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Avisauridae

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2230: 2236: 136: 2254: 577:, would qualify as members of the family. Nevertheless, enantiornithean taxonomy is notably difficult to resolve, and some analyses on enantiornitheans have not resolved the family. However, this may be due to such analyses focusing on early Cretaceous enantiornitheans (which are numerous and well-preserved) rather than fragmentary late Cretaceous taxa, such as most avisaurids. Another member of Avisauridae is, 113: 963: 457:
edge of this bone's trochlea (toe joint) has a bony tab which points downward, known as a plantar projection. The innermost bone of the tarsometatarsus, metatarsal I, is small, laterally compressed (flattened from side-to-side), and J-shaped from the side. It is connected to a reversed hallux sporting a very large and curved claw.
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Unlike in some prehistoric birds, avisaurid tarsometatarsals were not completely fused, with the distal (outer) parts of the metatarsals being separate from each other. The proximal (near) half of metatarsal III (the long bone in the middle of the tarsometatarsus) is convex from the front. The inside
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period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines (following Cau & Arduini, 2008), or a small family within that group, restricted to species from the Late Cretaceous of
510:, as well as various Cretaceous bird groups. Avisaurids also had a thin metatarsal IV (the outermost long bone of the tarsometatarsus) and a bony knob on the front of metatarsal II (the most innermost long bone of the tarsometatarsus) for the insertion of 483:(the only avisaurid known at that time), only the proximal parts of the metatarsals were fused, the proximal part of metatarsal III was wide, and the hypotarsus (a wide ridge extending down the back of the tarsometatarsus) was poorly developed. 476:
in 1985. At that time the family consisted of a few fossils that they believed belonged to small non-avian dinosaurs. They doubted that these fossils belonged to birds due to the presence of several features of the tarsometatarsus. In
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Avisaurids were among the largest and last enantiornithines to have lived, although they are also among the most poorly preserved. The majority of them are known primarily from fossilized
449:(the bones which comprise the foot in humans). As a result, members of this family are distinguished from other enantiornithines exclusively by features of the tarsometatarsal and pedal 942: 875:"Enantiophoenix electrophyla gen. et sp. nov. (Aves, Enantiornithes) from the Upper Cretaceous (Cenomanian) of Lebanon and its phylogenetic relationships" 1060:
Wang, Hu; Li, Zhiheng (2015). "A new small enantiornithine bird from the Jehol Biota, with implications for early evolution of avian skull morphology".
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with a large claw — that indicated the ability to perch in trees. They argued that an arboreal habit was most likely for all of the Avisauridae.
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to this group, although the erection of that name has been criticized by Cau. Under this broader definition, several other enantiornitheans, such as
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Chiappe and Calvo (1994) found that the Avisauridae shared adaptations of the foot — including a fully reversed and distally placed
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plus all its descendants. In 2008, the family was given a broader definition courtesy of Cau and Arduini. They redefined the group as
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Brett-Surman, Michael K.; Paul, Gregory S. (1985). "A new family of bird-like dinosaurs linking Laurasia and Gondwanaland".
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L.M. Chiappe. 1993. "Enantiornithine (Aves) tarsometatarsi from the Cretaceous Lecho Formation of northwestern Argentina".
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in 1992. He noted that the features used to exclude avisaurids from birds are in fact present in some early birds such as
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Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina
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Atti della Societa Italiana di Scienze Naturali e del Museo Ivico di Storia Naturale in Milano
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Gareth J. Dyke; Attila Ősi (2010). "A review of Late Cretaceous fossil birds from Hungary".
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Fernando Emilio Novas, Federico Lisandro Agnolín and Carlos Agustín Scanferla (2010).
2395: 2210: 1879: 1838: 1811: 1633: 1471: 1464: 1415: 874: 750: 582: 506: 367: 70: 967:, a New Late Cretaceous Bird (Enantiornithes: Avisauridae) from Patagonia, Argentina 2203: 2018: 1597: 1569: 1562: 1506: 1436: 1380: 996: 709: 529: 450: 360: 1073: 2306: 2116: 2032: 1963: 1942: 1893: 1865: 1845: 1748: 1697: 1640: 1534: 1527: 1499: 1485: 1478: 1450: 1429: 1387: 1241: 551: 302: 45: 2297: 913: 2146: 2139: 2066: 1997: 1990: 1790: 1655: 1626: 1583: 1492: 1457: 1422: 1153: 1144: 661: 557: 429: 419: 353: 332: 193: 90: 55: 2189: 2175: 2124: 2081: 2046: 1956: 1934: 1913: 1886: 1797: 1740: 1690: 1683: 1669: 1604: 1576: 1541: 1520: 1394: 1352: 1338: 1214: 1162: 733: 685: 589: 541: 535: 479: 425: 339: 310: 295: 206: 147: 112: 95: 39: 1037:"The Theropod Database Blog: Martyniuk's (2012) new definitions and clades" 2291: 2073: 2039: 1783: 1762: 1718: 1619: 1590: 1555: 1443: 1138: 610: 578: 563: 180: 167: 85: 80: 65: 60: 50: 2339: 2196: 2182: 1818: 1769: 1359: 1004: 922: 494: 346: 325: 219: 122: 100: 75: 1132: 546: 461: 157: 2268: 897: 846: 285: 1083: 2326: 490: 472:
Avisauridae was erected as a family by Michael Brett-Surman and
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Chiappe in 1993 defined the family as the common ancestor of
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However, Chiappe later reassigned the Avisauridae to the
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Matt Martyniuk gave the name 997:10.1080/02724634.1985.10011851 1: 2412:Prehistoric dinosaur families 1074:10.1080/14772019.2015.1073801 120:Skeletal reconstruction of 16:Extinct family of dinosaurs 2428: 1041:The Theropod Database Blog 914:10.1016/j.crpv.2010.09.005 468:History and classification 2248: 2226: 2111: 1333: 1318: 1299: 1249: 1222: 1195: 1182: 1121: 1019:American Museum Novitates 941:Chiappe, Luis M. (1992) " 747: 730: 723: 706: 699: 682: 675: 658: 651: 634: 627: 607: 600: 497:in modern usage) and the 383: 376: 291: 284: 131:Scientific classification 129: 119: 110: 23: 2103:Cau & Arduini, 2008) 902:Comptes Rendus Palevol 2374:Paleobiology Database 2361:Paleobiology Database 513:M. tibialis cranialis 542:Avisaurus archibaldi 536:Avisaurus archibaldi 522:M. tibialis anterior 1726:Piscivorenantiornis 965:Neuquenornis volans 588:The following is a 530:Neuquenornis volans 518:M. tibialis anticus 1830:Iberomesornithidae 835:Geological Journal 2389: 2388: 2348:Open Tree of Life 2275:Taxon identifiers 2266: 2265: 2244: 2243: 2224: 2223: 2220: 2219: 1409:Cuspirostrisornis 1314: 1313: 1310: 1309: 1295: 1294: 1291: 1290: 1287: 1286: 949:September, 1992. 816: 815: 807: 806: 798: 797: 789: 788: 780: 779: 771: 770: 762: 761: 409: 408: 403: 391: 280: 2419: 2407:Cretaceous birds 2382: 2381: 2369: 2368: 2356: 2355: 2343: 2342: 2330: 2329: 2317: 2316: 2315: 2302: 2301: 2300: 2270: 2256: 2255: 2239: 2238: 2232: 2168: 2167: 2109: 2105: 2104: 1663:Microenantiornis 1374:Castignovolucris 1331: 1320: 1247: 1220: 1193: 1184: 1177: 1176: 1123: 1108: 1101: 1094: 1085: 1078: 1077: 1057: 1051: 1050: 1048: 1047: 1032: 1026: 1015: 1009: 1008: 980: 974: 960: 954: 939: 928: 927: 925: 893: 887: 886: 870: 851: 850: 830: 726: 702: 678: 654: 630: 603: 596: 397: 389: 268: 263: 252:Euenantiornithes 250: 234: 139: 138: 115: 105: 42: 27:Temporal range: 21: 2427: 2426: 2422: 2421: 2420: 2418: 2417: 2416: 2392: 2391: 2390: 2385: 2377: 2372: 2364: 2359: 2351: 2346: 2338: 2333: 2325: 2320: 2311: 2310: 2305: 2296: 2295: 2290: 2277: 2267: 2262: 2240: 2233: 2216: 2161: 2160: 2152: 2098: 2097: 2096: 2088: 2052: 1982:Bohaiornithidae 1976: 1928:Dapingfangornis 1921:Camptodontornis 1905:Longipterygidae 1899: 1851: 1824: 1712:Paraprotopteryx 1677:Monoenantiornis 1613:Largirostrornis 1367:Brevirostruavis 1327: 1306: 1283: 1263: 1236: 1209: 1203:Avemetatarsalia 1188:Avemetatarsalia 1178: 1172:Ornithothoraces 1128: 1127: 1117: 1112: 1082: 1081: 1059: 1058: 1054: 1045: 1043: 1034: 1033: 1029: 1016: 1012: 982: 981: 977: 961: 957: 940: 931: 895: 894: 890: 872: 871: 854: 847:10.1002/gj.1209 832: 831: 827: 822: 817: 808: 799: 790: 781: 772: 763: 474:Gregory S. Paul 470: 443:tarsometatarsal 439: 423:enantiornithine 395:Mystiornithidae 390:Kurochkin, 1996 387:Concornithidae? 267: 261: 248: 232: 133: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 37: 36: 33:Late Cretaceous 25: 17: 12: 11: 5: 2425: 2423: 2415: 2414: 2409: 2404: 2394: 2393: 2387: 2386: 2384: 2383: 2370: 2357: 2344: 2331: 2318: 2303: 2287: 2285: 2279: 2278: 2273: 2264: 2263: 2261: 2260: 2249: 2246: 2245: 2242: 2241: 2227: 2225: 2222: 2221: 2218: 2217: 2215: 2214: 2207: 2200: 2193: 2186: 2179: 2171: 2169: 2166:Chiappe, 1992) 2154: 2153: 2151: 2150: 2143: 2136: 2133:Enantiophoenix 2129: 2121: 2112: 2106: 2090: 2089: 2087: 2086: 2078: 2070: 2062: 2060: 2058:Gobipterygidae 2054: 2053: 2051: 2050: 2043: 2036: 2029: 2026:Parabohaiornis 2022: 2015: 2008: 2001: 1994: 1986: 1984: 1978: 1977: 1975: 1974: 1971:Shengjingornis 1967: 1960: 1953: 1950:Longirostravis 1946: 1939: 1931: 1924: 1917: 1909: 1907: 1901: 1900: 1898: 1897: 1890: 1883: 1876: 1869: 1861: 1859: 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1115:Enantiornithes 1113: 1111: 1110: 1103: 1096: 1088: 1080: 1079: 1052: 1027: 1010: 991:(2): 133–138. 975: 955: 929: 908:(8): 499–503. 888: 852: 841:(4): 434–444. 824: 823: 821: 818: 814: 813: 810: 809: 805: 804: 801: 800: 796: 795: 792: 791: 787: 786: 783: 782: 778: 777: 774: 773: 769: 768: 765: 764: 760: 759: 756: 755: 746: 743: 742: 739: 738: 729: 724: 722: 719: 718: 715: 714: 705: 700: 698: 695: 694: 691: 690: 681: 676: 674: 671: 670: 667: 666: 657: 652: 650: 647: 646: 643: 642: 638:Enantiophoenix 633: 628: 626: 620: 619: 616: 615: 606: 601: 599: 594: 574:Enantiophoenix 502:Enantiornithes 469: 466: 438: 435: 407: 406: 405: 404: 392: 381: 380: 374: 373: 372: 371: 364: 357: 350: 343: 336: 329: 322: 319:Enantiophoenix 315: 307: 299: 289: 288: 282: 281: 259: 255: 254: 246: 239: 238: 236:Enantiornithes 230: 223: 222: 217: 210: 209: 204: 197: 196: 191: 184: 183: 178: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 117: 116: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 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bones). 452: 448: 444: 436: 434: 431: 427: 424: 421: 417: 413: 401: 396: 393: 388: 385: 384: 382: 379: 375: 370: 369: 368:Soroavisaurus 365: 363: 362: 358: 356: 355: 351: 349: 348: 344: 342: 341: 337: 335: 334: 330: 328: 327: 323: 321: 320: 316: 313: 312: 308: 305: 304: 300: 298: 297: 293: 292: 290: 287: 283: 279: 275: 271: 266: 260: 257: 256: 253: 247: 244: 241: 240: 237: 231: 228: 225: 224: 221: 218: 215: 212: 211: 208: 205: 202: 199: 198: 195: 192: 189: 186: 185: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 125: 124: 118: 114: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 34: 30: 22: 19: 2282: 2209: 2204:Neuquenornis 2202: 2195: 2188: 2181: 2174: 2163: 2157: 2145: 2138: 2131: 2123: 2115: 2100: 2093: 2080: 2072: 2065: 2045: 2038: 2031: 2024: 2019:Longusunguis 2017: 2010: 2003: 1996: 1989: 1969: 1962: 1955: 1948: 1941: 1933: 1926: 1919: 1912: 1892: 1885: 1878: 1871: 1864: 1844: 1837: 1817: 1810: 1803: 1796: 1789: 1782: 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Retrieved 1040: 1030: 1022: 1018: 1013: 988: 984: 978: 970: 964: 958: 953:(3): 344-350 950: 946: 905: 901: 891: 882: 878: 838: 834: 828: 749: 748: 732: 731: 710:Neuquenornis 708: 707: 684: 683: 660: 659: 636: 635: 623: 609: 608: 587: 572: 569:Avisauroidea 568: 562: 556: 550: 540: 534: 528: 526: 521: 517: 511: 505: 485: 478: 471: 459: 455: 440: 411: 410: 399: 394: 386: 366: 361:Neuquenornis 359: 352: 345: 338: 331: 324: 317: 309: 301: 294: 270:Brett-Surman 264: 242: 226: 213: 200: 187: 174: 121: 38:125–66  18: 2313:Avisauridae 2307:Wikispecies 2283:Avisauridae 2158:Avisauridae 2117:Bauxitornis 2094:Avisauridae 2033:Shenqiornis 2005:Gretcheniao 1964:Shanweiniao 1943:Longipteryx 1894:Yuanchuavis 1873:Eopengornis 1866:Chiappeavis 1846:Noguerornis 1756:Pterygornis 1749:Protopteryx 1698:Orienantius 1641:Liaoxiornis 1535:Gracilornis 1528:Grabauornis 1500:Flexomornis 1486:Falcatakely 1479:Explorornis 1451:Enantiornis 1430:Elbretornis 1388:Cathayornis 1257:Maniraptora 1242:Maniraptora 1068:(6): 1–17. 923:11336/60953 624:Avisauridae 552:Longipteryx 447:metatarsals 437:Description 412:Avisauridae 303:Bauxitornis 265:Avisauridae 24:Avisauridae 2402:Avisaurids 2396:Categories 2147:Mystiornis 2140:Halimornis 2067:Gobipteryx 2012:Linyiornis 1998:Bohaiornis 1991:Beiguornis 1791:Xiangornis 1733:Platanavis 1656:Martinavis 1627:Lenesornis 1584:Incolornis 1493:Feitianius 1458:Eoalulavis 1423:Dunhuangia 1278:see below↓ 1154:Dinosauria 1145:Sauropsida 1046:2018-02-03 820:References 662:Halimornis 558:Gobipteryx 430:Cretaceous 398:Kurochkin 354:Mystiornis 333:Halimornis 194:Saurischia 181:Dinosauria 2190:Intiornis 2176:Avisaurus 2125:Concornis 2082:Vescornis 2047:Zhouornis 1957:Rapaxavis 1935:Evgenavis 1914:Boluochia 1887:Pengornis 1798:Yatenavis 1777:Shangyang 1741:Praeornis 1705:Otogornis 1691:Nanantius 1684:Musivavis 1670:Mirusavis 1605:Kuszholia 1577:Imparavis 1542:Gurilynia 1521:Gobipipus 1395:Cratoavis 1353:Alexornis 1339:Abavornis 1230:Theropoda 1215:Theropoda 1163:Theropoda 1131:Kingdom: 734:Avisaurus 686:Concornis 590:cladogram 480:Avisaurus 451:phalanges 428:from the 426:dinosaurs 340:Intiornis 311:Concornis 296:Avisaurus 207:Theropoda 154:Kingdom: 148:Eukaryota 2298:Q4703066 2292:Wikidata 2258:Category 2074:Jibeinia 2040:Sulcavis 1784:Sinornis 1763:Qiliania 1719:Parvavis 1620:Lectavis 1591:Junornis 1556:Houornis 1549:Holbotia 1444:Elsornis 1139:Chordata 1137:Phylum: 1133:Animalia 611:Sinornis 579:Elsornis 564:Sinornis 545:and all 499:subclass 378:Synonyms 258:Family: 168:Chordata 164:Phylum: 158:Animalia 144:Domain: 2353:3605542 2340:4844232 2327:4434643 2197:Mirarce 2183:Gettyia 1819:Yuornis 1770:Sazavis 1360:Avimaia 1143:Class: 1025:, 39 pp 1005:4523037 495:Avialae 420:extinct 347:Mirarce 326:Gettyia 220:Avialae 123:Mirarce 2379:203286 1003:  547:genera 462:hallux 416:family 402:, 2011 400:et al. 286:Genera 2366:53285 2164:sensu 2101:sensu 1168:Clade 1159:Clade 1150:Clade 1001:JSTOR 488:class 414:is a 243:Clade 227:Clade 214:Clade 201:Clade 188:Clade 175:Clade 29:Early 2335:GBIF 1255:see 1228:see 1201:see 1023:3083 533:and 491:Aves 278:1985 274:Paul 272:and 46:PreꞒ 2322:EoL 1070:doi 993:doi 969:". 918:hdl 910:doi 883:149 843:doi 561:or 520:or 418:of 2398:: 2376:: 2363:: 2350:: 2337:: 2324:: 2309:: 2294:: 1170:: 1161:: 1152:: 1066:14 1064:. 1039:. 1021:. 999:. 987:. 951:12 945:" 932:^ 916:. 904:. 900:. 881:. 877:. 855:^ 839:45 837:. 585:. 555:, 276:, 245:: 229:: 216:: 203:: 190:: 177:: 96:Pg 40:Ma 35:, 2162:( 2128:? 2120:? 2099:( 2085:? 2077:? 1938:? 1744:? 1736:? 1644:? 1608:? 1107:e 1100:t 1093:v 1076:. 1072:: 1049:. 1007:. 995:: 989:5 926:. 920:: 912:: 906:9 849:. 845:: 314:? 306:? 262:† 249:† 233:† 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ 31:-

Index

Early
Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Mirarce
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Avialae
Enantiornithes
Euenantiornithes
Avisauridae
Brett-Surman

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