1047:. For monopartite viruses, all genes are on the single segment of the genome. Multipartite viruses typically package their genomes into a single virion so that the whole genome is in one virus particle, and the separate segments contain different genes. Monopartite viruses are found in all Baltimore groups, whereas multipartite viruses are usually RNA viruses. This is because most multipartite viruses infect plants or fungi, which are eukaryotes, and most eukaryotic viruses are RNA viruses. The family
142:
608:
91:(RNA), the strandedness of the genome, which can be either single- or double-stranded, and the sense of a single-stranded genome, which is either positive or negative. The primary advantage of Baltimore classification is that by classifying viruses according to the aforementioned characteristics, viruses that behave in the same manner can be studied as distinct groups. There are seven Baltimore groups numbered with Roman numerals, listed hereafter.
1117:
967:
685:
1233:
75:. Since then, it has become common among virologists to use Baltimore classification alongside standard virus taxonomy, which is based on evolutionary history. In 2018 and 2019, Baltimore classification was partially integrated into virus taxonomy based on evidence that certain groups were descended from common ancestors. Various realms, kingdoms, and phyla now correspond to specific Baltimore groups.
437:
777:, whereby a short sequence of host cell mRNA is taken and used as the 5′ cap of the viral mRNA. Genomic -ssRNA is replicated from the positive sense antigenome in a similar manner as transcription, except in reverse using the antigenome as a template for the genome. RdRp moves from the 3′-end to the 5′-end of the antigenome and ignores all transcription signals when synthesizing genomic -ssRNA.
1072:
animal and plant viruses, are RNA viruses, although eukaryotic DNA viruses are also common. By group, the vast majority of dsDNA viruses infect prokaryotes, ssDNA viruses are found in all three domains of life, dsRNA and +ssRNA viruses are primarily found in eukaryotes but also in bacteria, and -ssRNA and reverse transcribing viruses are only found in eukaryotes.
931:
genome in the cytoplasm, and the genome is replicated from these RNA strands. The dsDNA genome is produced from pregenomic RNA strands via the same general mechanism as ssRNA-RT viruses, but with replication occurring in a loop around the circular genome. After replication, the dsDNA genome may be packed or sent to the nucleus for further rounds of transcription.
796:-ssRNA viruses can be subdivided informally between those that have nonsegmented and segmented genomes. Nonsegmented -ssRNA viruses replicate in the cytoplasm, and segmented -ssRNA viruses replicate in the nucleus. During transcription, the RdRp produces one monocistronic mRNA strand from each segment of the genome. All -ssRNA viruses are classified in the phylum
3519:
7366:
228:
progressing in a loop around the circular genome is also common. Some dsDNA viruses use a strand displacement method whereby one strand is synthesized from a template strand, and a complementary strand is then synthesized from the prior synthesized strand, forming a dsDNA genome. Lastly, some dsDNA viruses are replicated as part of a process called
711:+ssRNA viruses can be subdivided between those that have polycistronic mRNA, which encodes a polyprotein that is cleaved to form multiple mature proteins, and those that produce subgenomic mRNAs and therefore undergo two or more rounds of translation. +ssRNA viruses are included in three phyla in the kingdom
1107:
Alternative splicing differs from RNA editing in that alternative splicing does not change the mRNA sequence like RNA editing but instead changes the coding capacity of an mRNA sequence as a result of alternative splicing sites. The two mechanisms otherwise have the same result: multiple proteins are
1080:
Viral genomes may be either linear with ends or circular in a loop. Whether a virus has a linear or circular genome varies from group to group. A significant percentage of dsDNA viruses are both, ssDNA viruses are primarily circular, RNA viruses and ssRNA-RT viruses are typically linear, and dsDNA-RT
1006:
is a mechanism by which different proteins can be produced from a single gene by means of using alternative splicing sites to produce different mRNAs. It is found in various DNA, -ssRNA, and reverse transcribing viruses. Viruses may make use of alternative splicing solely to produce multiple proteins
792:
and then moves back in the RNA sequence with the mRNA to transcribe it again, continuing this process numerous times until hundreds of adenines have been added to the 3′-end of the mRNA. Additionally, some -ssRNA viruses are ambisense, as both the positive and negative strands separately encode viral
768:
The fifth
Baltimore group contains viruses that have a negative sense, single-stranded RNA (-ssRNA) genome. mRNA, which is positive sense, is transcribed directly from the negative sense genome. The first process for -ssRNA transcription involves RdRp binding to a leader sequence on the 3′ end of the
235:
dsDNA viruses can be subdivided between those that replicate in the nucleus, and as such are relatively dependent on host cell machinery for transcription and replication, and those that replicate in the cytoplasm, in which case they have evolved or acquired their own means of executing transcription
2378:
Krupovic M, Blomberg J, Coffin JM, Dasgupta I, Fan H, Geering AD, Gifford R, Harrach B, Hull R, Johnson W, Kreuze JF, Lindemann D, Llorens C, Lockhart B, Mayer J, Muller E, Olszewski NE, Pappu HR, Pooggin MM, Richert-Poggeler KR, Sabanadzovic S, Sanfacon H, Schoelz JE, Seal S, Stavolone L, Stoye JP,
930:
to create a complete dsDNA genome prior to transcription. dsDNA-RT viruses are transcribed in the same manner as dsDNA viruses, but make use of reverse transcription to replicate their circular genome while it is still in the capsid. The host cell's RNA polymerase II transcribes RNA strands from the
844:
Reverse transcribing (RT) viruses have genomes made of either DNA or RNA and replicate via reverse transcription. Two groups of reverse transcribing viruses exist: single-stranded RNA-RT (ssRNA-RT) viruses, and double-stranded DNA-RT (dsDNA-RT) viruses. Reverse transcribing viruses are classified in
1255:
From the 1990s to the 2010s, virus taxonomy used a 5-rank system ranging from order to species with
Baltimore classification used in conjunction. Outside of the ICTV's official framework, various supergroups of viruses joining different families and orders were created over time based on increasing
695:
The fourth
Baltimore group contains viruses that have a positive sense single-stranded RNA (+ssRNA) genome. For +ssRNA viruses, the genome functions as mRNA, so no transcription is required for translation. +ssRNA viruses will also, however, produce positive sense copies of the genome from negative
622:
dsRNA is not a molecule made by cells, so cellular life has evolved antiviral systems to detect and inactivate viral dsRNA. To counteract this, many dsRNA genomes are constructed inside of capsids, thereby avoiding detection inside of the host cell's cytoplasm. mRNA is forced out from the capsid in
227:
dsDNA viruses make use of several mechanisms to replicate their genome. Bidirectional replication, in which two replication forks are established at a replication origin site and move in opposite directions of each other, is widely used. A rolling circle mechanism that produces linear strands while
1220:
Ribosomal skipping: also called stop-carry on, used by various dsRNA and +ssRNA viruses, a viral peptide, or amino acid sequence, may prevent a ribosome from covalently linking a new inserted amino acid, which blocks further translation. Consequently, the polyprotein is co-translationally cleaved,
699:
Many +ssRNA viruses are able to have only a portion of their genome transcribed. Typically, subgenomic RNA (sgRNA) strands are used for translation of structural and movement proteins needed during intermediate and late stages of infection. sgRNA transcription may occur by commencing RNA synthesis
1071:
Different
Baltimore groups tend to be found within different branches of cellular life. In prokaryotes, the large majority of viruses are dsDNA viruses, and a significant minority are ssDNA viruses. Prokaryotic RNA viruses, in contrast, are relatively rare. Most eukaryotic viruses, including most
467:
that bonds to and cleaves the positive strand, allowing a DNA polymerase to use the negative strand as a template for replication. Replication progresses in a loop around the genome by means of extending the 3′-end of the positive strand, displacing the prior positive strand, and the endonuclease
1099:
RNA editing is used by various ssRNA viruses to produce different proteins from a single gene. This can be done via polymerase slippage during transcription or by post-transcriptional editing. In polymerase slippage, the RNA polymerase slips one nucleotide back during transcription, inserting a
861:
The sixth
Baltimore group contains viruses that have a (positive-sense) single-stranded RNA genome that has a DNA intermediate ((+)ssRNA-RT) in its replication cycle. ssRNA-RT viruses are transcribed in the same manner as DNA viruses, but their linear genomes are first converted to a dsDNA form
707:
Because the process of replicating the viral genome produces intermediate dsRNA molecules, +ssRNA viruses can be targeted by the host cell's immune system. To avoid detection, +ssRNA viruses replicate in membrane-associated vesicles that are used as replication factories. From there, only viral
994:
A number of characteristics of viruses are not directly associated with
Baltimore classification but nonetheless closely correspond to multiple, specific Baltimore groups. This includes alternative splicing during transcription, whether the viral genome is segmented, the host range of viruses,
1158:
Viral initiation of translation: used primarily by +ssRNA and ssRNA-RT viruses, various viruses have evolved mechanisms to initiate translation, such as having internal ribosomal entry sites to allow for cap-independent translation, having downstream hairpin loops that allow for cap-dependent
486:
at each end of the genome that repeatedly unfold and refold during replication to change the direction of DNA synthesis to move back and forth along the genome, producing numerous copies of the genome in a continuous process. Individual genomes are then excised from this molecule by the viral
459:
upon entering a host cell. mRNA is then synthesized from the double-stranded form. The double-stranded form of ssDNA viruses may be produced either directly after entry into a cell or as a consequence of replication of the viral genome. Eukaryotic ssDNA viruses are replicated in the nucleus.
568:
RNA viruses have genomes made of ribonucleic acid (RNA) and comprise three groups: double-stranded RNA (dsRNA) viruses, positive sense single-stranded RNA (+ssRNA) viruses, and negative sense single-stranded RNA (-ssRNA) viruses. The majority of RNA viruses are classified in the kingdom
618:
The third
Baltimore group contains viruses that have a double-stranded RNA (dsRNA) genome. After entering a host cell, the dsRNA genome is transcribed to mRNA from the negative strand by the viral RNA-dependent RNA polymerase (RdRp). The mRNA may be used for translation or replication.
501:
package both the positive and negative linear strands. In any case, the sense of ssDNA viruses, unlike for ssRNA viruses, is not sufficient to separate ssDNA viruses into two groups since all ssDNA viral genomes are converted to dsDNA forms prior to transcription and replication.
121:
of the viral genome, and viruses within each group typically share the manners by which the mRNA synthesis occurs. While not the direct focus of
Baltimore classification, groups are organized in such a manner that viruses in each group also typically have the same mechanisms of
1248:. It initially contained the first six groups but was later expanded to include group VII. Because of the utility of Baltimore classification, it has come to be used alongside standard virus taxonomy, which is based on evolutionary relationships and governed by the
696:
sense strands of an intermediate dsRNA genome. This acts as both a transcription and a replication process since the replicated RNA is also mRNA. The 5′-end may be naked, capped, or covalently bound to a viral protein, and the 3′-end may be naked or polyadenylated.
454:
The second
Baltimore group contains viruses that have a single-stranded DNA (ssDNA) genome. ssDNA viruses have the same manner of transcription as dsDNA viruses. Because the genome is single-stranded, however, it is first made into a double-stranded form by a
33:(mRNA) synthesis. By organizing viruses based on their manner of mRNA production, it is possible to study viruses that behave similarly as a distinct group. Seven Baltimore groups are described that take into consideration whether the viral genome is made of
1204:
Suppression of termination: also called stop-codon readthrough, used by various dsRNA, +ssRNA, and ssRNA-RT viruses, certain viruses contain codons in their mRNA that would normally signal for termination of translation upon being recognized by a
1179:: used by various dsDNA, +ssRNA, -ssRNA, ssRNA-RT, a dsDNA-RT viruses, ribosomes will start scanning from a 5′-cap structure then bypass a leader structure in the mRNA, initiation translation downstream from the leader sequence.
165:
DNA viruses have genomes made of deoxyribonucleic acid (DNA) and are organized into two groups: double-stranded DNA (dsDNA) viruses, and single-stranded DNA (ssDNA) viruses. They are assigned to four separate realms:
220:. Second, once the RNA polymerase is recruited, it uses the negative strand as a template for synthesizing mRNA strands. Third, the RNA polymerase terminates transcription upon reaching a specific signal, such as a
1173:: used by various viruses in all Baltimore groups, the 40S ribosomal subunit may scan through a start codon, thereby skipping an ORF, only initiating translation with the 60S subunit at a subsequent start codon.
496:
is the only ssDNA family whose members have negative sense genomes, which are circular. Parvoviruses, as previously mentioned, may package either the positive or negative sense strand into virions. Lastly,
1256:
evidence of deeper evolutionary relations. Consequently, in 2016, the ICTV began to consider establishing ranks higher than order as well as how the
Baltimore groups would be treated among higher taxa.
925:
The seventh Baltimore group contains viruses that have a double-stranded DNA genome that has an RNA intermediate (dsDNA-RT) in its replication cycle. dsDNA-RT viruses have a gap in one strand, which is
1286:
Unlike RNA viruses and RT viruses, DNA viruses have not been united under a single realm but are instead dispersed across four realms and various taxa that are not assigned to a realm. The realms
1151:. Baltimore groups do not directly pertain to the translation of viral proteins, but various atypical types of translation used by viruses are usually found within specific Baltimore groups:
3551:
472:
into a circular loop. The new ssDNA may be packaged into virions or replicated by a DNA polymerase to form a double-stranded form for transcription or continuation of the replication cycle.
1213:
during translation, which allows for continued translation up to the next stop codon in order to produce an extended end of the viral protein. In viruses, this is often used to express
1328:
ssRNA-RT viruses are often called retroviruses, although this term is also used to refer to any reverse transcribing virus as well as specifically to viruses in the ssRNA-RT family
1100:
nucleotide not included in the template strand. Editing of a genomic template would impair gene expression, so RNA editing is only done during and after transcription. For
212:
The first Baltimore group contains viruses that have a double-stranded DNA (dsDNA) genome. All dsDNA viruses have their mRNA synthesized in a three-step process. First, a
1027:
use it as a potential anti-host defense mechanism to prevent synthesis of specific antiviral proteins. Furthermore, in addition to alternative splicing, because cellular
882:
then transcribes RNA in the nucleus from the proviral DNA. Some of this RNA may become mRNA whereas other strands will become copies of the viral genome for replication.
635:
have not been observed to form virions and as such apparently lack capsids. Endornaviruses are also unusual in that unlike other RNA viruses, they possess a single, long
83:
Baltimore classification groups viruses together based on their manner of mRNA synthesis. Characteristics directly related to this include whether the genome is made of
1182:
Termination-reinitiation: used by some dsRNA and +ssRNA viruses, ribosomes may translate an ORF, but following termination of translation of that ORF, a proportion of
1263:
to species was established by the ICTV. As part of this, the Baltimore groups for RNA viruses and RT viruses were incorporated into formal taxa. In 2018, the realm
138:
membrane that surrounds the capsid, have no direct relation to Baltimore groups, nor do the groups necessarily show genetic relation based on evolutionary history.
700:
within the genome rather than from the 5′-end, by stopping RNA synthesis at specific sequences in the genome, or by, as a part of both prior methods, synthesizing
3544:
1249:
870:
enzyme synthesizes a DNA strand from the ssRNA strand, and the RNA strand is degraded and replaced with a DNA strand to create a dsDNA genome. The genome is then
1197:: used by various dsDNA, dsRNA, +ssRNA, and ssRNA-RT viruses, produces merged proteins from overlapping ORFs. This is executed simply by ribosomes slipping one
7386:
623:
order to be translated or to be translocated from a mature capsid to a progeny capsid. While dsRNA viruses typically have capsids, viruses in the families
3537:
1183:
619:
Single-stranded mRNA is replicated to form the dsRNA genome. The 5′-end of the genome may be naked, capped, or covalently bound to a viral protein.
1632:"Create a megataxonomic framework, filling all principal taxonomic ranks, for DNA viruses encoding vertical jelly roll-type major capsid proteins"
704:
from the viral RNA that are then attached to sgRNA strands. Because replication is required for sgRNA synthesis, RdRp is always translated first.
1053:
varies as some viruses are monopartite ssDNA while others are bipartite with one segment being ssDNA and the other dsDNA. Viruses in the ssDNA
1597:"Create a megataxonomic framework, filling all principal/primary taxonomic ranks, for dsDNA viruses encoding HK97-type major capsid proteins"
60:
and replication. Certain subjects pertaining to viruses are associated with multiple, specific Baltimore groups, such as specific forms of
487:
endonuclease. For parvoviruses, either the positive or negative sense strand may be packaged into capsids, varying from virus to virus.
213:
126:
the viral genome. Because of this, Baltimore classification provides insights into both the transcription and replication parts of the
7391:
3493:
3466:
3439:
3412:
3523:
1855:
463:
Most ssDNA viruses contain circular genomes that are replicated via rolling circle replication (RCR). ssDNA RCR is initiated by an
773:, continuing until a stop signal is reached. The second manner is similar but instead of synthesizing a cap, RdRp may make use of
769:
genome, transcribing a 5′ triphosphate-leader RNA that is capped, then stopping and restarting on a transcription signal which is
793:
proteins, and these viruses produce two separate mRNA strands: one directly from the genome and one from a complementary strand.
130:. Structural characteristics of a virus particle, called a virion, such as the shape of the viral capsid and the presence of a
2258:
701:
68:, which stores the viral genome, and the evolutionary history of viruses are not necessarily related to Baltimore groups.
7356:
236:
and replication. dsDNA viruses are also commonly divided between tailed dsDNA viruses, referring to members of the realm
469:
1043:
Viral genomes can exist in a single, or monopartite, segment, or they may be split into more than one molecule, called
5337:
505:
ssDNA viruses are classified into one of the four realms and include several families that are unassigned to a realm:
479:
64:
of mRNA and the host range of different types of viruses. Structural characteristics such as the shape of the viral
7081:
6586:
5660:
780:
Various -ssRNA viruses use special mechanisms for transcription. The manner of producing the polyA tail may be via
251:
dsDNA viruses are classified into three of the four realms and include many taxa that are unassigned to a realm:
229:
46:
1279:. Furthermore, the three Baltimore groups for RNA viruses are used as defining characteristics of the phyla in
490:
Nearly all ssDNA viruses have positive sense genomes, but a few exceptions and peculiarities exist. The family
3373:
Gorbalenya, Alexander E.; Krupovic, Mart; Siddell, Stuart; Varsani, Arvind; Kuhn, Jens H. (15 October 2018).
2702:
Wolf YI, Kazlauskas D, Iranzo J, Lucia-Sanz A, Kuhn JH, Krupovic M, Dolja VV, Kooning EV (27 November 2018).
1221:
and a new amino acid sequence is started, leading to the production of two individual proteins from one ORF.
1194:
118:
57:
1007:
from a single pre-mRNA strand or for other specific purposes. For certain viruses, including the families
216:
binds to the DNA upstream of the site where transcription begins, allowing for the recruitment of a host
1140:
867:
863:
781:
84:
61:
34:
5848:
3375:"Riboviria: establishing a single taxon that comprises RNA viruses at the basal rank of virus taxonomy"
6322:
1271:
was expanded to also include both RT groups. Within the realm, RT viruses are included in the kingdom
6899:
6421:
5616:
3895:
3564:
2764:
2621:
2273:
1121:
1003:
971:
23:
6718:
6386:
6329:
6301:
6273:
5943:
5627:
5558:
4818:
4811:
3902:
3860:
3853:
1971:
Koonin EV, Dolja VV, Krupovic M, Varsani A, Wolf YI, Yutin N, Zerbini M, Kuhn JH (18 October 2019).
1723:
1722:
Harrison RL, Herniou EA, Jehle JA, Theilmann DA, Burand JP, Krell PJ, van Oers MM (28 August 2020).
1696:
Koonin EV, Dolja VV, Krupovic M, Varsani A, Wolf YI, Yutin N, Zerbini M, Kuhn JH (18 October 2019).
1630:
Koonin EV, Dolja VV, Krupovic M, Varsani A, Wolf YI, Yutin N, Zerbini M, Kuhn JH (18 October 2019).
1595:
Koonin EV, Dolja VV, Krupovic M, Varsani A, Wolf YI, Yutin N, Zerbini M, Kuhn JH (18 October 2019).
6913:
6767:
6546:
6294:
6287:
5107:
4305:
3923:
3909:
3867:
3780:
42:
7005:
6956:
6906:
6885:
6336:
3846:
3374:
639:(ORF), or translatable portion, and a site-specific nick in the 5′ region of the positive strand.
6725:
5798:
5791:
4864:
4499:
4485:
4077:
3499:
3445:
2297:
1176:
1083:
636:
1660:
1659:
Krupovic M, Kuhn JH, Wang F, Baquero DP, Egelman EH, Koonin EV, Prangishvili D (31 July 2020).
607:
141:
7021:
5980:
5855:
5834:
5727:
5383:
5041:
4683:
4246:
3839:
3766:
3489:
3462:
3435:
3408:
3355:
3299:
3014:
2792:
2733:
2651:
2599:
2531:
2410:
2289:
1973:"Create a megataxonomic framework, filling all principal taxonomic ranks, for realm Riboviria"
1851:
1129:
1125:
983:
975:
879:
590:
441:
402:
123:
53:
826:, whose members synthesize a cap structure on viral mRNA required for protein synthesis, and
7396:
6811:
6739:
6632:
6207:
6149:
6112:
5841:
5572:
5530:
5478:
5441:
5411:
5372:
5226:
5152:
5009:
4331:
4294:
4268:
4226:
4027:
4017:
3832:
3684:
3655:
3481:
3427:
3345:
3337:
3289:
3281:
3004:
2996:
2782:
2772:
2723:
2715:
2641:
2633:
2589:
2579:
2521:
2513:
2400:
2392:
2281:
1698:"Create a megataxonomic framework, filling all principal taxonomic ranks, for ssDNA viruses"
1260:
1015:
1009:
740:
671:
656:
232:
whereby a viral genome in a host cell's DNA is replicated to another part of a host genome.
127:
88:
38:
995:
whether the genome is linear or circular, and different methods of translating viral mRNA.
7271:
7142:
6977:
6925:
6838:
6794:
6783:
6621:
6473:
6414:
6238:
5959:
5889:
5777:
5760:
5695:
5292:
5247:
4968:
4475:
4257:
4168:
4088:
3984:
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3204:
3178:
3152:
3032:
2957:
2122:
1308:
1241:
1116:
1049:
1020:
966:
887:
839:
798:
763:
749:
537:
291:
221:
72:
3326:"The New Scope of Virus Taxonomy: Partitioning the Virosphere Into 15 Hierarchical Ranks"
2620:
Bamford DH, Pietilä MK, Roine E, Atanasova NS, Dienstbier A, Oksanen HM (December 2017).
1035:
contain their own proteins for exporting their unspliced genomic RNA out of the nucleus.
56:
the genome, so Baltimore classification is useful for grouping viruses together for both
2768:
2277:
684:
270:
are dsDNA viruses. Viruses in this realm belong to two groups: tailed bacteriophages in
7370:
7314:
7292:
7260:
7225:
7131:
7096:
7032:
6875:
6864:
6827:
6669:
6553:
6443:
6372:
6362:
6123:
5915:
5878:
5784:
5579:
5163:
5136:
4783:
4772:
4676:
4641:
4634:
4620:
4613:
4562:
4534:
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4453:
4418:
4202:
4005:
3874:
3811:
3745:
3723:
3645:
3485:
3431:
3350:
3325:
3009:
2984:
2787:
2752:
2728:
2703:
2646:
2594:
2567:
2526:
2501:
2405:
2380:
1206:
1170:
950:
940:
770:
631:
586:
456:
396:
366:
360:
348:
342:
334:
300:
217:
131:
3294:
3269:
2381:"Ortervirales: New Virus Order Unifying Five Families of Reverse-Transcribing Viruses"
7380:
7282:
7177:
7163:
6991:
6984:
6963:
6804:
6753:
6746:
6610:
6506:
6484:
6457:
6433:
6393:
6350:
6315:
6308:
6280:
6266:
6259:
6160:
6096:
6067:
6046:
6025:
6014:
5994:
5926:
5748:
5717:
5706:
5684:
5551:
5418:
5404:
5394:
5361:
5285:
5237:
5180:
5173:
5020:
4920:
4794:
4708:
4690:
4669:
4655:
4555:
4492:
4442:
4364:
4342:
4315:
4278:
4180:
4147:
3995:
3973:
3773:
3756:
3734:
3712:
3634:
3623:
2257:
Kuhn JH, Wolf YI, Krupovic M, Zhang YZ, Maes P, Dolja VV, Koonin EV (February 2019).
1059:
774:
725:
625:
571:
559:
522:
492:
422:
408:
390:
378:
329:
The following taxa that are unassigned to a realm exclusively contain dsDNA viruses:
278:
203:
174:
30:
3503:
3449:
2301:
1232:
145:
Visualization of the seven groups of virus according to the Baltimore Classification
7304:
7249:
7184:
7153:
7120:
6970:
6942:
6760:
6732:
6711:
6697:
6690:
6680:
6516:
6450:
6379:
6227:
6197:
6186:
6133:
6089:
6079:
6060:
6036:
5987:
5899:
5812:
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5215:
5126:
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4853:
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4701:
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4219:
4121:
4109:
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3916:
3822:
3674:
3529:
3285:
2751:
Yu C, Hernandez T, Zheng H, Yau SC, Huang HH, He RL, Yang J, Yau SS (22 May 2013).
1972:
1697:
1631:
1596:
1330:
1186:
remain attached to the mRNA as a way to reinitiate translation of a subsequent ORF.
1044:
1028:
979:
913:
734:
665:
528:
498:
464:
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416:
384:
372:
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304:
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186:
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113:
Group VII: double-stranded DNA viruses with an RNA intermediate in their life cycle
436:
2777:
2584:
1267:
was established and initially included the three RNA virus groups. A year later,
1091:, viruses contain both linear and circular genomes, varying from genus to genus.
110:
Group VI: single-stranded RNA viruses with a DNA intermediate in their life cycle
7170:
6949:
6935:
6848:
6704:
6494:
6407:
6400:
6343:
6249:
6170:
6053:
5970:
5936:
5862:
5824:
5805:
5741:
5734:
5565:
5425:
5114:
4942:
4842:
4737:
4587:
4569:
4545:
4408:
4386:
4212:
4190:
4158:
4099:
3888:
3800:
3662:
3612:
3588:
2517:
1054:
1024:
688:
595:
319:
3478:
Viruses: Molecular Biology, Host Interactions and Applications to Biotechnology
3424:
Viruses: Molecular Biology, Host Interactions and Applications to Biotechnology
3324:
International Committee on Taxonomy of Viruses Executive Committee (May 2020).
2285:
1665:) for classification of filamentous archaeal viruses with linear dsDNA genomes"
7042:
6998:
6892:
6653:
5462:
5452:
5204:
4804:
4131:
3881:
3341:
1198:
1101:
1032:
927:
475:
315:
7238:
7109:
6643:
6599:
5673:
5350:
5322:
4761:
3601:
3573:
1214:
871:
611:
577:
563:
483:
199:
168:
160:
3359:
3018:
3000:
2796:
2737:
2655:
2603:
2535:
2414:
2293:
3518:
3303:
2719:
2637:
2502:"Origins and Evolution of Viruses of Eukaryotes: The Ultimate Modularity"
2396:
1148:
875:
708:+ssRNA, which may be mRNA, enters the main cytoplasmic area of the cell.
323:
468:
cleaves the positive strand again to create a standalone genome that is
1164:
1163:
initiation factor, and initiation at a CUG or other start codon with a
1144:
785:
260:
7365:
1302:
exclusively contains dsDNA viruses, although some proposed members of
1298:
primarily contains ssDNA viruses but also contains dsDNA viruses, and
789:
582:
65:
3135:
3133:
3094:
3092:
3070:
2931:
2480:
2478:
2476:
2454:
2428:
2337:
2320:
2318:
2200:
2174:
2148:
2096:
2058:
2026:
2024:
2022:
2000:
1900:
1871:
1829:
1827:
1788:
1759:
1564:
1538:
1512:
1486:
1431:
3111:
2905:
2886:
2867:
2848:
2829:
2810:
2676:
1382:
814:
only contains -ssRNA viruses, so "-ssRNA virus" is synonymous with
52:
Baltimore classification also closely corresponds to the manner of
1231:
1115:
965:
683:
606:
435:
140:
135:
26:
1928:
1926:
1924:
1922:
2566:
Sicard A, Michalakis Y, Gutierrez S, Blanc S (3 November 2016).
1210:
1160:
1019:, alternative splicing acts as a way to regulate early and late
7222:
7093:
6583:
5657:
5334:
4734:
3585:
3533:
934:
dsDNA-RT viruses are, like ssRNA-RT, all included in the class
921:
Group VII: double-stranded DNA viruses with an RNA intermediate
830:, whose members instead obtain caps on mRNA via cap snatching.
642:
dsRNA viruses are classified into two phyla within the kingdom
1133:
987:
1259:
In two votes in 2018 and 2019, a 15-rank system ranging from
1031:
cannot be transported out of the nucleus, hepadnaviruses and
857:
Group VI: single-stranded RNA viruses with a DNA intermediate
2379:
Teycheney PY, Tristem M, Koonin EV, Kuhn JH (15 June 2018).
1732:) for four families of arthropod-specific large DNA viruses"
1240:
Baltimore classification was proposed in 1971 by virologist
1104:, RNA editing improves the ability to adapt to their hosts.
71:
Baltimore classification was created in 1971 by virologist
2549:
2547:
2545:
2240:
2238:
2236:
2234:
2201:"Ambisense transcription in negative stranded RNA viruses"
2041:
2039:
1814:
1812:
1810:
1358:
1356:
1120:
Life cycle of some viruses classified by Baltimore group:
1469:
1467:
1465:
1414:
1412:
1410:
1408:
1406:
1404:
1191:
Non-canonical elongation and termination of translation:
970:
Structure of some viruses classified by Baltimore group:
49:
of a single-stranded RNA genome is positive or negative.
874:
into the DNA of the host cell, where it is now called a
482:(RHR), which is similar to RCR. Parvovirus genomes have
244:, and tailless or non-tailed dsDNA viruses of the realm
3380:. International Committee on Taxonomy of Viruses (ICTV)
1978:. International Committee on Taxonomy of Viruses (ICTV)
1943:
1941:
1737:. International Committee on Taxonomy of Viruses (ICTV)
1670:. International Committee on Taxonomy of Viruses (ICTV)
1063:
likewise vary between being monopartite and bipartite.
2753:"Real Time Classification of Viruses in 12 Dimensions"
7354:
545:
is unassigned to a realm but is a proposed member of
478:
contain linear ssDNA genomes that are replicated via
1081:
viruses are typically circular. In the dsDNA family
680:
Group IV: positive sense single-stranded RNA viruses
104:
Group IV: positive sense single-stranded RNA viruses
7302:
7280:
7269:
7258:
7247:
7236:
7151:
7140:
7129:
7118:
7107:
7080:
7030:
7019:
6923:
6873:
6862:
6836:
6825:
6792:
6781:
6678:
6667:
6641:
6630:
6619:
6608:
6597:
6534:
6504:
6482:
6471:
6431:
6360:
6247:
6236:
6225:
6195:
6184:
6158:
6147:
6121:
6110:
6077:
6034:
6023:
6012:
5968:
5957:
5924:
5913:
5887:
5876:
5822:
5758:
5715:
5704:
5693:
5682:
5671:
5597:
5539:
5528:
5517:
5487:
5476:
5450:
5439:
5392:
5381:
5370:
5359:
5348:
5321:
5273:
5235:
5224:
5213:
5202:
5161:
5150:
5124:
5095:
5073:
5051:
5029:
5018:
5007:
4977:
4966:
4940:
4929:
4918:
4907:
4873:
4862:
4851:
4840:
4792:
4781:
4770:
4759:
4748:
4601:
4579:
4543:
4532:
4525:
4473:
4462:
4451:
4440:
4406:
4384:
4362:
4340:
4329:
4303:
4292:
4266:
4255:
4244:
4200:
4178:
4156:
4145:
4119:
4097:
4086:
4075:
4064:
4053:
4015:
3993:
3982:
3971:
3960:
3949:
3820:
3809:
3798:
3754:
3743:
3732:
3721:
3710:
3672:
3643:
3632:
3621:
3610:
3599:
3572:
2175:"Negative-stranded RNA virus polymerase stuttering"
938:. Two families of dsDNA-RT viruses are recognized:
758:
Group V: negative sense single-stranded RNA viruses
107:
Group V: negative sense single-stranded RNA viruses
16:
Virus classification system made by David Baltimore
907:, which belongs to Group VII, all members of the
314:are dsDNA viruses. Viruses in this realm include
240:, usually the tailed bacteriophages of the order
117:Baltimore classification is chiefly based on the
3229:
3139:
3098:
3057:
2704:"Origins and Evolution of the Global RNA Virome"
2484:
2324:
2030:
1932:
1846:Kerr J, Cotmore S, Bloom ME (25 November 2005).
1833:
1703:. International Committee on Taxonomy of Viruses
1637:. International Committee on Taxonomy of Viruses
1602:. International Committee on Taxonomy of Viruses
1456:
259:are dsDNA viruses. Viruses in this realm infect
2568:"The Strange Lifestyle of Multipartite Viruses"
2259:"Classify viruses - the gain is worth the pain"
885:ssRNA-RT viruses are all included in the class
2622:"ICTV Virus Taxonomy Profile: Pleolipoviridae"
1250:International Committee on Taxonomy of Viruses
3545:
1383:"Viral replication/transcription/translation"
8:
2500:Koonin EV, Dolja VV, Krupovic M (May 2015).
589:. Some of the latter category, such as the
2615:
2613:
2373:
2371:
2097:"Negative-stranded RNA virus transcription"
190:. Many have yet to be assigned to a realm.
7277:
7266:
7255:
7244:
7233:
7219:
7148:
7137:
7126:
7115:
7104:
7090:
7086:
7027:
6870:
6833:
6789:
6675:
6638:
6627:
6616:
6605:
6594:
6580:
6479:
6244:
6233:
6192:
6155:
6118:
6031:
6020:
5965:
5921:
5884:
5712:
5701:
5690:
5679:
5668:
5654:
5536:
5525:
5484:
5447:
5389:
5378:
5367:
5356:
5345:
5331:
5327:
5232:
5221:
5210:
5158:
5026:
5015:
4974:
4937:
4926:
4915:
4870:
4859:
4848:
4789:
4778:
4767:
4756:
4745:
4731:
4540:
4529:
4470:
4459:
4448:
4337:
4300:
4263:
4252:
4153:
4094:
4083:
4072:
4061:
3990:
3979:
3968:
3957:
3817:
3806:
3751:
3740:
3729:
3718:
3640:
3629:
3618:
3607:
3596:
3582:
3578:
3552:
3538:
3530:
2985:"Non-canonical translation in RNA viruses"
1783:
1781:
3459:Molecular and Cellular Biology of Viruses
3349:
3293:
3008:
2786:
2776:
2727:
2645:
2593:
2583:
2525:
2495:
2493:
2404:
2149:"Negative stranded RNA virus replication"
2001:"Positive stranded RNA virus replication"
1995:
1993:
1895:
1893:
1754:
1752:
1590:
1588:
1586:
1209:but are instead partially recognized by
954:, which is the sole family in the order
738:are +ssRNA viruses, excluding the class
41:(RNA), whether the genome is single- or
7361:
3319:
3317:
3315:
3313:
1966:
1964:
1962:
1960:
1958:
1956:
1901:"Double-stranded RNA virus replication"
1691:
1689:
1687:
1685:
1654:
1652:
1625:
1623:
1621:
1619:
1617:
1362:
1352:
1321:
1201:forward or backward during translation.
2553:
2244:
2045:
1947:
1818:
1473:
1155:Non-canonical translation initiation:
1023:during different stages of infection.
603:Group III: double-stranded RNA viruses
101:Group III: double-stranded RNA viruses
2429:"dsDNA(RT) replication/transcription"
2338:"ssRNA(RT) replication/transcription"
1539:"DNA strand displacement replication"
432:Group II: single-stranded DNA viruses
98:Group II: single-stranded DNA viruses
7:
3270:"Expression of animal virus genomes"
2362:
2225:
2083:
1418:
1377:
1375:
1373:
1371:
1294:exclusively contains dsDNA viruses,
194:Group I: double-stranded DNA viruses
95:Group I: double-stranded DNA viruses
7387:Viruses by Baltimore classification
3476:Rampersad, S.; Tennant, P. (2018).
3249:. Swiss Institute of Bioinformatics
3211:. Swiss Institute of Bioinformatics
3185:. Swiss Institute of Bioinformatics
3159:. Swiss Institute of Bioinformatics
3118:. Swiss Institute of Bioinformatics
3077:. Swiss Institute of Bioinformatics
3039:. Swiss Institute of Bioinformatics
2964:. Swiss Institute of Bioinformatics
2938:. Swiss Institute of Bioinformatics
2912:. Swiss Institute of Bioinformatics
2893:. Swiss Institute of Bioinformatics
2874:. Swiss Institute of Bioinformatics
2855:. Swiss Institute of Bioinformatics
2836:. Swiss Institute of Bioinformatics
2817:. Swiss Institute of Bioinformatics
2683:. Swiss Institute of Bioinformatics
2461:. Swiss Institute of Bioinformatics
2435:. Swiss Institute of Bioinformatics
2344:. Swiss Institute of Bioinformatics
2207:. Swiss Institute of Bioinformatics
2181:. Swiss Institute of Bioinformatics
2155:. Swiss Institute of Bioinformatics
2129:. Swiss Institute of Bioinformatics
2103:. Swiss Institute of Bioinformatics
2065:. Swiss Institute of Bioinformatics
2007:. Swiss Institute of Bioinformatics
1907:. Swiss Institute of Bioinformatics
1878:. Swiss Institute of Bioinformatics
1795:. Swiss Institute of Bioinformatics
1766:. Swiss Institute of Bioinformatics
1571:. Swiss Institute of Bioinformatics
1545:. Swiss Institute of Bioinformatics
1519:. Swiss Institute of Bioinformatics
1493:. Swiss Institute of Bioinformatics
1438:. Swiss Institute of Bioinformatics
1389:. Swiss Institute of Bioinformatics
784:, during which RdRp transcribes an
744:, whose members have dsRNA genomes.
214:transcription preinitiation complex
3486:10.1016/B978-0-12-811257-1.00003-6
3432:10.1016/B978-0-12-811257-1.00002-4
2983:Firth AE, Brierley I (July 2012).
1513:"dsDNA rolling circle replication"
1246:Expression of Animal Virus Genomes
14:
3033:"Viral initiation of translation"
1487:"dsDNA bidirectional replication"
1159:translation in the absence of an
7364:
3517:
3405:Principles of Molecular Virology
3205:"RNA suppression of termination"
513:, all members except viruses in
1850:. CRC Press. pp. 171–185.
1432:"dsDNA templated transcription"
1275:and RNA viruses in the kingdom
581:. The exceptions are generally
3407:. Elsevier. pp. 122–127.
3286:10.1128/MMBR.35.3.235-241.1971
3153:"RNA termination-reinitiation"
2906:"Reverse-transcribing viruses"
2059:"Subgenomic RNA transcription"
1108:expressed from a single gene.
822:is divided into two subphyla:
593:, are classified in the realm
295:are dsDNA viruses. Viruses in
1:
2887:"Negative strand RNA viruses"
2868:"Positive strand RNA viruses"
1789:"Rolling hairpin replication"
1147:are synthesized from mRNA by
944:, which belongs to the order
3480:. Elsevier. pp. 55–82.
3426:. Elsevier. pp. 35–46.
3230:Rampersad & Tennant 2018
3140:Rampersad & Tennant 2018
3099:Rampersad & Tennant 2018
3058:Rampersad & Tennant 2018
2778:10.1371/journal.pone.0064328
2585:10.1371/journal.ppat.1005819
2485:Rampersad & Tennant 2018
2325:Rampersad & Tennant 2018
2031:Rampersad & Tennant 2018
1933:Rampersad & Tennant 2018
1834:Rampersad & Tennant 2018
1457:Rampersad & Tennant 2018
1184:40S subunits of the ribosome
834:Reverse transcribing viruses
2849:"Double strand RNA viruses"
2830:"Single strand DNA viruses"
2811:"Double strand DNA viruses"
2518:10.1016/j.virol.2015.02.039
1565:"Replicative transposition"
962:Multi-group characteristics
669:, all members of the class
480:rolling hairpin replication
7413:
2286:10.1038/d41586-019-00599-8
1728:)including one new order (
1076:Linear vs circular genomes
837:
761:
557:
447:
197:
158:
7392:Systems of virus taxonomy
7232:
7218:
7103:
7089:
6593:
6579:
5667:
5653:
5344:
5330:
4744:
4730:
3595:
3581:
3342:10.1038/s41564-020-0709-x
3179:"Ribosomal frameshifting"
862:through a process called
532:are ssDNA virus families.
230:replicative transposition
29:based on their manner of
3524:Baltimore classification
1143:is the process by which
520:The unassigned families
20:Baltimore classification
1724:"Create one new class (
1661:"Create one new realm (
1195:Ribosomal frameshifting
541:contain ssDNA genomes.
299:constitute two groups:
289:, members of the class
3001:10.1099/vir.0.042499-0
2455:"Alternative splicing"
1760:"ssDNA Rolling circle"
1237:
1137:
991:
917:are ssRNA-RT viruses.
692:
615:
535:Viruses in the family
445:
146:
2720:10.1128/mBio.02329-18
2391:(12): e00515–e00518.
1312:, are ssDNA viruses.
1235:
1119:
969:
868:reverse transcriptase
864:reverse transcription
782:polymerase stuttering
687:
610:
439:
276:and herpesviruses in
144:
85:deoxyribonucleic acid
35:deoxyribonucleic acid
3565:virus classification
3526:at Wikimedia Commons
3457:Lostroh, P. (2019).
3397:General bibliography
3268:Baltimore D (1971).
3243:"Ribosomal skipping"
2932:"Sphaerolipoviridae"
2638:10.1099/jgv.0.000972
2397:10.1128/JVI.00515-18
1306:, namely the family
1087:, and in the family
1004:Alternative splicing
999:Alternative splicing
22:is a system used to
6547:Permutotetraviridae
4306:Orthopolintovirales
3781:Malacoherpesviridae
3461:. Garland Science.
3422:Fermin, G. (2018).
2995:(Pt 7): 1385–1409.
2769:2013PLoSO...864328Y
2278:2019Natur.566..318K
2228:, pp. 154–156.
2086:, pp. 151–154.
1421:, pp. 122–127.
1039:Genome segmentation
753:are +ssRNA viruses.
729:are +ssRNA viruses.
691:are +ssRNA viruses.
4865:Malgrandaviricetes
4500:Sphaerolipoviridae
4486:Matshushitaviridae
4078:Nucleocytoviricota
1244:in a paper titled
1238:
1177:Ribosomal shunting
1138:
1084:Sphaerolipoviridae
992:
878:. The host cell's
693:
675:are dsRNA viruses.
660:are dsRNA viruses.
637:open reading frame
616:
614:are dsRNA viruses.
517:are ssDNA viruses.
446:
444:is an ssDNA virus.
147:
45:, and whether the
7352:
7351:
7348:
7347:
7344:
7343:
7340:
7339:
7336:
7335:
7332:
7331:
7328:
7327:
7324:
7323:
7214:
7213:
7210:
7209:
7206:
7205:
7202:
7201:
7198:
7197:
7194:
7193:
7076:
7075:
7072:
7071:
7068:
7067:
7064:
7063:
7060:
7059:
7056:
7055:
7052:
7051:
7022:Yunchangviricetes
7015:
7014:
6858:
6857:
6821:
6820:
6777:
6776:
6663:
6662:
6575:
6574:
6571:
6570:
6567:
6566:
6563:
6562:
6530:
6529:
6526:
6525:
6467:
6466:
6221:
6220:
6217:
6216:
6180:
6179:
6143:
6142:
6106:
6105:
6008:
6007:
6004:
6003:
5981:Carmotetraviridae
5953:
5952:
5909:
5908:
5872:
5871:
5856:Gammaflexiviridae
5849:Deltaflexiviridae
5835:Alphaflexiviridae
5728:Alphatetraviridae
5649:
5648:
5645:
5644:
5641:
5640:
5637:
5636:
5593:
5592:
5589:
5588:
5513:
5512:
5509:
5508:
5472:
5471:
5435:
5434:
5384:Chrymotiviricetes
5317:
5316:
5313:
5312:
5309:
5308:
5269:
5268:
5265:
5264:
5261:
5260:
5257:
5256:
5198:
5197:
5194:
5193:
5190:
5189:
5146:
5145:
5042:Bacilladnaviridae
5003:
5002:
4999:
4998:
4962:
4961:
4903:
4902:
4899:
4898:
4895:
4894:
4836:
4835:
4832:
4831:
4828:
4827:
4726:
4725:
4722:
4721:
4718:
4717:
4684:Portogloboviridae
4597:
4596:
4521:
4520:
4517:
4516:
4513:
4512:
4509:
4508:
4436:
4435:
4432:
4431:
4428:
4427:
4325:
4324:
4288:
4287:
4247:Preplasmiviricota
4240:
4239:
4236:
4235:
4141:
4140:
4049:
4048:
4045:
4044:
4041:
4040:
4037:
4036:
3945:
3944:
3941:
3940:
3937:
3936:
3933:
3932:
3840:Autographiviridae
3794:
3793:
3790:
3789:
3767:Alloherpesviridae
3706:
3705:
3702:
3701:
3698:
3697:
3694:
3693:
3522:Media related to
3403:Cann, A. (2015).
3232:, pp. 72–73.
3142:, pp. 74–75.
3112:"Ribosomal shunt"
3101:, pp. 73–74.
3060:, pp. 69–70.
2632:(12): 2916–2917.
2556:, pp. 35–46.
2487:, pp. 71–72.
2327:, pp. 63–64.
2272:(7744): 318–320.
2247:, pp. 45–46.
2048:, pp. 43–44.
2033:, pp. 64–65.
1836:, pp. 61–62.
1821:, pp. 40–41.
1476:, pp. 36–40.
1365:, pp. 11–13.
1316:Explanatory notes
880:RNA polymerase II
828:Polyploviricotina
591:hepatitis D virus
442:canine parvovirus
403:Portogloboviridae
7404:
7369:
7368:
7360:
7278:
7267:
7256:
7245:
7234:
7226:dsDNA-RT viruses
7220:
7149:
7138:
7127:
7116:
7105:
7097:ssRNA-RT viruses
7091:
7087:
7028:
6871:
6839:Serpentovirales
6834:
6812:Orthomyxoviridae
6790:
6740:Peribunyaviridae
6676:
6639:
6633:Chunqiuviricetes
6628:
6617:
6606:
6595:
6587:(–)ssRNA viruses
6581:
6480:
6323:Nanghoshaviridae
6245:
6234:
6208:Botourmiaviridae
6193:
6156:
6150:Howeltoviricetes
6119:
6113:Amabiliviricetes
6032:
6021:
5966:
5922:
5885:
5842:Betaflexiviridae
5713:
5702:
5691:
5680:
5669:
5661:(+)ssRNA viruses
5655:
5573:Picobirnaviridae
5537:
5531:Duplopiviricetes
5526:
5485:
5479:Vidaverviricetes
5448:
5442:Resentoviricetes
5412:Megabirnaviridae
5390:
5379:
5373:Duplornaviricota
5368:
5357:
5346:
5332:
5328:
5233:
5227:Huolimaviricetes
5222:
5211:
5159:
5153:Repensiviricetes
5027:
5016:
5010:Cressdnaviricota
4975:
4938:
4927:
4916:
4871:
4860:
4849:
4790:
4779:
4768:
4757:
4746:
4732:
4541:
4530:
4471:
4460:
4449:
4338:
4332:Tectiliviricetes
4301:
4295:Polintoviricetes
4269:Priklausovirales
4264:
4253:
4227:Marseilleviridae
4154:
4095:
4084:
4073:
4062:
4028:Papillomaviridae
4018:Zurhausenvirales
3991:
3980:
3969:
3958:
3833:Ackermannviridae
3818:
3807:
3752:
3741:
3730:
3719:
3685:Tristromaviridae
3656:Lipothrixviridae
3641:
3630:
3619:
3608:
3597:
3583:
3579:
3554:
3547:
3540:
3531:
3521:
3507:
3472:
3453:
3418:
3390:
3389:
3387:
3385:
3379:
3370:
3364:
3363:
3353:
3321:
3308:
3307:
3297:
3265:
3259:
3258:
3256:
3254:
3239:
3233:
3227:
3221:
3220:
3218:
3216:
3201:
3195:
3194:
3192:
3190:
3175:
3169:
3168:
3166:
3164:
3149:
3143:
3137:
3128:
3127:
3125:
3123:
3108:
3102:
3096:
3087:
3086:
3084:
3082:
3071:"Leaky scanning"
3067:
3061:
3055:
3049:
3048:
3046:
3044:
3029:
3023:
3022:
3012:
2980:
2974:
2973:
2971:
2969:
2954:
2948:
2947:
2945:
2943:
2928:
2922:
2921:
2919:
2917:
2902:
2900:
2898:
2883:
2881:
2879:
2864:
2862:
2860:
2845:
2843:
2841:
2826:
2824:
2822:
2807:
2801:
2800:
2790:
2780:
2748:
2742:
2741:
2731:
2714:(6): e02329-18.
2699:
2693:
2692:
2690:
2688:
2673:
2667:
2666:
2664:
2662:
2649:
2617:
2608:
2607:
2597:
2587:
2578:(11): e1005819.
2563:
2557:
2551:
2540:
2539:
2529:
2497:
2488:
2482:
2471:
2470:
2468:
2466:
2451:
2445:
2444:
2442:
2440:
2425:
2419:
2418:
2408:
2375:
2366:
2360:
2354:
2353:
2351:
2349:
2334:
2328:
2322:
2313:
2312:
2310:
2308:
2263:
2254:
2248:
2242:
2229:
2223:
2217:
2216:
2214:
2212:
2197:
2191:
2190:
2188:
2186:
2171:
2165:
2164:
2162:
2160:
2145:
2139:
2138:
2136:
2134:
2119:
2113:
2112:
2110:
2108:
2093:
2087:
2081:
2075:
2074:
2072:
2070:
2055:
2049:
2043:
2034:
2028:
2017:
2016:
2014:
2012:
1997:
1988:
1987:
1985:
1983:
1977:
1968:
1951:
1945:
1936:
1930:
1917:
1916:
1914:
1912:
1897:
1888:
1887:
1885:
1883:
1868:
1862:
1861:
1843:
1837:
1831:
1822:
1816:
1805:
1804:
1802:
1800:
1785:
1776:
1775:
1773:
1771:
1756:
1747:
1746:
1744:
1742:
1736:
1719:
1713:
1712:
1710:
1708:
1702:
1693:
1680:
1679:
1677:
1675:
1669:
1656:
1647:
1646:
1644:
1642:
1636:
1627:
1612:
1611:
1609:
1607:
1601:
1592:
1581:
1580:
1578:
1576:
1561:
1555:
1554:
1552:
1550:
1535:
1529:
1528:
1526:
1524:
1509:
1503:
1502:
1500:
1498:
1483:
1477:
1471:
1460:
1454:
1448:
1447:
1445:
1443:
1428:
1422:
1416:
1399:
1398:
1396:
1394:
1379:
1366:
1360:
1335:
1326:
1016:Papillomaviridae
1010:Orthomyxoviridae
741:Duplopiviricetes
702:leader sequences
672:Duplopiviricetes
657:Duplornaviricota
301:papillomaviruses
128:viral life cycle
89:ribonucleic acid
39:ribonucleic acid
7412:
7411:
7407:
7406:
7405:
7403:
7402:
7401:
7377:
7376:
7375:
7363:
7355:
7353:
7320:
7298:
7272:Revtraviricetes
7228:
7190:
7143:Revtraviricetes
7099:
7048:
7011:
6978:Paramyxoviridae
6926:Mononegavirales
6919:
6900:Crepuscuviridae
6854:
6817:
6795:Articulavirales
6784:Insthoviricetes
6773:
6659:
6622:Negarnaviricota
6589:
6559:
6522:
6500:
6474:Stelpaviricetes
6463:
6427:
6422:Polycipiviridae
6415:Dicistroviridae
6356:
6239:Pisoniviricetes
6213:
6176:
6139:
6102:
6073:
6000:
5960:Tolucaviricetes
5949:
5905:
5890:Amarillovirales
5868:
5818:
5778:Closteroviridae
5761:Martellivirales
5754:
5696:Kitrinoviricota
5663:
5633:
5617:Polymycoviridae
5585:
5505:
5468:
5431:
5340:
5305:
5293:Finnlakeviridae
5253:
5248:Pleolipoviridae
5186:
5142:
5120:
5091:
5069:
5047:
4995:
4969:Quintoviricetes
4958:
4891:
4824:
4740:
4714:
4593:
4575:
4505:
4476:Halopanivirales
4424:
4402:
4380:
4358:
4321:
4284:
4258:Maveriviricetes
4232:
4196:
4174:
4169:Phycodnaviridae
4137:
4115:
4089:Pokkesviricetes
4033:
4011:
3985:Papovaviricetes
3929:
3896:Rountreeviridae
3786:
3690:
3668:
3591:
3568:
3558:
3514:
3496:
3475:
3469:
3456:
3442:
3421:
3415:
3402:
3399:
3394:
3393:
3383:
3381:
3377:
3372:
3371:
3367:
3323:
3322:
3311:
3267:
3266:
3262:
3252:
3250:
3241:
3240:
3236:
3228:
3224:
3214:
3212:
3203:
3202:
3198:
3188:
3186:
3177:
3176:
3172:
3162:
3160:
3151:
3150:
3146:
3138:
3131:
3121:
3119:
3110:
3109:
3105:
3097:
3090:
3080:
3078:
3069:
3068:
3064:
3056:
3052:
3042:
3040:
3031:
3030:
3026:
2982:
2981:
2977:
2967:
2965:
2956:
2955:
2951:
2941:
2939:
2930:
2929:
2925:
2915:
2913:
2904:
2903:
2896:
2894:
2885:
2884:
2877:
2875:
2866:
2865:
2858:
2856:
2847:
2846:
2839:
2837:
2828:
2827:
2820:
2818:
2809:
2808:
2804:
2750:
2749:
2745:
2701:
2700:
2696:
2686:
2684:
2677:"Geminiviridae"
2675:
2674:
2670:
2660:
2658:
2619:
2618:
2611:
2565:
2564:
2560:
2552:
2543:
2499:
2498:
2491:
2483:
2474:
2464:
2462:
2453:
2452:
2448:
2438:
2436:
2427:
2426:
2422:
2377:
2376:
2369:
2361:
2357:
2347:
2345:
2336:
2335:
2331:
2323:
2316:
2306:
2304:
2261:
2256:
2255:
2251:
2243:
2232:
2224:
2220:
2210:
2208:
2199:
2198:
2194:
2184:
2182:
2173:
2172:
2168:
2158:
2156:
2147:
2146:
2142:
2132:
2130:
2123:"Cap snatching"
2121:
2120:
2116:
2106:
2104:
2095:
2094:
2090:
2082:
2078:
2068:
2066:
2057:
2056:
2052:
2044:
2037:
2029:
2020:
2010:
2008:
1999:
1998:
1991:
1981:
1979:
1975:
1970:
1969:
1954:
1946:
1939:
1931:
1920:
1910:
1908:
1899:
1898:
1891:
1881:
1879:
1870:
1869:
1865:
1858:
1845:
1844:
1840:
1832:
1825:
1817:
1808:
1798:
1796:
1787:
1786:
1779:
1769:
1767:
1758:
1757:
1750:
1740:
1738:
1734:
1721:
1720:
1716:
1706:
1704:
1700:
1695:
1694:
1683:
1673:
1671:
1667:
1658:
1657:
1650:
1640:
1638:
1634:
1629:
1628:
1615:
1605:
1603:
1599:
1594:
1593:
1584:
1574:
1572:
1563:
1562:
1558:
1548:
1546:
1537:
1536:
1532:
1522:
1520:
1511:
1510:
1506:
1496:
1494:
1485:
1484:
1480:
1472:
1463:
1455:
1451:
1441:
1439:
1430:
1429:
1425:
1417:
1402:
1392:
1390:
1381:
1380:
1369:
1361:
1354:
1349:
1344:
1339:
1338:
1327:
1323:
1318:
1309:Finnlakeviridae
1242:David Baltimore
1236:David Baltimore
1230:
1132:(group V), and
1114:
1097:
1089:Pleolipoviridae
1078:
1069:
1050:Pleolipoviridae
1041:
1021:gene expression
1001:
986:(group V), and
964:
936:Revtraviricetes
923:
909:Revtraviricetes
888:Revtraviricetes
859:
842:
840:Revtraviricetes
836:
824:Haploviricotina
820:Negarnaviricota
816:Negarnaviricota
812:Negarnaviricota
802:in the kingdom
799:Negarnaviricota
766:
764:Negarnaviricota
760:
750:Kitrinoviricota
747:All viruses in
732:All viruses in
723:All viruses in
682:
654:All viruses in
605:
587:subviral agents
566:
558:Main articles:
556:
543:Finnlakeviridae
538:Finnlakeviridae
515:Papovaviricetes
452:
434:
310:All viruses in
297:Papovaviricetes
292:Papovaviricetes
266:All viruses in
255:All viruses in
222:polyadenylation
210:
196:
163:
157:
152:
81:
73:David Baltimore
43:double-stranded
17:
12:
11:
5:
7410:
7408:
7400:
7399:
7394:
7389:
7379:
7378:
7374:
7373:
7350:
7349:
7346:
7345:
7342:
7341:
7338:
7337:
7334:
7333:
7330:
7329:
7326:
7325:
7322:
7321:
7319:
7318:
7315:Caulimoviridae
7310:
7308:
7300:
7299:
7297:
7296:
7293:Hepadnaviridae
7288:
7286:
7275:
7264:
7261:Artverviricota
7253:
7242:
7230:
7229:
7223:
7216:
7215:
7212:
7211:
7208:
7207:
7204:
7203:
7200:
7199:
7196:
7195:
7192:
7191:
7189:
7188:
7181:
7174:
7167:
7159:
7157:
7146:
7135:
7132:Artverviricota
7124:
7113:
7101:
7100:
7094:
7084:
7078:
7077:
7074:
7073:
7070:
7069:
7066:
7065:
7062:
7061:
7058:
7057:
7054:
7053:
7050:
7049:
7047:
7046:
7038:
7036:
7033:Goujianvirales
7025:
7017:
7016:
7013:
7012:
7010:
7009:
7002:
6995:
6988:
6981:
6974:
6967:
6960:
6953:
6946:
6939:
6931:
6929:
6921:
6920:
6918:
6917:
6910:
6903:
6896:
6889:
6881:
6879:
6876:Jingchuvirales
6868:
6865:Monjiviricetes
6860:
6859:
6856:
6855:
6853:
6852:
6844:
6842:
6831:
6828:Milneviricetes
6823:
6822:
6819:
6818:
6816:
6815:
6808:
6800:
6798:
6787:
6779:
6778:
6775:
6774:
6772:
6771:
6764:
6757:
6750:
6743:
6736:
6729:
6722:
6719:Leishbuviridae
6715:
6708:
6701:
6694:
6686:
6684:
6673:
6670:Ellioviricetes
6665:
6664:
6661:
6660:
6658:
6657:
6649:
6647:
6636:
6625:
6614:
6603:
6591:
6590:
6584:
6577:
6576:
6573:
6572:
6569:
6568:
6565:
6564:
6561:
6560:
6558:
6557:
6554:Sarthroviridae
6550:
6538:
6536:
6532:
6531:
6528:
6527:
6524:
6523:
6521:
6520:
6512:
6510:
6502:
6501:
6499:
6498:
6490:
6488:
6477:
6469:
6468:
6465:
6464:
6462:
6461:
6454:
6447:
6444:Alvernaviridae
6439:
6437:
6429:
6428:
6426:
6425:
6418:
6411:
6404:
6397:
6390:
6387:Solinviviridae
6383:
6376:
6373:Picornaviridae
6368:
6366:
6363:Picornavirales
6358:
6357:
6355:
6354:
6347:
6340:
6333:
6330:Nanhypoviridae
6326:
6319:
6312:
6305:
6302:Medioniviridae
6298:
6291:
6284:
6277:
6274:Cremegaviridae
6270:
6263:
6255:
6253:
6242:
6231:
6223:
6222:
6219:
6218:
6215:
6214:
6212:
6211:
6203:
6201:
6190:
6182:
6181:
6178:
6177:
6175:
6174:
6166:
6164:
6153:
6145:
6144:
6141:
6140:
6138:
6137:
6129:
6127:
6124:Wolframvirales
6116:
6108:
6107:
6104:
6103:
6101:
6100:
6093:
6085:
6083:
6075:
6074:
6072:
6071:
6064:
6057:
6050:
6042:
6040:
6029:
6018:
6010:
6009:
6006:
6005:
6002:
6001:
5999:
5998:
5991:
5984:
5976:
5974:
5963:
5955:
5954:
5951:
5950:
5948:
5947:
5944:Sinhaliviridae
5940:
5932:
5930:
5919:
5916:Magsaviricetes
5911:
5910:
5907:
5906:
5904:
5903:
5895:
5893:
5882:
5879:Flasuviricetes
5874:
5873:
5870:
5869:
5867:
5866:
5859:
5852:
5845:
5838:
5830:
5828:
5820:
5819:
5817:
5816:
5809:
5802:
5795:
5788:
5785:Endornaviridae
5781:
5774:
5766:
5764:
5756:
5755:
5753:
5752:
5745:
5738:
5731:
5723:
5721:
5710:
5699:
5688:
5677:
5665:
5664:
5658:
5651:
5650:
5647:
5646:
5643:
5642:
5639:
5638:
5635:
5634:
5632:
5631:
5628:Botybirnavirus
5620:
5613:
5601:
5599:
5595:
5594:
5591:
5590:
5587:
5586:
5584:
5583:
5580:Partitiviridae
5576:
5569:
5562:
5559:Curvulaviridae
5555:
5547:
5545:
5534:
5523:
5515:
5514:
5511:
5510:
5507:
5506:
5504:
5503:
5495:
5493:
5482:
5474:
5473:
5470:
5469:
5467:
5466:
5458:
5456:
5445:
5437:
5436:
5433:
5432:
5430:
5429:
5422:
5415:
5408:
5400:
5398:
5387:
5376:
5365:
5354:
5342:
5341:
5335:
5325:
5319:
5318:
5315:
5314:
5311:
5310:
5307:
5306:
5304:
5303:
5296:
5289:
5277:
5275:
5271:
5270:
5267:
5266:
5263:
5262:
5259:
5258:
5255:
5254:
5252:
5251:
5243:
5241:
5230:
5219:
5208:
5200:
5199:
5196:
5195:
5192:
5191:
5188:
5187:
5185:
5184:
5177:
5169:
5167:
5164:Geplafuvirales
5156:
5148:
5147:
5144:
5143:
5141:
5140:
5137:Redondoviridae
5132:
5130:
5122:
5121:
5119:
5118:
5111:
5103:
5101:
5093:
5092:
5090:
5089:
5081:
5079:
5071:
5070:
5068:
5067:
5059:
5057:
5049:
5048:
5046:
5045:
5037:
5035:
5024:
5013:
5005:
5004:
5001:
5000:
4997:
4996:
4994:
4993:
4985:
4983:
4972:
4964:
4963:
4960:
4959:
4957:
4956:
4948:
4946:
4935:
4924:
4913:
4905:
4904:
4901:
4900:
4897:
4896:
4893:
4892:
4890:
4889:
4881:
4879:
4868:
4857:
4846:
4838:
4837:
4834:
4833:
4830:
4829:
4826:
4825:
4823:
4822:
4819:Plectroviridae
4815:
4812:Paulinoviridae
4808:
4800:
4798:
4787:
4784:Faserviricetes
4776:
4773:Hofneiviricota
4765:
4754:
4742:
4741:
4735:
4728:
4727:
4724:
4723:
4720:
4719:
4716:
4715:
4713:
4712:
4705:
4694:
4687:
4680:
4677:Polydnaviridae
4673:
4666:
4659:
4652:
4645:
4642:Globuloviridae
4638:
4635:Fuselloviridae
4631:
4624:
4621:Bicaudaviridae
4617:
4614:Ampullaviridae
4605:
4603:
4599:
4598:
4595:
4594:
4592:
4591:
4583:
4581:
4577:
4576:
4574:
4573:
4566:
4563:Hytrosaviridae
4559:
4551:
4549:
4538:
4535:Naldaviricetes
4527:
4523:
4522:
4519:
4518:
4515:
4514:
4511:
4510:
4507:
4506:
4504:
4503:
4496:
4489:
4481:
4479:
4468:
4465:Laserviricetes
4457:
4454:Dividoviricota
4446:
4438:
4437:
4434:
4433:
4430:
4429:
4426:
4425:
4423:
4422:
4419:Corticoviridae
4414:
4412:
4404:
4403:
4401:
4400:
4392:
4390:
4382:
4381:
4379:
4378:
4370:
4368:
4360:
4359:
4357:
4356:
4348:
4346:
4335:
4327:
4326:
4323:
4322:
4320:
4319:
4311:
4309:
4298:
4290:
4289:
4286:
4285:
4283:
4282:
4274:
4272:
4261:
4250:
4242:
4241:
4238:
4237:
4234:
4233:
4231:
4230:
4223:
4216:
4208:
4206:
4203:Pimascovirales
4198:
4197:
4195:
4194:
4186:
4184:
4176:
4175:
4173:
4172:
4164:
4162:
4151:
4143:
4142:
4139:
4138:
4136:
4135:
4127:
4125:
4117:
4116:
4114:
4113:
4105:
4103:
4092:
4081:
4070:
4059:
4051:
4050:
4047:
4046:
4043:
4042:
4039:
4038:
4035:
4034:
4032:
4031:
4023:
4021:
4013:
4012:
4010:
4009:
4006:Polyomaviridae
4001:
3999:
3988:
3977:
3966:
3955:
3947:
3946:
3943:
3942:
3939:
3938:
3935:
3934:
3931:
3930:
3928:
3927:
3920:
3913:
3906:
3903:Salasmaviridae
3899:
3892:
3885:
3878:
3875:Herelleviridae
3871:
3864:
3861:Drexlerviridae
3857:
3854:Demerecviridae
3850:
3843:
3836:
3828:
3826:
3815:
3812:Caudoviricetes
3804:
3796:
3795:
3792:
3791:
3788:
3787:
3785:
3784:
3777:
3770:
3762:
3760:
3749:
3746:Herviviricetes
3738:
3727:
3724:Heunggongvirae
3716:
3708:
3707:
3704:
3703:
3700:
3699:
3696:
3695:
3692:
3691:
3689:
3688:
3680:
3678:
3670:
3669:
3667:
3666:
3659:
3651:
3649:
3646:Ligamenvirales
3638:
3627:
3616:
3605:
3593:
3592:
3586:
3576:
3570:
3569:
3559:
3557:
3556:
3549:
3542:
3534:
3528:
3527:
3513:
3512:External links
3510:
3509:
3508:
3495:978-0128112571
3494:
3473:
3468:978-0429664304
3467:
3454:
3441:978-0128112571
3440:
3419:
3414:978-0128019559
3413:
3398:
3395:
3392:
3391:
3365:
3336:(5): 668–674.
3309:
3280:(3): 235–241.
3260:
3234:
3222:
3196:
3170:
3144:
3129:
3103:
3088:
3062:
3050:
3024:
2975:
2949:
2923:
2802:
2743:
2694:
2668:
2609:
2558:
2541:
2489:
2472:
2446:
2420:
2367:
2365:, p. 156.
2355:
2329:
2314:
2249:
2230:
2218:
2192:
2166:
2140:
2114:
2088:
2076:
2050:
2035:
2018:
1989:
1952:
1937:
1918:
1889:
1872:"Bidnaviridae"
1863:
1856:
1838:
1823:
1806:
1777:
1748:
1726:Naldaviricetes
1714:
1681:
1648:
1613:
1582:
1556:
1530:
1504:
1478:
1461:
1449:
1423:
1400:
1367:
1351:
1350:
1348:
1345:
1343:
1340:
1337:
1336:
1320:
1319:
1317:
1314:
1229:
1226:
1225:
1224:
1223:
1222:
1218:
1207:release factor
1202:
1189:
1188:
1187:
1180:
1174:
1171:Leaky scanning
1168:
1113:
1110:
1096:
1093:
1077:
1074:
1068:
1065:
1040:
1037:
1000:
997:
963:
960:
951:Hepadnaviridae
941:Caulimoviridae
922:
919:
905:Caulimoviridae
858:
855:
838:Main article:
835:
832:
762:Main article:
759:
756:
755:
754:
745:
730:
681:
678:
677:
676:
661:
632:Endornaviridae
604:
601:
555:
552:
551:
550:
533:
518:
457:DNA polymerase
433:
430:
429:
428:
427:
426:
412:
397:Polydnaviridae
367:Globuloviridae
361:Fuselloviridae
349:Bicaudaviridae
343:Ampullaviridae
338:
335:Naldaviricetes
327:
308:
305:polyomaviruses
283:
264:
218:RNA polymerase
195:
192:
159:Main article:
156:
153:
151:
150:Classification
148:
132:viral envelope
115:
114:
111:
108:
105:
102:
99:
96:
80:
77:
15:
13:
10:
9:
6:
4:
3:
2:
7409:
7398:
7395:
7393:
7390:
7388:
7385:
7384:
7382:
7372:
7367:
7362:
7358:
7317:
7316:
7312:
7311:
7309:
7307:
7306:
7301:
7295:
7294:
7290:
7289:
7287:
7285:
7284:
7283:Blubervirales
7279:
7276:
7274:
7273:
7268:
7265:
7263:
7262:
7257:
7254:
7252:
7251:
7246:
7243:
7241:
7240:
7235:
7231:
7227:
7221:
7217:
7187:
7186:
7182:
7180:
7179:
7178:Pseudoviridae
7175:
7173:
7172:
7168:
7166:
7165:
7164:Belpaoviridae
7161:
7160:
7158:
7156:
7155:
7150:
7147:
7145:
7144:
7139:
7136:
7134:
7133:
7128:
7125:
7123:
7122:
7117:
7114:
7112:
7111:
7106:
7102:
7098:
7092:
7088:
7085:
7083:
7079:
7045:
7044:
7040:
7039:
7037:
7035:
7034:
7029:
7026:
7024:
7023:
7018:
7008:
7007:
7003:
7001:
7000:
6996:
6994:
6993:
6992:Rhabdoviridae
6989:
6987:
6986:
6985:Pneumoviridae
6982:
6980:
6979:
6975:
6973:
6972:
6968:
6966:
6965:
6964:Mymonaviridae
6961:
6959:
6958:
6954:
6952:
6951:
6947:
6945:
6944:
6940:
6938:
6937:
6933:
6932:
6930:
6928:
6927:
6922:
6916:
6915:
6914:Natareviridae
6911:
6909:
6908:
6904:
6902:
6901:
6897:
6895:
6894:
6890:
6888:
6887:
6883:
6882:
6880:
6878:
6877:
6872:
6869:
6867:
6866:
6861:
6851:
6850:
6846:
6845:
6843:
6841:
6840:
6835:
6832:
6830:
6829:
6824:
6814:
6813:
6809:
6807:
6806:
6805:Amnoonviridae
6802:
6801:
6799:
6797:
6796:
6791:
6788:
6786:
6785:
6780:
6770:
6769:
6768:Wupedeviridae
6765:
6763:
6762:
6758:
6756:
6755:
6754:Phenuiviridae
6751:
6749:
6748:
6747:Phasmaviridae
6744:
6742:
6741:
6737:
6735:
6734:
6730:
6728:
6727:
6723:
6721:
6720:
6716:
6714:
6713:
6709:
6707:
6706:
6702:
6700:
6699:
6695:
6693:
6692:
6688:
6687:
6685:
6683:
6682:
6677:
6674:
6672:
6671:
6666:
6656:
6655:
6651:
6650:
6648:
6646:
6645:
6640:
6637:
6635:
6634:
6629:
6626:
6624:
6623:
6618:
6615:
6613:
6612:
6611:Orthornavirae
6607:
6604:
6602:
6601:
6596:
6592:
6588:
6582:
6578:
6556:
6555:
6551:
6549:
6548:
6543:
6540:
6539:
6537:
6533:
6519:
6518:
6514:
6513:
6511:
6509:
6508:
6507:Stellavirales
6503:
6497:
6496:
6492:
6491:
6489:
6487:
6486:
6485:Patatavirales
6481:
6478:
6476:
6475:
6470:
6460:
6459:
6458:Solemoviridae
6455:
6453:
6452:
6448:
6446:
6445:
6441:
6440:
6438:
6436:
6435:
6434:Sobelivirales
6430:
6424:
6423:
6419:
6417:
6416:
6412:
6410:
6409:
6405:
6403:
6402:
6398:
6396:
6395:
6394:Caliciviridae
6391:
6389:
6388:
6384:
6382:
6381:
6377:
6375:
6374:
6370:
6369:
6367:
6365:
6364:
6359:
6353:
6352:
6351:Tobaniviridae
6348:
6346:
6345:
6341:
6339:
6338:
6334:
6332:
6331:
6327:
6325:
6324:
6320:
6318:
6317:
6316:Mononiviridae
6313:
6311:
6310:
6309:Mesoniviridae
6306:
6304:
6303:
6299:
6297:
6296:
6295:Gresnaviridae
6292:
6290:
6289:
6288:Euroniviridae
6285:
6283:
6282:
6281:Coronaviridae
6278:
6276:
6275:
6271:
6269:
6268:
6267:Arteriviridae
6264:
6262:
6261:
6260:Abyssoviridae
6257:
6256:
6254:
6252:
6251:
6246:
6243:
6241:
6240:
6235:
6232:
6230:
6229:
6224:
6210:
6209:
6205:
6204:
6202:
6200:
6199:
6194:
6191:
6189:
6188:
6183:
6173:
6172:
6168:
6167:
6165:
6163:
6162:
6161:Cryppavirales
6157:
6154:
6152:
6151:
6146:
6136:
6135:
6131:
6130:
6128:
6126:
6125:
6120:
6117:
6115:
6114:
6109:
6099:
6098:
6097:Steitzviridae
6094:
6092:
6091:
6087:
6086:
6084:
6082:
6081:
6076:
6070:
6069:
6068:Solspiviridae
6065:
6063:
6062:
6058:
6056:
6055:
6051:
6049:
6048:
6047:Atkinsviridae
6044:
6043:
6041:
6039:
6038:
6033:
6030:
6028:
6027:
6026:Leviviricetes
6022:
6019:
6017:
6016:
6015:Lenarviricota
6011:
5997:
5996:
5995:Tombusviridae
5992:
5990:
5989:
5985:
5983:
5982:
5978:
5977:
5975:
5973:
5972:
5967:
5964:
5962:
5961:
5956:
5946:
5945:
5941:
5939:
5938:
5934:
5933:
5931:
5929:
5928:
5927:Nodamuvirales
5923:
5920:
5918:
5917:
5912:
5902:
5901:
5897:
5896:
5894:
5892:
5891:
5886:
5883:
5881:
5880:
5875:
5865:
5864:
5860:
5858:
5857:
5853:
5851:
5850:
5846:
5844:
5843:
5839:
5837:
5836:
5832:
5831:
5829:
5827:
5826:
5821:
5815:
5814:
5810:
5808:
5807:
5803:
5801:
5800:
5796:
5794:
5793:
5789:
5787:
5786:
5782:
5780:
5779:
5775:
5773:
5772:
5768:
5767:
5765:
5763:
5762:
5757:
5751:
5750:
5749:Matonaviridae
5746:
5744:
5743:
5739:
5737:
5736:
5732:
5730:
5729:
5725:
5724:
5722:
5720:
5719:
5718:Hepelivirales
5714:
5711:
5709:
5708:
5707:Alsuviricetes
5703:
5700:
5698:
5697:
5692:
5689:
5687:
5686:
5685:Orthornavirae
5681:
5678:
5676:
5675:
5670:
5666:
5662:
5656:
5652:
5630:
5629:
5624:
5621:
5619:
5618:
5614:
5612:
5611:
5606:
5603:
5602:
5600:
5596:
5582:
5581:
5577:
5575:
5574:
5570:
5568:
5567:
5563:
5561:
5560:
5556:
5554:
5553:
5552:Amalgaviridae
5549:
5548:
5546:
5544:
5543:
5538:
5535:
5533:
5532:
5527:
5524:
5522:
5521:
5516:
5502:
5501:
5497:
5496:
5494:
5492:
5491:
5486:
5483:
5481:
5480:
5475:
5465:
5464:
5460:
5459:
5457:
5455:
5454:
5449:
5446:
5444:
5443:
5438:
5428:
5427:
5423:
5421:
5420:
5419:Quadriviridae
5416:
5414:
5413:
5409:
5407:
5406:
5405:Chrysoviridae
5402:
5401:
5399:
5397:
5396:
5395:Ghabrivirales
5391:
5388:
5386:
5385:
5380:
5377:
5375:
5374:
5369:
5366:
5364:
5363:
5362:Orthornavirae
5358:
5355:
5353:
5352:
5347:
5343:
5339:
5338:dsRNA viruses
5333:
5329:
5326:
5324:
5320:
5302:
5301:
5297:
5295:
5294:
5290:
5288:
5287:
5286:Anelloviridae
5282:
5279:
5278:
5276:
5272:
5250:
5249:
5245:
5244:
5242:
5240:
5239:
5238:Haloruvirales
5234:
5231:
5229:
5228:
5223:
5220:
5218:
5217:
5212:
5209:
5207:
5206:
5201:
5183:
5182:
5181:Genomoviridae
5178:
5176:
5175:
5174:Geminiviridae
5171:
5170:
5168:
5166:
5165:
5160:
5157:
5155:
5154:
5149:
5139:
5138:
5134:
5133:
5131:
5129:
5128:
5123:
5117:
5116:
5112:
5110:
5109:
5108:Metaxyviridae
5105:
5104:
5102:
5100:
5099:
5094:
5088:
5087:
5083:
5082:
5080:
5078:
5077:
5072:
5066:
5065:
5061:
5060:
5058:
5056:
5055:
5050:
5044:
5043:
5039:
5038:
5036:
5034:
5033:
5028:
5025:
5023:
5022:
5021:Arfiviricetes
5017:
5014:
5012:
5011:
5006:
4992:
4991:
4987:
4986:
4984:
4982:
4981:
4976:
4973:
4971:
4970:
4965:
4955:
4954:
4950:
4949:
4947:
4945:
4944:
4939:
4936:
4934:
4933:
4928:
4925:
4923:
4922:
4921:Cossaviricota
4917:
4914:
4912:
4911:
4906:
4888:
4887:
4883:
4882:
4880:
4878:
4877:
4872:
4869:
4867:
4866:
4861:
4858:
4856:
4855:
4850:
4847:
4845:
4844:
4839:
4821:
4820:
4816:
4814:
4813:
4809:
4807:
4806:
4802:
4801:
4799:
4797:
4796:
4795:Tubulavirales
4791:
4788:
4786:
4785:
4780:
4777:
4775:
4774:
4769:
4766:
4764:
4763:
4758:
4755:
4753:
4752:
4747:
4743:
4739:
4738:ssDNA viruses
4733:
4729:
4711:
4710:
4709:Rhizidiovirus
4706:
4704:
4703:
4698:
4695:
4693:
4692:
4691:Thaspiviridae
4688:
4686:
4685:
4681:
4679:
4678:
4674:
4672:
4671:
4670:Plasmaviridae
4667:
4665:
4664:
4660:
4658:
4657:
4656:Halspiviridae
4653:
4651:
4650:
4646:
4644:
4643:
4639:
4637:
4636:
4632:
4630:
4629:
4625:
4623:
4622:
4618:
4616:
4615:
4610:
4607:
4606:
4604:
4600:
4590:
4589:
4585:
4584:
4582:
4578:
4572:
4571:
4567:
4565:
4564:
4560:
4558:
4557:
4556:Baculoviridae
4553:
4552:
4550:
4548:
4547:
4542:
4539:
4537:
4536:
4531:
4528:
4524:
4502:
4501:
4497:
4495:
4494:
4493:Simuloviridae
4490:
4488:
4487:
4483:
4482:
4480:
4478:
4477:
4472:
4469:
4467:
4466:
4461:
4458:
4456:
4455:
4450:
4447:
4445:
4444:
4443:Helvetiavirae
4439:
4421:
4420:
4416:
4415:
4413:
4411:
4410:
4405:
4399:
4398:
4394:
4393:
4391:
4389:
4388:
4383:
4377:
4376:
4372:
4371:
4369:
4367:
4366:
4365:Kalamavirales
4361:
4355:
4354:
4350:
4349:
4347:
4345:
4344:
4343:Belfryvirales
4339:
4336:
4334:
4333:
4328:
4318:
4317:
4316:Adintoviridae
4313:
4312:
4310:
4308:
4307:
4302:
4299:
4297:
4296:
4291:
4281:
4280:
4279:Lavidaviridae
4276:
4275:
4273:
4271:
4270:
4265:
4262:
4260:
4259:
4254:
4251:
4249:
4248:
4243:
4229:
4228:
4224:
4222:
4221:
4217:
4215:
4214:
4210:
4209:
4207:
4205:
4204:
4199:
4193:
4192:
4188:
4187:
4185:
4183:
4182:
4181:Imitervirales
4177:
4171:
4170:
4166:
4165:
4163:
4161:
4160:
4155:
4152:
4150:
4149:
4148:Megaviricetes
4144:
4134:
4133:
4129:
4128:
4126:
4124:
4123:
4118:
4112:
4111:
4107:
4106:
4104:
4102:
4101:
4096:
4093:
4091:
4090:
4085:
4082:
4080:
4079:
4074:
4071:
4069:
4068:
4063:
4060:
4058:
4057:
4052:
4030:
4029:
4025:
4024:
4022:
4020:
4019:
4014:
4008:
4007:
4003:
4002:
4000:
3998:
3997:
3996:Sepolyvirales
3992:
3989:
3987:
3986:
3981:
3978:
3976:
3975:
3974:Cossaviricota
3970:
3967:
3965:
3964:
3959:
3956:
3954:
3953:
3948:
3926:
3925:
3924:Zobellviridae
3921:
3919:
3918:
3914:
3912:
3911:
3910:Schitoviridae
3907:
3905:
3904:
3900:
3898:
3897:
3893:
3891:
3890:
3886:
3884:
3883:
3879:
3877:
3876:
3872:
3870:
3869:
3868:Guelinviridae
3865:
3863:
3862:
3858:
3856:
3855:
3851:
3849:
3848:
3844:
3842:
3841:
3837:
3835:
3834:
3830:
3829:
3827:
3825:
3824:
3819:
3816:
3814:
3813:
3808:
3805:
3803:
3802:
3797:
3783:
3782:
3778:
3776:
3775:
3774:Herpesviridae
3771:
3769:
3768:
3764:
3763:
3761:
3759:
3758:
3757:Herpesvirales
3753:
3750:
3748:
3747:
3742:
3739:
3737:
3736:
3735:Peploviricota
3731:
3728:
3726:
3725:
3720:
3717:
3715:
3714:
3713:Duplodnaviria
3709:
3687:
3686:
3682:
3681:
3679:
3677:
3676:
3671:
3665:
3664:
3660:
3658:
3657:
3653:
3652:
3650:
3648:
3647:
3642:
3639:
3637:
3636:
3635:Tokiviricetes
3631:
3628:
3626:
3625:
3624:Taleaviricota
3620:
3617:
3615:
3614:
3609:
3606:
3604:
3603:
3598:
3594:
3590:
3589:dsDNA viruses
3584:
3580:
3577:
3575:
3571:
3566:
3562:
3555:
3550:
3548:
3543:
3541:
3536:
3535:
3532:
3525:
3520:
3516:
3515:
3511:
3505:
3501:
3497:
3491:
3487:
3483:
3479:
3474:
3470:
3464:
3460:
3455:
3451:
3447:
3443:
3437:
3433:
3429:
3425:
3420:
3416:
3410:
3406:
3401:
3400:
3396:
3376:
3369:
3366:
3361:
3357:
3352:
3347:
3343:
3339:
3335:
3331:
3330:Nat Microbiol
3327:
3320:
3318:
3316:
3314:
3310:
3305:
3301:
3296:
3291:
3287:
3283:
3279:
3275:
3274:Bacteriol Rev
3271:
3264:
3261:
3248:
3244:
3238:
3235:
3231:
3226:
3223:
3210:
3206:
3200:
3197:
3184:
3180:
3174:
3171:
3158:
3154:
3148:
3145:
3141:
3136:
3134:
3130:
3117:
3113:
3107:
3104:
3100:
3095:
3093:
3089:
3076:
3072:
3066:
3063:
3059:
3054:
3051:
3038:
3034:
3028:
3025:
3020:
3016:
3011:
3006:
3002:
2998:
2994:
2990:
2986:
2979:
2976:
2963:
2959:
2958:"RNA editing"
2953:
2950:
2937:
2933:
2927:
2924:
2911:
2907:
2892:
2888:
2873:
2869:
2854:
2850:
2835:
2831:
2816:
2812:
2806:
2803:
2798:
2794:
2789:
2784:
2779:
2774:
2770:
2766:
2763:(5): e64328.
2762:
2758:
2754:
2747:
2744:
2739:
2735:
2730:
2725:
2721:
2717:
2713:
2709:
2705:
2698:
2695:
2682:
2678:
2672:
2669:
2657:
2653:
2648:
2643:
2639:
2635:
2631:
2627:
2623:
2616:
2614:
2610:
2605:
2601:
2596:
2591:
2586:
2581:
2577:
2573:
2569:
2562:
2559:
2555:
2550:
2548:
2546:
2542:
2537:
2533:
2528:
2523:
2519:
2515:
2511:
2507:
2503:
2496:
2494:
2490:
2486:
2481:
2479:
2477:
2473:
2460:
2456:
2450:
2447:
2434:
2430:
2424:
2421:
2416:
2412:
2407:
2402:
2398:
2394:
2390:
2386:
2382:
2374:
2372:
2368:
2364:
2359:
2356:
2343:
2339:
2333:
2330:
2326:
2321:
2319:
2315:
2303:
2299:
2295:
2291:
2287:
2283:
2279:
2275:
2271:
2267:
2260:
2253:
2250:
2246:
2241:
2239:
2237:
2235:
2231:
2227:
2222:
2219:
2206:
2202:
2196:
2193:
2180:
2176:
2170:
2167:
2154:
2150:
2144:
2141:
2128:
2124:
2118:
2115:
2102:
2098:
2092:
2089:
2085:
2080:
2077:
2064:
2060:
2054:
2051:
2047:
2042:
2040:
2036:
2032:
2027:
2025:
2023:
2019:
2006:
2002:
1996:
1994:
1990:
1974:
1967:
1965:
1963:
1961:
1959:
1957:
1953:
1950:, p. 42.
1949:
1944:
1942:
1938:
1935:, p. 65.
1934:
1929:
1927:
1925:
1923:
1919:
1906:
1902:
1896:
1894:
1890:
1877:
1873:
1867:
1864:
1859:
1857:9781444114782
1853:
1849:
1842:
1839:
1835:
1830:
1828:
1824:
1820:
1815:
1813:
1811:
1807:
1794:
1790:
1784:
1782:
1778:
1765:
1761:
1755:
1753:
1749:
1733:
1731:
1727:
1718:
1715:
1699:
1692:
1690:
1688:
1686:
1682:
1666:
1664:
1655:
1653:
1649:
1633:
1626:
1624:
1622:
1620:
1618:
1614:
1598:
1591:
1589:
1587:
1583:
1570:
1566:
1560:
1557:
1544:
1540:
1534:
1531:
1518:
1514:
1508:
1505:
1492:
1488:
1482:
1479:
1475:
1470:
1468:
1466:
1462:
1459:, p. 66.
1458:
1453:
1450:
1437:
1433:
1427:
1424:
1420:
1415:
1413:
1411:
1409:
1407:
1405:
1401:
1388:
1384:
1378:
1376:
1374:
1372:
1368:
1364:
1359:
1357:
1353:
1346:
1341:
1333:
1332:
1325:
1322:
1315:
1313:
1311:
1310:
1305:
1301:
1297:
1293:
1292:Duplodnaviria
1289:
1284:
1282:
1281:Orthornavirae
1278:
1277:Orthornavirae
1274:
1270:
1266:
1262:
1257:
1253:
1251:
1247:
1243:
1234:
1227:
1219:
1216:
1212:
1208:
1203:
1200:
1196:
1193:
1192:
1190:
1185:
1181:
1178:
1175:
1172:
1169:
1166:
1162:
1157:
1156:
1154:
1153:
1152:
1150:
1146:
1142:
1135:
1131:
1127:
1123:
1118:
1111:
1109:
1105:
1103:
1102:ebola viruses
1094:
1092:
1090:
1086:
1085:
1075:
1073:
1066:
1064:
1062:
1061:
1060:Geminiviridae
1056:
1052:
1051:
1046:
1038:
1036:
1034:
1030:
1029:unspliced RNA
1026:
1025:Herpesviruses
1022:
1018:
1017:
1012:
1011:
1005:
998:
996:
989:
985:
981:
977:
973:
968:
961:
959:
957:
956:Blubervirales
953:
952:
947:
943:
942:
937:
932:
929:
920:
918:
916:
915:
910:
906:
902:
899:of the realm
898:
894:
893:Arterviricota
890:
889:
883:
881:
877:
873:
869:
865:
856:
854:
852:
849:in the realm
848:
841:
833:
831:
829:
825:
821:
817:
813:
809:
806:in the realm
805:
804:Orthornavirae
801:
800:
794:
791:
787:
783:
778:
776:
775:cap snatching
772:
765:
757:
752:
751:
746:
743:
742:
737:
736:
731:
728:
727:
726:Lenarviricota
722:
721:
720:
718:
715:in the realm
714:
713:Orthornavirae
709:
705:
703:
697:
690:
689:Coronaviruses
686:
679:
674:
673:
668:
667:
662:
659:
658:
653:
652:
651:
649:
646:of the realm
645:
644:Orthornavirae
640:
638:
634:
633:
628:
627:
626:Amalgaviridae
620:
613:
609:
602:
600:
598:
597:
592:
588:
584:
580:
579:
575:in the realm
574:
573:
572:Orthornavirae
565:
561:
560:Orthornavirae
553:
548:
544:
540:
539:
534:
531:
530:
525:
524:
523:Anelloviridae
519:
516:
512:
508:
507:
506:
503:
500:
495:
494:
493:Anelloviridae
488:
485:
484:hairpin loops
481:
477:
473:
471:
466:
461:
458:
451:
443:
438:
431:
425:
424:
423:Rhizidiovirus
419:
418:
413:
411:
410:
409:Thaspiviridae
405:
404:
399:
398:
393:
392:
391:Plasmaviridae
387:
386:
381:
380:
379:Halspiviridae
375:
374:
369:
368:
363:
362:
357:
356:
351:
350:
345:
344:
339:
337:
336:
331:
330:
328:
325:
321:
320:giant viruses
317:
313:
309:
306:
302:
298:
294:
293:
288:
284:
281:
280:
279:Herpesvirales
275:
274:
269:
268:Duplodnaviria
265:
262:
258:
254:
253:
252:
249:
247:
243:
239:
238:Duplodnaviria
233:
231:
225:
223:
219:
215:
209:
205:
204:Duplodnaviria
201:
193:
191:
189:
188:
183:
182:
177:
176:
175:Duplodnaviria
171:
170:
162:
154:
149:
143:
139:
137:
133:
129:
125:
120:
119:transcription
112:
109:
106:
103:
100:
97:
94:
93:
92:
90:
86:
78:
76:
74:
69:
67:
63:
59:
58:transcription
55:
50:
48:
44:
40:
36:
32:
31:messenger RNA
28:
25:
21:
7313:
7305:Ortervirales
7303:
7291:
7281:
7270:
7259:
7250:Pararnavirae
7248:
7237:
7185:Retroviridae
7183:
7176:
7169:
7162:
7154:Ortervirales
7152:
7141:
7130:
7121:Pararnavirae
7119:
7108:
7041:
7031:
7020:
7006:Xinmoviridae
7004:
6997:
6990:
6983:
6976:
6971:Nyamiviridae
6969:
6962:
6957:Lispiviridae
6955:
6948:
6943:Bornaviridae
6941:
6934:
6924:
6912:
6907:Myriaviridae
6905:
6898:
6891:
6886:Aliusviridae
6884:
6874:
6863:
6847:
6837:
6826:
6810:
6803:
6793:
6782:
6766:
6761:Tospoviridae
6759:
6752:
6745:
6738:
6733:Nairoviridae
6731:
6724:
6717:
6712:Hantaviridae
6710:
6703:
6698:Arenaviridae
6696:
6691:Cruliviridae
6689:
6681:Bunyavirales
6679:
6668:
6652:
6642:
6631:
6620:
6609:
6598:
6552:
6545:
6541:
6517:Astroviridae
6515:
6505:
6493:
6483:
6472:
6456:
6451:Barnaviridae
6449:
6442:
6432:
6420:
6413:
6406:
6399:
6392:
6385:
6380:Marnaviridae
6378:
6371:
6361:
6349:
6342:
6337:Olifoviridae
6335:
6328:
6321:
6314:
6307:
6300:
6293:
6286:
6279:
6272:
6265:
6258:
6248:
6237:
6228:Pisuviricota
6226:
6206:
6198:Ourlivirales
6196:
6187:Miaviricetes
6185:
6169:
6159:
6148:
6134:Narnaviridae
6132:
6122:
6111:
6095:
6090:Blumeviridae
6088:
6080:Timlovirales
6078:
6066:
6061:Fiersviridae
6059:
6052:
6045:
6037:Norzivirales
6035:
6024:
6013:
5993:
5988:Luteoviridae
5986:
5979:
5969:
5958:
5942:
5935:
5925:
5914:
5900:Flaviviridae
5898:
5888:
5877:
5861:
5854:
5847:
5840:
5833:
5823:
5813:Virgaviridae
5811:
5804:
5797:
5790:
5783:
5776:
5771:Bromoviridae
5769:
5759:
5747:
5740:
5733:
5726:
5716:
5705:
5694:
5683:
5672:
5626:
5622:
5615:
5610:Birnaviridae
5608:
5604:
5578:
5571:
5564:
5557:
5550:
5542:Durnavirales
5540:
5529:
5520:Pisuviricota
5518:
5500:Cystoviridae
5498:
5490:Mindivirales
5488:
5477:
5461:
5451:
5440:
5424:
5417:
5410:
5403:
5393:
5382:
5371:
5360:
5349:
5300:Spiraviridae
5298:
5291:
5284:
5280:
5246:
5236:
5225:
5216:Saleviricota
5214:
5203:
5179:
5172:
5162:
5151:
5135:
5127:Recrevirales
5125:
5113:
5106:
5098:Mulpavirales
5096:
5086:Smacoviridae
5084:
5076:Cremevirales
5074:
5064:Circoviridae
5062:
5054:Cirlivirales
5052:
5040:
5032:Baphyvirales
5030:
5019:
5008:
4990:Parvoviridae
4988:
4980:Piccovirales
4978:
4967:
4953:Bidnaviridae
4951:
4941:
4932:Mouviricetes
4930:
4919:
4910:Shotokuvirae
4908:
4886:Microviridae
4884:
4876:Petitvirales
4874:
4863:
4854:Phixviricota
4852:
4841:
4817:
4810:
4803:
4793:
4782:
4771:
4760:
4751:Monodnaviria
4749:
4707:
4702:Dinodnavirus
4700:
4696:
4689:
4682:
4675:
4668:
4663:Ovaliviridae
4661:
4654:
4649:Guttaviridae
4647:
4640:
4633:
4628:Clavaviridae
4626:
4619:
4612:
4608:
4586:
4568:
4561:
4554:
4544:
4533:
4498:
4491:
4484:
4474:
4463:
4452:
4441:
4417:
4407:
4397:Adenoviridae
4395:
4385:
4375:Tectiviridae
4373:
4363:
4353:Turriviridae
4351:
4341:
4330:
4314:
4304:
4293:
4277:
4267:
4256:
4245:
4225:
4220:Iridoviridae
4218:
4211:
4201:
4189:
4179:
4167:
4157:
4146:
4130:
4122:Chitovirales
4120:
4110:Asfarviridae
4108:
4098:
4087:
4076:
4067:Bamfordvirae
4065:
4056:Varidnaviria
4054:
4026:
4016:
4004:
3994:
3983:
3972:
3963:Shotokuvirae
3961:
3952:Monodnaviria
3950:
3922:
3917:Siphoviridae
3915:
3908:
3901:
3894:
3887:
3880:
3873:
3866:
3859:
3852:
3847:Chaseviridae
3845:
3838:
3831:
3823:Caudovirales
3821:
3810:
3799:
3779:
3772:
3765:
3755:
3744:
3733:
3722:
3711:
3683:
3675:Primavirales
3673:
3661:
3654:
3644:
3633:
3622:
3611:
3600:
3560:
3477:
3458:
3423:
3404:
3382:. Retrieved
3368:
3333:
3329:
3277:
3273:
3263:
3251:. Retrieved
3246:
3237:
3225:
3213:. Retrieved
3208:
3199:
3187:. Retrieved
3182:
3173:
3161:. Retrieved
3156:
3147:
3120:. Retrieved
3115:
3106:
3079:. Retrieved
3074:
3065:
3053:
3041:. Retrieved
3036:
3027:
2992:
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2126:
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2100:
2091:
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2062:
2053:
2009:. Retrieved
2004:
1980:. Retrieved
1909:. Retrieved
1904:
1880:. Retrieved
1875:
1866:
1848:Parvoviruses
1847:
1841:
1797:. Retrieved
1792:
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1763:
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1729:
1725:
1717:
1705:. Retrieved
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1490:
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1435:
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1391:. Retrieved
1386:
1363:Lostroh 2019
1331:Retroviridae
1329:
1324:
1307:
1304:Varidnaviria
1303:
1300:Varidnaviria
1299:
1296:Monodnaviria
1295:
1291:
1287:
1285:
1280:
1276:
1273:Pararnavirae
1272:
1268:
1264:
1258:
1254:
1245:
1239:
1139:
1128:(group IV),
1106:
1098:
1088:
1082:
1079:
1070:
1058:
1048:
1045:multipartite
1042:
1033:retroviruses
1014:
1008:
1002:
993:
982:(group IV),
978:(group IV),
955:
949:
946:Ortervirales
945:
939:
935:
933:
924:
914:Ortervirales
912:
908:
904:
903:. Excluding
900:
897:Pararnavirae
896:
892:
886:
884:
866:. The viral
860:
850:
847:Pararnavirae
846:
845:the kingdom
843:
827:
823:
819:
815:
811:
807:
803:
797:
795:
779:
767:
748:
739:
735:Pisuviricota
733:
724:
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710:
706:
698:
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664:
655:
647:
643:
641:
630:
624:
621:
617:
594:
576:
570:
567:
547:Varidnaviria
546:
542:
536:
529:Spiraviridae
527:
521:
514:
511:Monodnaviria
510:
504:
499:bidnaviruses
491:
489:
476:Parvoviruses
474:
465:endonuclease
462:
453:
450:Monodnaviria
421:
417:Dinodnavirus
415:
407:
401:
395:
389:
385:Ovaliviridae
383:
377:
373:Guttaviridae
371:
365:
359:
355:Clavaviridae
353:
347:
341:
333:
316:adenoviruses
312:Varidnaviria
311:
296:
290:
287:Monodnaviria
286:
277:
273:Caudovirales
271:
267:
256:
250:
246:Varidnaviria
245:
242:Caudovirales
241:
237:
234:
226:
211:
208:Varidnaviria
187:Varidnaviria
185:
181:Monodnaviria
179:
173:
167:
164:
116:
82:
70:
51:
19:
18:
7171:Metaviridae
6950:Filoviridae
6936:Artoviridae
6849:Aspiviridae
6726:Mypoviridae
6705:Fimoviridae
6495:Potyviridae
6408:Secoviridae
6401:Iflaviridae
6344:Roniviridae
6250:Nidovirales
6171:Mitoviridae
6054:Duinviridae
5971:Tolivirales
5937:Nodaviridae
5863:Tymoviridae
5825:Tymovirales
5806:Togaviridae
5799:Mayoviridae
5792:Kitaviridae
5742:Hepeviridae
5735:Benyviridae
5566:Hypoviridae
5426:Totiviridae
5115:Nanoviridae
4943:Polivirales
4843:Sangervirae
4588:Nimaviridae
4570:Nudiviridae
4546:Lefavirales
4409:Vinavirales
4387:Rowavirales
4213:Ascoviridae
4191:Mimiviridae
4159:Algavirales
4100:Asfuvirales
3889:Podoviridae
3801:Uroviricota
3663:Rudiviridae
3613:Zilligvirae
2989:J Gen Virol
2626:J Gen Virol
2572:PLOS Pathog
2554:Fermin 2018
2245:Fermin 2018
2046:Fermin 2018
1948:Fermin 2018
1819:Fermin 2018
1730:Lefavirales
1474:Fermin 2018
1167:amino acid.
1141:Translation
1124:(group I),
1112:Translation
1095:RNA editing
1055:plant virus
990:(group VI)
974:(group I),
612:Rotaviruses
596:Ribozyviria
554:RNA viruses
155:DNA viruses
124:replicating
62:translation
54:replicating
7381:Categories
7043:Yueviridae
6999:Sunviridae
6893:Chuviridae
6654:Qinviridae
6535:Unassigned
5598:Unassigned
5463:Reoviridae
5453:Reovirales
5274:Unassigned
5205:Trapavirae
4805:Inoviridae
4602:Unassigned
4580:Unassigned
4526:Unassigned
4132:Poxviridae
3882:Myoviridae
1342:References
1199:nucleobase
1136:(group VI)
1067:Host range
895:, kingdom
872:integrated
585:and other
448:See also:
340:Families:
324:poxviruses
198:See also:
7239:Riboviria
7110:Riboviria
6644:Muvirales
6600:Riboviria
5674:Riboviria
5351:Riboviria
4762:Loebvirae
3602:Adnaviria
3561:Baltimore
3247:ViralZone
3209:ViralZone
3183:ViralZone
3157:ViralZone
3116:ViralZone
3075:ViralZone
3037:ViralZone
2962:ViralZone
2936:ViralZone
2910:ViralZone
2891:ViralZone
2872:ViralZone
2853:ViralZone
2834:ViralZone
2815:ViralZone
2681:ViralZone
2459:ViralZone
2433:ViralZone
2363:Cann 2015
2342:ViralZone
2226:Cann 2015
2205:ViralZone
2179:ViralZone
2153:ViralZone
2127:ViralZone
2101:ViralZone
2084:Cann 2015
2063:ViralZone
2005:ViralZone
1905:ViralZone
1876:ViralZone
1793:ViralZone
1764:ViralZone
1663:Adnaviria
1569:ViralZone
1543:ViralZone
1517:ViralZone
1491:ViralZone
1436:ViralZone
1419:Cann 2015
1387:ViralZone
1347:Citations
1288:Adnaviria
1269:Riboviria
1265:Riboviria
1215:replicase
1149:ribosomes
901:Riboviria
891:, phylum
851:Riboviria
808:Riboviria
717:Riboviria
648:Riboviria
578:Riboviria
564:RNA virus
332:Classes:
257:Adnaviria
200:Adnaviria
169:Adnaviria
161:DNA virus
87:(DNA) or
37:(DNA) or
6542:Families
5605:Families
5281:Families
4609:Families
3504:90170103
3450:89706800
3384:6 August
3360:32341570
3253:6 August
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3019:22535777
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2821:6 August
2797:23717598
2757:PLOS ONE
2738:30482837
2687:6 August
2661:6 August
2656:29125455
2604:27812219
2536:25771806
2512:: 2–25.
2506:Virology
2465:6 August
2439:6 August
2415:29618642
2348:6 August
2307:6 August
2302:67769904
2294:30787460
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2011:6 August
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1606:6 August
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1393:6 August
1252:(ICTV).
1217:enzymes.
1145:proteins
928:repaired
876:provirus
414:Genera:
79:Overview
24:classify
7397:Viruses
7371:Viruses
3351:7186216
3304:4329869
3010:3542737
2788:3661469
2765:Bibcode
2729:6282212
2647:5882103
2595:5094692
2527:5898234
2406:5974489
2385:J Virol
2274:Bibcode
1228:History
1165:leucine
1057:family
786:adenine
583:viroids
470:ligated
261:archaea
27:viruses
7357:Portal
5623:Genera
4697:Genera
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3492:
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1976:(docx)
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1735:(docx)
1701:(docx)
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1668:(docx)
1635:(docx)
1600:(docx)
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911:order
790:uracil
771:capped
322:, and
224:site.
206:, and
184:, and
66:capsid
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3500:S2CID
3446:S2CID
2298:S2CID
2262:(PDF)
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1134:HIV-1
988:HIV-1
788:from
136:lipid
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3490:ISBN
3463:ISBN
3436:ISBN
3409:ISBN
3386:2020
3356:PMID
3300:PMID
3255:2020
3217:2020
3191:2020
3165:2020
3124:2020
3083:2020
3045:2020
3015:PMID
2970:2020
2944:2020
2918:2020
2899:2020
2880:2020
2861:2020
2842:2020
2823:2020
2793:PMID
2734:PMID
2708:mBio
2689:2020
2663:2020
2652:PMID
2600:PMID
2532:PMID
2467:2020
2441:2020
2411:PMID
2350:2020
2309:2020
2290:PMID
2213:2020
2187:2020
2161:2020
2135:2020
2109:2020
2071:2020
2013:2020
1984:2020
1913:2020
1884:2020
1852:ISBN
1801:2020
1772:2020
1743:2021
1709:2020
1676:2021
1643:2020
1608:2020
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1551:2020
1525:2020
1499:2020
1444:2020
1395:2020
1290:and
1211:tRNA
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980:DENV
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