413:
2465:
shovel-like, since it doesn't have a "dorsoventrally narrowed cutting edge" on it, but rather has a "relatively expansive anterior surface" which is almost completely flat. The lower jaw also wouldn't be "shielded" to a significant extent in response to the reaction force during burrowing; even the shielded portion, the premaxillary teeth which is crucial for feeding, would not have been able to endure the "extreme loading regimes associated with head-first burrowing." The upper jaw was also suggested to be not adapted for keeping its mouth shut when faced by reaction forces associated with burrowing. Its relatively large orbits were also inconsistent with a burrowing lifestyle, since large eyes have a risk of being damaged in head-first burrowing.
3299:
139:
118:
3957:
2407:
4529:
3976:
4522:
4536:
445:
4544:
3969:
2457:
2830:
424:, which served as the basis for its initial description in 2000, included a complete skull and lower jaw, the front of the postcranial skeleton, and parts of the posterior postcranial skeleton. Five more specimens were later described, representing the majority of the skeleton. Many isolated teeth have also been found in the Mahajanga Basin. Most remains of
3963:
495:
2473:
was adept at burrowing, since many extant crocodylians that are able to burrow also don't show any of the specific morphological specializations. Georgi and Krause, also in the same year, agreed that the burrowing hypothesis couldn't be confidently excluded, and claimed that the hypothesis needs more
2464:
In 2010, Kley and colleagues have argued against this hypothesis on the basis of its morphology which doesn't show an adaptation as head-first burrowers, with its relatively large, wide head being disadvantageous and contrasting modern head-first burrowers. They also pointed out that the snout is not
2452:
has been suggested in its initial description based on the robust limbs and short snout, which appears shovel-like, and the underslung lower jaw that would prevent friction when the animal opens its jaws during burrowing. There are also areas on the skull that may have attached to strong neck muscles
590:
has a tetraserial paravertebral shield over its back, meaning that there are four rows of tightly locking paramedial osteoderms (osteoderms to either side of the midline of the back). To either side of the shield, there are four rows of accessory parasagittal osteoderms. These accessory osteoderms
2474:
extensive comparative and functional analyses. They also suggested that its relatively short neck would have been a potential advantage for moving through a dense medium like a tunnel, and that it may have likely used its head on occasions to aid scratch-digging with its limbs.
2431:
as a possible adaptation to an entirely terrestrial lifestyle. Robust legs are also consistent with terrestrial locomotion. The deltopectoral crest on the humerus and the anterior crest on the femur served as attachment points for strong limb muscles. The hindlimbs of
581:
also has osteoderms covering much of the limbs. Osteoderms covering the back, tail, and limbs are light and porous, while the osteoderms covering the belly are plate-like and have an inner structure resembling spongy
2468:
In the same year, Sertich and
Groenke noted that while the appendicular skeleton including the scapula and the forelimb doesn't show specific morphological adaptation for digging, it doesn't exclude the idea that
3103:
Agustín G. Martinelli; Joseph J.W. Sertich; Alberto C. Garrido & Ángel M. Praderio (2012). "A new peirosaurid from the Upper
Cretaceous of Argentina: Implications for specimens referred to
412:
535:
of the forearm fit tightly together. The front feet are small with large claws, and the back feet are also reduced in size. There is a small crest along the anterior edge of the
2622:
Krause, D.W.; Sertich, J.J.W.; Rogers, R.R.; Kast, S.C.; Rasoamiaramanana, A.H.; Buckley, G.A. (2010). "Overview of the discovery, distribution, and geological context of
455:
was small, about 0.75 metres (2.5 ft) long based on the skeletons of mature individuals. In contrast to most other crocodyliforms, which have long, low skulls,
4637:
512:
is broad and tripartite (three-pronged). On its surface, there is a laterally directed prominence. The deltopectoral crest, a crest on the upper end of the
467:. The shape of skulls differs considerably between specimens, with variation in ornamentation and bony projections. These differences may be indications of
508:
resembles that of other terrestrial crocodyliforms. There are several differences, however, that have been used to distinguish it from related forms. The
562:
at the hip, and no more than twenty caudal vertebrae in the tail. The number of vertebrae in the tail is less than that of most crocodyliforms, giving
4721:
3285:
4696:
4624:
475:
likely held its head so that the preorbital area was angled about 45° from horizontal. The teeth line the front of the jaws and are shaped like
4706:
3332:
4701:
4571:
2718:
2744:
Buckley, G.A.; Brochu, C.A.; Krause, D.W.; Pol, D. (2000). "A pug-nosed crocodyliform from the Late
Cretaceous of Madagascar".
2411:
429:
2540:. Unresolved relationships among notosuchians along with an incomplete fossil record have made it difficult to determine the
436:. Material was usually found in clays that were part of flow deposits in the Anembalemba Member of the Maevarano Formation.
2891:
Hill, R.V. (2010). "Osteoderms of
Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2798:
Kley, N.J.; Sertich, J.J.W.; Turner, A.H.; Krause, D.W.; O'Connor, P.M.; Georgi, J.A. (2010). "Craniofacial morphology of
2585:"A new Late Cretaceous vertebrate fauna from the Cauvery Basin, South India: implications for Gondwanan paleobiogeography"
355:. It is named for its unusually short skull. Fully grown individuals were about 0.75 metres (2.5 ft) in length. The
2488:
1750:
2708:
4716:
1781:
4528:
138:
3956:
2080:
3325:
3043:
1805:
3975:
2939:
1377:(2004), based on different character values than previous studies, produced a very different relationship among
4711:
2424:. Like other notosuchians, it was fully terrestrial, and the short tail would have had little use in swimming.
1417:
has a short snout and was probably herbivorous. Both genera were placed outside
Notosuchia in the larger clade
4521:
3192:
4535:
2926:
Turner, A.H.; Calvo, J.O. (2005). "A new sebecosuchian crocodyliform from the Late
Cretaceous of Patagonia".
4592:
1726:
433:
4543:
4691:
4663:
3417:
1822:
1266:
3177:"Late Cretaceous terrestrial vertebrates from Madagascar: Implications for Latin American biogeography"
523:
of the humerus, which connects to the pectoral girdle in the shoulder joint, has a distinctive rounded
3254:
3968:
3318:
3062:
2753:
289:
126:
3304:
2584:
2063:
1448:
The following cladogram simplified after a comprehensive analysis of notosuchians which focused on
374:
366:
4271:
3235:
3196:
3024:
2982:
2961:
Diego Pol; Juan M. Leardi; Agustina
Lecuona & Marcelo Krause (2012). "Postcranial anatomy of
2943:
2908:
2873:
2819:
2777:
2689:
2643:
2604:
2564:
517:
370:
133:
4668:
3962:
3175:
Krause, D.W.; O'Connor, P.M.; Rogers, K.C.; Sampson, S.D.; Buckley, G.A.; Rogers, R.R. (2006).
4650:
4642:
3831:
3561:
3157:
3078:
2769:
2714:
1500:
520:
480:
468:
4655:
4415:
3377:
3262:
3227:
3188:
3147:
3116:
3070:
3016:
2974:
2935:
2900:
2865:
2811:
2761:
2681:
2635:
2596:
2177:
1406:
1324:
1290:
498:
4474:
4449:
4287:
4260:
4205:
4180:
4173:
4127:
4080:
4054:
3866:
3838:
3577:
2194:
1649:
1574:
1531:
1430:
892:
397:
348:
232:
219:
30:
1358:
and other genera within
Notosuchia. Turner and Calvo (2005) also found a clade including
577:. These form shields over the back, underside, and tail. Unusually among crocodyliforms,
3290:
3066:
2757:
2406:
4490:
4440:
4402:
4348:
4326:
4212:
4150:
4097:
3824:
3748:
3727:
3625:
3542:
3176:
2505:
2039:
1438:
1418:
1135:
484:
464:
117:
3074:
4685:
4483:
4456:
4422:
4383:
4368:
4341:
4334:
4280:
4246:
4239:
4230:
4192:
4157:
4069:
4039:
4032:
3901:
3817:
3810:
3794:
3720:
3692:
3135:
2647:
2608:
2541:
2509:
2129:
1999:
1864:
1666:
1553:
1275:
1086:
1069:
907:
766:
700:
344:
68:
3239:
3028:
2986:
2947:
2912:
2877:
2823:
2693:
2420:
was probably a herbivore, with its complex dentition resembling that of herbivorous
4463:
4359:
4313:
4104:
4047:
3873:
3845:
3803:
3762:
3755:
3739:
3712:
3705:
3667:
3604:
3402:
3200:
2781:
2558:
2234:
1968:
1948:
1898:
1625:
1340:
1331:
1327:
1308:
1303:
1298:
1285:
1177:
1152:
1014:
966:
826:
742:
356:
316:
206:
3120:
3020:
2978:
2904:
2869:
2815:
2685:
2639:
2600:
1354:. Most of the following phylogenetic analyses resulted in a similar placement of
4376:
4294:
4253:
4165:
4137:
4024:
3894:
3880:
3783:
3681:
3632:
3618:
3611:
3597:
3570:
3528:
3497:
3479:
3466:
3429:
3341:
2583:
Prasad, Guntupalli V. R.; Verma, Omkar; Flynn, John J.; Goswami, Anjali (2013).
2393:
2248:
1877:
1514:
1473:
1442:
952:
683:
264:
43:
2834:
444:
4429:
4391:
4111:
4090:
4006:
3993:
3924:
3887:
3859:
3852:
3776:
3660:
3653:
3646:
3550:
3386:
3371:
3294:
3231:
3152:
3136:"Repeated Evolution of Herbivorous Crocodyliforms during the Age of Dinosaurs"
2525:
2497:
2153:
1982:
1592:
1459:
1350:
1045:
939:
872:
659:
630:
604:
476:
389:
352:
341:
248:
88:
53:
4615:
3218:(Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar".
3082:
4219:
3769:
3639:
3588:
3535:
2521:
2445:
2421:
574:
524:
401:
393:
193:
150:
93:
37:
3267:
3161:
2773:
2456:
2436:
were semierect, unlike the fully erect posture of most other notosuchians.
583:
1425:, previously considered to be a basal notosuchian and a close relative of
373:, although one isolated multicuspid tooth of this genus was discovered in
4609:
3365:
2501:
2500:
in various phylogenetic analyses. Nearly all notosuchians are known from
555:
170:
83:
78:
63:
58:
48:
3048:, a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil"
4629:
3674:
3514:
3007:(Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2856:(Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2829:
2802:(Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2672:(Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2626:(Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar".
2264:
1280:
544:
513:
509:
494:
396:
like most other crocodylomorphs. In fact, these features have led many
334:
297:
180:
98:
73:
3086:
3359:
2765:
2513:
2496:
is unclear. It has been classified as both a notosuchian and a basal
2427:
The osteoderm shield was inflexible, restricting lateral movement in
1294:
559:
540:
528:
160:
4586:
392:, which coupled with its short and deep snout suggests it was not a
428:
were found as part of the
Mahajanga Basin Project, directed by the
3214:
Turner, A.H. (2006). "Osteology and phylogeny of a new species of
2668:
Georgi, J.A.; Krause, D.W. (2010). "Postcranial axial skeleton of
2537:
2517:
2492:, is also known from the Maevarano Formation, but its relation to
2455:
2405:
1398:
1270:
536:
493:
471:. The front (preorbita) portion of the skull is angled downwards.
463:, giving the genus its name, which means "pug-nosed crocodile" in
443:
411:
378:
337:
2852:
Sertich, J.J.W.; Groenke, J.R. (2010). "Appendicular skeleton of
2460:
Outdated restoration done before discovery of whole-body skeleton
558:
in the neck, at least fifteen dorsal vertebrae in the back, two
532:
4590:
4560:
3991:
3464:
3352:
3314:
459:
has a distinctively short snout. The snout resembles that of a
3310:
1452:
presented by Alan H. Turner and Joseph J. W. Sertich in 2010.
1273:
Notosuchia, and was often considered to be closely related to
483:(which articulates with the neck vertebrae) is downturned. 45
460:
3003:
Turner, A.H.; Sertich, J.W. (2010). "Phylogenetic history of
2833:
Text was copied from this source, which is available under a
2508:
that existed throughout much of the
Mesozoic and encompassed
2965:(Crocodyliformes, Sebecidae) from the Eocene of Patagonia".
2940:
10.1671/0272-4634(2005)025[0087:ANSCFT]2.0.CO;2
3193:
10.3417/0026-6493(2006)93[178:LCTVFM]2.0.CO;2
3042:
Carvalho, I.S.; Ribeiro, L.C.B.; Avilla, L.S. (2004).
2835:
Creative Commons Attribution 4.0 International License
1344:. These taxa were placed in Notosuchidae along with
306:
4599:
4473:
4439:
4401:
4358:
4312:
4270:
4229:
4191:
4136:
4125:
4079:
4004:
3923:
3793:
3738:
3691:
3587:
3560:
3512:
3477:
3428:
3401:
2453:that would have been well suited for burrowing.
2486:arrived in Madagascar. A similar crocodyliform,
3134:Melstrom, Keegan; Irmis, Randall (2019-06-01).
2391:: Based on a specimen that was reassigned from
2532:, regarded as one of the closest relatives of
1311:position than some other notosuchians such as
504:In most respects, the postcranial skeleton of
3326:
8:
3255:"Cretaceous crocodyliforms from the Sahara"
2713:. John Wiley & Sons, Ltd. p. 249.
1301:studies have placed it closer to the genus
4587:
4557:
4407:
4318:
4197:
4142:
4133:
4016:
4012:
4001:
3988:
3920:
3697:
3520:
3489:
3485:
3474:
3461:
3407:
3398:
3349:
3333:
3319:
3311:
2793:
2791:
116:
20:
3266:
3253:Sereno, P. C.; Larsson, H. C. E. (2009).
3151:
3107:Price (Crocodyliformes: Peirosauridae)".
2739:
2737:
1397:was found to be the sister taxon of the
1315:. In its initial description by Buckley
3181:Annals of the Missouri Botanical Garden
2998:
2996:
2575:
2847:
2845:
2843:
2663:
2661:
2659:
2657:
1441:, specifically a member of the family
1373:The phylogenetic analysis of Carvalho
7:
591:tightly interlock with one another.
2240:
2169:
2145:
2121:
2114:
2055:
2031:
2024:
1974:
1964:
1940:
1890:
1883:
1856:
1797:
1773:
1766:
1742:
1718:
1641:
1617:
1566:
1559:
1506:
1496:
1489:
1465:
1455:
1127:
1120:
1061:
1037:
1030:
1006:
982:
958:
931:
885:
878:
868:
758:
734:
675:
651:
644:
637:
626:
554:is known. There are eight cervical
491:, can be found in the skull alone.
3009:Journal of Vertebrate Paleontology
2967:Journal of Vertebrate Paleontology
2928:Journal of Vertebrate Paleontology
2893:Journal of Vertebrate Paleontology
2858:Journal of Vertebrate Paleontology
2804:Journal of Vertebrate Paleontology
2674:Journal of Vertebrate Paleontology
2628:Journal of Vertebrate Paleontology
2589:Journal of Vertebrate Paleontology
1393:, were placed outside Notosuchia.
573:was covered in bony plates called
14:
3046:Uberabasuchus terrificus sp. nov.
527:shape. The limbs are robust. The
4722:Prehistoric pseudosuchian genera
4542:
4534:
4527:
4520:
3974:
3967:
3961:
3955:
3297:
2828:
1429:, was placed in its own family,
479:. At the back of the skull, the
137:
4697:Late Cretaceous crocodylomorphs
2412:Field Museum of Natural History
1338:, and the two were allied with
2448:, or burrowing, lifestyle for
1751:"Araripesuchus" tsangatsangana
543:, the anterior process of the
1:
4707:Crocodylomorphs of Madagascar
3121:10.1016/j.cretres.2012.03.017
3075:10.1016/S1342-937X(05)71079-0
1265:was first considered to be a
550:Most of the spinal column of
3284:Mikko's Phylogenic Archives
3021:10.1080/02724634.2010.532348
2979:10.1080/02724634.2012.646833
2905:10.1080/02724634.2010.518110
2870:10.1080/02724634.2010.516902
2816:10.1080/02724634.2010.532674
2686:10.1080/02724634.2010.519172
2640:10.1080/02724634.2010.516784
2601:10.1080/02724634.2013.777348
2489:Araripesuchus tsangatsangana
2440:Possible fossorial lifestyle
1437:was found to be a member of
1279:from the Late Cretaceous of
4702:Terrestrial crocodylomorphs
4572:Terrestrial crocodylomorphs
2410:Reconstructed skull in the
1782:Araripesuchus buitreraensis
1433:, also outside Notosuchia.
598:
569:Like other crocodyliforms,
4738:
3015:(6, Supplement): 177–236.
2899:(6, Supplement): 154–176.
2864:(6, Supplement): 122–153.
2707:Michael J. Benton (2015).
2081:Sphagesaurus montealtensis
448:Size comparison to a human
125:Reconstructed skeleton in
4567:
4556:
4518:
4410:
4321:
4200:
4145:
4019:
4015:
4000:
3987:
3953:
3919:
3700:
3523:
3492:
3488:
3473:
3460:
3410:
3397:
3348:
3232:10.1080/08912960500516112
3153:10.1016/j.cub.2019.05.076
2680:(6, Supplement): 99–121.
2262:
2245:
2238:
2191:
2174:
2167:
2150:
2143:
2126:
2119:
2112:
2077:
2060:
2053:
2036:
2029:
2022:
1996:
1979:
1972:
1962:
1945:
1938:
1912:
1895:
1888:
1881:
1861:
1854:
1819:
1806:Araripesuchus patagonicus
1802:
1795:
1778:
1771:
1764:
1747:
1740:
1723:
1716:
1663:
1646:
1639:
1622:
1615:
1589:
1571:
1564:
1557:
1528:
1511:
1504:
1494:
1487:
1470:
1463:
1175:
1149:
1132:
1125:
1118:
1083:
1066:
1059:
1042:
1035:
1028:
1011:
1004:
987:
980:
963:
956:
936:
929:
905:
890:
883:
876:
824:
780:
763:
756:
739:
732:
697:
680:
673:
656:
649:
642:
635:
430:Université d'Antananarivo
303:
296:
134:Scientific classification
132:
124:
115:
23:
16:Extinct genus of reptiles
2810:(6, Supplement): 13–98.
2710:Vertebrate Palaeontology
1727:"Araripesuchus" wegeneri
1381:and other notosuchians.
487:, or features unique to
2634:(6, Supplement): 4–12.
3268:10.3897/zookeys.28.325
2461:
2414:
619:Turner and Calvo, 2005
501:
449:
434:Stony Brook University
420:The first specimen of
417:
315:Buckley et al., 2000 (
4664:Paleobiology Database
2459:
2409:
1823:Araripesuchus gomesii
497:
447:
415:
3146:(14): 2389–2395.e3.
3105:Peirosaurus torminni
127:Royal Ontario Museum
3305:Paleontology portal
3109:Cretaceous Research
3067:2004GondR...7..975C
2963:Sebecus icaeorhinus
2758:2000Natur.405..941B
2064:Sphagesaurus huenei
566:a very short tail.
375:Kallamedu Formation
367:Maevarano Formation
4717:Maastrichtian life
4562:Related categories
4272:Pissarrachampsinae
3220:Historical Biology
2565:Smallest organisms
2482:It is unknown how
2462:
2415:
1323:was placed in the
502:
450:
418:
416:Skull FMNH PR 2597
371:Mahajanga Province
4679:
4678:
4651:Open Tree of Life
4593:Taxon identifiers
4584:
4583:
4580:
4579:
4552:
4551:
4516:
4515:
4512:
4511:
4508:
4507:
4504:
4503:
4500:
4499:
4308:
4307:
4304:
4303:
4121:
4120:
3983:
3982:
3951:
3950:
3947:
3946:
3943:
3942:
3915:
3914:
3911:
3910:
3832:Colhuehuapisuchus
3562:Mahajangasuchidae
3508:
3507:
3456:
3455:
3452:
3451:
3448:
3447:
3055:Gondwana Research
3005:Simosuchus clarki
2854:Simosuchus clarki
2800:Simosuchus clarki
2752:(6789): 941–944.
2670:Simosuchus clarki
2624:Simosuchus clarki
2478:Paleobiogeography
2398:
2383:
2382:
2374:
2373:
2365:
2364:
2356:
2355:
2347:
2346:
2338:
2337:
2329:
2328:
2320:
2319:
2311:
2310:
2302:
2301:
2293:
2292:
2284:
2283:
2275:
2274:
2224:
2223:
2215:
2214:
2206:
2205:
2101:
2100:
2092:
2091:
2011:
2010:
1927:
1926:
1843:
1842:
1834:
1833:
1705:
1704:
1696:
1695:
1687:
1686:
1678:
1677:
1604:
1603:
1543:
1542:
1501:Mahajangasuchidae
1450:Simosuchus clarki
1260:
1259:
1256:
1255:
1251:
1250:
1242:
1241:
1233:
1232:
1224:
1223:
1215:
1214:
1206:
1205:
1197:
1196:
1188:
1187:
1164:
1163:
1107:
1106:
1098:
1097:
918:
917:
851:
850:
846:
845:
837:
836:
813:
812:
804:
803:
795:
794:
721:
720:
712:
711:
481:occipital condyle
469:sexual dimorphism
422:Simosuchus clarki
400:to consider it a
388:were shaped like
365:, found from the
362:Simosuchus clarki
326:
325:
320:
292:
4729:
4672:
4671:
4659:
4658:
4646:
4645:
4633:
4632:
4620:
4619:
4618:
4588:
4558:
4547:
4546:
4539:
4538:
4532:
4531:
4524:
4416:Adamantinasuchus
4408:
4319:
4198:
4143:
4134:
4017:
4013:
4002:
3989:
3978:
3972:
3971:
3965:
3959:
3921:
3698:
3521:
3490:
3486:
3475:
3462:
3408:
3399:
3392:
3391:
3378:Mesoeucrocodylia
3350:
3335:
3328:
3321:
3312:
3307:
3302:
3301:
3300:
3289:Digimorph Entry
3273:
3272:
3270:
3250:
3244:
3243:
3211:
3205:
3204:
3172:
3166:
3165:
3155:
3131:
3125:
3124:
3100:
3094:
3093:
3091:
3085:. Archived from
3052:
3039:
3033:
3032:
3000:
2991:
2990:
2958:
2952:
2951:
2923:
2917:
2916:
2888:
2882:
2881:
2849:
2838:
2832:
2827:
2795:
2786:
2785:
2766:10.1038/35016061
2741:
2732:
2731:
2729:
2727:
2704:
2698:
2697:
2665:
2652:
2651:
2619:
2613:
2612:
2595:(6): 1260–1268.
2580:
2386:
2241:
2178:Adamantinasuchus
2170:
2146:
2122:
2115:
2056:
2032:
2025:
1975:
1965:
1941:
1891:
1884:
1857:
1798:
1774:
1767:
1743:
1719:
1642:
1618:
1567:
1560:
1507:
1497:
1490:
1466:
1456:
1407:Chimaerasuchidae
1370:in their study.
1291:Early Cretaceous
1128:
1121:
1062:
1038:
1031:
1007:
983:
959:
932:
886:
879:
869:
861:Pol et al., 2012
857:
856:
759:
735:
676:
652:
645:
638:
627:
615:
614:
599:
560:sacral vertebrae
516:, is small. The
499:Life restoration
398:palaeontologists
314:
308:
283:
276:
263:
247:
142:
141:
120:
110:
40:
29:Temporal range:
21:
4737:
4736:
4732:
4731:
4730:
4728:
4727:
4726:
4712:Maevarano fauna
4682:
4681:
4680:
4675:
4667:
4662:
4654:
4649:
4641:
4636:
4628:
4623:
4614:
4613:
4608:
4595:
4585:
4576:
4563:
4548:
4541:
4533:
4526:
4496:
4475:Caipirasuchinae
4469:
4450:Armadillosuchus
4435:
4397:
4354:
4300:
4288:Pissarrachampsa
4266:
4261:Stratiotosuchus
4225:
4206:Cynodontosuchus
4187:
4181:Razanandrongobe
4174:Pehuenchesuchus
4130:
4128:Xenodontosuchia
4117:
4081:Candidodontidae
4075:
4055:Razanandrongobe
4009:
3996:
3979:
3966:
3939:
3907:
3867:Montealtosuchus
3839:Gasparinisuchus
3789:
3734:
3687:
3583:
3578:Mahajangasuchus
3556:
3517:
3504:
3482:
3469:
3444:
3424:
3393:
3355:
3354:
3344:
3339:
3303:
3298:
3296:
3281:
3276:
3252:
3251:
3247:
3213:
3212:
3208:
3174:
3173:
3169:
3140:Current Biology
3133:
3132:
3128:
3102:
3101:
3097:
3089:
3061:(4): 975–1002.
3050:
3041:
3040:
3036:
3002:
3001:
2994:
2960:
2959:
2955:
2925:
2924:
2920:
2890:
2889:
2885:
2851:
2850:
2841:
2797:
2796:
2789:
2743:
2742:
2735:
2725:
2723:
2721:
2706:
2705:
2701:
2667:
2666:
2655:
2621:
2620:
2616:
2582:
2581:
2577:
2573:
2554:
2504:, the southern
2480:
2442:
2404:
2384:
2375:
2366:
2357:
2348:
2339:
2330:
2321:
2312:
2303:
2294:
2285:
2276:
2225:
2216:
2207:
2195:Armadillosuchus
2102:
2093:
2012:
1928:
1844:
1835:
1706:
1697:
1688:
1679:
1650:Montealtosuchus
1605:
1575:Gasparinisuchus
1544:
1532:Mahajangasuchus
1431:Uruguaysuchidae
1252:
1243:
1234:
1225:
1216:
1207:
1198:
1189:
1165:
1108:
1099:
919:
893:Uruguaysuchidae
847:
838:
814:
805:
796:
722:
713:
609:
597:
442:
410:
349:Late Cretaceous
282:
274:
261:
245:
233:Crocodyliformes
220:Crocodylomorpha
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
4735:
4733:
4725:
4724:
4719:
4714:
4709:
4704:
4699:
4694:
4684:
4683:
4677:
4676:
4674:
4673:
4660:
4647:
4634:
4621:
4605:
4603:
4597:
4596:
4591:
4582:
4581:
4578:
4577:
4575:
4574:
4568:
4565:
4564:
4561:
4554:
4553:
4550:
4549:
4519:
4517:
4514:
4513:
4510:
4509:
4506:
4505:
4502:
4501:
4498:
4497:
4495:
4494:
4491:Morrinhosuchus
4487:
4479:
4477:
4471:
4470:
4468:
4467:
4460:
4453:
4445:
4443:
4441:Sphagesaurinae
4437:
4436:
4434:
4433:
4426:
4419:
4411:
4405:
4403:Sphagesauridae
4399:
4398:
4396:
4395:
4388:
4380:
4373:
4364:
4362:
4356:
4355:
4353:
4352:
4349:Eptalofosuchus
4345:
4338:
4331:
4327:Brasileosaurus
4322:
4316:
4310:
4309:
4306:
4305:
4302:
4301:
4299:
4298:
4291:
4284:
4276:
4274:
4268:
4267:
4265:
4264:
4257:
4250:
4243:
4235:
4233:
4227:
4226:
4224:
4223:
4216:
4213:Gondwanasuchus
4209:
4201:
4195:
4189:
4188:
4186:
4185:
4177:
4170:
4162:
4154:
4151:Chimaerasuchus
4146:
4140:
4131:
4126:
4123:
4122:
4119:
4118:
4116:
4115:
4108:
4101:
4098:Lavocatchampsa
4094:
4086:
4084:
4077:
4076:
4074:
4073:
4066:
4059:
4051:
4044:
4036:
4029:
4020:
4010:
4005:
3998:
3997:
3992:
3985:
3984:
3981:
3980:
3954:
3952:
3949:
3948:
3945:
3944:
3941:
3940:
3938:
3937:
3936:
3935:
3929:
3927:
3917:
3916:
3913:
3912:
3909:
3908:
3906:
3905:
3898:
3891:
3884:
3877:
3870:
3863:
3856:
3849:
3842:
3835:
3828:
3825:Bayomesasuchus
3821:
3814:
3807:
3799:
3797:
3791:
3790:
3788:
3787:
3780:
3773:
3766:
3759:
3752:
3749:Barreirosuchus
3744:
3742:
3736:
3735:
3733:
3732:
3728:Stolokrosuchus
3724:
3717:
3709:
3701:
3695:
3689:
3688:
3686:
3685:
3678:
3671:
3664:
3657:
3650:
3643:
3636:
3629:
3626:Dentaneosuchus
3622:
3615:
3608:
3601:
3593:
3591:
3585:
3584:
3582:
3581:
3574:
3566:
3564:
3558:
3557:
3555:
3554:
3547:
3543:Trematochampsa
3539:
3532:
3524:
3518:
3513:
3510:
3509:
3506:
3505:
3503:
3502:
3493:
3483:
3478:
3471:
3470:
3465:
3458:
3457:
3454:
3453:
3450:
3449:
3446:
3445:
3443:
3442:
3441:
3440:
3434:
3432:
3426:
3425:
3423:
3422:
3421:
3420:
3411:
3405:
3395:
3394:
3390:
3389:
3380:
3374:
3368:
3362:
3353:
3346:
3345:
3340:
3338:
3337:
3330:
3323:
3315:
3309:
3308:
3293:
3292:
3287:
3280:
3279:External links
3277:
3275:
3274:
3245:
3226:(3): 255–369.
3206:
3167:
3126:
3095:
3092:on 2011-07-06.
3034:
2992:
2973:(2): 328–354.
2953:
2918:
2883:
2839:
2787:
2733:
2719:
2699:
2653:
2614:
2574:
2572:
2569:
2568:
2567:
2562:
2553:
2550:
2506:supercontinent
2479:
2476:
2441:
2438:
2403:
2400:
2381:
2380:
2377:
2376:
2372:
2371:
2368:
2367:
2363:
2362:
2359:
2358:
2354:
2353:
2350:
2349:
2345:
2344:
2341:
2340:
2336:
2335:
2332:
2331:
2327:
2326:
2323:
2322:
2318:
2317:
2314:
2313:
2309:
2308:
2305:
2304:
2300:
2299:
2296:
2295:
2291:
2290:
2287:
2286:
2282:
2281:
2278:
2277:
2273:
2272:
2269:
2268:
2261:
2258:
2257:
2254:
2253:
2244:
2239:
2237:
2231:
2230:
2227:
2226:
2222:
2221:
2218:
2217:
2213:
2212:
2209:
2208:
2204:
2203:
2200:
2199:
2190:
2187:
2186:
2183:
2182:
2173:
2168:
2166:
2163:
2162:
2159:
2158:
2149:
2144:
2142:
2139:
2138:
2135:
2134:
2125:
2120:
2118:
2113:
2111:
2108:
2107:
2104:
2103:
2099:
2098:
2095:
2094:
2090:
2089:
2086:
2085:
2076:
2073:
2072:
2069:
2068:
2059:
2054:
2052:
2049:
2048:
2045:
2044:
2040:Chimaerasuchus
2035:
2030:
2028:
2023:
2021:
2018:
2017:
2014:
2013:
2009:
2008:
2005:
2004:
1995:
1992:
1991:
1988:
1987:
1978:
1973:
1971:
1963:
1961:
1958:
1957:
1954:
1953:
1944:
1939:
1937:
1934:
1933:
1930:
1929:
1925:
1924:
1921:
1920:
1911:
1908:
1907:
1904:
1903:
1894:
1889:
1887:
1882:
1880:
1874:
1873:
1870:
1869:
1860:
1855:
1853:
1850:
1849:
1846:
1845:
1841:
1840:
1837:
1836:
1832:
1831:
1828:
1827:
1818:
1815:
1814:
1811:
1810:
1801:
1796:
1794:
1791:
1790:
1787:
1786:
1777:
1772:
1770:
1765:
1763:
1760:
1759:
1756:
1755:
1746:
1741:
1739:
1736:
1735:
1732:
1731:
1722:
1717:
1715:
1712:
1711:
1708:
1707:
1703:
1702:
1699:
1698:
1694:
1693:
1690:
1689:
1685:
1684:
1681:
1680:
1676:
1675:
1672:
1671:
1662:
1659:
1658:
1655:
1654:
1645:
1640:
1638:
1635:
1634:
1631:
1630:
1621:
1616:
1614:
1611:
1610:
1607:
1606:
1602:
1601:
1598:
1597:
1588:
1585:
1584:
1581:
1580:
1570:
1565:
1563:
1558:
1556:
1550:
1549:
1546:
1545:
1541:
1540:
1537:
1536:
1527:
1524:
1523:
1520:
1519:
1510:
1505:
1503:
1495:
1493:
1488:
1486:
1483:
1482:
1479:
1478:
1469:
1464:
1462:
1454:
1439:Peirosauroidea
1419:Gondwanasuchia
1415:Chimaerasuchus
1405:in the family
1403:Chimaerasuchus
1307:and in a more
1269:member of the
1258:
1257:
1254:
1253:
1249:
1248:
1245:
1244:
1240:
1239:
1236:
1235:
1231:
1230:
1227:
1226:
1222:
1221:
1218:
1217:
1213:
1212:
1209:
1208:
1204:
1203:
1200:
1199:
1195:
1194:
1191:
1190:
1186:
1185:
1182:
1181:
1174:
1171:
1170:
1167:
1166:
1162:
1161:
1158:
1157:
1148:
1145:
1144:
1141:
1140:
1136:Chimaerasuchus
1131:
1126:
1124:
1119:
1117:
1114:
1113:
1110:
1109:
1105:
1104:
1101:
1100:
1096:
1095:
1092:
1091:
1082:
1079:
1078:
1075:
1074:
1065:
1060:
1058:
1055:
1054:
1051:
1050:
1041:
1036:
1034:
1029:
1027:
1024:
1023:
1020:
1019:
1010:
1005:
1003:
1000:
999:
996:
995:
986:
981:
979:
976:
975:
972:
971:
962:
957:
955:
949:
948:
945:
944:
935:
930:
928:
925:
924:
921:
920:
916:
915:
912:
911:
904:
901:
900:
897:
896:
889:
884:
882:
877:
875:
867:
864:
863:
853:
852:
849:
848:
844:
843:
840:
839:
835:
834:
831:
830:
823:
820:
819:
816:
815:
811:
810:
807:
806:
802:
801:
798:
797:
793:
792:
789:
788:
779:
776:
775:
772:
771:
762:
757:
755:
752:
751:
748:
747:
738:
733:
731:
728:
727:
724:
723:
719:
718:
715:
714:
710:
709:
706:
705:
696:
693:
692:
689:
688:
679:
674:
672:
669:
668:
665:
664:
655:
650:
648:
643:
641:
636:
634:
625:
622:
621:
611:
610:
602:
596:
595:Classification
593:
547:is spur-like.
485:autapomorphies
441:
438:
409:
406:
345:crocodyliforms
324:
323:
322:
321:
301:
300:
294:
293:
272:
268:
267:
259:
252:
251:
243:
236:
235:
230:
223:
222:
217:
210:
209:
204:
197:
196:
191:
184:
183:
178:
174:
173:
168:
164:
163:
158:
154:
153:
148:
144:
143:
130:
129:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4734:
4723:
4720:
4718:
4715:
4713:
4710:
4708:
4705:
4703:
4700:
4698:
4695:
4693:
4690:
4689:
4687:
4670:
4665:
4661:
4657:
4652:
4648:
4644:
4639:
4635:
4631:
4626:
4622:
4617:
4611:
4607:
4606:
4604:
4602:
4598:
4594:
4589:
4573:
4570:
4569:
4566:
4559:
4555:
4545:
4540:
4537:
4530:
4523:
4493:
4492:
4488:
4486:
4485:
4484:Caipirasuchus
4481:
4480:
4478:
4476:
4472:
4466:
4465:
4461:
4459:
4458:
4457:Caryonosuchus
4454:
4452:
4451:
4447:
4446:
4444:
4442:
4438:
4432:
4431:
4427:
4425:
4424:
4423:Labidiosuchus
4420:
4418:
4417:
4413:
4412:
4409:
4406:
4404:
4400:
4394:
4393:
4389:
4386:
4385:
4384:Mariliasuchus
4381:
4379:
4378:
4374:
4371:
4370:
4369:Coronelsuchus
4366:
4365:
4363:
4361:
4357:
4351:
4350:
4346:
4344:
4343:
4342:Coronelsuchus
4339:
4337:
4336:
4335:Coringasuchus
4332:
4329:
4328:
4324:
4323:
4320:
4317:
4315:
4311:
4297:
4296:
4292:
4290:
4289:
4285:
4283:
4282:
4281:Campinasuchus
4278:
4277:
4275:
4273:
4269:
4263:
4262:
4258:
4256:
4255:
4251:
4249:
4248:
4247:Aplestosuchus
4244:
4242:
4241:
4240:Aphaurosuchus
4237:
4236:
4234:
4232:
4231:Baurusuchinae
4228:
4222:
4221:
4217:
4215:
4214:
4210:
4208:
4207:
4203:
4202:
4199:
4196:
4194:
4193:Baurusuchidae
4190:
4183:
4182:
4178:
4176:
4175:
4171:
4168:
4167:
4163:
4160:
4159:
4158:Comahuesuchus
4155:
4153:
4152:
4148:
4147:
4144:
4141:
4139:
4135:
4132:
4129:
4124:
4114:
4113:
4109:
4107:
4106:
4102:
4100:
4099:
4095:
4093:
4092:
4088:
4087:
4085:
4082:
4078:
4072:
4071:
4070:Uruguaysuchus
4067:
4065:
4064:
4060:
4057:
4056:
4052:
4050:
4049:
4045:
4042:
4041:
4040:Comahuesuchus
4037:
4035:
4034:
4033:Araripesuchus
4030:
4027:
4026:
4022:
4021:
4018:
4014:
4011:
4008:
4003:
3999:
3995:
3990:
3986:
3977:
3973:
3970:
3964:
3958:
3933:
3932:
3931:
3930:
3928:
3926:
3922:
3918:
3904:
3903:
3902:Uberabasuchus
3899:
3897:
3896:
3892:
3890:
3889:
3885:
3883:
3882:
3878:
3876:
3875:
3871:
3869:
3868:
3864:
3862:
3861:
3857:
3855:
3854:
3850:
3848:
3847:
3843:
3841:
3840:
3836:
3834:
3833:
3829:
3827:
3826:
3822:
3820:
3819:
3818:Barrosasuchus
3815:
3813:
3812:
3811:Barcinosuchus
3808:
3806:
3805:
3801:
3800:
3798:
3796:
3795:Peirosaurinae
3792:
3786:
3785:
3781:
3779:
3778:
3774:
3772:
3771:
3767:
3765:
3764:
3760:
3758:
3757:
3753:
3751:
3750:
3746:
3745:
3743:
3741:
3737:
3730:
3729:
3725:
3723:
3722:
3721:Miadanasuchus
3718:
3715:
3714:
3710:
3708:
3707:
3703:
3702:
3699:
3696:
3694:
3693:Peirosauridae
3690:
3684:
3683:
3679:
3677:
3676:
3672:
3670:
3669:
3665:
3663:
3662:
3658:
3656:
3655:
3651:
3649:
3648:
3644:
3642:
3641:
3637:
3635:
3634:
3630:
3628:
3627:
3623:
3621:
3620:
3616:
3614:
3613:
3609:
3607:
3606:
3602:
3600:
3599:
3595:
3594:
3592:
3590:
3586:
3580:
3579:
3575:
3573:
3572:
3568:
3567:
3565:
3563:
3559:
3553:
3552:
3548:
3545:
3544:
3540:
3538:
3537:
3533:
3531:
3530:
3526:
3525:
3522:
3519:
3516:
3511:
3500:
3499:
3495:
3494:
3491:
3487:
3484:
3481:
3476:
3472:
3468:
3463:
3459:
3438:
3437:
3436:
3435:
3433:
3431:
3427:
3419:
3415:
3414:
3413:
3412:
3409:
3406:
3404:
3400:
3396:
3388:
3384:
3381:
3379:
3375:
3373:
3369:
3367:
3363:
3361:
3357:
3356:
3351:
3347:
3343:
3336:
3331:
3329:
3324:
3322:
3317:
3316:
3313:
3306:
3295:
3291:
3288:
3286:
3283:
3282:
3278:
3269:
3264:
3261:(28): 1–143.
3260:
3256:
3249:
3246:
3241:
3237:
3233:
3229:
3225:
3221:
3217:
3216:Araripesuchus
3210:
3207:
3202:
3198:
3194:
3190:
3187:(2): 178–20.
3186:
3182:
3178:
3171:
3168:
3163:
3159:
3154:
3149:
3145:
3141:
3137:
3130:
3127:
3122:
3118:
3114:
3110:
3106:
3099:
3096:
3088:
3084:
3080:
3076:
3072:
3068:
3064:
3060:
3056:
3049:
3047:
3038:
3035:
3030:
3026:
3022:
3018:
3014:
3010:
3006:
2999:
2997:
2993:
2988:
2984:
2980:
2976:
2972:
2968:
2964:
2957:
2954:
2949:
2945:
2941:
2937:
2933:
2929:
2922:
2919:
2914:
2910:
2906:
2902:
2898:
2894:
2887:
2884:
2879:
2875:
2871:
2867:
2863:
2859:
2855:
2848:
2846:
2844:
2840:
2836:
2831:
2825:
2821:
2817:
2813:
2809:
2805:
2801:
2794:
2792:
2788:
2783:
2779:
2775:
2771:
2767:
2763:
2759:
2755:
2751:
2747:
2740:
2738:
2734:
2722:
2720:9781118407554
2716:
2712:
2711:
2703:
2700:
2695:
2691:
2687:
2683:
2679:
2675:
2671:
2664:
2662:
2660:
2658:
2654:
2649:
2645:
2641:
2637:
2633:
2629:
2625:
2618:
2615:
2610:
2606:
2602:
2598:
2594:
2590:
2586:
2579:
2576:
2570:
2566:
2563:
2561:
2560:
2556:
2555:
2551:
2549:
2547:
2543:
2542:biogeographic
2539:
2535:
2531:
2527:
2523:
2519:
2515:
2511:
2510:South America
2507:
2503:
2499:
2495:
2491:
2490:
2485:
2477:
2475:
2472:
2466:
2458:
2454:
2451:
2447:
2439:
2437:
2435:
2430:
2425:
2423:
2419:
2413:
2408:
2401:
2399:
2396:
2395:
2390:
2379:
2378:
2370:
2369:
2361:
2360:
2352:
2351:
2343:
2342:
2334:
2333:
2325:
2324:
2316:
2315:
2307:
2306:
2298:
2297:
2289:
2288:
2280:
2279:
2271:
2270:
2267:
2266:
2260:
2259:
2256:
2255:
2252:
2251:
2250:
2243:
2242:
2236:
2233:
2232:
2229:
2228:
2220:
2219:
2211:
2210:
2202:
2201:
2198:
2197:
2196:
2189:
2188:
2185:
2184:
2181:
2180:
2179:
2172:
2171:
2165:
2164:
2161:
2160:
2157:
2156:
2155:
2148:
2147:
2141:
2140:
2137:
2136:
2133:
2132:
2131:
2130:Comahuesuchus
2124:
2123:
2117:
2116:
2110:
2109:
2106:
2105:
2097:
2096:
2088:
2087:
2084:
2083:
2082:
2075:
2074:
2071:
2070:
2067:
2066:
2065:
2058:
2057:
2051:
2050:
2047:
2046:
2043:
2042:
2041:
2034:
2033:
2027:
2026:
2020:
2019:
2016:
2015:
2007:
2006:
2003:
2002:
2001:
2000:Mariliasuchus
1994:
1993:
1990:
1989:
1986:
1985:
1984:
1977:
1976:
1970:
1967:
1966:
1960:
1959:
1956:
1955:
1952:
1951:
1950:
1943:
1942:
1936:
1935:
1932:
1931:
1923:
1922:
1919:
1918:
1917:
1910:
1909:
1906:
1905:
1902:
1901:
1900:
1893:
1892:
1886:
1885:
1879:
1876:
1875:
1872:
1871:
1868:
1867:
1866:
1865:Uruguaysuchus
1859:
1858:
1852:
1851:
1848:
1847:
1839:
1838:
1830:
1829:
1826:
1825:
1824:
1817:
1816:
1813:
1812:
1809:
1808:
1807:
1800:
1799:
1793:
1792:
1789:
1788:
1785:
1784:
1783:
1776:
1775:
1769:
1768:
1762:
1761:
1758:
1757:
1754:
1753:
1752:
1745:
1744:
1738:
1737:
1734:
1733:
1730:
1729:
1728:
1721:
1720:
1714:
1713:
1710:
1709:
1701:
1700:
1692:
1691:
1683:
1682:
1674:
1673:
1670:
1669:
1668:
1667:Uberabasuchus
1661:
1660:
1657:
1656:
1653:
1652:
1651:
1644:
1643:
1637:
1636:
1633:
1632:
1629:
1628:
1627:
1620:
1619:
1613:
1612:
1609:
1608:
1600:
1599:
1596:
1595:
1594:
1587:
1586:
1583:
1582:
1579:
1577:
1576:
1569:
1568:
1562:
1561:
1555:
1554:Peirosauridae
1552:
1551:
1548:
1547:
1539:
1538:
1535:
1534:
1533:
1526:
1525:
1522:
1521:
1518:
1517:
1516:
1509:
1508:
1502:
1499:
1498:
1492:
1491:
1485:
1484:
1481:
1480:
1477:
1476:
1475:
1468:
1467:
1461:
1458:
1457:
1453:
1451:
1446:
1444:
1440:
1436:
1432:
1428:
1424:
1423:Uruguaysuchus
1420:
1416:
1412:
1408:
1404:
1400:
1396:
1392:
1391:Comahuesuchus
1388:
1387:Uruguaysuchus
1385:, along with
1384:
1380:
1376:
1371:
1369:
1365:
1364:Uruguaysuchus
1361:
1357:
1353:
1352:
1347:
1343:
1342:
1337:
1336:Uruguaysuchus
1333:
1329:
1326:
1322:
1318:
1314:
1313:Uruguaysuchus
1310:
1306:
1305:
1300:
1296:
1292:
1288:
1287:
1282:
1278:
1277:
1276:Uruguaysuchus
1272:
1268:
1264:
1247:
1246:
1238:
1237:
1229:
1228:
1220:
1219:
1211:
1210:
1202:
1201:
1193:
1192:
1184:
1183:
1180:
1179:
1173:
1172:
1169:
1168:
1160:
1159:
1156:
1155:
1154:
1147:
1146:
1143:
1142:
1139:
1138:
1137:
1130:
1129:
1123:
1122:
1116:
1115:
1112:
1111:
1103:
1102:
1094:
1093:
1090:
1089:
1088:
1087:Mariliasuchus
1081:
1080:
1077:
1076:
1073:
1072:
1071:
1070:Comahuesuchus
1064:
1063:
1057:
1056:
1053:
1052:
1049:
1048:
1047:
1040:
1039:
1033:
1032:
1026:
1025:
1022:
1021:
1018:
1017:
1016:
1009:
1008:
1002:
1001:
998:
997:
994:
993:
992:
985:
984:
978:
977:
974:
973:
970:
969:
968:
961:
960:
954:
951:
950:
947:
946:
943:
942:
941:
934:
933:
927:
926:
923:
922:
914:
913:
910:
909:
908:Peirosauridae
903:
902:
899:
898:
895:
894:
888:
887:
881:
880:
874:
871:
870:
866:
865:
862:
859:
858:
855:
854:
842:
841:
833:
832:
829:
828:
822:
821:
818:
817:
809:
808:
800:
799:
791:
790:
787:
786:
785:
778:
777:
774:
773:
770:
769:
768:
767:Uruguaysuchus
761:
760:
754:
753:
750:
749:
746:
745:
744:
737:
736:
730:
729:
726:
725:
717:
716:
708:
707:
704:
703:
702:
701:Comahuesuchus
695:
694:
691:
690:
687:
686:
685:
678:
677:
671:
670:
667:
666:
663:
662:
661:
654:
653:
647:
646:
640:
639:
632:
629:
628:
624:
623:
620:
617:
616:
613:
612:
608:
607:of Notosuchia
606:
601:
600:
594:
592:
589:
585:
580:
576:
572:
567:
565:
561:
557:
553:
548:
546:
542:
538:
534:
530:
526:
522:
519:
515:
511:
507:
500:
496:
492:
490:
486:
482:
478:
474:
470:
466:
462:
458:
454:
446:
439:
437:
435:
431:
427:
423:
414:
407:
405:
403:
399:
395:
391:
387:
384:The teeth of
382:
380:
376:
372:
368:
364:
363:
358:
354:
350:
346:
343:
339:
336:
332:
331:
318:
313:
312:
305:
304:
302:
299:
295:
291:
287:
281:
280:
273:
270:
269:
266:
260:
257:
254:
253:
250:
244:
241:
238:
237:
234:
231:
228:
225:
224:
221:
218:
215:
212:
211:
208:
205:
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
4692:Notosuchians
4600:
4525:
4489:
4482:
4464:Sphagesaurus
4462:
4455:
4448:
4428:
4421:
4414:
4390:
4382:
4375:
4367:
4360:Notosuchidae
4347:
4340:
4333:
4325:
4314:Sphagesauria
4293:
4286:
4279:
4259:
4252:
4245:
4238:
4218:
4211:
4204:
4179:
4172:
4164:
4156:
4149:
4110:
4105:Malawisuchus
4103:
4096:
4089:
4068:
4062:
4061:
4053:
4048:Libycosuchus
4046:
4038:
4031:
4023:
3960:
3900:
3893:
3886:
3879:
3874:Patagosuchus
3872:
3865:
3858:
3851:
3846:Hamadasuchus
3844:
3837:
3830:
3823:
3816:
3809:
3804:Antaeusuchus
3802:
3782:
3775:
3768:
3763:Epoidesuchus
3761:
3756:Caririsuchus
3754:
3747:
3740:Pepesuchinae
3726:
3719:
3713:Fortignathus
3711:
3706:Amargasuchus
3704:
3680:
3673:
3668:Sahitisuchus
3666:
3659:
3652:
3645:
3638:
3631:
3624:
3617:
3610:
3605:Barinasuchus
3603:
3596:
3576:
3569:
3549:
3541:
3534:
3527:
3496:
3418:Pseudosuchia
3403:Pseudosuchia
3382:
3258:
3248:
3223:
3219:
3215:
3209:
3184:
3180:
3170:
3143:
3139:
3129:
3112:
3108:
3104:
3098:
3087:the original
3058:
3054:
3045:
3037:
3012:
3008:
3004:
2970:
2966:
2962:
2956:
2934:(1): 87–98.
2931:
2927:
2921:
2896:
2892:
2886:
2861:
2857:
2853:
2807:
2803:
2799:
2749:
2745:
2726:25 September
2724:. Retrieved
2709:
2702:
2677:
2673:
2669:
2631:
2627:
2623:
2617:
2592:
2588:
2578:
2559:Malawisuchus
2557:
2545:
2533:
2530:Libycosuchus
2529:
2493:
2487:
2483:
2481:
2470:
2467:
2463:
2449:
2443:
2433:
2428:
2426:
2417:
2416:
2402:Paleobiology
2392:
2388:
2385:
2263:
2247:
2246:
2235:Sebecosuchia
2193:
2192:
2176:
2175:
2152:
2151:
2128:
2127:
2079:
2078:
2062:
2061:
2038:
2037:
1998:
1997:
1981:
1980:
1969:Notosuchidae
1949:Malawisuchus
1947:
1946:
1915:
1914:
1913:
1899:Libycosuchus
1897:
1896:
1863:
1862:
1821:
1820:
1804:
1803:
1780:
1779:
1749:
1748:
1725:
1724:
1665:
1664:
1648:
1647:
1626:Hamadasuchus
1624:
1623:
1591:
1590:
1573:
1572:
1530:
1529:
1513:
1512:
1472:
1471:
1449:
1447:
1435:Malawisuchus
1434:
1426:
1422:
1414:
1410:
1402:
1394:
1390:
1386:
1382:
1378:
1374:
1372:
1368:Malawisuchus
1367:
1363:
1359:
1355:
1349:
1346:Libycosuchus
1345:
1341:Malawisuchus
1339:
1335:
1332:sister taxon
1328:Notosuchidae
1320:
1316:
1312:
1304:Libycosuchus
1302:
1299:phylogenetic
1286:Malawisuchus
1284:
1274:
1262:
1261:
1178:Sebecosuchia
1176:
1153:Sphagesaurus
1151:
1150:
1134:
1133:
1085:
1084:
1068:
1067:
1044:
1043:
1015:Malawisuchus
1013:
1012:
990:
989:
988:
967:Libycosuchus
965:
964:
938:
937:
906:
891:
860:
827:Sebecosuchia
825:
783:
782:
781:
765:
764:
743:Malawisuchus
741:
740:
699:
698:
682:
681:
658:
657:
618:
603:
587:
578:
570:
568:
563:
551:
549:
518:glenohumeral
505:
503:
488:
477:maple leaves
472:
456:
452:
451:
425:
421:
419:
390:maple leaves
385:
383:
361:
360:
357:type species
329:
328:
327:
310:
309:
285:
278:
277:
255:
239:
226:
213:
207:Pseudosuchia
200:
187:
24:
18:
4377:Llanosuchus
4295:Wargosuchus
4254:Baurusuchus
4166:Eremosuchus
4138:Baurusuchia
4025:Anatosuchus
3895:Rukwasuchus
3881:Peirosaurus
3784:Roxochampsa
3682:Zulmasuchus
3633:Iberosuchus
3619:Bretesuchus
3612:Bergisuchus
3598:Ayllusuchus
3571:Kaprosuchus
3529:Baharijodon
3498:Microsuchus
3480:Ziphosuchia
3467:Ziphosuchia
3430:Ziphosuchia
3342:Ziphosuchia
3115:: 191–200.
2544:origins of
2536:, lived in
2394:Peirosaurus
2249:Baurusuchus
1878:Ziphosuchia
1515:Kaprosuchus
1474:Anatosuchus
1443:Itasuchidae
953:Ziphosuchia
684:Anatosuchus
440:Description
342:notosuchian
265:Ziphosuchia
194:Archosauria
36:70–66
4686:Categories
4601:Simosuchus
4430:Yacarerani
4392:Notosuchus
4112:Pakasuchus
4091:Candidodon
4063:Simosuchus
4007:Notosuchia
3994:Notosuchia
3934:see below↓
3925:Notosuchia
3888:Pepesuchus
3860:Lomasuchus
3853:Kinesuchus
3777:Pepesuchus
3661:Ogresuchus
3654:Lorosuchus
3647:Langstonia
3551:Wanosuchus
3439:see below↓
3387:Metasuchia
3372:Sauropsida
2571:References
2546:Simosuchus
2534:Simosuchus
2526:Antarctica
2498:neosuchian
2494:Simosuchus
2484:Simosuchus
2471:Simosuchus
2450:Simosuchus
2434:Simosuchus
2429:Simosuchus
2418:Simosuchus
2154:Yacarerani
1983:Notosuchus
1916:Simosuchus
1593:Lomasuchus
1460:Notosuchia
1427:Simosuchus
1411:Simosuchus
1395:Simosuchus
1383:Simosuchus
1379:Simosuchus
1360:Simosuchus
1356:Simosuchus
1351:Notosuchus
1321:Simosuchus
1263:Simosuchus
1046:Notosuchus
991:Simosuchus
940:Candidodon
873:Notosuchia
784:Simosuchus
660:Notosuchus
631:Notosuchia
605:Cladograms
588:Simosuchus
579:Simosuchus
575:osteoderms
571:Simosuchus
564:Simosuchus
552:Simosuchus
506:Simosuchus
489:Simosuchus
473:Simosuchus
457:Simosuchus
453:Simosuchus
426:Simosuchus
353:Madagascar
330:Simosuchus
279:Simosuchus
249:Notosuchia
25:Simosuchus
4220:Pabwehshi
3770:Itasuchus
3640:Ilchunaia
3589:Sebecidae
3536:Doratodon
3358:Kingdom:
3083:1342-937X
2648:129124296
2609:131192480
2522:Australia
2446:fossorial
1289:from the
556:vertebrae
539:. On the
525:ellipsoid
402:herbivore
394:carnivore
386:S. clarki
347:from the
311:S. clarki
157:Kingdom:
151:Eukaryota
4610:Wikidata
3366:Chordata
3364:Phylum:
3360:Animalia
3240:83717721
3162:31257139
3029:86737170
2987:86565764
2948:86257810
2913:85720441
2878:86113454
2824:84437972
2774:10879533
2694:84710186
2552:See also
2502:Gondwana
2422:iguanids
1319:(2000),
1297:. Later
284:Buckley
181:Reptilia
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
4656:4132224
4643:1186416
4630:4575536
4616:Q135263
3675:Sebecus
3515:Sebecia
3376:Order:
3370:Class:
3259:ZooKeys
3201:9166607
3063:Bibcode
2782:4426288
2754:Bibcode
2265:Sebecia
1409:. Like
1399:Chinese
1309:derived
1281:Uruguay
633:
545:ischium
521:condyle
514:humerus
510:scapula
408:History
335:extinct
298:Species
271:Genus:
177:Class:
105:↓
4669:114380
3238:
3199:
3160:
3081:
3027:
2985:
2946:
2911:
2876:
2822:
2780:
2772:
2746:Nature
2717:
2692:
2646:
2607:
2524:, and
2514:Africa
1401:genus
1375:et al.
1366:, and
1330:. Its
1325:family
1317:et al.
1295:Malawi
584:diploë
541:pelvis
529:radius
333:is an
286:et al.
4638:IRMNG
3383:Clade
3236:S2CID
3197:S2CID
3090:(PDF)
3051:(PDF)
3025:S2CID
2983:S2CID
2944:S2CID
2909:S2CID
2874:S2CID
2820:S2CID
2778:S2CID
2690:S2CID
2644:S2CID
2605:S2CID
2538:Egypt
2518:India
1271:clade
1267:basal
537:femur
465:Greek
379:India
338:genus
256:Clade
240:Clade
227:Clade
214:Clade
201:Clade
188:Clade
4625:GBIF
3416:see
3158:PMID
3079:ISSN
2770:PMID
2728:2022
2715:ISBN
2389:Note
1389:and
1348:and
1334:was
1283:and
533:ulna
531:and
432:and
317:type
290:2000
44:PreꞒ
3263:doi
3228:doi
3189:doi
3148:doi
3117:doi
3071:doi
3017:doi
2975:doi
2936:doi
2901:doi
2866:doi
2812:doi
2762:doi
2750:405
2682:doi
2636:doi
2597:doi
1293:of
461:pug
377:of
369:in
359:is
351:of
340:of
4688::
4666::
4653::
4640::
4627::
4612::
3385::
3257:.
3234:.
3224:18
3222:.
3195:.
3185:93
3183:.
3179:.
3156:.
3144:29
3142:.
3138:.
3113:37
3111:.
3077:.
3069:.
3057:.
3053:.
3023:.
3013:30
3011:.
2995:^
2981:.
2971:32
2969:.
2942:.
2932:25
2930:.
2907:.
2897:30
2895:.
2872:.
2862:30
2860:.
2842:^
2818:.
2808:30
2806:.
2790:^
2776:.
2768:.
2760:.
2748:.
2736:^
2688:.
2678:30
2676:.
2656:^
2642:.
2632:30
2630:.
2603:.
2593:33
2591:.
2587:.
2548:.
2528:.
2520:,
2516:,
2512:,
2444:A
1578:*
1445:.
1421:.
1413:,
1362:,
586:.
404:.
381:.
288:,
258::
242::
229::
216::
203::
190::
94:Pg
38:Ma
33:,
4387:?
4372:?
4330:?
4184:?
4169:?
4161:?
4083:?
4058:?
4043:?
4028:?
3731:?
3716:?
3546:?
3501:?
3334:e
3327:t
3320:v
3271:.
3265::
3242:.
3230::
3203:.
3191::
3164:.
3150::
3123:.
3119::
3073::
3065::
3059:7
3044:"
3031:.
3019::
2989:.
2977::
2950:.
2938::
2915:.
2903::
2880:.
2868::
2837:.
2826:.
2814::
2784:.
2764::
2756::
2730:.
2696:.
2684::
2650:.
2638::
2611:.
2599::
2397:.
2387:*
319:)
307:†
275:†
262:†
246:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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