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is one of several enigmatic early therapsids from Russia that may be based on juvenile material. Its phylogenetic affinities are controversial; it has been classified as a biarmosuchian, dinocephalian, or anomodont, and it has also been suggested to belong to a lineage of its own.
364:, but subsequently proposed a monotypic order, Niaftasuchida, for it. In 2000, Battail and Surkov argued it was a biarmosuchian. In 2001, Ivakhnenko suggested it was an anomodont, but in 2003 returned to regarding it as a dinocephalian on the basis of new material.
316:
has a distinctive dentition composed of large, procumbent, leaf-shaped teeth. There are three pairs of incisors, but the canines are not particularly distinct from the other maxillary teeth. There are a few palatal teeth. Like biarmosuchians,
410:
is interpreted as an herbivore, with its teeth adapted primarily for tearing off soft plant parts. Based on possible coprolites found in the mouth of a juvenile specimen, Ivakhnenko suggested that juvenile
360:). It was named in 1990 by M. F. Ivakhnenko, with the name referring to the place where it was found and honoring the geologist I. D. Zekkel. Ivakhnenko initially interpreted it as a
556:
Battail, Bernard; Surkov, Mikhail V. (2000). "Mammal-like reptiles from Russia". In Benton, Michael J.; Shishkin, Mikhail A.; Unwin, David M.; Kurochkin, Evgenii N. (eds.).
399:, but a specimen that may represent a second species is known. It differs from the type species in having a straight tooth row without enlarged maxillary teeth.
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are very small; the holotype skull is only 9 centimetres (3.5 in) long. It has been interpreted as being based on probable juvenile material.
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Ivakhnenko, M. F.; Golubev, V. K.; Gubin, Yu. M.; Kalandadze, N. N.; Novikov, I. V.; Sennikov, A. G.; Rautian, A. S. (1997).
266:
had distinctive dentition. It has been interpreted as one of the earliest known herbivorous therapsids. It lived during the
2476:
116:
2012:
2000:
376:
is considered problematic. It is assigned to a family of its own, Niaftasuchidae. Ivakhnenko, who first described
577:
Ivakhnenko, M. F. (1990). "Позднепалеозойский фаунистический комплекс тетрапод из отложени бассейна р. мезень".
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392:. It is also possible that it belongs to neither lineage, and forms a distinctive therapsid group of its own.
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2006:
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has exceptionally large eye sockets, though this may be a juvenile character. The skull is low and broad.
278:, and inhabited an environment alongside some of the last non-therapsid synapsids, such as the caseasaur
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Kammerer, Christian F. (2011). "Systematics of the
Anteosauria (Therapsida: Dinocephalia)".
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601:"Cranial morphology and evolution of Permian Dinomorpha (Eotherapsida) of eastern Europe"
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Ivakhnenko, M. F. (2003). "Eotherapsids from the East
European placket".
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708:. Oxford biology. Oxford ; New York: Oxford University Press.
340:, an incomplete skull without the mandible, was collected near the
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650:Permian and Triassic tetrapods of Eastern Europe
562:. Cambridge University Press. pp. 86–119.
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559:The Age of Dinosaurs in Russia and Mongolia
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415:would ingest coprolites to develop their
478:
384:, whereas Battail and Surkov classified
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254:. Its type and only named species is
7:
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514:
2533:Monotypic prehistoric animal genera
2523:Guadalupian genus first appearances
706:The origin and evolution of mammals
669:Journal of Systematic Palaeontology
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395:There is only one named species,
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2334:
2301:
2011:
2005:
1999:
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429:
419:, as in many modern herbivores.
115:
1:
2528:Guadalupian genus extinctions
2518:Fossil taxa described in 1990
2503:Prehistoric therapsid genera
2456:. You can help Knowledge by
681:10.1080/14772019.2010.492645
579:Paleontologicheskii Zhurnal
16:Extinct genus of therapsids
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599:Ivakhnenko, M. F. (2008).
2508:Extinct animals of Russia
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617:10.1134/S0031030108090013
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336:The holotype specimen of
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112:Scientific classification
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101:
23:
581:(in Russian) (4): 81–90.
380:, regards it as a basal
605:Paleontological Journal
588:Paleontological Journal
324:The known specimens of
2452:-related article is a
503:Ivakhnenko et al. 1997
372:The classification of
310:
2513:Guadalupian synapsids
2415:Paleobiology Database
308:
2365:Niaftasuchus zekkeli
704:Kemp, T. S. (2005).
397:Niaftasuchus zekkeli
338:Niaftasuchus zekkeli
257:Niaftasuchus zekkeli
228:Niaftasuchus zekkeli
437:Paleontology portal
445:List of therapsids
346:Arkhangelsk Oblast
311:
284:and the varanopid
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2402:Open Tree of Life
2357:Taxon identifiers
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2020:
2019:
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1827:Sphenacodontoidea
1800:Palaeohatteriidae
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1857:Sphenacodontidae
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1099:Lalieudorhynchus
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652:. Moscow: GEOS.
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332:History of study
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29:Temporal range:
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1923:Secodontosaurus
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810:Tetrapodomorpha
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541:, p. S371.
539:Ivakhnenko 2003
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529:, p. S372.
527:Ivakhnenko 2003
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521:
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493:, p. S359.
491:Ivakhnenko 2003
489:
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464:Ivakhnenko 1990
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362:tapinocephalian
352:, near Nyafta (
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274:in what is now
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1849:
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1829:
1823:
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1809:Palaeohatteria
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1802:
1793:
1792:
1790:
1789:
1779:
1769:
1759:
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1729:Sphenacodontia
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1393:Incertae sedis
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1220:
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1154:Eupelycosauria
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1123:
1116:
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1102:
1095:
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1081:
1074:
1067:
1064:Cotylorhynchus
1060:
1053:
1046:
1039:
1032:
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852:Reptiliomorpha
845:
839:
837:Reptiliomorpha
833:
832:
830:
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743:
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728:
721:
720:
714:
701:
675:(2): 261–304.
664:
658:
645:
611:(9): 859–995.
596:
594:(S4): 339–465.
583:
574:
568:
552:
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519:
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483:
481:, p. 276.
468:
455:
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417:gut microbiota
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368:Classification
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246:is an extinct
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60:
55:
50:
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40:
35:
34:
32:Guadalupian
28:
15:
13:
10:
9:
6:
4:
3:
2:
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2534:
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2326:
2325:Phthinosuchia
2323:
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2317:
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2286:
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2206:
2205:Therocephalia
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2194:Therocephalia
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2134:
2133:
2132:
2130:
2128:
2121:
2113:
2109:
2108:
2107:
2106:
2104:
2102:
2095:
2087:
2083:
2082:
2081:
2080:
2078:
2076:
2069:
2063:
2062:
2055:
2054:
2051:
2048:
2045:
2040:
2036:
2032:
2027:
2023:
2014:
2010:
2008:
2002:
1973:
1972:
1971:
1970:
1968:
1966:
1962:
1955:
1954:
1953:Tetraceratops
1950:
1947:
1946:
1945:Tappenosaurus
1942:
1939:
1938:
1934:
1932:
1931:
1927:
1925:
1924:
1920:
1918:
1917:
1913:
1911:
1910:
1906:
1904:
1903:
1899:
1897:
1896:
1892:
1889:
1888:
1884:
1882:
1881:
1877:
1875:
1874:
1870:
1868:
1867:
1866:Cryptovenator
1863:
1862:
1860:
1858:
1851:
1845:
1844:
1837:
1836:
1833:
1830:
1828:
1824:
1818:
1817:
1816:Pantelosaurus
1813:
1811:
1810:
1806:
1805:
1803:
1801:
1794:
1788:
1787:
1786:Tetraceratops
1780:
1778:
1777:
1776:Kenomagnathus
1770:
1768:
1767:
1760:
1758:
1757:
1750:
1748:
1747:
1740:
1739:
1736:
1733:
1730:
1725:
1715:
1714:
1713:Xyrospondylus
1710:
1708:
1707:
1703:
1701:
1700:
1696:
1694:
1693:
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1686:
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1610:
1607:
1603:
1600:
1597:
1592:
1582:
1581:
1577:
1575:
1574:
1573:Stereorhachis
1570:
1568:
1567:
1563:
1561:
1560:
1556:
1554:
1553:
1549:
1547:
1546:
1542:
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1535:
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1531:Archaeothyris
1528:
1527:
1525:
1523:
1516:
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1508:
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1499:
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1351:
1350:
1346:
1344:
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1339:
1337:
1336:
1332:
1330:
1329:
1328:Tambacarnifex
1325:
1323:
1322:
1318:
1315:
1314:
1313:Mycterosaurus
1310:
1308:
1307:
1303:
1302:
1300:
1298:
1294:
1288:
1287:
1286:Microvaranops
1283:
1281:
1280:
1276:
1274:
1273:
1269:
1267:
1266:
1265:Elliotsmithia
1262:
1260:
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1255:
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1199:
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1177:
1174:
1171:
1168:
1161:
1158:
1155:
1150:
1136:
1135:
1131:
1129:
1128:
1127:Ruthenosaurus
1124:
1122:
1121:
1120:Phreatophasma
1117:
1115:
1114:
1110:
1108:
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1103:
1101:
1100:
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1038:
1037:
1036:Callibrachion
1033:
1031:
1030:
1026:
1024:
1023:
1019:
1017:
1016:
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1009:
1007:
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659:5-89118-029-4
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569:9780521545822
565:
561:
560:
554:
553:
548:
540:
535:
532:
528:
523:
520:
517:, p. 31.
516:
511:
508:
505:, p. 30.
504:
499:
496:
492:
487:
484:
480:
479:Kammerer 2011
475:
473:
469:
466:, p. 88.
465:
460:
457:
450:
446:
443:
442:
438:
427:
422:
420:
418:
414:
409:
402:
400:
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393:
391:
387:
383:
382:dinocephalian
379:
375:
367:
365:
363:
355:
351:
347:
343:
339:
331:
329:
327:
322:
320:
315:
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291:
289:
288:
283:
282:
277:
273:
270:epoch of the
269:
265:
261:
259:
258:
253:
249:
245:
244:
231:
229:
220:
217:
216:Binomial name
213:
209:
208:
202:
199:
198:
191:
190:
183:
180:
179:
176:
173:
170:
167:
166:
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160:
157:
154:
153:
150:
147:
144:
143:
140:
137:
134:
133:
130:
127:
124:
123:
118:
113:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
2458:expanding it
2447:
2432:
2364:
2330:Theriodontia
2320:Pelycosauria
2284:Niaftasuchus
2283:
2282:
2272:
2262:
2249:
2112:Dinocephalia
2101:Dinocephalia
2086:Biarmosuchia
2075:Biarmosuchia
2059:
2004:
1951:
1943:
1937:Steppesaurus
1935:
1928:
1921:
1914:
1907:
1900:
1893:
1885:
1878:
1873:Ctenorhachis
1871:
1864:
1841:
1814:
1807:
1784:
1774:
1764:
1754:
1744:
1711:
1704:
1697:
1690:
1683:
1678:Ianthasaurus
1676:
1669:
1664:Glaucosaurus
1662:
1657:Edaphosaurus
1655:
1648:
1625:
1615:
1580:Varanosaurus
1578:
1571:
1564:
1557:
1552:Echinerpeton
1550:
1543:
1536:
1529:
1454:
1444:
1434:
1424:
1414:
1404:
1391:
1347:
1340:
1333:
1326:
1319:
1311:
1304:
1284:
1279:Mesenosaurus
1277:
1270:
1263:
1256:
1249:
1229:
1222:
1214:
1207:
1200:
1195:Ascendonanus
1193:
1186:
1179:
1134:Trichasaurus
1132:
1125:
1118:
1111:
1104:
1097:
1090:
1083:
1078:Ennatosaurus
1076:
1071:Datheosaurus
1069:
1062:
1055:
1048:
1041:
1034:
1029:Arisierpeton
1027:
1022:Angelosaurus
1020:
1015:Alierasaurus
1013:
993:
986:
979:
971:
964:Eothyrididae
937:
789:
783:Superclass:
705:
672:
668:
649:
608:
604:
591:
587:
578:
558:
549:Bibliography
534:
522:
510:
498:
486:
459:
413:Niaftasuchus
412:
408:Niaftasuchus
407:
406:
403:Paleobiology
396:
394:
390:Biarmosuchia
386:Niaftasuchus
385:
378:Niaftasuchus
377:
374:Niaftasuchus
373:
371:
337:
335:
326:Niaftasuchus
325:
323:
319:Niaftasuchus
318:
314:Niaftasuchus
313:
312:
294:Niaftasuchus
293:
292:
287:Mesenosaurus
285:
281:Ennatosaurus
279:
264:Niaftasuchus
263:
262:
256:
255:
243:Niaftasuchus
242:
241:
240:
227:
222:
206:
205:
189:Niaftasuchus
188:
187:
168:
155:
25:Niaftasuchus
24:
18:
2170:Gorgonopsia
2159:Gorgonopsia
2144:dicynodonts
2138:Anomodontia
2127:Anomodontia
1930:Sphenacodon
1909:Macromerion
1699:Ramodendron
1685:Lupeosaurus
1545:Clepsydrops
1538:Baldwinonus
1503:Metopophora
1446:Mastersonia
1436:Knoxosaurus
1364:Metopophora
1272:Heleosaurus
1167:Varanopidae
995:Vaughnictis
973:Asaphestera
954:Caseasauria
309:Restoration
301:Description
268:Guadalupian
2497:Categories
2318:See also:
2274:Kamagorgon
2264:Dinosaurus
2228:Cynodontia
2217:Cynodontia
2044:Therapsida
2031:Therapsida
1974:see below↓
1965:Therapsida
1895:Dimetrodon
1843:Shashajaia
1627:Milosaurus
1559:Ophiacodon
1406:Dimacrodon
1373:see below↓
1321:Ruthiromia
1306:Aerosaurus
1216:Nitosaurus
1209:Dendromaia
1181:Apsisaurus
1106:Martensius
1092:Euromycter
873:see below↓
451:References
252:therapsids
175:Therapsida
2450:therapsid
2232:includes
2142:includes
2061:Raranimus
1916:Neosaurus
1617:Ianthodon
1342:Varanodon
1202:Eoscansor
1113:Oromycter
1057:Caseopsis
1050:Caseoides
988:Oedaleops
918:Synapsida
905:Synapsida
864:Synapsida
785:Tetrapoda
771:Kingdom:
755:Synapsida
689:1477-2019
633:0031-0301
625:1555-6174
515:Kemp 2005
344:river in
200:Species:
162:Synapsida
135:Kingdom:
129:Eukaryota
2380:Q4162565
2374:Wikidata
2340:Category
1902:Eosyodon
1887:Cutleria
1756:Haptodus
1746:Cutleria
1671:Gordodon
1426:Gorgodon
1416:Driveria
1349:Watongia
1335:Varanops
1258:Cabarzia
1251:Anningia
1006:Caseidae
981:Eothyris
779:Chordata
777:Phylum:
773:Animalia
697:84799772
641:85114195
423:See also
149:Chordata
145:Phylum:
139:Animalia
125:Domain:
2407:6149440
2394:8680465
2234:mammals
1085:Eocasea
794:Amniota
354:Russian
272:Permian
225:†
204:†
181:Genus:
2420:335883
1224:Pyozia
712:
695:
687:
656:
639:
631:
623:
566:
350:Russia
276:Russia
2448:This
1043:Casea
790:Clade
693:S2CID
637:S2CID
621:eISSN
358:Няфта
342:Pyoza
248:genus
169:Clade
156:Clade
2454:stub
2389:GBIF
2226:see
2203:see
2168:see
2136:see
2110:see
2084:see
850:see
823:see
710:ISBN
685:ISSN
654:ISBN
629:ISSN
564:ISBN
37:PreꞒ
677:doi
613:doi
388:in
250:of
2499::
2417::
2404::
2391::
2376::
792::
691:.
683:.
671:.
635:.
627:.
619:.
609:42
607:.
603:.
592:37
590:.
471:^
356::
348:,
290:.
260:.
171::
158::
87:Pg
2485:e
2478:t
2471:v
2460:.
2280:†
2270:†
2260:†
2191:†
2156:†
2124:†
2098:†
2072:†
2057:†
1956:?
1948:?
1940:?
1890:?
1854:†
1839:†
1797:†
1782:†
1772:†
1762:†
1752:†
1742:†
1638:†
1623:†
1613:†
1519:†
1459:?
1452:†
1442:†
1432:†
1422:†
1412:†
1402:†
1316:?
1219:?
1169:?
1164:†
976:?
956:?
951:†
942:?
935:†
747:e
740:t
733:v
718:.
699:.
679::
673:9
662:.
643:.
615::
572:.
185:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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