399:
396:type species
375:
373:
349:
348:
301:
300:
299:
286:
281:
265:
264:
247:
246:
214:
201:
188:
136:
24:
18:
3945:Wikispecies
3788:Linheraptor
3766:Deinonychus
3717:Yurgovuchia
3673:Deinonychus
3630:Bambiraptor
3595:Vectiraptor
3573:Bambiraptor
3512:Microraptor
3423:Pamparaptor
3358:Natovenator
3197:Maniraptora
3182:Maniraptora
2149:10810/56781
1508:Pamparaptor
1288:unenlagiine
1148:Deinonychus
1076:Bambiraptor
843:Microraptor
698:Microraptor
651:Microraptor
613:Description
406:comes from
324:, from the
4008:Categories
3951:Pyroraptor
3921:Pyroraptor
3884:See also:
3758:Boreonykus
3710:Utahraptor
3461:Variraptor
3430:Pyroraptor
3308:Variraptor
3286:Pyroraptor
3218:see below↓
3094:Dinosauria
3085:Sauropsida
2970:Pyroraptor
2958:Pyroraptor
2946:Pyroraptor
2644:: 105123.
2550:2022-03-17
2301:: 135–144.
2006:References
1982:urodeles,
1858:Pyroraptor
1826:lineages.
1814:than with
1801:Pyroraptor
1774:Pyroraptor
1770:Pyroraptor
1768:rules out
1760:Pyroraptor
1755:Pyroraptor
1748:Pyroraptor
1744:Pyroraptor
1740:Pyroraptor
1714:Pyroraptor
1706:Pyroraptor
1575:sensu lato
1491:Pyroraptor
1298:species.
1284:Pyroraptor
1026:Utahraptor
776:Pyroraptor
703:Pyroraptor
693:Pyroraptor
686:Pyroraptor
682:Pyroraptor
646:Pyroraptor
642:Pyroraptor
635:serrations
607:Variraptor
603:Pyroraptor
592:Pyroraptor
588:Pyroraptor
580:Variraptor
576:Variraptor
572:Variraptor
565:Pyroraptor
561:Pyroraptor
557:Pyroraptor
542:Pyroraptor
530:Variraptor
494:Variraptor
482:Pyroraptor
478:Pyroraptor
474:Pyroraptor
461:, a right
459:metacarpal
451:Pyroraptor
443:metatarsal
336:and early
322:unenlagiid
302:Pyroraptor
248:Pyroraptor
208:Saurischia
195:Dinosauria
137:Pyroraptor
25:Pyroraptor
4014:Paravians
3751:Adasaurus
3446:Unenlagia
3438:Rahonavis
3344:Hulsanpes
3170:Theropoda
3155:Theropoda
3103:Theropoda
3071:Kingdom:
2994:Dinosaurs
2878:238854675
2870:2056-2799
2823:0001-3765
2764:0567-7920
2713:247378741
2705:0891-2963
2666:245324247
2658:0195-6671
2600:0028-0836
2487:2167-8359
2471:: e7247.
2427:1476-4687
2372:0003-0082
2262:0003-0090
2248:: 1–206.
2212:1777-5817
2166:246028305
2158:2056-2799
2102:1886-7995
1978:anurans,
1972:zhelestid
1970:turtles,
1968:Solemydid
1824:Gondwanan
1809:Gondwanan
1718:laurasian
1710:Gondwanan
1702:laurasian
1573:Unenlagia
1532:Rahonavis
1353:ISMDVP09
1296:Gondwanan
1286:to be an
1182:Adasaurus
935:Rahonavis
809:Unenlagia
678:paravians
668:Although
439:paratypes
334:Campanian
259:Species:
221:Theropoda
168:Kingdom:
162:Eukaryota
3930:Wikidata
3896:Category
3886:Timeline
3803:Nuthetes
3724:Zapsalis
3695:Itemirus
3469:Ypupiara
3351:Mahakala
3279:Daurlong
3079:Chordata
3077:Phylum:
3073:Animalia
2928:25610343
2831:21437380
2772:62811731
2616:27078534
2608:16222297
2505:31333906
2435:24990749
2380:38984583
2286:(1998).
2270:83572446
1994:See also
1990:snakes.
1863:Zalmoxes
1805:Gondwana
1384:Mahakala
760:Mahakala
711:polytomy
707:polytomy
630:theropod
526:holotype
520:to be a
380:Provence
345:Provence
314:dinosaur
311:paravian
227:Family:
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
37:72
3978:4822956
3965:4433557
3936:Q132542
3826:Tsaagan
3112:Paraves
3083:Class:
2980:Portals
2919:4296572
2900:ZooKeys
2580:Bibcode
2496:6626525
2443:4464659
2407:Bibcode
1963:Dortoka
1100:Tsaagan
284:†
263:†
240:Genus:
106:↓
3565:Balaur
3534:Wulong
3401:Diuqin
3293:Shanag
3018:France
2952:French
2926:
2916:
2876:
2868:
2829:
2821:
2770:
2762:
2711:
2703:
2664:
2656:
2614:
2606:
2598:
2572:Nature
2541:
2503:
2493:
2485:
2441:
2433:
2425:
2399:Nature
2378:
2370:
2268:
2260:
2210:
2164:
2156:
2100:
1966:, and
1199:Balaur
826:Shanag
680:where
463:radius
455:digits
420:raptor
402:. The
394:, the
355:Taquet
341:stages
330:France
3991:65133
3108:Clade
3099:Clade
3090:Clade
2950:, in
2874:S2CID
2768:S2CID
2709:S2CID
2662:S2CID
2612:S2CID
2465:PeerJ
2439:S2CID
2376:S2CID
2291:(PDF)
2266:S2CID
2162:S2CID
2134:(1).
424:Latin
416:Greek
307:genus
215:Clade
202:Clade
189:Clade
3973:GBIF
3819:Shri
3781:Kuru
3195:see
3168:see
3141:see
2924:PMID
2866:ISSN
2827:PMID
2819:ISSN
2760:ISSN
2701:ISSN
2654:ISSN
2604:PMID
2596:ISSN
2539:ISBN
2501:PMID
2483:ISSN
2431:PMID
2423:ISSN
2368:ISSN
2295:Gaia
2258:ISSN
2208:ISSN
2154:ISSN
2098:ISSN
1954:and
1937:and
654:and
605:and
447:ulna
433:The
392:2000
359:2000
45:PreꞒ
3960:EoL
2914:PMC
2904:doi
2858:doi
2809:hdl
2799:doi
2750:hdl
2740:doi
2693:doi
2646:doi
2642:134
2588:doi
2576:437
2531:doi
2491:PMC
2473:doi
2415:doi
2403:511
2360:doi
2330:in
2313:doi
2250:doi
2246:371
2200:doi
2196:180
2144:hdl
2136:doi
2088:hdl
2078:doi
2044:doi
1853:.
1738:as
412:pûr
408:πῦρ
390:in
357:in
320:or
309:of
4010::
3988::
3975::
3962::
3947::
3932::
3110::
3101::
3092::
2922:.
2912:.
2898:.
2886:^
2872:.
2864:.
2852:.
2848:.
2825:.
2817:.
2807:.
2795:83
2793:.
2789:.
2766:.
2758:.
2748:.
2736:56
2734:.
2730:.
2707:.
2699:.
2689:34
2687:.
2683:.
2660:.
2652:.
2640:.
2636:.
2624:^
2610:.
2602:.
2594:.
2586:.
2574:.
2570:.
2558:^
2537:,
2525:,
2513:^
2499:.
2489:.
2481:.
2467:.
2463:.
2451:^
2437:.
2429:.
2421:.
2413:.
2401:.
2397:.
2374:.
2366:.
2354:.
2299:15
2297:.
2293:.
2264:.
2256:.
2244:.
2240:.
2220:^
2206:.
2194:.
2190:.
2174:^
2160:.
2152:.
2142:.
2130:.
2126:.
2110:^
2096:.
2086:.
2074:41
2072:.
2068:.
2056:^
2040:20
2038:.
2014:^
1948:,
1919:,
1913:,
1906:,
1900:,
1894:,
1884:,
1878:,
1872:,
1866:,
488:.
414:,
361:.
217::
204::
191::
95:Pg
39:Ma
3814:?
3806:?
3798:?
3769:?
3761:?
3705:?
3676:?
3668:?
3633:?
3605:?
3590:?
3568:?
3529:?
3507:?
3482:?
3464:?
3456:?
3441:?
3433:?
3418:?
3396:?
3371:?
3329:?
3311:?
3296:?
3047:e
3040:t
3033:v
2982::
2930:.
2906::
2880:.
2860::
2854:7
2833:.
2811::
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