1041:
1225:
1151:
147:
528:
1135:
126:
1249:
attacked its prey from below and behind, much like the modern great white shark does. The deepest bite marks on the dolphin's ribs indicate the shark aimed for the abdomen of its prey to inflict a fatal bite quickly and incapacitate its prey, and that when the dolphin was attacked a second time, it
1515:
Dana J. Ehret; Bruce J. Macfadden; Douglas S. Jones; Thomas J. Devries; David A. Foster; Rodolfo Salas-Gismondi (2012). "Origin of the white shark
Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru". Palaeontology. 55 (6): 1139–1153.
1062:
teeth being found worldwide. The species is divided into two forms based on tooth morphology, each with a unique evolutionary line. The maximum adult length is estimated between 5 and 7 m (16 and 23 ft). Smaller individuals were about 2.6–4.5 metres (8.5–14.8 ft) long.
1116:
and their roots are more uneven in thickness and shape, with more narrow builds and rounder lobes, somewhat making the appearance of a heart. But the crowns of the lower teeth of this species are described to be more straight and identical to the lower teeth of
1275:
1167:
The broad form is characterized by broad-shaped teeth often described as being identical to that of modern great whites besides the lack of serrations. Fossil evidence shows that the broad-form is the direct ancestor of the genus
944:, which was a move that was first proposed by Glikman (1964) but was long rejected beforehand. Later discoveries supports Ward & Bonavi's conclusion of its ancestry to the great white. An analysis of a newly discovered
1239:. It most likely would have been one of the major predators in its ecosystem, preying upon small whales and other mammals. Trace fossils in the form of tooth marks on the bones of a Pliocene dolphin of the species
1892:
McCormack, Jeremy; Griffiths, Michael L.; Kim, Sora L.; Shimada, Kenshu; Karnes, Molly; Maisch, Harry; Pederzani, Sarah; Bourgon, Nicolas; Jaouen, Klervia; Becker, Martin A.; Jöns, Niels (May 31, 2022).
1250:
was bitten near the dorsal fin, suggesting that the dolphin rolled over while injured. The size of the bites indicates further that the shark responsible was estimated to be 4 m (13 ft) long.
1257:
from Peru described in 2017 showcases that the shark also included fish in its diet. Stomach contents preserved with the fossil show this shark also fed on pilchards such as ancient sardines (genus
1054:
teeth can grow up to 8.9 cm (3.5 in) in length, suggesting a very large shark. Its body was probably very similar to that of modern great whites. It is also believed to have a
646:. Although this conclusion is widely accepted, some scientists disagree, with a study by Whitenack and Gottfried (2010) demonstrating geometrically morphological differences between
454:
can reach lengths up to 3.5 in (7.5 cm) and are found worldwide. It is believed to be an ancestor to the great white shark, an argument supported by the transitional species
2026:
Collareta, Alberto; Landini, Walter; Chacaltana Budiel, César
Augusto; Valdivia Vera, Waldir; Altamirano Sierra, Ali J.; Urbina Schmitt, Mario; Bianucci, Giovanni (March 2017).
1725:
Cione, Alberto Luis; Cabrera, Daniel
Alfredo; Barla, María Julia (2012). "Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic".
1017:
and the great white "have been documented from the same deposits, hence the former cannot be a chronospecific ancestor of the latter." He also criticized the fact that the "
1092:. True to its name, each tooth is "hooked", the point of which is shifted away from the middle axis, in the direction of the corners of the mouth. In this, the teeth of
2159:
2092:
1314:
565:, although some indications show that he had coined the taxon as early as 1838. Throughout the early and mid 20th century, different genera and species of other
609:). Eventually, later studies would begin to show that much of the sharks within the genus and two species are distinct from each other, discarding the genus
2028:"A well preserved skeleton of the fossil shark Cosmopolitodus hastalis from the late Miocene of Peru, featuring fish remains as fossilized stomach contents"
1536:"A Well Preserved Skeleton of the Fossil Shark Cosmopolitodus Hastalis from the Late Miocene of Peru, Featuring Fish Remains as Fossilized Stomach Contents"
2200:
2146:
2079:
1980:
Ciampaglio, C., Ricketts,D. Miocene
Madness: Fossil Finder ID and Information Guide. Bakersfield, California. Buena Vista Museum of Natural History.
1822:
2240:
2215:
1293:
632:, citing the similarities between the teeth of the two and that of modern mako sharks. In 2001, a study by Ward & Bonavi (2001) considered
2225:
613:
altogether and creating another issue on what new taxa the sharks should be placed in. As of now, the genus is still uncertain and debated.
2245:
1535:
2220:
2210:
1626:"A morphometric approach for addressing toothbased species delimitation in fossil mako sharks, Isurus (Elasmobranchii: Lamniformes)"
1678:"Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru"
1121:. The hooked teeth, which probably would be efficient for gripping prey, would suggest a diet of smaller and medium-sized animals.
1040:
1676:
Ehret, Dana J.; MacFadden, Bruce J.; Jones, Douglas S.; Devries, Thomas J.; Foster, David A.; Salas-Gismondi, Rodolfo (2012).
1199:
teeth. Unlike the broad-form, the narrow form is believed to be the ancestor of two extinct sharks, the hooked-tooth "mako" (
940:
is a direct ancestor of the great white and is more related to it than other makos. They moved the species into the genus
146:
1386:
1835:
1055:
1224:
994:. However, the study also noted that this would only be done if the putative sister species is conspecific with
974:
teeth showing basal serrations. Cione et al. (2012) also noted a possibility of moving all species in the genus
2235:
1772:"New example of Cosmopolitodus hastalis (Lamniformes, Lamnidae) from the Miocene of South Korea | Zoodiversity"
1472:
2230:
1341:"The occurrence of early Pleistocene marine fish remains from the Gulf Coast of Mobile County, Alabama, USA"
493:
1771:
2047:
1021:
1808:
1024:
has never been tested through phylogenetic analyses," and denoted that as of 2021, the argument that the
2172:
2105:
1204:
794:
599:
593:
543:
1431:
Zevallos-Valdivia, L.; Martínez-Pérez, C.; García-Flores, V.; Chávez-Valencia, A.; Botella, H. (2023).
1821:
Patrick L. Jambura; Julia Türtscher; Faviel A. López-Romero; Catalina
Pimiento; Jürgen Kriwet (2019).
1906:
1734:
1689:
1637:
946:
837:
456:
1895:"Trophic position of Otodus megalodon and great white sharks through time revealed by zinc isotopes"
1080:
587:
2012:
1948:
1874:
1707:
1655:
1454:
1413:
1362:
1241:
1193:
The narrow form has more slender teeth than its broad-form counterpart, which resemble broadened
317:
141:
2177:
2110:
1150:
2164:
2205:
2097:
2027:
1940:
1922:
1866:
1858:
933:
854:
574:
45:
1990:
1930:
1914:
1850:
1783:
1750:
1742:
1697:
1645:
1555:
1547:
1540:
Rivista
Italiana di Paleontologia e Stratigrafia (Research in Paleontology and Stratigraphy)
1444:
1405:
1352:
527:
1175:
1391:
comb. nov. (Chondrichthyes, Elasmobranchii) from the
Miocene of Pohang City, South Korea"
1185:
to reflect its difference from the narrow-form, although this is scientifically invalid.
1074:
teeth are somewhat similar to those of other mako sharks, especially the extinct species
1910:
1738:
1693:
1641:
1498:
1134:
1935:
1894:
1288:
1089:
962:, remarking that the two genera were separated solely due to the lack of serrations in
198:
188:
658:
is considered as the "traditional view" in the debate regarding the shark's taxonomy.
125:
2194:
1952:
1711:
1702:
1677:
1606:
1581:
1458:
1417:
1366:
1315:"Skeletal remains of the Miocene lamniform shark, Cosmopolitodus hastalis, from Peru"
1280:
1263:), suggesting bony fish featured prominently in the shark's diet while it was young.
1236:
1195:
558:
481:
461:
271:
83:
1878:
1659:
1517:
1274:
642:
542:, especially the status of its genus, has long been subject to debate. The initial
254:
1834:
Pimiento, C.; Cantalapiedra, J.L.; Shimada, K.; Field, D.J.; Smaers, J.B. (2019).
1746:
17:
2070:
1170:
1109:
566:
451:
443:
405:
218:
58:
2137:
1918:
1650:
1625:
1551:
1409:
1270:
103:
68:
1926:
1862:
1788:
1259:
1085:
983:
605:
548:
401:
158:
108:
52:
1944:
1870:
1228:
Specimen CPI-7899, featuring aggregate of skeletal and dermal fish remains
557:
for the narrow and broad-form variations respectively by Swiss naturalist
2131:
2064:
1357:
1340:
397:
228:
178:
98:
93:
78:
73:
63:
41:
1755:
998:. The study concluded that there is indeed a putative sister species of
2151:
2084:
1607:"Additions to and a review of the Miocene Shark and Ray fauna of Malta"
986:
that would occur if one of them, which was traditionally identified as
113:
88:
1854:
1449:
1432:
932:
and noted a strong morphological similarity between it and the extant
168:
2041:
400:
that lived between thirty and one million years ago during the late
1223:
1039:
526:
500:
408:
393:
208:
954:'s ancestry to the great white, however, they also proposed that
433:
was reassigned to the genus, and thus became the second species
2045:
2013:"Jaws -- 4 million BC: How an extinct shark attacked its prey"
1823:
On the origin of the great white shark
Carcharodon carcharias.
1084:. Adult upper teeth are generally 2–4 cm long with an un
1836:"Evolutionary pathways toward gigantism in sharks and rays"
437:. However, some researchers still consider both species of
1181:
This form is also commonly labeled by its species synonym
1582:"Central Paratethyan shark fauna (Ipolytarnóc, Hungary)"
1339:
Ebersole, J.A.; Ebersole, S.M.; Cicimurri, D.J. (2017).
950:
by Ehret et al. (2012) further cemented the theory of
1013:
However, Yun argued that the tooth fossil remains of
620:, a review by Holec et al. (1995) placed the species
1968:
Fossil Shark Teeth of the World: A collector's guide
1624:
Whitenack †, Lisa B.; Gottfried, Michael D. (2010).
300:
287:
261:
2121:
2054:
1433:"First record of palaeozoic vertebrates from Peru"
1178:in Peru show an evolutionary mosaic between them.
1809:"Great White Sharks and Prehistoric White Sharks"
1473:"Carcharodon hastalis Agassiz 1843 (white shark)"
1028:lineage with narrow, serrated teeth evolved from
928:Ward & Bonavi (2001) reexamined the teeth of
585:included two species of extinct ancestral makos,
1518:https://doi.org/10.1111/j.1475-4983.2012.01201.x
577:. Sharks previously identified as variations of
569:would be clumped into the two taxa as the genus
8:
1803:
1801:
1799:
1530:
1528:
1526:
1032:with broad, unserrated teeth is uncertain.
488:" meaning "citizen of the world" and ὀδών "
2042:
970:and pointing out examples of Late Miocene
616:Shortly after the discarding of the genus
124:
31:
1934:
1787:
1754:
1701:
1649:
1575:
1573:
1448:
1356:
640:as conspecific and declared the latter a
511:is derived from the Ancient Greek ξίφος "
1671:
1669:
1253:A well-preserved skeleton of a juvenile
1010:for it, resolving the paraphyly issue.
1601:
1599:
1305:
1387:"A tooth of the extinct lamnid shark,
1294:List of prehistoric cartilaginous fish
507:meaning "spear". The disputed species
1380:
1378:
1376:
628:as a species of mako under the genus
7:
1497:Crumpton, Nick (November 14, 2012).
966:and the lack of lateral cusplets in
563:Recherches sur les poissons fossiles
1499:"Great whites 'not from megashark'"
829:
771:
763:
756:
732:
710:
688:
666:
464:analyses have been done for proof.
2032:Repositorio Institucional Ingemmet
1630:Journal of Vertebrate Paleontology
990:, is a putative sister species of
25:
2201:Prehistoric fish of South America
1322:Erlanger Geologische Abhandlungen
1770:Yun, Chan-gyu (August 7, 2020).
1703:10.1111/j.1475-4983.2012.01201.x
1580:Kocsis, László (February 2007).
1437:Spanish Journal of Palaeontology
1273:
1149:
1133:
421:(other common names include the
145:
603:), and the false-toothed mako (
27:Extinct genus of mackerel shark
1:
2241:Fossil taxa described in 1843
2216:Fossil taxa described in 1964
1747:10.1016/j.geobios.2011.06.002
1398:Acta Palaeontologica Romaniae
2226:Pliocene species extinctions
1201:Cosmopolitodus/Isurus planus
1088:cutting edge and no lateral
515:" meaning "sword" and ὀδών "
2246:Taxa named by Louis Agassiz
936:. The study concluded that
2262:
1919:10.1038/s41467-022-30528-9
1235:was a confirmed hunter of
2221:Miocene first appearances
1651:10.1080/02724630903409055
1410:10.35463/j.apr.2022.01.02
1174:, and specimens from the
1056:cosmopolitan distribution
921:and its close relatives.
851:
834:
827:
791:
776:
769:
761:
754:
737:
730:
715:
708:
693:
686:
671:
323:
316:
284:
279:
260:
253:
142:Scientific classification
140:
132:
123:
34:
2211:Prehistoric shark genera
499:may be derived from the
1789:10.15407/zoo2020.05.433
1552:10.13130/2039-4942/8005
1006:and proposed the taxon
741:Cosmopolitodus hastalis
492:" meaning "tooth". The
427:broad-tooth white shark
414:Cosmopolitodus hastalis
266:Cosmopolitodus hastalis
1313:Klug, S.; Kriwet, J.,
1229:
1048:
1008:Carcharodon plicatilis
855:Carcharodon carcharias
654:. The scientific name
573:began to be used as a
535:
411:. Its type species is
2173:Paleobiology Database
2106:Paleobiology Database
1991:"Great White Lineage"
1899:Nature Communications
1389:Cosmopolitodus planus
1242:Astadelphis gastaldii
1227:
1209:Carcharomodus escheri
1043:
917:Relationship between
795:Carcharomodus escheri
779:Cosmopolitodus planus
600:Carcharomodus escheri
597:, the serrated mako (
530:
460:, but as of 2021, no
2123:Carcharodon hastalis
1586:Geologica Carpathica
1358:10.18476/pale.v10.a6
1108:teeth are much more
982:to avoid a possible
947:Carcharodon hubbelli
838:Carcharodon hubbelli
480:is derived from the
457:Carcharodon hubbelli
327:Carcharodon hastalis
1966:Cocke, Joe (2002).
1911:2022NatCo..13.2980M
1739:2012Geobi..45..167C
1694:2012Palgy..55.1139E
1642:2010JVPal..30...17W
958:should be moved to
606:Parotodus benedenii
546:was first given as
519:" meaning "tooth".
1230:
1196:Isurus oxyrhinchus
1049:
674:Isurus schoutedeni
594:Isurus retroflexus
561:in his 1843 paper
536:
423:extinct giant mako
18:Broad-toothed mako
2188:
2187:
2048:Taxon identifiers
1855:10.1111/evo.13680
1450:10.7203/sjp.25691
934:great white shark
925:
924:
911:
910:
902:
901:
893:
892:
884:
883:
875:
874:
866:
865:
815:
814:
806:
805:
767:Narrow form
696:Isurus praecursor
583:Oxyrhina xiphodon
579:Oxyrhina hastalis
575:wastebasket taxon
555:Oxyrhina xiphodon
523:Taxonomic history
385:
384:
379:
371:
363:
359:Oxyrhina xiphodon
355:
351:Oxyrhina hastalis
347:
339:
331:
310:
297:
249:
16:(Redirected from
2253:
2181:
2180:
2168:
2167:
2155:
2154:
2142:
2141:
2140:
2114:
2113:
2101:
2100:
2088:
2087:
2075:
2074:
2073:
2043:
2036:
2035:
2023:
2017:
2016:
2009:
2003:
2002:
2000:
1998:
1987:
1981:
1978:
1972:
1971:
1963:
1957:
1956:
1938:
1889:
1883:
1882:
1840:
1831:
1825:
1819:
1813:
1812:
1805:
1794:
1793:
1791:
1767:
1761:
1760:
1758:
1722:
1716:
1715:
1705:
1688:(6): 1139–1153.
1673:
1664:
1663:
1653:
1621:
1615:
1614:
1603:
1594:
1593:
1577:
1568:
1567:
1565:
1563:
1554:. Archived from
1532:
1521:
1513:
1507:
1506:
1494:
1488:
1487:
1485:
1483:
1469:
1463:
1462:
1452:
1428:
1422:
1421:
1395:
1385:Yun, C. (2021).
1382:
1371:
1370:
1360:
1336:
1330:
1329:
1319:
1310:
1283:
1278:
1277:
1220:Hunting behavior
1153:
1137:
830:
825:Broad form
772:
764:
757:
733:
711:
689:
667:
662:
661:
538:The taxonomy of
531:Fossil teeth of
419:broad-tooth mako
377:
369:
361:
353:
345:
337:
329:
308:
302:
295:
289:
263:
247:
240:
150:
149:
133:Tooth series of
128:
118:
55:
40:Temporal range:
32:
21:
2261:
2260:
2256:
2255:
2254:
2252:
2251:
2250:
2236:Pliocene sharks
2191:
2190:
2189:
2184:
2176:
2171:
2163:
2158:
2150:
2145:
2136:
2135:
2130:
2117:
2109:
2104:
2096:
2091:
2083:
2078:
2069:
2068:
2063:
2050:
2040:
2039:
2025:
2024:
2020:
2011:
2010:
2006:
1996:
1994:
1989:
1988:
1984:
1979:
1975:
1965:
1964:
1960:
1891:
1890:
1886:
1838:
1833:
1832:
1828:
1820:
1816:
1807:
1806:
1797:
1769:
1768:
1764:
1724:
1723:
1719:
1675:
1674:
1667:
1623:
1622:
1618:
1605:
1604:
1597:
1579:
1578:
1571:
1561:
1559:
1558:on June 1, 2022
1534:
1533:
1524:
1514:
1510:
1496:
1495:
1491:
1481:
1479:
1471:
1470:
1466:
1430:
1429:
1425:
1393:
1384:
1383:
1374:
1345:Palaeodiversity
1338:
1337:
1333:
1317:
1312:
1311:
1307:
1302:
1279:
1272:
1269:
1222:
1217:
1205:serrated "mako"
1191:
1176:Pisco Formation
1165:
1164:
1163:
1162:
1161:
1154:
1146:
1145:
1138:
1127:
1069:
1038:
978:into the genus
926:
912:
903:
894:
885:
876:
867:
816:
807:
656:Isurus hastalis
638:Isurus xiphodon
634:Isurus hastalis
525:
475:
470:
370:(Agassiz, 1856)
362:(Agassiz, 1843)
354:(Agassiz, 1843)
346:(Agassiz, 1843)
343:Isurus xiphodon
338:(Agassiz, 1843)
335:Isurus hastalis
330:(Agassiz, 1843)
296:(Agassiz, 1843)
275:
269:
246:
238:
144:
119:
117:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
50:
49:
38:
28:
23:
22:
15:
12:
11:
5:
2259:
2257:
2249:
2248:
2243:
2238:
2233:
2231:Miocene sharks
2228:
2223:
2218:
2213:
2208:
2203:
2193:
2192:
2186:
2185:
2183:
2182:
2169:
2156:
2143:
2127:
2125:
2119:
2118:
2116:
2115:
2102:
2089:
2076:
2060:
2058:
2056:Cosmopolitodus
2052:
2051:
2046:
2038:
2037:
2018:
2004:
1982:
1973:
1958:
1884:
1849:(3): 588–599.
1826:
1814:
1795:
1782:(5): 433–438.
1762:
1733:(2): 167–172.
1717:
1665:
1616:
1595:
1569:
1522:
1508:
1489:
1464:
1423:
1372:
1331:
1304:
1303:
1301:
1298:
1297:
1296:
1291:
1289:List of sharks
1285:
1284:
1268:
1265:
1237:marine mammals
1221:
1218:
1216:
1213:
1190:
1187:
1156:A narrow form
1155:
1148:
1147:
1139:
1132:
1131:
1130:
1129:
1128:
1126:
1123:
1068:
1065:
1037:
1034:
1030:Cosmopolitodus
1019:Cosmopolitodus
1015:Cosmopolitodus
1002:distinct from
976:Cosmopolitodus
956:Cosmopolitodus
942:Cosmopolitodus
923:
922:
914:
913:
909:
908:
905:
904:
900:
899:
896:
895:
891:
890:
887:
886:
882:
881:
878:
877:
873:
872:
869:
868:
864:
863:
860:
859:
850:
847:
846:
843:
842:
833:
828:
826:
822:
821:
818:
817:
813:
812:
809:
808:
804:
803:
800:
799:
790:
787:
786:
783:
782:
775:
770:
768:
762:
760:
755:
753:
750:
749:
746:
745:
736:
731:
729:
726:
725:
722:
721:
714:
709:
707:
704:
703:
700:
699:
692:
687:
685:
682:
681:
678:
677:
670:
665:
660:
524:
521:
484:κοσμοπολίτης "
478:Cosmopolitodus
474:
471:
469:
466:
441:as species of
439:Cosmopolitodus
398:mackerel shark
392:is an extinct
389:Cosmopolitodus
383:
382:
381:
380:
378:(Jordan, 1907)
372:
367:Oxyrhina plana
364:
356:
348:
340:
332:
321:
320:
314:
313:
312:
311:
298:
282:
281:
277:
276:
270:
258:
257:
251:
250:
243:Cosmopolitodus
236:
232:
231:
226:
222:
221:
216:
212:
211:
206:
202:
201:
199:Elasmobranchii
196:
192:
191:
189:Chondrichthyes
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
130:
129:
121:
120:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
56:
39:
36:Cosmopolitodus
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2258:
2247:
2244:
2242:
2239:
2237:
2234:
2232:
2229:
2227:
2224:
2222:
2219:
2217:
2214:
2212:
2209:
2207:
2204:
2202:
2199:
2198:
2196:
2179:
2174:
2170:
2166:
2161:
2157:
2153:
2148:
2144:
2139:
2133:
2129:
2128:
2126:
2124:
2120:
2112:
2107:
2103:
2099:
2094:
2090:
2086:
2081:
2077:
2072:
2066:
2062:
2061:
2059:
2057:
2053:
2049:
2044:
2033:
2029:
2022:
2019:
2014:
2008:
2005:
1992:
1986:
1983:
1977:
1974:
1969:
1962:
1959:
1954:
1950:
1946:
1942:
1937:
1932:
1928:
1924:
1920:
1916:
1912:
1908:
1904:
1900:
1896:
1888:
1885:
1880:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1848:
1844:
1837:
1830:
1827:
1824:
1818:
1815:
1810:
1804:
1802:
1800:
1796:
1790:
1785:
1781:
1777:
1773:
1766:
1763:
1757:
1752:
1748:
1744:
1740:
1736:
1732:
1728:
1721:
1718:
1713:
1709:
1704:
1699:
1695:
1691:
1687:
1683:
1682:Palaeontology
1679:
1672:
1670:
1666:
1661:
1657:
1652:
1647:
1643:
1639:
1635:
1631:
1627:
1620:
1617:
1612:
1608:
1602:
1600:
1596:
1591:
1587:
1583:
1576:
1574:
1570:
1557:
1553:
1549:
1545:
1541:
1537:
1531:
1529:
1527:
1523:
1519:
1512:
1509:
1504:
1500:
1493:
1490:
1478:
1474:
1468:
1465:
1460:
1456:
1451:
1446:
1443:(1): 95–100.
1442:
1438:
1434:
1427:
1424:
1419:
1415:
1411:
1407:
1403:
1399:
1392:
1390:
1381:
1379:
1377:
1373:
1368:
1364:
1359:
1354:
1351:(1): 97–115.
1350:
1346:
1342:
1335:
1332:
1327:
1323:
1316:
1309:
1306:
1299:
1295:
1292:
1290:
1287:
1286:
1282:
1281:Sharks portal
1276:
1271:
1266:
1264:
1262:
1261:
1256:
1251:
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1243:
1238:
1234:
1226:
1219:
1214:
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1197:
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1186:
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1179:
1177:
1173:
1172:
1159:
1152:
1143:
1140:A broad form
1136:
1124:
1122:
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1115:
1111:
1107:
1103:
1099:
1095:
1091:
1087:
1083:
1082:
1081:Isurus desori
1077:
1073:
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1064:
1061:
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1042:
1035:
1033:
1031:
1027:
1023:
1020:
1016:
1011:
1009:
1005:
1001:
1000:C. carcharias
997:
993:
992:C. carcharias
989:
985:
981:
977:
973:
969:
968:C. carcharias
965:
961:
957:
953:
949:
948:
943:
939:
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931:
920:
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898:
897:
889:
888:
880:
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871:
870:
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861:
858:
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841:
840:
839:
832:
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810:
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748:
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744:
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735:
734:
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727:
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723:
720:
719:
718:Isurus desori
713:
712:
706:
705:
702:
701:
698:
697:
691:
690:
684:
683:
680:
679:
676:
675:
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653:
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645:
644:
639:
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631:
627:
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614:
612:
608:
607:
602:
601:
596:
595:
590:
589:
588:Isurus desori
584:
580:
576:
572:
568:
564:
560:
559:Louis Agassiz
556:
552:
550:
545:
544:binomial name
541:
534:
529:
522:
520:
518:
514:
510:
506:
502:
498:
495:
494:specific name
491:
487:
483:
482:Ancient Greek
479:
472:
467:
465:
463:
459:
458:
453:
448:
446:
445:
440:
436:
432:
431:Isurus planus
428:
424:
420:
416:
415:
410:
407:
404:to the Early
403:
399:
395:
391:
390:
376:
375:Isurus planus
373:
368:
365:
360:
357:
352:
349:
344:
341:
336:
333:
328:
325:
324:
322:
319:
315:
309:Agassiz, 1856
307:
306:
299:
294:
293:
286:
285:
283:
278:
273:
268:
267:
259:
256:
252:
248:Glikman, 1964
245:
244:
237:
234:
233:
230:
227:
224:
223:
220:
217:
214:
213:
210:
209:Selachimorpha
207:
205:Subdivision:
204:
203:
200:
197:
194:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
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148:
143:
139:
136:
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127:
122:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
54:
51:30–1.07
47:
43:
37:
33:
30:
19:
2122:
2055:
2031:
2021:
2007:
1995:. Retrieved
1993:. elasmo.com
1985:
1976:
1967:
1961:
1902:
1898:
1887:
1846:
1842:
1829:
1817:
1779:
1776:Zoodiversity
1775:
1765:
1756:11336/139163
1730:
1726:
1720:
1685:
1681:
1636:(1): 17–25.
1633:
1629:
1619:
1611:ResearchGate
1610:
1589:
1585:
1560:. Retrieved
1556:the original
1543:
1539:
1511:
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1482:December 17,
1480:. Retrieved
1476:
1467:
1440:
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1321:
1308:
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1254:
1252:
1246:
1245:reveal that
1240:
1232:
1231:
1215:Paleobiology
1208:
1200:
1194:
1192:
1182:
1180:
1169:
1166:
1157:
1141:
1118:
1113:
1110:heterodontic
1105:
1101:
1100:differ from
1097:
1093:
1079:
1075:
1071:
1070:
1059:
1051:
1050:
1044:
1029:
1025:
1018:
1014:
1012:
1007:
1003:
999:
995:
991:
987:
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971:
967:
963:
959:
955:
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937:
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927:
918:
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852:
836:
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793:
792:
778:
777:
740:
739:
738:
717:
716:
695:
694:
673:
672:
655:
651:
647:
643:nomen dubium
641:
637:
633:
629:
625:
621:
617:
615:
610:
604:
598:
592:
586:
582:
578:
570:
562:
554:
547:
539:
537:
532:
516:
512:
508:
504:
496:
489:
486:kosmopolítēs
485:
477:
476:
462:phylogenetic
455:
449:
442:
438:
434:
430:
429:). In 2021,
426:
422:
418:
413:
412:
388:
387:
386:
374:
366:
358:
350:
342:
334:
326:
304:
303:
291:
290:
265:
264:
255:Type species
242:
241:
134:
35:
29:
1905:(1): 2980.
1546:(1). 2017.
1477:Fossilworks
1404:(1): 9–16.
1255:C. hastalis
1247:C. hastalis
1233:C. hastalis
1203:), and the
1189:Narrow-form
1171:Carcharodon
1158:C. hastalis
1142:C. hastalis
1119:C. hastalis
1102:C. hastalis
1076:C. hastalis
1060:C. hastalis
1052:C. hastalis
1045:C. hastalis
1036:Description
1026:Carcharodon
1004:C. hastalis
996:C. hastalis
988:C. xiphodon
980:Carcharodon
972:C. hastalis
964:C. hastalis
960:Carcharodon
952:C. hastalis
938:I. hastalis
930:I. hastalis
919:C. hastalis
652:I. xiphodon
648:I. hastalis
567:lamniformes
540:C. hastalis
533:C. hastalis
444:Carcharodon
406:Pleistocene
292:C. hastalis
219:Lamniformes
135:C. hastalis
2195:Categories
1300:References
1125:Broad-form
1022:morphotype
195:Subclass:
2071:Q20686304
1997:April 20,
1953:249235478
1927:2041-1723
1863:1558-5646
1843:Evolution
1712:128666594
1562:August 8,
1459:256585278
1418:242113412
1367:134476316
1260:Sardinops
1114:C. planus
1106:I. desori
1098:I. desori
1094:C. planus
1072:C. planus
984:paraphyly
473:Etymology
435:C. planus
402:Oligocene
305:C. planus
165:Kingdom:
159:Eukaryota
46:Calabrian
2206:Lamnidae
2165:11083619
2138:Q3803933
2132:Wikidata
2065:Wikidata
1945:35641494
1879:59224442
1871:30675721
1660:85821523
1592:: 27–40.
1503:BBC News
1267:See also
1183:xiphodon
1090:cusplets
1086:serrated
626:xiphodon
622:hastalis
618:Oxyrhina
611:Oxyrhina
571:Oxyrhina
551:hastalis
549:Oxyrhina
509:xiphodon
497:hastalis
468:Taxonomy
318:Synonyms
280:Species
229:Lamnidae
225:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
42:Rupelian
2152:5816397
2098:1223316
2085:4824543
1936:9156768
1907:Bibcode
1735:Bibcode
1727:Geobios
1690:Bibcode
1638:Bibcode
1058:, with
272:Agassiz
235:Genus:
215:Order:
185:Class:
2178:265174
2111:118027
1951:
1943:
1933:
1925:
1877:
1869:
1861:
1710:
1658:
1457:
1416:
1365:
630:Isurus
513:xíphos
417:, the
409:epochs
274:, 1843
2160:IRMNG
2093:IRMNG
1949:S2CID
1875:S2CID
1839:(PDF)
1708:S2CID
1656:S2CID
1455:S2CID
1414:S2CID
1394:(PDF)
1363:S2CID
1318:(PDF)
1160:tooth
1144:tooth
1112:than
1067:Teeth
505:hasta
503:word
501:Latin
452:teeth
394:genus
2147:GBIF
2080:GBIF
1999:2022
1941:PMID
1923:ISSN
1867:PMID
1859:ISSN
1564:2019
1484:2021
1328:: 99
1096:and
1078:and
1047:jaws
650:and
636:and
624:and
591:and
553:and
517:odṓn
490:odṓn
450:Its
425:and
59:PreꞒ
1931:PMC
1915:doi
1851:doi
1784:doi
1751:hdl
1743:doi
1698:doi
1646:doi
1548:doi
1544:123
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1406:doi
1353:doi
581:or
396:of
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1375:^
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1347:.
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109:Pg
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2034:.
2015:.
2001:.
1970:.
1955:.
1917::
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301:†
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