717:
147:
1556:
694:
323:
1436:
128:
1001:. Two-segmented chelicerae is the ancestral condition in Acariformes, but in more derived groups they are single-segmented. And three-segmented chelicerae is the ancestral condition in Parasitiformes, but has been reduced to just two segments in more derived groups. The pedipalps differ between taxa depending on diet; in some species the appendages resemble legs while in others they are modified into chelicerae-like structures. The oral cavity connects posteriorly to the mouth and
1174:
521:
857:
170:
4536:
1345:
662:
974:
805:
1779:
1107:
1160:, or wherever the mite happens to live. They take up to six weeks to hatch, according to species, with the next stage being the six-legged larvae. After three moults, the larvae become nymphs, with eight legs, and after a further three moults, they become adults. Longevity varies between species, but the lifespan of mites is short compared to many other arachnids.
827:
1427:; the mite, often equipped with suitable claspers or suckers, grips onto an insect or other animal, and gets transported to another place. A phoretic mite is just a hitch-hiker and does not feed during the time it is carried by its temporary host. These travelling mites are mostly species that reproduce rapidly and are quick to colonise new habitats.
1068:= middle), have just one pair of stigmata in the region of the fourth pair of legs. They also have peritremes: in Ixodida these consist of paired encircling plates around the stigmata, while the peritremes in Mesostigmata and Holothyrida are grooves extending from the stigmata anteriorly (sometimes also posteriorly).
1082:
sinuses and most mites lacks a heart, movement of fluid being driven by the contraction of body muscles. But ticks, and some of the larger species of mites, have a dorsal, longitudinal heart. Gas exchange is carried out across the body surface, but many species additionally have between one and four pairs of
1058:= before/in front) have stigmata at the front of the body, usually on the lateral margins or between the chelicerae. These are associated with peritremes that may be on the prodorsum near the cheliceral bases, or be horn-like and emergent, or form a line or network on the dorsum of the gnathosomal capsule.
1419:
Being unable to fly, mites need some other means of dispersal. On a small scale, walking is used to access other suitable locations in the immediate vicinity. Some species mount to a high point and adopt a dispersal posture and get carried away by the wind, while others waft a thread of silk aloft to
1253:
The mites living in soil consist of a range of taxa. Oribatida and
Prostigmata are more numerous in soil than Mesostigmata, and have more soil-dwelling species. When soil is affected by an ecological disturbance such as agriculture, most mites (Astigmata, Mesostigmata and Prostigmata) recolonise it
1081:
that open into the preoral space rather than the foregut. Most species carry two to six pairs of salivary glands that empty at various points into the subcheliceral space. A few mite species lack an anus: they do not defecate during their short lives. The circulatory system consists of a network of
936:
However, a few phylogenomic studies have found strong support for monophyly of Acari and a sister relationship between
Acariformes and Parasitiformes, although this finding has been questioned, with other studies suggesting that this likely represents a
1454:
Mites are tiny and apart from those that are of economic concern to humans, little studied. The majority are beneficial, living in the soil or aqueous environments and assisting in the decomposition of decaying organic material, as part of the
4356:
1233:), in the soil, in forests, pastures, agricultural crops, ornamental plants, thermal springs and caves. They inhabit organic debris of all kinds and are extremely numerous in leaf litter. They feed on animals, plants and fungi and some are
1243:
is one of the strongest animals in the world, relative to its mass (100 ÎĽg): It lifts up to 1,182 times its own weight, over five times more than would be expected of such a minute animal. A mite also holds a speed record: for its length,
1031:
Many mites have stigmata (openings used in respiration). In some mites, the stigmata are associated with peritremes: paired, tubular, elaborated extensions of the tracheal system. The higher taxa of mites are defined by these structures:
3922:
Pham XD, Otsuka Y, Suzuki H, Takaoka H (March 2001). "Detection of
Orientia tsutsugamushi (Rickettsiales: rickettsiaceae) in unengorged chiggers (Acari: Trombiculidae) from Oita Prefecture, Japan, by nested polymerase chain reaction".
1024:(genital opening) is located on the ventral surface between the fourth pair of legs. Some species have one to five median or lateral eyes but many species are blind, and slit and pit sense organs are common. Both body and limbs bear
2378:
Arribas P, AndĂşjar C, Moraza ML, Linard B, Emerson BC, Vogler AP (March 2020). Teeling E (ed.). "Mitochondrial
Metagenomics Reveals the Ancient Origin and Phylodiversity of Soil Mites and Provides a Phylogeny of the Acari".
4020:
van den Broek AH, Huntley JF, MacHell J, Taylor M, Bates P, Groves B, Miller HR (August 2000). "Cutaneous and systemic responses during primary and challenge infestations of sheep with the sheep scab mite, Psoroptes ovis".
277:. Most mites are tiny, less than 1 mm (0.04 in) in length, and have a simple, unsegmented body plan. The small size of most species makes them easily overlooked; some species live in water, many live in soil as
2233:
Dabert M, Witalinski W, Kazmierski A, Olszanowski Z, Dabert J (July 2010). "Molecular phylogeny of acariform mites (Acari, Arachnida): strong conflict between phylogenetic signal and long-branch attraction artifacts".
3451:"Stingless bees (Meliponidae) of the Western Hemisphere: Lestrimelitta and the following subgenera of Trigona: Trigona, Paratrigona, Schwarziana, Parapartamona, Cephalotrigona, Oxytrigona, Scaura, and Mourella"
2964:
Magud BD, Stanisavljević LZ, Petanović RU (2007). "Morphological variation in different populations of Aceria anthocoptes (Acari: Eriophyoidea) associated with the Canada thistle, Cirsium arvense, in Serbia".
2415:
Taxonomically, the Acari can readily be separated into two superorders, the
Acariformes and Parasitiformes, .... most acarologists would agree that both lineages are not closely related and thus Acari are not
2485:
Howard RJ, Puttick MN, Edgecombe GD, Lozano-Fernandez J (November 2020). "Arachnid monophyly: Morphological, palaeontological and molecular support for a single terrestrialization within
Chelicerata".
2799:
Shatrov AB (January 2005). "Ultrastructural investigations of the salivary glands in adults of the microtrombidiid mite
Platytrombidium fasciatum (CL Koch, 1836)(Acariformes: Microtrombidiidae)".
1028:(bristles) which may be simple, flattened, club-shaped or sensory. Mites are usually some shade of brown, but some species are red, orange, black or green, or some combination of these colours.
977:
1 Chelicerae, 2 Palps, 3 Salivary glands, 4 Gut, 5 Excretory (Malpighian) tubules, 6 Anus, 7 Ovary or testes, 8 Air-breathing tubes (tracheae), 9 Central ganglion, 10 Legs, 11 Hypostome.
2590:
958:; most are in the size range 250 to 750 ÎĽm (0.01 to 0.03 in) but some are larger and some are no bigger than 100 ÎĽm (0.004 in) as adults. The body plan has two
3830:
4206:"Comparing the Life Table and Population Projection of Gaeolaelaps aculeifer and Stratiolaelaps scimitus (Acari: Laelapidae) Based on the Age-Stage, Two-Sex Life Table Theory"
2835:
970:
or abdomen. Segmentation has almost entirely been lost and the prosoma and opisthosoma are fused, only the positioning of the limbs indicating the location of the segments.
1496:
for disease transmission, and causing or contributing to allergenic diseases. Mites which colonize human skin are the cause of several types of itchy skin rashes, such as
716:
1817:
1040:= hidden), and Endeostigmata (endeo- = internal) lack primary stigmata and peritremes but may have secondary respiratory systems. For example, oribatids in the suborder
3008:
Arroyo J, Keith AM, Schmidt O, Bolger T (2013). "Mite abundance and richness in an Irish survey of soil biodiversith with comments on some newly recorded species".
1656:. This organism is an obligate external parasite, able to reproduce only in bee colonies. It directly weakens its host by sucking up the bee's fat, and can spread
476:– plant parasitic mites (spider mites, peacock mites, red-legged earth mites, etc.), snout mites, chiggers, hair follicle mites, velvet mites, water mites, etc.
4816:
4314:
4064:
Carp RI, Meeker HC, Rubenstein R, Sigurdarson S, Papini M, Kascsak RJ, et al. (April 2000). "Characteristics of scrapie isolates derived from hay mites".
1149:
on a surface from which it is picked up by the female, or he uses his chelicerae or third pair of legs to insert it into the female's gonopore. In some of the
1237:
of plants and animals. Some 48,200 species of mites have been described, but there may be a million or more species as yet undescribed. The tropical species
4322:
1290:, which feed mostly on dead skin and hair shed from humans instead of consuming them from the organism directly, evolved from these parasitic ancestors.
1094:. Several families of mites, such as Tetranychidae, Eriophyidae, Camerobiidae, Cunaxidae, Trombidiidae, Trombiculidae, Erythraeidae and Bdellidae have
3994:
4803:
3690:
Theis J, Lavoipierre MM, LaPerriere R, Kroese H (June 1981). "Tropical rat mite dermatitis. Report of six cases and review of mite infestations".
2911:
2864:
1867:"Superorder Parasitiformes: In: Zhang, Z-Q. (ed.) Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness"
1327:
workers have been found with mites attached to the outer face of their hind tibiae. Some are thought to be parasites, while others are beneficial
146:
3383:
4266:
3745:
3639:
3607:
3575:
3332:
3299:
3137:
2885:
2758:
2709:
2623:
2556:
1890:
1849:
539:
The mite fossil record is sparse, due to their small size and low preservation potential. The oldest fossils of acariform mites are from the
273:. However, most recent genetic analyses do not recover the two as each other's closest relative within Arachnida, rendering the group non-
1294:
are a prominent group of mites that are parasitic on vertebrates, mostly mammal and birds, feeding on blood with specialised mouthparts.
1209:
in many habitats. They eat a wide variety of material including living and dead plant and fungal material, lichens and carrion; some are
3871:
1664:. Heavy infestation causes the death of a colony, generally over the winter. Since 2006, more than 10 million beehives have been lost.
1098:
glands used to produce silk for various purposes. Additionally, water mites (Hydrachnidia) produce long thin threads that may be silk.
3761:
Andrews RM, McCarthy J, Carapetis JR, Currie BJ (December 2009). "Skin disorders, including pyoderma, scabies, and tinea infections".
1555:
1321:. Hundreds of species are associated with other bees, mostly poorly described. They attach to bees in a variety of ways. For example,
985:
or capitulum. This is not a head and does not contain the eyes or the brain, but is a retractable feeding apparatus consisting of the
1916:"Ordered phylogenomic subsampling enables diagnosis of systematic errors in the placement of the enigmatic arachnid order Palpigradi"
356:
is being extensively used to understand relationships between groups. The 18 S rRNA gene provides information on relationships among
3813:
3170:
1012:), each with six segments, which may be modified for swimming or other purposes. The dorsal surface of the body is clad in hardened
2843:
4865:
3188:"Small but powerful: the oribatid mite Archegozetes longisetosus Aoki (Acari, Oribatida) produces disproportionately high forces"
2329:
Ballesteros JA, Santibáñez-López CE, Baker CM, Benavides LR, Cunha TJ, Gainett G, et al. (February 2022). Teeling E (ed.).
4903:
4410:
2115:"Mitochondrial Metagenomics Reveals the Ancient Origin and Phylodiversity of Soil Mites and Provides a Phylogeny of the Acari"
4808:
3076:"Molecular phylogeny of marine mites (Acariformes: Halacaridae), the oldest radiation of extant secondarily marine animals"
4160:
3282:
Coleman DC, Crossley DA, Hendrix PF (2004). "Secondary
Production: Activities of Heterotrophic Organisms—The Soil Fauna".
2430:"Increasing species sampling in chelicerate genomic-scale datasets provides support for monophyly of Acari and Arachnida"
4898:
1549:
307:
mites of humans. Most species are harmless to humans, but a few are associated with allergies or may transmit diseases.
693:
4728:
361:
322:
1435:
4782:
2900:
1246:
169:
3450:
4821:
1615:
1423:
Parasitic mites use their hosts to disperse, and spread from host to host by direct contact. Another strategy is
1239:
4371:
4332:
1649:
1522:
is a parasitic mite responsible for scabies, which is one of the three most common skin disorders in children.
1044:
have stigmata on the ventral plate of the body that are difficult to see (thus the former name
Cryptostigmata).
587:(of multiple origins). A study using molecular data from the mitochondria and nucleus recovered Acariformes as
1225:, and because of their small size go largely unnoticed. They are found in freshwater (e.g. the water mites or
1113:(Trombiculidae) life cycle: the larvae and nymphs resemble small adults, though the larvae have only six legs.
3481:
3437:
Ticks (Acari: Ixodida) are large parasitiform mites characterized by mouthparts specialized for blood feeding
1965:"A new opilioacarid species (Parasitiformes: Opilioacarida) from Crete (Greece) with notes on its karyotype"
1741:
1706:
1622:
which lives on the skin, causing hypersensitivity and inflammation. Hay mites are a suspected reservoir for
1190:
136:
2774:
2331:"Comprehensive Species Sampling and Sophisticated Algorithmic Approaches Refute the Monophyly of Arachnida"
1700:
4668:
3998:
3159:
Nature and human society: the quest for a sustainable world: proceedings of the 1997 Forum on
Biodiversity
2725:
1501:
938:
2673:
1341:
are ectoparasites of arthropods, while later life stages in this group tend to shift to being predators.
4842:
4318:
1964:
1493:
31:
3655:
Schulze KE, Cohen PR (February 1994). "Dove-associated gamasoidosis: a case of avian mite dermatitis".
1061:
Opilioacaridae have four pairs of dorsolateral stigmata that are added sequentially during development.
599:, with other arachnid orders separating these two groupings on the phylogenetic tree, as shown below.
4870:
4764:
4715:
4305:
3493:
2808:
2494:
2441:
2428:
Lozano-Fernandez J, Tanner AR, Giacomelli M, Carton R, Vinther J, Edgecombe GD, Pisani D (May 2019).
2282:
2164:
de la Fuente J (2003). "The fossil record and the origin of ticks (Acari: Parasitiformes: Ixodida)".
2022:
2010:
1715:
1464:
1318:
1138:
4475:
4403:
3737:
3315:
Behan-Pelletier VM (1999). "Oribatid mite biodiversity in agroecosystems: role for bioindication".
1661:
1596:, found in warm and humid places such as beds, cause several forms of allergic diseases, including
1350:
1157:
4470:
4108:
973:
4857:
4141:
4089:
4046:
3948:
3509:
3428:
3350:"Is permanent parasitism reversible?--critical evidence from early evolution of house dust mites"
3262:
3162:
3105:
3056:
2990:
2946:
2518:
2189:
2046:
1986:
1745:
1424:
1372:
1270:, or nest mites, live primarily in the nests of birds and other animals. These mites are largely
1179:
1153:, insemination is direct using the male's penis. The spermatophora in all mites are aflagellate.
164:
4847:
4465:
4460:
4124:
2901:
Observation on Silk Production and Morphology of Silk in Water Mites (Acariformes: Hydrachnidia)
2113:
Arribas P, AndĂşjar C, Moraza ML, Linard B, Emerson BC, Vogler AP (March 2020). Teeling E (ed.).
1173:
856:
520:
127:
4180:
3627:
4829:
4647:
4639:
4365:
4262:
4227:
4081:
4038:
3975:
3940:
3863:
3809:
3778:
3741:
3707:
3672:
3635:
3603:
3595:
3571:
3563:
3420:
3371:
3328:
3295:
3250:
3209:
3166:
3154:
3133:
3097:
3048:
2982:
2881:
2754:
2705:
2619:
2552:
2510:
2467:
2406:
2360:
2308:
2251:
2181:
2146:
2095:
2038:
1945:
1896:
1886:
1845:
1837:
1784:
1640:
1609:
1518:
1333:
1299:
1222:
1198:
1091:
652:
596:
572:
369:
365:
155:
3797:
4834:
4624:
4289:
4217:
4133:
4073:
4030:
3966:
Diaz JH (2010). "Endemic mite-transmitted dermatoses and infectious diseases in the South".
3932:
3853:
3845:
3798:"The Role of Demodex Mites in the Pathogenesis of Rosacea and Blepharitis and Their Control"
3770:
3729:
3699:
3664:
3501:
3462:
3412:
3361:
3320:
3287:
3240:
3199:
3127:
3087:
3040:
2974:
2936:
2865:
Medical Entomology: A Textbook on Public Health and Veterinary Problems Caused by Arthropods
2816:
2749:
Norton RA, Behan-Pelletier VM (2009). "Suborder Oribatida". In Krantz GW, Walter DE (eds.).
2611:
2502:
2457:
2449:
2396:
2388:
2350:
2342:
2298:
2290:
2243:
2173:
2136:
2126:
2085:
2077:
2030:
1976:
1935:
1927:
1878:
1812:
1585:
1509:
1356:
4535:
1544:
contributes to the disease is unclear. Ticks are well known for carrying diseases, such as
1145:. In most mites, sperm is transferred to the female indirectly; the male either deposits a
583:(sometimes known as Anactinotrichida). Recent genetic research has suggested that Acari is
547:, around 410 million years ago while the earliest fossils of Parasitiformes are known from
4310:
2269:
Sanggaard KW, Bechsgaard JS, Fang X, Duan J, Dyrlund TF, Gupta V, et al. (May 2014).
1593:
1489:
1323:
1185:
959:
328:
3730:
3703:
1914:
Ballesteros JA, Santibáñez López CE, Kováč Ľ, Gavish-Regev E, Sharma PP (December 2019).
340:
The mites are not a defined taxon, but is used for two distinct groups of arachnids, the
3497:
2812:
2498:
2445:
2286:
2070:
Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences
2026:
1672:
Various mites prey on other invertebrates and can be used to control their populations.
1344:
661:
4893:
4777:
4544:
4521:
4508:
4485:
4396:
3858:
3480:
Berghoff SM, Wurst E, Ebermann E, Sendova-Franks AB, Rettenmeyer CW, Franks NR (2009).
3416:
3324:
3291:
2462:
2429:
2355:
2330:
2303:
2270:
2090:
2065:
1940:
1915:
1756:
1589:
1470:
1309:
1078:
821:
804:
799:
675:
580:
493:
479:
473:
383:
345:
266:
240:
46:
3668:
2603:
1866:
4887:
4619:
4493:
4252:
4113:
is the main culprit for the death and reduced populations of overwintered honey bee (
4034:
3505:
3028:
2950:
2522:
1990:
1963:
Vázquez MM, Herrera IM, Just P, Lerma AC, Chatzaki M, Heller T, Král J (2021-09-30).
1439:
1404:
1368:
1338:
1267:
1146:
963:
457:
432:
390:
353:
84:
4283:
4145:
4093:
4050:
3513:
3432:
3109:
3060:
2994:
2193:
2050:
1536:
and other domesticated animals, have also been implicated in the human skin disease
4629:
4611:
4601:
4566:
4561:
4455:
4450:
4248:
4165:
4107:
Guzmán-Novoa E, Eccles L, Calvete Y, Mcgowan J, Kelly PG, Correa-BenĂtez A (2009).
3952:
2644:
2572:
Balashov YS (1972). "Bloodsucking Ticks - Vectors of Diseases of Man and Animals".
1981:
1802:
1792:
1778:
1736:
1731:
1690:
1673:
1581:
1577:
1565:
1560:
1545:
1497:
1456:
1443:
1396:
1376:
1314:
1226:
1218:
1122:
1110:
1041:
1009:
588:
540:
533:
529:
467:
461:
426:
420:
274:
2034:
1106:
1064:
The other three orders of Parasitiformes, Holothyrida, Ixodida, and Mesostigmata (
4706:
4376:
4328:
4256:
3027:
Di Sabatino A, Smit H, Gerecke R, Goldschmidt T, Matsumoto N, Cicolani B (2008).
2271:"Spider genomes provide insight into composition and evolution of venom and silk"
4790:
4700:
4571:
4553:
4516:
4498:
4433:
3482:"Symbionts of societies that fission: Mites as guests or parasites of army ants"
3092:
3075:
3029:"Global diversity of water mites (Acari, Hydrachnidia; Arachnida) in freshwater"
2247:
1760:
1684:
1400:
1392:
1384:
1364:
1230:
1206:
1150:
1037:
967:
786:
576:
504:
501:– oribatid mites, beetle mites, armored mites (formerly known as Cryptostigmata)
485:
450:
404:
352:
of the Acari has been relatively little studied, but molecular information from
341:
262:
235:
211:
59:
3936:
3403:
Beati L, Klompen H (January 2019). "Phylogeography of Ticks (Acari: Ixodida)".
2941:
2924:
2615:
2453:
1882:
4720:
4077:
3774:
3044:
2978:
2820:
2506:
2177:
1807:
1797:
1774:
1764:
1727:
1695:
1678:
1653:
1505:
1287:
1271:
1263:
1234:
1214:
1087:
986:
982:
626:
552:
394:
290:
278:
104:
69:
4733:
4231:
4222:
4205:
4161:"Fears for crops as shock figures from America show scale of bee catastrophe"
3052:
2346:
2042:
1900:
4591:
4442:
4137:
3366:
3349:
2392:
2131:
2114:
1711:
1657:
1645:
1597:
1483:
1380:
1328:
1304:
1210:
1202:
1065:
848:
606:
592:
584:
498:
398:
349:
311:
300:
286:
258:
201:
181:
109:
53:
4691:
4085:
4042:
3979:
3944:
3867:
3782:
3424:
3375:
3254:
3213:
3101:
2986:
2514:
2471:
2410:
2364:
2312:
2255:
2185:
2150:
2099:
2081:
1949:
1931:
1763:, seen under the microscope; the short film was shown in London's Alhambra
826:
4388:
3849:
3711:
3676:
1580:, but they can also spread disease in some limited circumstances, such as
1055:
555:, around 100 million years ago. Most fossil acarids are no older than the
4751:
4743:
4685:
4596:
1573:
1118:
1021:
1017:
994:
990:
955:
886:
556:
544:
408:
333:
254:
221:
99:
94:
79:
74:
64:
4348:
3538:
2401:
2141:
1759:
in which they all lived. The world's first science documentary featured
4756:
3245:
3228:
3204:
3187:
2700:
Krantz GW (2009). "Form and Function". In Krantz GW, Walter DE (eds.).
2294:
1753:
1749:
1623:
1537:
1524:
1513:
1447:
1283:
1134:
1083:
1002:
998:
688:
414:
304:
114:
89:
17:
3896:
1048:
579:(sometimes known as Actinotrichida) are mites, as well as some of the
4795:
4360:
3796:
Mumcuoglu KY, Akilov OE (March 2010). Whitehead J, Barrows B (eds.).
3539:"Parasitengona - velvet mites (including chiggers) & water mites"
3466:
3384:"Genetic study of house dust mites demonstrates reversible evolution"
1605:
1601:
1408:
1013:
993:
and the oral cavity. It is covered above by an extension of the body
882:
711:
438:
357:
295:
191:
4662:
3317:
Invertebrate Biodiversity as Bioindicators of Sustainable Landscapes
1121:
in the mid-region of the body, each connected to the gonopore by a
3517:
2574:
Miscellaneous Publications of the Entomological Society of America
1627:
1554:
1529:
1434:
1388:
1343:
1275:
1172:
1142:
1130:
1126:
1105:
972:
548:
519:
321:
310:
The scientific discipline devoted to the study of mites is called
4769:
4582:
4420:
3227:
Rubin S, Young MH, Wright JC, Whitaker DL, Ahn AN (March 2016).
1291:
1279:
1095:
1025:
560:
507:– stored product, fur, feather, dust, and human itch mites, etc.
282:
4666:
4392:
2912:
How the sperm lost its tail: The evolution of aflagellate sperm
2208:
1744:
from dirt, they were "very prettily shap'd Insects". In 1898,
1533:
1189:, the Canada thistle, across the world. It may be usable as a
4204:
Park J, Mostafiz MM, Hwang HS, Jung DO, Lee KY (2021-05-25).
1492:, but a few species can colonize mammals directly, acting as
1767:
in 1903, causing a boom in the sales of simple microscopes.
1694:, are used to control pests such as spider mites. Among the
1450:, an often overlooked condition especially among the elderly
1387:). Among the species that attack animals are members of the
1254:
within a few months, whereas Oribatida take multiple years.
4325:
National Public Health Pesticide Applicator Training Manual
3802:
Journal of the Rosacea Research & Development Institute
2704:(3rd ed.). Lubbock, Tex: Texas Tech University Press.
2610:, Dordrecht: Kluwer Academic Publishers, pp. 913–919,
1051:= without) lack stigmata and respire through their cuticle.
393:– a small order of large mites that superficially resemble
269:, which were historically grouped together in the subclass
2066:"Terrestrial invertebrates in the Rhynie chert ecosystem"
1217:. Mites are among the most diverse and successful of all
1488:
The majority of mite species are harmless to humans and
3634:. Springer Science & Business Media. p. 2425.
3229:"Exceptional running and turning performance in a mite"
2228:
2226:
293:. This last type includes the commercially destructive
2753:(3rd ed.). Lubbock, Tex: Texas Tech Univ. Press.
1117:
The sexes are separate in mites; males have a pair of
997:
and is connected to the body by a flexible section of
3263:"Mite sets new record as world's fastest land animal"
2207:
Walter DE, Krantz G, Lindquist E (13 December 1996).
4338:
2880:(7th ed.). Cengage Learning. pp. 590–595.
2551:(7th ed.). Cengage Learning. pp. 590–595.
1474:; both are frequently referred to as eyelash mites.
595:(camel spiders) and Parasitiformes as sister to the
4675:
4638:
4610:
4580:
4552:
4543:
4507:
4484:
4441:
4432:
4342:
4181:"twospotted spider mite - Tetranychus urticae Koch"
3732:
Andrews' Diseases of the Skin: Clinical Dermatology
1865:Beaulieu, Frédéric (2011). Zhang, Zhi-Qiang (ed.).
227:
163:
41:
3968:The Journal of the Louisiana State Medical Society
3455:Bulletin of the American Museum of Natural History
3074:Pepato AR, Vidigal TH, Klimov PB (December 2018).
2324:
2322:
1331:. Mites also parasitize some ant species, such as
407:- small group of predatory mites native to former
2604:"Four-Legged Mites (Eriophyoidea or Tetrapodili)"
1818:List of mites associated with cutaneous reactions
1752:of some cheese mites disputing the origin of the
1008:Most mites have four pairs of legs (two pairs in
423:– a large order of predatory and parasitic mites
261:). Mites span two large orders of arachnids, the
3132:. University of California Press. pp. 1–3.
435:- diverse order of parasitic and predatory mites
2925:"Closeups of a not-so-domestic mite tritonymph"
441:- small order of mites containing five families
4285:Pen and pencil: A souvenir of the Press Bazaar
3657:Journal of the American Academy of Dermatology
2836:"You Almost Certainly Have Mites On Your Face"
1748:wrote a satirical poem, "A Parable", with the
1020:; sometimes these form transverse ridges. The
482:- small order of mites containing two families
4404:
4315:United States Environmental Protection Agency
4243:
4241:
3564:"Chapter 7: Mites and other non-insect pests"
1036:Oribatida, formerly known as Cryptostigmata (
8:
3736:(10th ed.). Saunders Elsevier. p.
1266:on plants and animals. One family of mites,
1221:groups. They have exploited a wide array of
281:, others live on plants, sometimes creating
4323:Institute of Food and Agricultural Sciences
3621:
3619:
3602:. Cambridge University Press. p. 260.
3153:Halliday RB, O'Connor BM, Baker AS (2000).
1125:, and in some species there is a chitinous
966:(with no separate head) or prosoma, and an
4663:
4549:
4438:
4411:
4397:
4389:
4339:
3723:
3721:
3121:
3119:
2775:"Astigmata Sarcoptiformes - Urban Insects"
2542:
2540:
2538:
2536:
2534:
2532:
1297:Parasitic mites sometimes infest insects.
145:
126:
38:
4331:at the U.S. National Library of Medicine
4261:. Chatto & Windus. pp. 122–125.
4221:
3857:
3594:Harrison S, Knott H, Bergfeld WF (2009).
3365:
3244:
3203:
3091:
2940:
2461:
2400:
2354:
2302:
2140:
2130:
2089:
1980:
1939:
1403:) are parasites that live in or near the
3600:Skin Infections: Diagnosis and Treatment
3186:Heethoff M, Koerner L (September 2007).
3126:Jeppson LR, Keifer HH, Baker EW (1975).
2923:Uzbekov R, Bouakaz A, Postema M (2023).
1588:is the only known vector of the disease
3589:
3587:
2064:Dunlop JA, Garwood RJ (February 2018).
1828:
372:genes, provide clues at deeper levels.
2876:Ruppert EE, Fox RS, Barnes RD (2004).
2695:
2693:
2668:
2666:
2664:
2591:Parasite Diversity and Diversification
2547:Ruppert EE, Fox RS, Barnes RD (2004).
3897:"Rocky Mountain Spotted Fever (RMSF)"
3080:Molecular Phylogenetics and Evolution
2801:Arthropod Structure & Development
2487:Arthropod Structure & Development
2236:Molecular Phylogenetics and Evolution
543:, Scotland, which dates to the early
7:
3704:10.1001/archderm.1981.01650060031018
2967:Experimental & Applied Acarology
2166:Experimental & Applied Acarology
2004:
2002:
2000:
1726:Mites were first observed under the
1016:and the ventral surface by hardened
954:Mites are tiny members of the class
4306:Bitingmites.org: What's biting you?
3838:The New England Journal of Medicine
3233:The Journal of Experimental Biology
3192:The Journal of Experimental Biology
1462:Two species live on humans, namely
792:
782:
775:
768:
681:
648:
641:
619:
612:
602:
228:Mites are found in two superorders
3763:Pediatric Clinics of North America
3417:10.1146/annurev-ento-020117-043027
3325:10.1016/b978-0-444-50019-9.50023-6
3157:. In Raven PH, Williams T (eds.).
3129:Mites Injurious to Economic Plants
2015:International Journal of Acarology
2011:"Mites as fossils: forever small?"
1540:, although the mechanism by which
1363:Plant pests include the so-called
1086:. The excretory system includes a
454:– the most diverse group of mites
25:
3831:"Clinical practice. Lyme disease"
3348:Klimov PB, OConnor B (May 2013).
1842:Fundamentals of Applied Acarology
1838:"Molecular Biology and Acarology"
1618:, sheep are affected by the mite
1528:mites, which are common cause of
4534:
4035:10.1046/j.1365-3024.2000.00318.x
3506:10.1111/j.1365-2311.2009.01125.x
3292:10.1016/b978-012179726-3/50005-8
3269:(Press release). April 27, 2014.
1920:Proceedings. Biological Sciences
1777:
1740:, he stated that far from being
1395:), which burrow under the skin.
1250:is the fastest animal on Earth.
1183:, is found on the invasive weed
1133:connected to the gonopore by an
981:At the front of the body is the
855:
825:
803:
715:
692:
660:
168:
2381:Molecular Biology and Evolution
2335:Molecular Biology and Evolution
2119:Molecular Biology and Evolution
1559:Mites and their eggs, drawn by
1442:worker Stefania Lanzia using a
1213:, though no oribatid mites are
387:– ticks and a variety of mites
4117:) colonies in Ontario, Canada"
3598:. In Hall JC, Hall BJ (eds.).
3319:. Elsevier. pp. 411–423.
1576:are known primarily for their
802:(chiggers, velvet mites, etc)
488:- large order of sucking mites
49: – Present,
1:
3925:Journal of Medical Entomology
3669:10.1016/S0190-9622(08)81930-5
3628:"Mite Pests of Crops in Asia"
3568:Plant Pests and Their Control
3449:Schwarz HF, Bacon AL (1948).
3286:. Elsevier. pp. 79–185.
2674:"All mites have a small head"
2035:10.1080/01647954.2018.1497085
1608:, and are known to aggravate
1197:Mites occupy a wide range of
3728:James WD, Berger TG (2006).
3284:Fundamentals of Soil Ecology
1982:10.24349/acarologia/20214449
1550:Rocky Mountain spotted fever
1077:Mite digestive systems have
824:(dust & fur mites, etc)
551:specimens dating to the mid-
27:Small eight-legged arthropod
3997:. NetDoctor. Archived from
3405:Annual Review of Entomology
3155:"Global Diversity of Mites"
3093:10.1016/j.ympev.2018.08.012
2248:10.1016/j.ympev.2009.12.020
2009:Sidorchuk EA (2018-11-17).
1420:balloon to a new position.
1102:Reproduction and life cycle
571:Members of the superorders
4920:
4282:Doyle AC (June 28, 1898).
3937:10.1603/0022-2585-38.2.308
3632:Encyclopedia of Entomology
3010:Irish Naturalists' Journal
2942:10.1007/s40629-023-00249-6
2616:10.1007/0-306-48380-7_1689
2608:Encyclopedia of Entomology
2454:10.1038/s41467-019-10244-7
2211:. Tree of Life Web Project
1883:10.11646/zootaxa.3148.1.23
1481:
1247:Paratarsotomus macropalpis
881:other Arachnids including
470:– gall mites and relatives
360:and superphyla, while the
29:
4532:
4159:Benjamin A (2 May 2010).
4078:10.3109/13550280009013157
3995:"House dust mite allergy"
3993:Klenerman P, Lipworth B.
3775:10.1016/j.pcl.2009.09.002
3596:"Infections of the Scalp"
3570:. Elsevier. p. 112.
3045:10.1007/s10750-007-9025-1
2979:10.1007/s10493-007-9085-y
2834:Yong E (27 August 2014).
2821:10.1016/j.asd.2004.09.001
2507:10.1016/j.asd.2020.100997
1844:. Springer. p. 176.
1240:Archegozetes longisetosus
1156:The eggs are laid in the
918:
879:
852:
819:
797:
790:
780:
773:
766:
707:
686:
679:
656:
646:
639:
624:
617:
610:
232:
165:Scientific classification
153:
144:
134:
125:
4333:Medical Subject Headings
4223:10.3390/agronomy11061062
4066:Journal of Neurovirology
3630:. In Capinera JL (ed.).
1730:by the English polymath
1718:and various soil pests.
1676:, especially members of
1650:colony collapse disorder
1431:Relationship with humans
1303:attaches to the body of
1129:; females have a single
1090:and one or two pairs of
3974:(3): 140–145, 147–149.
3829:Shapiro ED (May 2014).
3692:Archives of Dermatology
2178:10.1023/A:1025824702816
1742:spontaneously generated
1707:Stratiolaelaps scimitus
1668:Biological pest control
1191:biological pest control
137:Trombidium holosericeum
4904:Arthropod common names
2347:10.1093/molbev/msac021
2082:10.1098/rstb.2016.0493
1932:10.1098/rspb.2019.2426
1616:Among domestic animals
1570:
1502:rodent mite dermatitis
1451:
1360:
1229:) and saltwater (most
1194:
1114:
978:
939:long branch attraction
709:Parasitic mites, inc.
536:
429:- large, diverse order
337:
318:Evolution and taxonomy
4843:Paleobiology Database
4319:University of Florida
4138:10.1051/apido/2009076
3850:10.1056/NEJMcp1314325
3486:Ecological Entomology
3367:10.1093/sysbio/syt008
2846:on September 11, 2014
2751:A manual of acarology
2730:keys.lucidcentral.org
2702:A Manual of Acarology
2678:keys.lucidcentral.org
2434:Nature Communications
2393:10.1093/molbev/msz255
2275:Nature Communications
2132:10.1093/molbev/msz255
1701:Gaeolaelaps aculeifer
1644:is a serious pest of
1558:
1482:Further information:
1438:
1354:, caused by the mite
1347:
1176:
1109:
976:
523:
326:The microscopic mite
325:
32:Mite (disambiguation)
4001:on February 11, 2008
3562:Fenemore PG (2016).
3198:(Pt 17): 3036–3042.
3165:. pp. 192–212.
2878:Invertebrate Zoology
2549:Invertebrate Zoology
1710:are used to control
1478:Medical significance
1465:Demodex folliculorum
1411:, including humans.
1205:mites are important
1193:agent for this weed.
526:Glaesacarus rhombeus
30:For other uses, see
4899:Paraphyletic groups
4023:Parasite Immunology
3663:(2 Pt 1): 278–280.
3498:2009EcoEn..34..684B
2840:National Geographic
2813:2005ArtSD..34...49S
2499:2020ArtSD..5900997H
2446:2019NatCo..10.2295L
2287:2014NatCo...5.3765S
2027:2018IJAca..44..349S
1836:Dhooria MS (2016).
1734:. In his 1665 book
1662:deformed wing virus
1373:thread-footed mites
1348:Lime nail galls on
1141:for the storage of
401:), hence their name
285:, while others are
140:mite (Acariformes)
4185:entnemdept.ufl.edu
3903:. 15 November 2018
3354:Systematic Biology
3246:10.1242/jeb.128652
3205:10.1242/jeb.008276
3163:National Academies
2295:10.1038/ncomms4765
2209:"Acari: The mites"
2076:(1739): 20160493.
1926:(1917): 20192426.
1746:Arthur Conan Doyle
1648:, contributing to
1630:disease of sheep.
1571:
1452:
1361:
1195:
1139:seminal receptacle
1115:
1092:Malpighian tubules
979:
925:(mites and ticks)
537:
338:
4881:
4880:
4830:Open Tree of Life
4669:Taxon identifiers
4660:
4659:
4656:
4655:
4640:Opilioacariformes
4530:
4529:
4386:
4385:
4268:978-0-7011-8180-2
4111:Varroa destructor
3877:on 21 August 2016
3844:(18): 1724–1731.
3747:978-0-7216-2921-6
3641:978-1-4020-6242-1
3609:978-0-521-89729-7
3577:978-1-4831-8286-5
3334:978-0-444-50019-9
3301:978-0-12-179726-3
3239:(Pt 5): 676–685.
3139:978-0-520-02381-9
2887:978-81-315-0104-7
2760:978-0-89672-620-8
2711:978-0-89672-620-8
2625:978-0-7923-8670-4
2558:978-81-315-0104-7
1892:978-1-86977-849-1
1851:978-981-10-1594-6
1785:Arthropods portal
1716:poultry red mites
1641:Varroa destructor
1610:atopic dermatitis
1519:Sarcoptes scabiei
1337:. Most larvae of
1334:Eciton burchellii
1317:) lives in their
1300:Varroa destructor
1199:ecological niches
933:
932:
915:
914:
906:
905:
897:
896:
868:
867:
838:
837:
755:
754:
746:
745:
737:
736:
728:
727:
653:Pseudoscorpionida
597:Pseudoscorpionida
573:Opilioacariformes
370:28S ribosomal RNA
366:18S ribosomal RNA
248:
247:
159:(Parasitiformes)
156:Varroa destructor
120:
16:(Redirected from
4911:
4874:
4873:
4861:
4860:
4851:
4850:
4838:
4837:
4825:
4824:
4812:
4811:
4799:
4798:
4786:
4785:
4773:
4772:
4760:
4759:
4747:
4746:
4737:
4736:
4724:
4723:
4711:
4710:
4709:
4696:
4695:
4694:
4664:
4550:
4538:
4439:
4413:
4406:
4399:
4390:
4340:
4294:
4293:
4290:Punch_(magazine)
4279:
4273:
4272:
4245:
4236:
4235:
4225:
4201:
4195:
4194:
4192:
4191:
4177:
4171:
4170:
4156:
4150:
4149:
4121:
4104:
4098:
4097:
4061:
4055:
4054:
4017:
4011:
4010:
4008:
4006:
3990:
3984:
3983:
3963:
3957:
3956:
3919:
3913:
3912:
3910:
3908:
3893:
3887:
3886:
3884:
3882:
3876:
3870:. Archived from
3861:
3835:
3826:
3820:
3819:
3793:
3787:
3786:
3769:(6): 1421–1440.
3758:
3752:
3751:
3735:
3725:
3716:
3715:
3687:
3681:
3680:
3652:
3646:
3645:
3623:
3614:
3613:
3591:
3582:
3581:
3559:
3553:
3552:
3550:
3549:
3535:
3529:
3528:
3526:
3525:
3516:. Archived from
3477:
3471:
3470:
3446:
3440:
3439:
3400:
3394:
3391:
3390:. March 8, 2013.
3379:
3369:
3345:
3339:
3338:
3312:
3306:
3305:
3279:
3273:
3270:
3258:
3248:
3224:
3218:
3217:
3207:
3183:
3177:
3176:
3150:
3144:
3143:
3123:
3114:
3113:
3095:
3071:
3065:
3064:
3024:
3018:
3017:
3005:
2999:
2998:
2961:
2955:
2954:
2944:
2920:
2914:
2909:
2903:
2898:
2892:
2891:
2873:
2867:
2862:
2856:
2855:
2853:
2851:
2842:. Archived from
2831:
2825:
2824:
2796:
2790:
2789:
2787:
2786:
2771:
2765:
2764:
2746:
2740:
2739:
2737:
2736:
2722:
2716:
2715:
2697:
2688:
2687:
2685:
2684:
2670:
2659:
2658:
2656:
2655:
2641:
2635:
2634:
2633:
2632:
2599:
2593:
2588:
2582:
2581:
2569:
2563:
2562:
2544:
2527:
2526:
2482:
2476:
2475:
2465:
2425:
2419:
2418:
2404:
2375:
2369:
2368:
2358:
2326:
2317:
2316:
2306:
2266:
2260:
2259:
2230:
2221:
2220:
2218:
2216:
2204:
2198:
2197:
2172:(3–4): 331–344.
2161:
2155:
2154:
2144:
2134:
2110:
2104:
2103:
2093:
2061:
2055:
2054:
2006:
1995:
1994:
1984:
1960:
1954:
1953:
1943:
1911:
1905:
1904:
1862:
1856:
1855:
1833:
1787:
1782:
1781:
1654:commercial hives
1594:House dust mites
1586:house-mouse mite
1490:domestic animals
1357:Eriophyes tiliae
1351:Tilia Ă— europaea
859:
829:
807:
793:
783:
776:
769:
719:
696:
682:
665:
664:
658:False scorpions
649:
642:
620:
613:
603:
528:, fossilised in
173:
172:
149:
130:
119:
56:
45:Temporal range:
44:
39:
21:
4919:
4918:
4914:
4913:
4912:
4910:
4909:
4908:
4884:
4883:
4882:
4877:
4869:
4864:
4856:
4854:
4846:
4841:
4833:
4828:
4820:
4815:
4807:
4802:
4794:
4789:
4781:
4776:
4768:
4763:
4755:
4750:
4742:
4740:
4732:
4727:
4719:
4714:
4705:
4704:
4699:
4690:
4689:
4684:
4671:
4661:
4652:
4634:
4606:
4602:Nuttalliellidae
4576:
4539:
4526:
4503:
4480:
4428:
4417:
4387:
4382:
4381:
4351:
4311:Mites and Ticks
4302:
4297:
4281:
4280:
4276:
4269:
4258:Bugs Britannica
4247:
4246:
4239:
4203:
4202:
4198:
4189:
4187:
4179:
4178:
4174:
4158:
4157:
4153:
4119:
4106:
4105:
4101:
4063:
4062:
4058:
4019:
4018:
4014:
4004:
4002:
3992:
3991:
3987:
3965:
3964:
3960:
3921:
3920:
3916:
3906:
3904:
3895:
3894:
3890:
3880:
3878:
3874:
3833:
3828:
3827:
3823:
3816:
3795:
3794:
3790:
3760:
3759:
3755:
3748:
3727:
3726:
3719:
3689:
3688:
3684:
3654:
3653:
3649:
3642:
3625:
3624:
3617:
3610:
3593:
3592:
3585:
3578:
3561:
3560:
3556:
3547:
3545:
3537:
3536:
3532:
3523:
3521:
3479:
3478:
3474:
3448:
3447:
3443:
3402:
3401:
3397:
3382:
3347:
3346:
3342:
3335:
3314:
3313:
3309:
3302:
3281:
3280:
3276:
3261:
3226:
3225:
3221:
3185:
3184:
3180:
3173:
3152:
3151:
3147:
3140:
3125:
3124:
3117:
3073:
3072:
3068:
3026:
3025:
3021:
3007:
3006:
3002:
2963:
2962:
2958:
2922:
2921:
2917:
2910:
2906:
2899:
2895:
2888:
2875:
2874:
2870:
2863:
2859:
2849:
2847:
2833:
2832:
2828:
2798:
2797:
2793:
2784:
2782:
2773:
2772:
2768:
2761:
2748:
2747:
2743:
2734:
2732:
2724:
2723:
2719:
2712:
2699:
2698:
2691:
2682:
2680:
2672:
2671:
2662:
2653:
2651:
2643:
2642:
2638:
2630:
2628:
2626:
2602:Hoy MA (2004),
2601:
2600:
2596:
2589:
2585:
2571:
2570:
2566:
2559:
2546:
2545:
2530:
2484:
2483:
2479:
2427:
2426:
2422:
2377:
2376:
2372:
2328:
2327:
2320:
2268:
2267:
2263:
2232:
2231:
2224:
2214:
2212:
2206:
2205:
2201:
2163:
2162:
2158:
2112:
2111:
2107:
2063:
2062:
2058:
2008:
2007:
1998:
1962:
1961:
1957:
1913:
1912:
1908:
1893:
1864:
1863:
1859:
1852:
1835:
1834:
1830:
1826:
1783:
1776:
1773:
1724:
1670:
1636:
1486:
1480:
1433:
1417:
1389:sarcoptic mange
1324:Trigona corvina
1262:Many mites are
1260:
1201:. For example,
1186:Cirsium arvense
1171:
1166:
1137:, as well as a
1104:
1079:salivary glands
1075:
952:
947:
934:
924:
916:
907:
898:
869:
839:
756:
747:
738:
729:
659:
569:
518:
378:
329:Lorryia formosa
320:
167:
121:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
51:
50:
35:
28:
23:
22:
15:
12:
11:
5:
4917:
4915:
4907:
4906:
4901:
4896:
4886:
4885:
4879:
4878:
4876:
4875:
4862:
4852:
4839:
4826:
4813:
4800:
4787:
4778:Fauna Europaea
4774:
4761:
4748:
4738:
4725:
4712:
4697:
4681:
4679:
4673:
4672:
4667:
4658:
4657:
4654:
4653:
4651:
4650:
4648:Opilioacaridae
4644:
4642:
4636:
4635:
4633:
4632:
4627:
4622:
4616:
4614:
4608:
4607:
4605:
4604:
4599:
4594:
4588:
4586:
4578:
4577:
4575:
4574:
4569:
4564:
4558:
4556:
4547:
4545:Parasitiformes
4541:
4540:
4533:
4531:
4528:
4527:
4525:
4524:
4522:Sphaerolichida
4519:
4513:
4511:
4509:Trombidiformes
4505:
4504:
4502:
4501:
4496:
4490:
4488:
4486:Sarcoptiformes
4482:
4481:
4479:
4478:
4473:
4468:
4463:
4458:
4453:
4447:
4445:
4436:
4430:
4429:
4418:
4416:
4415:
4408:
4401:
4393:
4384:
4383:
4380:
4379:
4368:
4352:
4347:
4346:
4344:
4343:Classification
4337:
4336:
4326:
4308:
4301:
4300:External links
4298:
4296:
4295:
4274:
4267:
4237:
4196:
4172:
4151:
4132:(4): 443–450.
4115:Apis mellifera
4099:
4072:(2): 137–144.
4056:
4029:(8): 407–414.
4012:
3985:
3958:
3931:(2): 308–311.
3914:
3888:
3821:
3814:
3788:
3753:
3746:
3717:
3698:(6): 341–343.
3682:
3647:
3640:
3626:Ho CC (2008).
3615:
3608:
3583:
3576:
3554:
3530:
3492:(6): 684–695.
3472:
3441:
3411:(1): 379–397.
3395:
3393:
3392:
3360:(3): 411–423.
3340:
3333:
3307:
3300:
3274:
3272:
3271:
3219:
3178:
3171:
3145:
3138:
3115:
3066:
3039:(1): 303–315.
3019:
3000:
2973:(3): 173–183.
2956:
2935:(8): 337–339.
2915:
2904:
2893:
2886:
2868:
2857:
2826:
2791:
2766:
2759:
2741:
2717:
2710:
2689:
2660:
2636:
2624:
2594:
2583:
2564:
2557:
2528:
2477:
2420:
2387:(3): 683–694.
2370:
2318:
2261:
2242:(1): 222–241.
2222:
2199:
2156:
2125:(3): 683–694.
2105:
2056:
2021:(8): 349–359.
1996:
1975:(3): 548–563.
1955:
1906:
1891:
1857:
1850:
1827:
1825:
1822:
1821:
1820:
1815:
1810:
1805:
1800:
1795:
1789:
1788:
1772:
1769:
1757:cheddar cheese
1723:
1720:
1669:
1666:
1635:
1632:
1620:Psoroptes ovis
1590:rickettsialpox
1479:
1476:
1471:Demodex brevis
1432:
1429:
1416:
1413:
1405:hair follicles
1391:mites (family
1310:Acarapis woodi
1259:
1256:
1180:A. anthocoptes
1170:
1167:
1165:
1162:
1103:
1100:
1074:
1071:
1070:
1069:
1062:
1059:
1052:
1045:
951:
948:
946:
943:
931:
930:
927:
926:
917:
913:
912:
909:
908:
904:
903:
900:
899:
895:
894:
891:
890:
878:
875:
874:
871:
870:
866:
865:
862:
861:
854:Camel spiders
851:
845:
844:
841:
840:
836:
835:
832:
831:
822:Sarcoptiformes
818:
815:
814:
811:
810:
800:Trombidiformes
796:
791:
789:
781:
779:
774:
772:
767:
765:
762:
761:
758:
757:
753:
752:
749:
748:
744:
743:
740:
739:
735:
734:
731:
730:
726:
725:
722:
721:
706:
703:
702:
699:
698:
685:
680:
678:
676:Parasitiformes
672:
671:
668:
667:
655:
647:
645:
640:
638:
635:
634:
631:
630:
623:
618:
616:
611:
609:
601:
581:Parasitiformes
568:
565:
517:
514:
513:
512:
511:
510:
509:
508:
502:
494:Sarcoptiformes
491:
490:
489:
483:
480:Sphaerolichida
474:Trombidiformes
471:
465:
446:
445:
444:
443:
442:
436:
430:
418:
412:
402:
384:Parasitiformes
377:
374:
346:Parasitiformes
319:
316:
267:Parasitiformes
257:(eight-legged
246:
245:
244:
243:
241:Parasitiformes
238:
230:
229:
225:
224:
219:
215:
214:
209:
205:
204:
199:
195:
194:
189:
185:
184:
179:
175:
174:
161:
160:
151:
150:
142:
141:
132:
131:
123:
122:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
57:
47:Early Devonian
43:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4916:
4905:
4902:
4900:
4897:
4895:
4892:
4891:
4889:
4872:
4867:
4863:
4859:
4853:
4849:
4844:
4840:
4836:
4831:
4827:
4823:
4818:
4814:
4810:
4805:
4801:
4797:
4792:
4788:
4784:
4779:
4775:
4771:
4766:
4762:
4758:
4753:
4749:
4745:
4739:
4735:
4730:
4726:
4722:
4717:
4713:
4708:
4702:
4698:
4693:
4687:
4683:
4682:
4680:
4678:
4674:
4670:
4665:
4649:
4646:
4645:
4643:
4641:
4637:
4631:
4628:
4626:
4623:
4621:
4620:Monogynaspida
4618:
4617:
4615:
4613:
4609:
4603:
4600:
4598:
4595:
4593:
4590:
4589:
4587:
4584:
4579:
4573:
4570:
4568:
4565:
4563:
4560:
4559:
4557:
4555:
4551:
4548:
4546:
4542:
4537:
4523:
4520:
4518:
4515:
4514:
4512:
4510:
4506:
4500:
4497:
4495:
4494:Endeostigmata
4492:
4491:
4489:
4487:
4483:
4477:
4476:Parhyposomata
4474:
4472:
4469:
4467:
4464:
4462:
4459:
4457:
4454:
4452:
4449:
4448:
4446:
4444:
4440:
4437:
4435:
4431:
4426:
4422:
4414:
4409:
4407:
4402:
4400:
4395:
4394:
4391:
4378:
4374:
4373:
4369:
4367:
4363:
4362:
4358:
4354:
4353:
4350:
4345:
4341:
4334:
4330:
4327:
4324:
4320:
4316:
4312:
4309:
4307:
4304:
4303:
4299:
4292:. p. 58.
4291:
4287:
4286:
4278:
4275:
4270:
4264:
4260:
4259:
4254:
4250:
4244:
4242:
4238:
4233:
4229:
4224:
4219:
4215:
4211:
4207:
4200:
4197:
4186:
4182:
4176:
4173:
4168:
4167:
4162:
4155:
4152:
4147:
4143:
4139:
4135:
4131:
4127:
4126:
4118:
4116:
4112:
4103:
4100:
4095:
4091:
4087:
4083:
4079:
4075:
4071:
4067:
4060:
4057:
4052:
4048:
4044:
4040:
4036:
4032:
4028:
4024:
4016:
4013:
4000:
3996:
3989:
3986:
3981:
3977:
3973:
3969:
3962:
3959:
3954:
3950:
3946:
3942:
3938:
3934:
3930:
3926:
3918:
3915:
3902:
3898:
3892:
3889:
3873:
3869:
3865:
3860:
3855:
3851:
3847:
3843:
3839:
3832:
3825:
3822:
3817:
3815:9781450203449
3811:
3807:
3803:
3799:
3792:
3789:
3784:
3780:
3776:
3772:
3768:
3764:
3757:
3754:
3749:
3743:
3739:
3734:
3733:
3724:
3722:
3718:
3713:
3709:
3705:
3701:
3697:
3693:
3686:
3683:
3678:
3674:
3670:
3666:
3662:
3658:
3651:
3648:
3643:
3637:
3633:
3629:
3622:
3620:
3616:
3611:
3605:
3601:
3597:
3590:
3588:
3584:
3579:
3573:
3569:
3565:
3558:
3555:
3544:
3540:
3534:
3531:
3520:on 2017-02-06
3519:
3515:
3511:
3507:
3503:
3499:
3495:
3491:
3487:
3483:
3476:
3473:
3468:
3464:
3460:
3456:
3452:
3445:
3442:
3438:
3434:
3430:
3426:
3422:
3418:
3414:
3410:
3406:
3399:
3396:
3389:
3388:Michigan News
3385:
3381:
3380:
3377:
3373:
3368:
3363:
3359:
3355:
3351:
3344:
3341:
3336:
3330:
3326:
3322:
3318:
3311:
3308:
3303:
3297:
3293:
3289:
3285:
3278:
3275:
3268:
3264:
3260:
3259:
3256:
3252:
3247:
3242:
3238:
3234:
3230:
3223:
3220:
3215:
3211:
3206:
3201:
3197:
3193:
3189:
3182:
3179:
3174:
3172:9780309065559
3168:
3164:
3160:
3156:
3149:
3146:
3141:
3135:
3131:
3130:
3122:
3120:
3116:
3111:
3107:
3103:
3099:
3094:
3089:
3085:
3081:
3077:
3070:
3067:
3062:
3058:
3054:
3050:
3046:
3042:
3038:
3034:
3033:Hydrobiologia
3030:
3023:
3020:
3015:
3011:
3004:
3001:
2996:
2992:
2988:
2984:
2980:
2976:
2972:
2968:
2960:
2957:
2952:
2948:
2943:
2938:
2934:
2930:
2929:Allergo J Int
2926:
2919:
2916:
2913:
2908:
2905:
2902:
2897:
2894:
2889:
2883:
2879:
2872:
2869:
2866:
2861:
2858:
2845:
2841:
2837:
2830:
2827:
2822:
2818:
2814:
2810:
2806:
2802:
2795:
2792:
2780:
2776:
2770:
2767:
2762:
2756:
2752:
2745:
2742:
2731:
2727:
2726:"Acariformes"
2721:
2718:
2713:
2707:
2703:
2696:
2694:
2690:
2679:
2675:
2669:
2667:
2665:
2661:
2650:
2646:
2640:
2637:
2627:
2621:
2617:
2613:
2609:
2605:
2598:
2595:
2592:
2587:
2584:
2579:
2575:
2568:
2565:
2560:
2554:
2550:
2543:
2541:
2539:
2537:
2535:
2533:
2529:
2524:
2520:
2516:
2512:
2508:
2504:
2500:
2496:
2492:
2488:
2481:
2478:
2473:
2469:
2464:
2459:
2455:
2451:
2447:
2443:
2439:
2435:
2431:
2424:
2421:
2417:
2412:
2408:
2403:
2398:
2394:
2390:
2386:
2382:
2374:
2371:
2366:
2362:
2357:
2352:
2348:
2344:
2340:
2336:
2332:
2325:
2323:
2319:
2314:
2310:
2305:
2300:
2296:
2292:
2288:
2284:
2280:
2276:
2272:
2265:
2262:
2257:
2253:
2249:
2245:
2241:
2237:
2229:
2227:
2223:
2210:
2203:
2200:
2195:
2191:
2187:
2183:
2179:
2175:
2171:
2167:
2160:
2157:
2152:
2148:
2143:
2138:
2133:
2128:
2124:
2120:
2116:
2109:
2106:
2101:
2097:
2092:
2087:
2083:
2079:
2075:
2071:
2067:
2060:
2057:
2052:
2048:
2044:
2040:
2036:
2032:
2028:
2024:
2020:
2016:
2012:
2005:
2003:
2001:
1997:
1992:
1988:
1983:
1978:
1974:
1970:
1966:
1959:
1956:
1951:
1947:
1942:
1937:
1933:
1929:
1925:
1921:
1917:
1910:
1907:
1902:
1898:
1894:
1888:
1884:
1880:
1876:
1872:
1868:
1861:
1858:
1853:
1847:
1843:
1839:
1832:
1829:
1823:
1819:
1816:
1814:
1813:Grocer's itch
1811:
1809:
1806:
1804:
1801:
1799:
1796:
1794:
1791:
1790:
1786:
1780:
1775:
1770:
1768:
1766:
1762:
1758:
1755:
1751:
1747:
1743:
1739:
1738:
1733:
1729:
1721:
1719:
1717:
1713:
1709:
1708:
1703:
1702:
1697:
1693:
1692:
1687:
1686:
1681:
1680:
1675:
1667:
1665:
1663:
1659:
1655:
1651:
1647:
1643:
1642:
1634:In beekeeping
1633:
1631:
1629:
1625:
1621:
1617:
1613:
1611:
1607:
1603:
1599:
1595:
1591:
1587:
1583:
1579:
1575:
1568:
1567:
1562:
1557:
1553:
1551:
1547:
1543:
1539:
1535:
1531:
1527:
1526:
1521:
1520:
1515:
1511:
1510:grocer's itch
1507:
1503:
1499:
1495:
1491:
1485:
1477:
1475:
1473:
1472:
1467:
1466:
1460:
1458:
1449:
1446:to publicise
1445:
1441:
1440:Public health
1437:
1430:
1428:
1426:
1421:
1414:
1412:
1410:
1406:
1402:
1398:
1397:Demodex mites
1394:
1390:
1386:
1382:
1378:
1374:
1370:
1369:Tetranychidae
1366:
1359:
1358:
1353:
1352:
1346:
1342:
1340:
1339:Parasitengona
1336:
1335:
1330:
1326:
1325:
1320:
1316:
1312:
1311:
1306:
1302:
1301:
1295:
1293:
1289:
1285:
1281:
1277:
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458:Endeostigmata
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433:Monogynaspida
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391:Opilioacarida
389:
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363:
359:
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302:
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4612:Mesostigmata
4567:Holothyridae
4562:Allothyridae
4471:Palaeosomata
4456:Enarthronota
4451:Brachypylina
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3872:the original
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2839:
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1831:
1803:Gamasoidosis
1793:Chigger bite
1761:cheese mites
1737:Micrographia
1735:
1732:Robert Hooke
1725:
1712:fungus gnats
1705:
1699:
1691:Phytoseiulus
1689:
1683:
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1674:Phytoseiidae
1671:
1639:
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1582:scrub typhus
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1566:Micrographia
1564:
1561:Robert Hooke
1546:Lyme disease
1541:
1523:
1517:
1498:gamasoidosis
1487:
1469:
1463:
1461:
1457:carbon cycle
1453:
1444:scabies mite
1422:
1418:
1377:Tarsonemidae
1365:spider mites
1362:
1355:
1349:
1332:
1322:
1315:Tarsonemidae
1308:
1298:
1296:
1274:and consume
1261:
1252:
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1227:Hydrachnidia
1219:invertebrate
1196:
1184:
1178:
1155:
1123:vas deferens
1116:
1111:Harvest mite
1076:
1042:Brachypylina
1030:
1010:Eriophyoidea
1007:
980:
953:
935:
920:
880:
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820:
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710:
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585:polyphyletic
570:
541:Rhynie Chert
538:
534:Upper Eocene
530:Baltic amber
525:
468:Eriophyoidea
462:paraphyletic
449:
427:Trigynaspida
421:Mesostigmata
382:
339:
327:
309:
299:parasite of
294:
275:monophyletic
270:
250:
249:
154:
135:
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4791:iNaturalist
4701:Wikispecies
4572:Neothyridae
4554:Holothyrida
4517:Prostigmata
4499:Psoroptidia
4434:Acariformes
4313:chapter in
4216:(6): 1062.
3086:: 182–188.
2850:23 November
2649:Bee Mite ID
2440:(1): 2295.
1685:Metaseiulus
1658:RNA viruses
1401:Demodecidae
1393:Sarcoptidae
1385:Eriophyidae
1379:), and the
1231:Halacaridae
1207:decomposers
1151:Acariformes
1047:Astigmata (
968:opisthosoma
787:Acariformes
577:Acariformes
505:Astigmatina
486:Prostigmata
451:Acariformes
448:Superorder
405:Holothyrida
381:Superorder
342:Acariformes
279:decomposers
263:Acariformes
236:Acariformes
212:Chelicerata
208:Subphylum:
52:410–0
4888:Categories
4466:Mixonomata
4461:Holosomata
4288:. London:
4190:2023-02-09
4125:Apidologie
3907:20 January
3548:2023-02-09
3524:2019-08-19
2785:2023-06-26
2735:2023-06-27
2683:2023-06-26
2654:2023-06-26
2645:"Glossary"
2631:2023-02-08
2580:: 161–376.
2493:: 100997.
1969:Acarologia
1824:References
1808:Grain itch
1798:Copra itch
1765:music hall
1728:microscope
1722:In culture
1696:Laelapidae
1679:Amblyseius
1660:including
1646:honey bees
1578:itchy bite
1506:grain itch
1381:gall mites
1305:honey bees
1288:Dust mites
1258:Parasitism
1088:nephridium
987:chelicerae
983:gnathosoma
941:artefact.
627:Palpigradi
559:(up to 65
553:Cretaceous
460:(probably
411:landmasses
395:harvestmen
364:, and the
301:honey bees
259:arthropods
253:are small
202:Arthropoda
4592:Argasidae
4581:Ixodida (
4443:Oribatida
4232:2073-4395
4169:. London.
3467:2246/1231
3053:0018-8158
2951:258581966
2523:222302964
2215:6 October
2043:0164-7954
1991:236270478
1901:1175-5326
1638:The mite
1598:hay fever
1484:Acariasis
1415:Dispersal
1329:symbionts
1272:parasitic
1264:parasitic
1235:parasites
1215:parasitic
1211:predatory
1203:Oribatida
1158:substrate
1018:sclerites
991:pedipalps
956:Arachnida
887:scorpions
849:Solifugae
607:Arachnida
593:Solifugae
567:Phylogeny
524:Mite, cf
499:Oribatida
399:Opiliones
350:phylogeny
312:acarology
291:parasites
287:predators
255:arachnids
222:Arachnida
188:Kingdom:
182:Eukaryota
4752:BugGuide
4741:BioLib:
4686:Wikidata
4597:Ixodidae
4255:(2010).
4249:Marren P
4210:Agronomy
4146:10898654
4094:16441609
4086:10822327
4051:41549010
4043:10972847
3980:20666166
3945:11296840
3868:24785207
3783:19962029
3514:84324830
3433:53023797
3425:30354695
3376:23417682
3255:26787481
3214:17704078
3110:52145427
3102:30172010
3061:10262035
3016:: 19–27.
2995:25895062
2987:17611806
2515:33039753
2472:31127117
2411:31670799
2365:35137183
2313:24801114
2281:: 3765.
2256:20060051
2194:11271627
2186:14635818
2151:31670799
2100:29254958
2051:92357151
1950:31847768
1771:See also
1574:Chiggers
1399:(family
1383:(family
1375:(family
1367:(family
1319:tracheae
1313:(family
1223:habitats
1084:tracheae
1073:Internal
1022:gonopore
1014:tergites
995:carapace
950:External
691:(ticks)
557:Tertiary
545:Devonian
409:Gondwana
376:Taxonomy
344:and the
334:Tydeidae
265:and the
198:Phylum:
192:Animalia
178:Domain:
4770:2586535
4419:Acari (
4377:D008925
4253:Mabey R
3953:8133110
3859:4487875
3712:7247425
3677:8288795
3494:Bibcode
2809:Bibcode
2495:Bibcode
2463:6534568
2442:Bibcode
2356:8845124
2304:4273655
2283:Bibcode
2091:5745329
2023:Bibcode
1941:6939912
1871:Zootaxa
1750:conceit
1624:scrapie
1542:demodex
1538:rosacea
1525:Demodex
1514:scabies
1494:vectors
1448:scabies
1425:phoresy
1409:mammals
1284:keratin
1164:Ecology
1135:oviduct
1038:crypto-
1003:pharynx
999:cuticle
960:regions
945:Anatomy
883:spiders
808:
689:Ixodida
591:to the
417:– ticks
415:Ixodida
305:scabies
218:Class:
18:Acarine
4871:292684
4848:155665
4835:150772
4809:733321
4692:Q19137
4625:Sejina
4335:(MeSH)
4265:
4230:
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1602:asthma
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1307:, and
1169:Niches
1119:testes
989:, the
712:Varroa
589:sister
439:Sejida
348:. The
296:Varroa
4894:Acari
4866:WoRMS
4858:acari
4855:PPE:
4796:52788
4783:10616
4757:91197
4744:16798
4734:Acari
4721:Acari
4707:Acari
4677:Acari
4583:ticks
4425:mites
4421:ticks
4329:Mites
4142:S2CID
4120:(PDF)
4090:S2CID
4047:S2CID
3949:S2CID
3875:(PDF)
3834:(PDF)
3510:S2CID
3429:S2CID
3106:S2CID
3057:S2CID
2991:S2CID
2947:S2CID
2519:S2CID
2341:(2).
2190:S2CID
2047:S2CID
1987:S2CID
1754:round
1628:prion
1530:mange
1292:Ticks
1276:blood
1143:sperm
1131:ovary
1127:penis
1066:meso-
1026:setae
921:Acari
549:amber
358:phyla
283:galls
271:Acari
251:Mites
42:Mites
4822:6933
4817:NCBI
4804:ITIS
4423:and
4372:MeSH
4366:1G07
4317:and
4263:ISBN
4228:ISSN
4082:PMID
4039:PMID
4007:2008
3976:PMID
3941:PMID
3909:2019
3883:2016
3864:PMID
3810:ISBN
3779:PMID
3742:ISBN
3708:PMID
3673:PMID
3636:ISBN
3604:ISBN
3572:ISBN
3421:PMID
3372:PMID
3329:ISBN
3296:ISBN
3251:PMID
3210:PMID
3167:ISBN
3134:ISBN
3098:PMID
3049:ISSN
2983:PMID
2882:ISBN
2852:2017
2755:ISBN
2706:ISBN
2620:ISBN
2553:ISBN
2511:PMID
2468:PMID
2407:PMID
2361:PMID
2309:PMID
2252:PMID
2217:2017
2182:PMID
2147:PMID
2096:PMID
2039:ISSN
1946:PMID
1897:ISSN
1887:ISBN
1875:3148
1846:ISBN
1704:and
1626:, a
1604:and
1548:and
1534:dogs
1468:and
1282:and
1280:skin
1096:silk
1056:pro-
962:, a
885:and
575:and
368:and
362:ITS2
60:Preęž’
4765:EoL
4729:AFD
4716:ADW
4357:ICD
4218:doi
4134:doi
4074:doi
4031:doi
3972:162
3933:doi
3901:CDC
3854:PMC
3846:doi
3842:370
3771:doi
3738:454
3700:doi
3696:117
3665:doi
3502:doi
3463:hdl
3413:doi
3362:doi
3321:doi
3288:doi
3241:doi
3237:219
3200:doi
3196:210
3088:doi
3084:129
3041:doi
3037:595
2975:doi
2937:doi
2817:doi
2612:doi
2503:doi
2458:PMC
2450:doi
2397:hdl
2389:doi
2351:PMC
2343:doi
2299:PMC
2291:doi
2244:doi
2174:doi
2137:hdl
2127:doi
2086:PMC
2078:doi
2074:373
2031:doi
1977:doi
1936:PMC
1928:doi
1924:286
1879:doi
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