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Avisauridae

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2241: 2247: 147: 2265: 588:, would qualify as members of the family. Nevertheless, enantiornithean taxonomy is notably difficult to resolve, and some analyses on enantiornitheans have not resolved the family. However, this may be due to such analyses focusing on early Cretaceous enantiornitheans (which are numerous and well-preserved) rather than fragmentary late Cretaceous taxa, such as most avisaurids. Another member of Avisauridae is, 124: 974: 468:
edge of this bone's trochlea (toe joint) has a bony tab which points downward, known as a plantar projection. The innermost bone of the tarsometatarsus, metatarsal I, is small, laterally compressed (flattened from side-to-side), and J-shaped from the side. It is connected to a reversed hallux sporting a very large and curved claw.
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Unlike in some prehistoric birds, avisaurid tarsometatarsals were not completely fused, with the distal (outer) parts of the metatarsals being separate from each other. The proximal (near) half of metatarsal III (the long bone in the middle of the tarsometatarsus) is convex from the front. The inside
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period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines (following Cau & Arduini, 2008), or a small family within that group, restricted to species from the Late Cretaceous of
521:, as well as various Cretaceous bird groups. Avisaurids also had a thin metatarsal IV (the outermost long bone of the tarsometatarsus) and a bony knob on the front of metatarsal II (the most innermost long bone of the tarsometatarsus) for the insertion of 494:(the only avisaurid known at that time), only the proximal parts of the metatarsals were fused, the proximal part of metatarsal III was wide, and the hypotarsus (a wide ridge extending down the back of the tarsometatarsus) was poorly developed. 487:
in 1985. At that time the family consisted of a few fossils that they believed belonged to small non-avian dinosaurs. They doubted that these fossils belonged to birds due to the presence of several features of the tarsometatarsus. In
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Avisaurids were among the largest and last enantiornithines to have lived, although they are also among the most poorly preserved. The majority of them are known primarily from fossilized
460:(the bones which comprise the foot in humans). As a result, members of this family are distinguished from other enantiornithines exclusively by features of the tarsometatarsal and pedal 953: 886:"Enantiophoenix electrophyla gen. et sp. nov. (Aves, Enantiornithes) from the Upper Cretaceous (Cenomanian) of Lebanon and its phylogenetic relationships" 1071:
Wang, Hu; Li, Zhiheng (2015). "A new small enantiornithine bird from the Jehol Biota, with implications for early evolution of avian skull morphology".
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with a large claw — that indicated the ability to perch in trees. They argued that an arboreal habit was most likely for all of the Avisauridae.
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to this group, although the erection of that name has been criticized by Cau. Under this broader definition, several other enantiornitheans, such as
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Chiappe and Calvo (1994) found that the Avisauridae shared adaptations of the foot — including a fully reversed and distally placed
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plus all its descendants. In 2008, the family was given a broader definition courtesy of Cau and Arduini. They redefined the group as
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Brett-Surman, Michael K.; Paul, Gregory S. (1985). "A new family of bird-like dinosaurs linking Laurasia and Gondwanaland".
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L.M. Chiappe. 1993. "Enantiornithine (Aves) tarsometatarsi from the Cretaceous Lecho Formation of northwestern Argentina".
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in 1992. He noted that the features used to exclude avisaurids from birds are in fact present in some early birds such as
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Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina
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Atti della Societa Italiana di Scienze Naturali e del Museo Ivico di Storia Naturale in Milano
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Gareth J. Dyke; Attila Ősi (2010). "A review of Late Cretaceous fossil birds from Hungary".
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Fernando Emilio Novas, Federico Lisandro Agnolín and Carlos Agustín Scanferla (2010).
2406: 2221: 1890: 1849: 1822: 1644: 1482: 1475: 1426: 885: 761: 593: 517: 378: 81: 978:, a New Late Cretaceous Bird (Enantiornithes: Avisauridae) from Patagonia, Argentina 2214: 2029: 1608: 1580: 1573: 1517: 1447: 1391: 1007: 720: 540: 461: 371: 1084: 2317: 2127: 2043: 1974: 1953: 1904: 1876: 1856: 1759: 1708: 1651: 1545: 1538: 1510: 1496: 1489: 1461: 1440: 1398: 1252: 562: 313: 56: 2308: 924: 2157: 2150: 2077: 2008: 2001: 1801: 1666: 1637: 1594: 1503: 1468: 1433: 1164: 1155: 672: 568: 440: 430: 364: 343: 204: 101: 66: 2200: 2186: 2135: 2092: 2057: 1967: 1945: 1924: 1897: 1808: 1751: 1701: 1694: 1680: 1615: 1587: 1552: 1531: 1405: 1363: 1349: 1225: 1173: 744: 696: 600: 552: 546: 490: 436: 350: 321: 306: 217: 158: 123: 106: 50: 1048:"The Theropod Database Blog: Martyniuk's (2012) new definitions and clades" 2302: 2084: 2050: 1794: 1773: 1729: 1630: 1601: 1566: 1454: 1149: 621: 589: 574: 191: 178: 96: 91: 76: 71: 61: 17: 2350: 2207: 2193: 1829: 1780: 1370: 1015: 933: 505: 357: 336: 230: 133: 111: 86: 1143: 557: 472: 168: 2279: 908: 857: 296: 1094: 2337: 501: 483:
Avisauridae was erected as a family by Michael Brett-Surman and
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Chiappe in 1993 defined the family as the common ancestor of
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However, Chiappe later reassigned the Avisauridae to the
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Matt Martyniuk gave the name 1008:10.1080/02724634.1985.10011851 1: 2423:Prehistoric dinosaur families 1085:10.1080/14772019.2015.1073801 131:Skeletal reconstruction of 27:Extinct family of dinosaurs 2439: 1052:The Theropod Database Blog 925:10.1016/j.crpv.2010.09.005 479:History and classification 2259: 2237: 2122: 1344: 1329: 1310: 1260: 1233: 1206: 1193: 1132: 1030:American Museum Novitates 952:Chiappe, Luis M. (1992) " 758: 741: 734: 717: 710: 693: 686: 669: 662: 645: 638: 618: 611: 508:in modern usage) and the 394: 387: 302: 295: 142:Scientific classification 140: 130: 121: 34: 2114:Cau & Arduini, 2008) 913:Comptes Rendus Palevol 2385:Paleobiology Database 2372:Paleobiology Database 524:M. tibialis cranialis 553:Avisaurus archibaldi 547:Avisaurus archibaldi 533:M. tibialis anterior 1737:Piscivorenantiornis 976:Neuquenornis volans 599:The following is a 541:Neuquenornis volans 529:M. tibialis anticus 1841:Iberomesornithidae 846:Geological Journal 2400: 2399: 2359:Open Tree of Life 2286:Taxon identifiers 2277: 2276: 2255: 2254: 2235: 2234: 2231: 2230: 1420:Cuspirostrisornis 1325: 1324: 1321: 1320: 1306: 1305: 1302: 1301: 1298: 1297: 960:September, 1992. 827: 826: 818: 817: 809: 808: 800: 799: 791: 790: 782: 781: 773: 772: 420: 419: 414: 402: 291: 16:(Redirected from 2430: 2418:Cretaceous birds 2393: 2392: 2380: 2379: 2367: 2366: 2354: 2353: 2341: 2340: 2328: 2327: 2326: 2313: 2312: 2311: 2281: 2267: 2266: 2250: 2249: 2243: 2179: 2178: 2120: 2116: 2115: 1674:Microenantiornis 1385:Castignovolucris 1342: 1331: 1258: 1231: 1204: 1195: 1188: 1187: 1134: 1119: 1112: 1105: 1096: 1089: 1088: 1068: 1062: 1061: 1059: 1058: 1043: 1037: 1026: 1020: 1019: 991: 985: 971: 965: 950: 939: 938: 936: 904: 898: 897: 881: 862: 861: 841: 737: 713: 689: 665: 641: 614: 607: 408: 400: 279: 274: 263:Euenantiornithes 261: 245: 150: 149: 126: 116: 53: 38:Temporal range: 32: 21: 2438: 2437: 2433: 2432: 2431: 2429: 2428: 2427: 2403: 2402: 2401: 2396: 2388: 2383: 2375: 2370: 2362: 2357: 2349: 2344: 2336: 2331: 2322: 2321: 2316: 2307: 2306: 2301: 2288: 2278: 2273: 2251: 2244: 2227: 2172: 2171: 2163: 2109: 2108: 2107: 2099: 2063: 1993:Bohaiornithidae 1987: 1939:Dapingfangornis 1932:Camptodontornis 1916:Longipterygidae 1910: 1862: 1835: 1723:Paraprotopteryx 1688:Monoenantiornis 1624:Largirostrornis 1378:Brevirostruavis 1338: 1317: 1294: 1274: 1247: 1220: 1214:Avemetatarsalia 1199:Avemetatarsalia 1189: 1183:Ornithothoraces 1139: 1138: 1128: 1123: 1093: 1092: 1070: 1069: 1065: 1056: 1054: 1045: 1044: 1040: 1027: 1023: 993: 992: 988: 972: 968: 951: 942: 906: 905: 901: 883: 882: 865: 858:10.1002/gj.1209 843: 842: 838: 833: 828: 819: 810: 801: 792: 783: 774: 485:Gregory S. Paul 481: 454:tarsometatarsal 450: 434:enantiornithine 406:Mystiornithidae 401:Kurochkin, 1996 398:Concornithidae? 278: 272: 259: 243: 144: 117: 115: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 48: 47: 44:Late Cretaceous 36: 28: 23: 22: 15: 12: 11: 5: 2436: 2434: 2426: 2425: 2420: 2415: 2405: 2404: 2398: 2397: 2395: 2394: 2381: 2368: 2355: 2342: 2329: 2314: 2298: 2296: 2290: 2289: 2284: 2275: 2274: 2272: 2271: 2260: 2257: 2256: 2253: 2252: 2238: 2236: 2233: 2232: 2229: 2228: 2226: 2225: 2218: 2211: 2204: 2197: 2190: 2182: 2180: 2177:Chiappe, 1992) 2165: 2164: 2162: 2161: 2154: 2147: 2144:Enantiophoenix 2140: 2132: 2123: 2117: 2101: 2100: 2098: 2097: 2089: 2081: 2073: 2071: 2069:Gobipterygidae 2065: 2064: 2062: 2061: 2054: 2047: 2040: 2037:Parabohaiornis 2033: 2026: 2019: 2012: 2005: 1997: 1995: 1989: 1988: 1986: 1985: 1982:Shengjingornis 1978: 1971: 1964: 1961:Longirostravis 1957: 1950: 1942: 1935: 1928: 1920: 1918: 1912: 1911: 1909: 1908: 1901: 1894: 1887: 1880: 1872: 1870: 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1126:Enantiornithes 1124: 1122: 1121: 1114: 1107: 1099: 1091: 1090: 1063: 1038: 1021: 1002:(2): 133–138. 986: 966: 940: 919:(8): 499–503. 899: 863: 852:(4): 434–444. 835: 834: 832: 829: 825: 824: 821: 820: 816: 815: 812: 811: 807: 806: 803: 802: 798: 797: 794: 793: 789: 788: 785: 784: 780: 779: 776: 775: 771: 770: 767: 766: 757: 754: 753: 750: 749: 740: 735: 733: 730: 729: 726: 725: 716: 711: 709: 706: 705: 702: 701: 692: 687: 685: 682: 681: 678: 677: 668: 663: 661: 658: 657: 654: 653: 649:Enantiophoenix 644: 639: 637: 631: 630: 627: 626: 617: 612: 610: 605: 585:Enantiophoenix 513:Enantiornithes 480: 477: 449: 446: 418: 417: 416: 415: 403: 392: 391: 385: 384: 383: 382: 375: 368: 361: 354: 347: 340: 333: 330:Enantiophoenix 326: 318: 310: 300: 299: 293: 292: 270: 266: 265: 257: 250: 249: 247:Enantiornithes 241: 234: 233: 228: 221: 220: 215: 208: 207: 202: 195: 194: 189: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 138: 137: 128: 127: 119: 118: 110: 105: 100: 95: 90: 85: 80: 75: 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464:(toe bones). 463: 459: 455: 447: 445: 442: 438: 435: 432: 428: 424: 412: 407: 404: 399: 396: 395: 393: 390: 386: 381: 380: 379:Soroavisaurus 376: 374: 373: 369: 367: 366: 362: 360: 359: 355: 353: 352: 348: 346: 345: 341: 339: 338: 334: 332: 331: 327: 324: 323: 319: 316: 315: 311: 309: 308: 304: 303: 301: 298: 294: 290: 286: 282: 277: 271: 268: 267: 264: 258: 255: 252: 251: 248: 242: 239: 236: 235: 232: 229: 226: 223: 222: 219: 216: 213: 210: 209: 206: 203: 200: 197: 196: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 148: 143: 139: 136: 135: 129: 125: 120: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 52: 45: 41: 33: 30: 19: 2293: 2220: 2215:Neuquenornis 2213: 2206: 2199: 2192: 2185: 2174: 2168: 2156: 2149: 2142: 2134: 2126: 2111: 2104: 2091: 2083: 2076: 2056: 2049: 2042: 2035: 2030:Longusunguis 2028: 2021: 2014: 2007: 2000: 1980: 1973: 1966: 1959: 1952: 1944: 1937: 1930: 1923: 1903: 1896: 1889: 1882: 1875: 1855: 1848: 1828: 1821: 1814: 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Retrieved 1051: 1041: 1033: 1029: 1024: 999: 995: 989: 981: 975: 969: 964:(3): 344-350 961: 957: 916: 912: 902: 893: 889: 849: 845: 839: 760: 759: 743: 742: 721:Neuquenornis 719: 718: 695: 694: 671: 670: 647: 646: 634: 620: 619: 598: 583: 580:Avisauroidea 579: 573: 567: 561: 551: 545: 539: 537: 532: 528: 522: 516: 496: 489: 482: 470: 466: 451: 422: 421: 410: 405: 397: 377: 372:Neuquenornis 370: 363: 356: 349: 342: 335: 328: 320: 312: 305: 281:Brett-Surman 275: 253: 237: 224: 211: 198: 185: 132: 49:125–66  29: 2324:Avisauridae 2318:Wikispecies 2294:Avisauridae 2169:Avisauridae 2128:Bauxitornis 2105:Avisauridae 2044:Shenqiornis 2016:Gretcheniao 1975:Shanweiniao 1954:Longipteryx 1905:Yuanchuavis 1884:Eopengornis 1877:Chiappeavis 1857:Noguerornis 1767:Pterygornis 1760:Protopteryx 1709:Orienantius 1652:Liaoxiornis 1546:Gracilornis 1539:Grabauornis 1511:Flexomornis 1497:Falcatakely 1490:Explorornis 1462:Enantiornis 1441:Elbretornis 1399:Cathayornis 1268:Maniraptora 1253:Maniraptora 1079:(6): 1–17. 934:11336/60953 635:Avisauridae 563:Longipteryx 458:metatarsals 448:Description 423:Avisauridae 314:Bauxitornis 276:Avisauridae 35:Avisauridae 2413:Avisaurids 2407:Categories 2158:Mystiornis 2151:Halimornis 2078:Gobipteryx 2023:Linyiornis 2009:Bohaiornis 2002:Beiguornis 1802:Xiangornis 1744:Platanavis 1667:Martinavis 1638:Lenesornis 1595:Incolornis 1504:Feitianius 1469:Eoalulavis 1434:Dunhuangia 1289:see below↓ 1165:Dinosauria 1156:Sauropsida 1057:2018-02-03 831:References 673:Halimornis 569:Gobipteryx 441:Cretaceous 409:Kurochkin 365:Mystiornis 344:Halimornis 205:Saurischia 192:Dinosauria 2201:Intiornis 2187:Avisaurus 2136:Concornis 2093:Vescornis 2058:Zhouornis 1968:Rapaxavis 1946:Evgenavis 1925:Boluochia 1898:Pengornis 1809:Yatenavis 1788:Shangyang 1752:Praeornis 1716:Otogornis 1702:Nanantius 1695:Musivavis 1681:Mirusavis 1616:Kuszholia 1588:Imparavis 1553:Gurilynia 1532:Gobipipus 1406:Cratoavis 1364:Alexornis 1350:Abavornis 1241:Theropoda 1226:Theropoda 1174:Theropoda 1142:Kingdom: 745:Avisaurus 697:Concornis 601:cladogram 491:Avisaurus 462:phalanges 439:from the 437:dinosaurs 351:Intiornis 322:Concornis 307:Avisaurus 218:Theropoda 165:Kingdom: 159:Eukaryota 18:Avisaurid 2309:Q4703066 2303:Wikidata 2269:Category 2085:Jibeinia 2051:Sulcavis 1795:Sinornis 1774:Qiliania 1730:Parvavis 1631:Lectavis 1602:Junornis 1567:Houornis 1560:Holbotia 1455:Elsornis 1150:Chordata 1148:Phylum: 1144:Animalia 622:Sinornis 590:Elsornis 575:Sinornis 556:and all 510:subclass 389:Synonyms 269:Family: 179:Chordata 175:Phylum: 169:Animalia 155:Domain: 2364:3605542 2351:4844232 2338:4434643 2208:Mirarce 2194:Gettyia 1830:Yuornis 1781:Sazavis 1371:Avimaia 1154:Class: 1036:, 39 pp 1016:4523037 506:Avialae 431:extinct 358:Mirarce 337:Gettyia 231:Avialae 134:Mirarce 2390:203286 1014:  558:genera 473:hallux 427:family 413:, 2011 411:et al. 297:Genera 2377:53285 2175:sensu 2112:sensu 1179:Clade 1170:Clade 1161:Clade 1012:JSTOR 499:class 425:is a 254:Clade 238:Clade 225:Clade 212:Clade 199:Clade 186:Clade 40:Early 2346:GBIF 1266:see 1239:see 1212:see 1034:3083 544:and 502:Aves 289:1985 285:Paul 283:and 57:PreꞒ 2333:EoL 1081:doi 1004:doi 980:". 929:hdl 921:doi 894:149 854:doi 572:or 531:or 429:of 2409:: 2387:: 2374:: 2361:: 2348:: 2335:: 2320:: 2305:: 1181:: 1172:: 1163:: 1077:14 1075:. 1050:. 1032:. 1010:. 998:. 962:12 956:" 943:^ 927:. 915:. 911:. 892:. 888:. 866:^ 850:45 848:. 596:. 566:, 287:, 256:: 240:: 227:: 214:: 201:: 188:: 107:Pg 51:Ma 46:, 2173:( 2139:? 2131:? 2110:( 2096:? 2088:? 1949:? 1755:? 1747:? 1655:? 1619:? 1118:e 1111:t 1104:v 1087:. 1083:: 1060:. 1018:. 1006:: 1000:5 937:. 931:: 923:: 917:9 860:. 856:: 325:? 317:? 273:† 260:† 244:† 112:N 102:K 97:J 92:T 87:P 82:C 77:D 72:S 67:O 62:Ꞓ 42:- 20:)

Index

Avisaurid
Early
Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Mirarce
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Avialae
Enantiornithes
Euenantiornithes
Avisauridae

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