Knowledge (XXG)

Cat communication

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another. Once they are close enough to attack, they pause slightly, and then one cat leaps and tries to bite the nape of the other cat. The other cat has no choice but to retaliate and both cats roll aggressively on the ground. During such confrontations both cats produce loud intense screams. After some time, the cats separate and stand face to face to begin the attack all over again. This can go on for some time until one remains seated, showing defeat. The defeated cat does not move until the victor completes sniffing the area and moves outside the fighting area. Once this happens, the defeated cat leaves the area, ending the fight.
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side (defensive posture), and retreating when the dominant cat approaches. Dominant cats present a specific body posture as well. The cat displays ears straight up, the base of its tail will be arched, and it looks directly at subordinate cats. These dominant cats are usually not aggressive, but if a subordinate cat blocks the food source they may become aggressive. When this aggressive behavior occurs, it could also lead to the dominant cat preventing subordinate cats from eating and using the litter box. This can cause the subordinate cat to defecate somewhere else and create problems with human interaction.
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for many hours a week. The process is made easier if there is another socialized cat present but not necessarily in the same space as the one being socialized. If the handler can get a cat to urinate in the litter tray, then the others in a litter will usually follow. Initial contact with thick gloves is highly recommended until trust is established, usually within the first week. On the other hand, it is a challenge to socialize an adult cat. This is because socialized adult feral cats tend to trust only those who they trusted in their socialization period, and therefore can be very fearful around strangers.
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half-raised tail can indicate less pleasure, and discontent is indicated with a tail held low. A cat's tail may swing from side to side, and if this motion is slow and "lazy", it generally indicates that the cat is in a relaxed state. Cats will twitch the tip of their tails when hunting, alert, or playful. A stalking domestic cat will typically hold its tail low to the ground while in a crouch, and twitch it quickly from side to side. This tail behavior is also observed when a cat becomes "irritated" and is about to lash out and attack typically done by biting or scratching with the claws extended.
797: 895: 933:. After this period, socialization can be less effective. Studies have shown that the earlier the kitten is handled by people, the less fearful the kitten will be toward people. Other factors that can enhance socialization are having many people handle the kitten frequently, the presence of the mother, and feeding. The presence of the mother is important because cats are observational learners. For example, a mother that is comfortable around humans can reduce anxiety in the kitten and promote the kitten-human relationship. 431: 953:. This type of learning emerges early in a cat's life, and has been shown in many laboratory studies. Young kittens learn to hunt from their mothers by observing their techniques when catching prey. The mother ensures their kittens learn hunting techniques by first bringing dead prey to the litter, followed by live prey. With the live prey, she demonstrates the techniques required for successful capture. Prey-catching behavior of kittens improves at higher levels over time when their mothers are present. 529:– The cat is lying on its belly or crouching directly on top of its paws. Its entire body may be shaking and very near the ground when standing up; Breathing is also fast, with its legs bent near the surface, and its tail curled and very close to its body when standing on all fours. As such, a fearful and a defensive cat makes itself smaller, lowers itself toward the ground, arches its back and leans its body away from the threat rather than forward. Fighting usually occurs only when escape is impossible. 887: 88: 541:– The hind legs stiffen, the rump elevated, but the back stays flat; while tail hairs are erected. The nose is pushed forward and the ears are pulled back slightly. And because cats have both claws and teeth, they can easily cause injury if they become involved in a fight, so this posture is an attempt to elicit deference from a competitor without fighting. The aggressor may attempt to make the challengers retreat and will pursue them if they do not flee. 565:– The cat is lying on its belly, while the back of the body is more visibly lower than the front part when the cat is standing or moving. Its breathing may be fast, and its legs are tucked under its body. The tail is close to the body and maybe curled forward (or close to the body when standing), with the tip of the tail moving up and down (or side to side). The tail is also moved to the side when ready to be mounted by the male cat. 459: 477: 974: 495: 535:– The cat is crouched directly on top of its paws, with visible shaking seen in some parts of the body. Its tail is close to the body, and might be propped up, together with its hair on the back. The legs are very stiff or bent to increase their size. Typically, cats avoid contact when they feel threatened, although they can resort to varying degrees of aggression when they feel cornered, or when escape is impossible. 4194: 767:". This communication might have an olfactory component as there are scent glands in this area of the body, and is possibly for seeking attention when the cat turns its head down or to the side. Some cats also rub their faces on humans as a friendly greeting or indicating affection. This tactile action is combined with olfactory communication as the contact leaves scent around the mouth and cheeks. 626: 282: 445: 189: 571:– This posture also indicates that the cat is relaxed. When cats lie on their back with their bellies exposed, they are in a position of vulnerability. Therefore, this position may communicate a feeling of trust or comfort. They may also roll onto their backs to defend themselves with their claws or to bask in areas of bright sunlight. 613:
When greeting their owners, cats often hold their tails straight up with a quivering motion that indicates extreme happiness. A scared or surprised cat may erect the hairs on its tail and back. It may stand more upright and turn its body sideways to increase its apparent size as a threat. Tailless cats, such as the
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dogs. If the hiss and growl warning does not remove the threat, an attack may follow. It's also used to scare off intruders into their territory. Kittens as young as two to three weeks may hiss and spit when first picked up by a human. Spitting is a shorter but louder and more emphatic version of hissing.
559:– The cat is lying on its belly, with the back of its body lower than its upper body (slinking) when standing or moving back; Its hind legs are bent and front legs are extended when standing, and the tail is close to the body, tensed or curled downward; there can be twitching when the cat is standing up. 508:(as perceived by humans) such as slowly blinking to signal relaxation and comfort in their environment. Domestic cats frequently use visual communication with their eyes, ears, mouths, tails, coats and body postures. The change in a cat's facial features can be a strong indicator of their communication. 1013:
Dominance can be seen among domestic cats in multi-cat households. "Subordinate" cats submit to the "dominant" cat. Dominance includes such behaviors as the submissive cats walking around the dominant cat, waiting for the dominant cat to walk past them, avoiding eye contact, crouching, lying on their
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Cats carry these infantile behaviors beyond nursing and into adulthood. Some cats "nurse", that is, suck on clothing or bedding during kneading. The cat exerts firm downward pressure with its paw, spreading its toes to expose its claws, then curls its toes as it lifts its paw. The process takes place
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Usually, when strange cats meet, one of them makes a sudden move that puts the other cat into a defensive mode. The cat will then draw in on itself and prepare to attack if needed. The submissive cat will usually run away before a physical altercation ensues. This is not always the case, and what is
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Observational learning is not limited to kitten-hood – it can also be observed during adulthood. Studies have shown that adult cats that see others performing a task, such as pressing a lever after a visual cue, learn to perform the same task faster than those who did not witness another cat at that
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Kittens also show observational learning when they are socializing with humans. They are more likely to initiate socialization with humans when their mothers are exhibiting non-aggressive and non-defensive behaviors. Even though mothers spend more time with their kittens, male cats play an important
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Observational learning for cats can be described in terms of the drive to complete the behavior, the cue that initiates the behavior, the response to the cue, and the reward for completing the behavior. This is shown when cats learn predatory behavior from their mothers. The drive is hunger, the cue
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rubbing the scent glands, by urine and fecal deposits, spraying seems to be the "loudest" feline olfactory communication. It is most frequently observed in intact male cats in competition with other males. Males neutered in adulthood may still spray after neutering. Female cats also sometimes spray.
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Cats have nine different scent glands in their body. These are the pinna (outer ear flaps), temporal (on their temples), cheek (on the sides of their face), perioral (on the mouth corners), submandibular (under the jaw), interdigital (between toes), anal (on the sides of the anus), caudal (all along
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from the mother during nursing. Kittens "knead" the breast while suckling, using the forelimbs one at a time in an alternating pattern to push against the mammary glands to stimulate lactation. Cats have scent glands on the underside of their paws and when they knead or scratch objects or people, it
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Cats often use their tails to communicate. For example, a cat holding its tail vertically generally indicates positive emotions such as happiness or confidence; the vertical tail is often used as a friendly gesture toward people or other cats. Additionally, a cat may twitch its tail when playing. A
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Cats use body language and movement to communicate a wide range of feelings and information. There are various responses such as when cats arch their backs, erect their hairs and adopt a sideward posture to communicate fear or aggression. Other visual communication can be a single behavioral change
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The growl, spit, and hiss are sounds associated with either offensive or defensive aggression. They are usually accompanied by a postural display intended to have a visual effect on the perceived threat. Cats growl, hiss, and spit as a display of defense against both cats and other species, such as
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The chirr or chirrup sounds like a meow rolled on the tongue. It is commonly used by mother cats to call their kittens inside the nest. As such, kittens recognize their own mother's chirp, but they do not respond to the chirps of other mothers. It is also used in a friendly manner by cats when they
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Brown et al. categorized the vocal responses of cats based on the behavioral context. These contexts include situations such as the separation of kittens from mother cats, instances of food deprivation, pain responses, occurrences before or during threatening or aggressive behaviors (e.g., disputes
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communicate for a variety of reasons, including to show happiness, express anger, solicit attention, and observe potential prey. Additionally, they collaborate, play, and share resources. When cats communicate with humans, they do so to get what they need or want, such as food, water, attention, or
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The first sign of an imminent tomcat duel is when both cats draw themselves up high on their legs, all hair along the middle of their backs is raised straight up, and they mew and howl loudly as they approach one another. The steps the cats make become slower and shorter, as they get closer to one
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Social conflict among cats depends solely on the behavior of the cats. Some research has shown that cats rarely pick fights, but when they do, it is usually for protecting food and/or litters, and defending their territory. Fights can happen between two females or between a male and a female. Cats
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Feral kittens around two to seven weeks old can be socialized usually within a month of capture. Some species of cats cannot be socialized because of factors such as genetic influence and in some cases specific learning experiences. The best way to get a kitten to socialize is to handle the kitten
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Grooming is extremely important not only to clean themselves but also to control ectoparasites. Fleas tend to be the most common ectoparasite in cats and some studies show indirect evidence that grooming in cats is effective in dislodging fleas from the head and neck. Cats may also use grooming to
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Owens et al. categorized cat vocalizations based on their acoustic structures. There are three categories: tonal sounds, pulse sounds, and broadband sounds. Tonal sounds are further categorized into groups of harmonically structured sounds or regular tonal sounds. Pulse vocalizations are separated
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There are a number of problematic behaviors that affect the human-cat relationship. One behavior is when cats attack people by scratching and biting. This often occurs spontaneously or could be triggered by sudden movements. Another problematic behavior is the "petting and biting syndrome", which
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Gentle biting (often accompanied by purring and kneading) can communicate affection or playfulness, directed at the human owner or another cat. Stronger bites that are often accompanied by hissing or growling usually communicate aggression. When cats mate, the tom bites the scruff of the female's
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while kneading, usually taken to indicate contentment and affection. They also purr mostly when newborn, when feeding, or when trying to feed on their mother's teat. The common association between the two behaviors may confirm the evidence in favor of the origin of kneading as a remnant instinct.
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When playing, a cat, usually a kitten, may raise the base of its tail high and stiffen all but the tip into a shape like an upside-down "U". This signals great excitement, to the point of hyperactivity. It may also be seen when younger cats chase each other, or when they run around by themselves.
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from glands located around the mouth, chin, forehead, cheeks, lower back, tail and paws. Their rubbing and head-bumping behaviors are methods of depositing these scents on substrates, including humans. The cat rubs its cheeks on prominent objects in the preferred territory, depositing a chemical
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is a rare occurrence in cats, except in warm weather environments, or after delivery. Some cats may pant in response to anxiety, fear or excitement. It can also be caused by play, exercise, or stress from stimuli, such as car rides. Panting in cat-moms after delivery is normal and not related to
523:– The cat is seen lying on the side or sitting. Its breathing is slow to normal, with legs bent, or hind legs laid out or extended. The tail is loosely wrapped, extended, or held up. It also hangs down loosely when it is standing. When they are calm, they tend to stand relaxed with a still tail. 841:
points backward. Although cats may mark with both sprayed and non-sprayed urine, the spray is usually more thick and oily than normally deposited urine, and may contain additional secretions from anal sacs that help the cat make a stronger communication. While cats mark their territory both by
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Cats can change the position of their ears very quickly, and continuously. They are erect when the cat is alert and focused, slightly relaxed when the cat is calm, and flattened against the head when extremely defensive or aggressive. In cats, flattened ears generally indicate that it feels
675: 209:. However, the reason why cats purr is still uncertain. Cats may purr for a variety of reasons, including when they are hungry, happy, or anxious. In some cases, purring is thought to be a sign of contentment and encouragement for further interaction. Purring is believed to indicate a 154:
Moelk used a phonetic alphabet to transcribe or write down the different sounds. She claimed that cats had six different forms of meows to represent friendliness, confidence, dissatisfaction, anger, fear, and pain. Moelk classified eight other sounds involved in mating and fighting.
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into pulse bursts and hybrid pulse bursts with tonal endings. Broadband sounds are separated into four groups: non-tonal broadband sounds, broadband sounds with tonal beginnings, broadband sounds with short tonal elements, and broadband sounds with long tonal endings.
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involves the cat being petted and then suddenly attacking and running away. Other problems are house soiling, scratching furniture, and bringing dead prey into the house. It is these kinds of behaviors that put a strain on the relationship between cats and people.
553:– The cat is lying on its belly, or it may be sitting; Its back is almost horizontal when standing and moving; Its breathing normal, with its legs bent or extended (when standing); Its tail is curved back or straight upward and may twitch when positioned downward. 695:
or to bond (this grooming is usually done between familiar cats). They also sometimes lick humans, which may indicate affection. Oral grooming for domestic and feral cats is a common behavior. Domestic cats spend about 8% of waking time grooming themselves.
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A mew is a high-pitched meow that is often produced by domestic kittens. It is apparently used to solicit attention from their mother, but they are also used by adult cats. By around three to four weeks of age, kittens do not mew when at least one
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Cats, domestic or wild, participate in social behaviors, even though it is thought that most cat species (besides lions) are solitary, antisocial animals. These include socialization between humans and other cats, social learning, and conflicts.
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Purring is sometimes accompanied by other sounds, though this varies between individual cats. Some may only purr, while others emit low-level outbursts, sometimes called "lurps" or "yowps". It was once believed that only the cats of the
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There are 52 measured cat personality traits in cats, with one study showing that five reliable personality factors were found using principal-axis factor-analysis: neuroticism, extroversion, dominance, impulsiveness and agreeableness.
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Cat vocalizations have been categorized according to a range of characteristics. In 1944, Mildred Moelk published the first phonetic study of cat sounds and classified the 16 different vocal patterns into three main classes:
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is present, and at four to five months of age, kittens stop mewing altogether. Adult cats rarely meow to each other, and so adult meowing to human beings is likely to be a post-domestication extension of mewing by kittens.
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A direct stare by a cat usually communicates a challenge or threat and is more likely to be seen in high-ranking cats; lower-ranking cats usually withdraw in response. The direct stare is often used during
547:– The cat may walk around in a more comfortable manner with its tail up to the sky. Cats often walk through houses with their tails standing up high above them, making them look grander and more elegant. 324:
Although videos which seemingly show cats speaking in human language are frequently shared on the internet, differences in cats' vocal tract prevent them from vocalising human language exactly. Instead,
312:). The meow can be assertive, plaintive, friendly, bold, welcoming, attention-soliciting, demanding, or complaining. It can even be silent, where the cat opens its mouth but does not produce any sound. 361:
Cats sometimes make excited chirping or chattering noises when observing or stalking prey. These sounds range from quiet clicking sounds to a loud but sustained chirping mixed with an occasional meow.
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The call is a loud, rhythmic sound, that is made with the mouth closed. It is primarily associated with female cats soliciting males, and sometimes occurs in males when fighting with each other. A
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A cat that urinates outside the litter box may indicate dissatisfaction with the box, due to a variety of factors such as substrate texture, cleanliness, and privacy. It can also be a sign of
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explain they are modifying the "meow" vocalisation to mimic certain human words. For example, a cat which frequently hears its owner say "no" may learn to use "mow" in a low tone.
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made a purring sound when air was passed through them, meaning that muscle contraction is not required. Instead, the sounds were made possible by connective tissue embedded in the
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that lowered the frequency of the sounds they produced. When an animal purrs, its vocal cords vibrate at a low frequency, which creates a distinctive rumbling sound produced with
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Szenczi, P.; Bánszegi, O.; Urrutia, A.; Faragó, T.; Hudson, R. (2016). "Mother–offspring recognition in the domestic cat: Kittens recognize their own mother's call".
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Casey, Rachel; Bradshaw, John (November 2008). "The effects of additional socialisation for kittens in a rescue centre on their behaviour and suitability as a pet".
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Miller classified vocalizations into categories according to the sound produced: the purr, meow, chirrup, chirp, call, and growl/snarl/hiss, and the howl/moan/wail.
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pheromone produced in glands in the cheeks. This is known as a contentment pheromone. Synthetic versions of the feline facial pheromone are available commercially.
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Herbst, Christian T.; Prigge, Tamara; Garcia, Maxime; Hampala, Vit; Hofer, Riccardo; Weissengruber, Gerald E.; Svec, Jan G.; Fitch, W. Tecumseh (November 2023).
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are more tonal, while moans are long and slowly modulated. On the other hand, anger wails are combined with growls, while yowls are similar to howls but longer.
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greeting calls or murmurs, extended vocal dialogues between cats in separate cages, "frustration" calls during training, or extinction of conditioned responses.
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with alternate paws at intervals of one to two seconds. They may knead while sitting on their owner's lap, which may be painful if the cat has sharp claws.
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or for territorial reasons. In contrast to a direct stare, cats will lower their eyelids or slowly blink them to show trust and affection to their owners.
4005: 1941: 3540: 679: 3520: 3159: 577:– Sometimes combined with a stretch this is another posture of a relaxed cat. Having the mouth open and no teeth exposed indicates playfulness. 1001: 2638: 1463: 1085: 983: 641:
However, if panting is excessive or the cat appears in distress, it may be a symptom of a more serious condition, such as a nasal blockage,
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Brown, Kenneth A.; Buchwald, Jennifer S.; Johnson, Judith R.; Mikolich, Dennis J. (November 1978). "Vocalization in the cat and kitten".
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Head-bumping and cheek rubbing may be displays of social dominance as they are often exhibited by a dominant cat toward a subordinate.
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or juvenile traits that persist in the adults, kneading may be a relic juvenile behavior retained in adult domestic cats. It may also
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A cat's posture can be friendly or aggressive, depending on the situation. Some of the most basic and familiar cat postures include:
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Male cats on poor diets are susceptible to crystal formation in the urine which can block the urethra and lead to health problems.
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sounds produced with the mouth held tensely open in the same position (growls, snarls, hisses, spits, chattering, and chirping).
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Litchfield, Carla A.; Quinton, Gillian; Tindle, Hayley; Chiera, Belinda; Kikillus, K. Heidy; Roetman, Philip (23 August 2017).
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Two cats lay with each other to conserve body heat, notice how they lay on their stomach and side indicating relaxation
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Owens, Jessica L.; Olsen, Mariana; Fontaine, Amy; Kloth, Christopher; Kershenbaum, Arik; Waller, Sara (June 2017).
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Cats sometimes "head-bump" humans or other cats with the front part of the head; this action is referred to as "
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are greeted by another cat or a human. Therefore, people can mimic the sound to reassure and greet pet cats.
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Touching noses, sometimes referred to as "sniffing noses", as a sign of affection and to mark territory.
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Crowell-Davis, Sharon L. (2007). "Cat Behaviour: Social Organization, Communication & Development".
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injections), and instances of kitten deprivation. Less common calls from mature cats included purring,
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Caro, T. M. (1980). "Effects of the Mother, Object Play, and Adult Experience on Predation in Cats".
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Miyazaki, M.; Yamashita, T.; Suzuki, Y.; Saito, Y.; Soeta, S.; Taira, H.; Suzuki, A. (October 2006).
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known as a "tomcat duel" may follow. Dominance is also seen as an underlying factor that depict how
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sounds produced when the mouth is first opened and then gradually closed (meowing, howling, yowling)
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is the prey, the response is to catch the prey, and the reward is to relieve the hunger sensation.
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Eckstein, Robert A.; Hart, Benjamin L. (2000). "Grooming and Control of Fleas in Cats".
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The howl, moan, and wail sounds are commonly vocalized during threatening situations.
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The purr is a continuous, soft, vibrating sound made in the throat by most species of
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A cat hissing and arching its back to make itself appear larger to ward off a threat
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1st Intl. Workshop on Vocal interactivity in-and-between Humans, Animals and Robots
973: 833:. Cats urinate by squatting onto a horizontal surface, while standing up. Unlike a 414: 160: 1886:
Schötz, S. (June 2015). "Agonistic vocalisations in domestic cats: A case study".
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Tavernier, Chloé; Ahmed, Sohail; Houpt, Katherine Albro; Yeon, Seong Chan (2020).
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Cats sometimes repeatedly tread their front paws on humans or soft objects with a
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Schwartz, Stefanie (June 2003). "Separation anxiety syndrome in dogs and cats".
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Brown, K. A.; Buchwald, J. S.; Johnson, J. R.; Mikolich, D. J. (November 1978).
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may need to be reintroduced or separated to avoid fights in a closed household.
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Cat Behaviour: The Predatory & Social Behaviour of Domestic & Wild Cats
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Bradshaw, John W.S. (January 2016). "Sociality in cats: A comparative review".
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McPherson, F.J; Chenoweth, P.J (April 2012). "Mammalian sexual dimorphism".
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has erect hairs on her tail and back. This indicates excitement or curiosity
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Little, Susan E. (2011). "Female Reproduction". In Little, Susan E. (ed.).
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temperature, female cats may pant for several days up to weeks postpartum.
468:; the Cyprus cat is arched with erect hair to appear larger and threatening 444: 3053: 1652: 1246: 1230: 4020: 4015: 3942: 3867: 3762: 3652: 3480: 3413: 3403: 3269: 3209: 1660: 1388: 1278:"Visual classification of feral cat Felis silvestris catus vocalizations" 1262: 857: 450:
Surprised cats have enlarged pupils and erect ears held slightly backward
289:
The most familiar sounds of adult cats are "meow" or "miaow" (pronounced
263: 229: 159:
over territory or food), episodes of acute stress or pain (e.g., routine
1602:
Schötz, Susanne; van de Weijer, Joost; Eklund, Robert (25 August 2017).
1293: 54:
Cats can use a range of communication methods, including vocal, visual,
3970: 3913: 3812: 1215: 1183: 735: 661:), is considered to be abnormal and is treated as a medical emergency. 658: 245: 206: 1742: 3965: 3882: 3782: 3637: 3455: 3314: 3199: 2864:. Wallingford, Oxfordshire: CAB International. pp. 78, 198–200. 1679:"People Are Seriously Convinced That Their Cats Are Speaking English" 992:
A conflict situation of one tomcat meeting another, making loud calls
868: 650: 377: 241: 221: 70:
or biting each other. They also use olfactory communication, such as
1199: 727:
are transferred to the person or object being kneaded or scratched.
43:. Studies have shown that domestic cats tend to meow much more than 1009:
Domestic cats fighting shortly after being introduced to each other
3752: 3742: 2699:. Chichester, West Sussex: John Wiley & Sons. pp. 49–59. 1815: 995: 980: 972: 872: 811: 807: 795: 747: 673: 654: 624: 345: 280: 257: 253: 248:
rapidly, causing air vibrations during inhalation and exhalation.
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Alder, H. E. (1955). "Some Factors of Observational Learning".
2722:"The 'Feline Five': An exploration of personality in pet cats ( 587:
threatened and may attack used as a defense or attack posture.
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Turner, Dennis C.; Bateson, Paul Patrick Gordon (2000-06-08).
1356:. Association of Animal Behavior Professionals. Archived from 1184:"Vocalizing in the House-Cat; A Phonetic and Functional Study" 1074:
Turner, Dennis C.; Bateson, Paul Patrick Gordon (2000-06-08).
929:
Cats between the age of three and nine weeks are sensitive to
31: 2599:(2nd ed.). University Press, Cambridge. pp. 69–70. 786:
position which communicates that she is receptive to mating.
304: 236:
studies, is that cats produce the purring noise by using the
1379:
Herbst, Christian T.; Svec, Jan G.; Fitch, Tecumseh (2023).
875:
into the more-volatile chemical MMB. Felinine is a possible
86: 3120:. New York: Garland Publishing. pp. 164–216, 227–231. 298: 2885:
Heath, Sarah E. (2007). "Behaviour Problems and Welfare".
2826:
Bernstein, Penny L. (2007). "The Human–Cat Relationship".
2024:. Wallingford, Oxfordshire: CAB International. p. 58. 1902:"Test to determine how well you know feline body language" 2101:
American Society for the Prevention of Cruelty to Animals
852:
The urine of mature male cats in particular contains the
2697:
Free-ranging Cats: Behaviour, Ecology & Management
2341:
Journal of the American Veterinary Medical Association
2116:"Cat Slow Blinking: Why Your Cat Blinks Slowly at You" 1928:
An Ethogram for Behavioral Studies of the Domestic Cat
1605:
Phonetic Characteristics of Domestic Cat Vocalisations
1533:
Anatomy by Dyce, K.M., Sack, W.O. and Wensing, C.J.G."
2543:"Cat Pheromones – Synthetic Feline Facial Pheromones" 2830:. Dordrecht, Netherlands: Springer. pp. 47–89. 1888:
Proceedings from Fonetik 2015: Lund, June 8–10, 2015
4162: 4119: 3979: 3941: 3901: 3598: 3579: 3506: 3372: 3295: 3250: 3175: 1981:. Saint Louis: W. B. Saunders. pp. 1195–1227. 301: 295: 3071: 2789: 2593:Turner, Dennis C.; Bateman, Patrick, eds. (2000). 2036: 742:the cat and make it feel good, similar to a human 734:Since most of the preferred "domestic traits" are 140:sounds produced with the mouth closed (murmurs – 1106:Library of Congress, Washington, D.C. 20540 USA 823:tail), and supra-caudal (at the base of tail). 2792:The Domestic Cat: The Biology of Its Behaviour 1455:The Domestic Cat: The Biology of Its Behaviour 1077:The Domestic Cat: The Biology of Its Behaviour 961:role by breaking up fights among littermates. 3153: 2627:Sunquist, Mel; Sunquist, Fiona (2017-05-15). 220:researching cat vocalizations found that the 8: 2788:Turner, Dennis C.; Bateson, Patrick (1988). 2523:. VetWest Animal Hospitals. 2 February 2008 2281:"Why Does My Cat Lick Me? 7 Common Reasons" 1567: 1565: 39:methods have been significantly altered by 3595: 3160: 3146: 3138: 2521:"Communication – How do cats communicate?" 1428:"The complicated truth about a cat's purr" 1102:"How do cats communicate with each other?" 368:argues that chattering and chirping mimic 3078:. New York: Simon and Schuster. pp.  2759: 2749: 2671: 2004: 1979:The Cat: Clinical Medicine and Management 1672: 1670: 1597: 1595: 1504: 1309: 1158: 3074:The Cat: History, Biology, and Behaviour 2912: 2910: 2908: 2906: 2889:. Springer, Dordrecht. pp. 91–107. 1347: 1345: 1343: 1341: 1339: 1337: 1335: 1333: 1331: 1329: 871:. It then slowly degrades via bacterial 244:to alternately dilate and constrict the 2796:. Cambridge University Press. pp.  2467:"Cat Nose Tap: Why Do Cats Touch Noses" 2172:. www.best-cat-tips.com. Archived from 1066: 23:A primary form of cat communication is 3521:Associazione Nazionale Felina Italiana 3111: 3109: 3107: 3105: 3103: 3101: 3099: 3065: 3063: 2956: 2954: 2952: 2950: 2948: 2946: 2944: 2942: 2940: 2855: 2853: 2851: 2849: 2847: 2821: 2819: 2817: 2198:Common Sense Book of Complete Cat Care 1405: 1394: 125: 16:Feline means of exchanging information 2984: 2982: 2783: 2781: 2779: 1703:Laurent, Anaëlle (December 8, 2020). 1677:Rennex, Michelle (12 February 2020). 1421: 1419: 806:Cats communicate through scent using 7: 2965:. Dordrecht, Netherlands: Springer. 2201:. Quill/William Morrow. p. 66. 1231:"Vocalization in the cat and kitten" 380:. It is used as a hunting strategy. 2417:Johnson-Bennett, Pam (3 May 2012). 746:. Kneading is often a precursor to 27:, such as the position of the ears. 3546:Governing Council of the Cat Fancy 1987:10.1016/b978-1-4377-0660-4.00040-5 1548:10.1111/j.2042-3306.1988.tb01554.x 1188:The American Journal of Psychology 14: 4006:Congenital sensorineural deafness 3561:The International Cat Association 3516:American Cat Fanciers Association 2991:The Journal of Genetic Psychology 2862:The Behaviour of the Domestic Cat 2396:10.1016/j.anireprosci.2012.02.007 700:scratch itchy areas of the body. 4193: 4192: 3541:Fédération Internationale Féline 2919:Applied Animal Behaviour Science 2465:McGonagle, Joanne (2024-01-12). 2306:Applied Animal Behaviour Science 2231:"Panting in Cats: Is It Normal?" 2120:Veterinary Healthcare Associates 893: 885: 493: 475: 457: 443: 429: 291: 126:Problems playing this file? See 102: 2633:. University of Chicago Press. 2043:. Barron's Educational Series. 1784:"Why Do Cats Chatter at Birds?" 232:. One hypothesis, supported by 3003:10.1080/00221325.1955.10532903 2931:10.1016/j.applanim.2008.01.003 2673:10.1016/j.chembiol.2006.08.013 2549:. 20 June 2018. Archived from 2140:"Cat articles on Iams website" 1862:"Stages of Kitten Development" 1574:Journal of Veterinary Behavior 1458:. Cambridge University Press. 1080:. Cambridge University Press. 691:Cats often lick other cats as 1: 3092:– via Internet Archive. 3046:10.1016/S0163-1047(80)92456-5 3034:Behavioral and Neural Biology 2860:Bradshaw, John W. S. (1992). 2810:– via Internet Archive. 2318:10.1016/s0168-1591(00)00095-2 2053:– via Internet Archive. 1131:Journal of Veterinary Science 862:3-mercapto-3-methylbutan-1-ol 2751:10.1371/journal.pone.0183455 2022:Feline Behaviour and Welfare 1127:"Feline vocal communication" 261:could purr. Now people know 4175:List of longest-living cats 4056:Lower urinary tract disease 4036:Hypertrophic cardiomyopathy 3556:Southern Africa Cat Council 2384:Animal Reproduction Science 2353:10.2460/javma.2003.222.1526 2039:Communicating with Your Cat 1731:Developmental Psychobiology 1641:Developmental Psychobiology 1235:Developmental Psychobiology 285:A cat meowing for attention 4242: 3531:Cat Aficionado Association 1586:10.1016/j.jveb.2015.09.004 1542:(5): 387. September 1988. 1022:interact with each other. 707: 274: 240:and/or the muscles of the 181: 4188: 4011:Feline corneal sequestrum 3536:Cat Fanciers' Association 2035:Helgren, J. Anne (1999). 1536:Equine Veterinary Journal 1506:10.1016/j.cub.2023.09.014 464:A Tabby cat hissing at a 357:A cat chattering at birds 3526:Canadian Cat Association 3116:Leyhausen, Paul (1979). 2695:Spotte, Stephen (2014). 2573:"Scent Glands on Cats |" 860:which is a precursor to 327:animal behaviour experts 211:positive emotional state 4170:List of individual cats 3070:Beadle, Muriel (1977). 2660:Chemistry & Biology 2256:"Cat Panting Explained" 2195:Vine, Louis L. (1992). 2020:Fraser, Andrew (2012). 1812:Virtual Pet Behaviorist 1182:Moelk, Mildred (1944). 1143:10.4142/jvs.2020.21.e18 847:urinary tract problems. 790:Olfactory communication 72:marking their territory 60:olfactory communication 4046:Infectious peritonitis 4041:Immunodeficiency virus 2630:Wild Cats of the World 2166:"Common Cat Behaviors" 2071:. Three Rivers Press. 1952:Warner Bros. Discovery 1531:Textbook of Veterinary 1404:Cite journal requires 1010: 993: 978: 951:observational learners 803: 782:neck as she assumes a 688: 657:or shallow breathing ( 630: 358: 286: 202: 201:Video of a cat purring 91: 28: 4111:Viral rhinotracheitis 4086:Tritrichomonas foetus 3551:New Zealand Cat Fancy 3419:Cats in ancient Egypt 2114:JacobP (2023-06-28). 1808:"Meowing and Yowling" 1653:10.1002/dev.420110605 1247:10.1002/dev.420110605 1008: 991: 976: 799: 686: 665:Tactile communication 628: 356: 284: 200: 90: 22: 4219:Animal communication 3673:Colorpoint Shorthair 3571:World Cat Federation 3305:Cat–dog relationship 2491:"Play Therapy Pt. 2" 2260:The Cat Health Guide 2093:"Aggression in Cats" 1491:(21): 4727–4732.e4. 1389:10.25365/phaidra.420 1045:Animal communication 421:Visual communication 409:Howl, moan, and wail 392:is the cry of a cat 218:University of Vienna 216:Scientists from the 3873:Traditional Persian 3658:California Spangled 3628:Brazilian Shorthair 3424:Cultural depictions 3409:Cat-scratch disease 2963:The Welfare of Cats 2887:The Welfare of Cats 2828:The Welfare of Cats 2742:2017PLoSO..1283455L 2441:"Cat Behavior Tips" 2146:on 25 February 2010 1958:on 21 November 2011 1764:Schötz, S. (2013). 1497:2023CBio...33E4727H 1352:Miller, P. (2000). 931:human socialization 831:territorial marking 814:, and chemicals or 687:Cat grooming itself 78:Vocal communication 3793:Oriental Shorthair 3703:European Shorthair 3618:American Shorthair 3566:World Cat Congress 2495:Cats International 2419:"Bunting Behavior" 1942:"Reading Your Cat" 1426:Dowling, Stephen. 1360:on 5 November 2013 1354:"Whisker whispers" 1011: 994: 979: 804: 689: 631: 359: 287: 203: 92: 29: 4206: 4205: 4031:Hepatic lipidosis 3937: 3936: 3633:British Shorthair 3242:Tortoiseshell cat 2640:978-0-226-51823-7 2471:The Tiniest Tiger 1743:10.1002/dev.21402 1465:978-0-521-63648-3 1294:10.1093/cz/zox013 1087:978-0-521-63648-3 1055:Dog communication 1006: 989: 977:Two cats fighting 909: 908: 684: 563:Anxious/ovulating 484:tortoiseshell cat 400:Growl, spit, hiss 354: 234:electromyographic 198: 116: 37:cat communication 4231: 4196: 4195: 4180:Cats in folklore 3778:Norwegian Forest 3748:Kurilian Bobtail 3728:Japanese Bobtail 3708:Exotic Shorthair 3596: 3461:National Cat Day 3218: 3162: 3155: 3148: 3139: 3132: 3131: 3113: 3094: 3093: 3077: 3067: 3058: 3057: 3029: 3023: 3022: 2986: 2977: 2976: 2958: 2935: 2934: 2925:(1–2): 196–205. 2914: 2901: 2900: 2882: 2876: 2875: 2857: 2842: 2841: 2823: 2812: 2811: 2798:112–113, 159–168 2795: 2785: 2774: 2773: 2763: 2753: 2717: 2711: 2710: 2692: 2686: 2685: 2675: 2651: 2645: 2644: 2624: 2618: 2617: 2615: 2613: 2596:The Domestic Cat 2590: 2584: 2583: 2581: 2579: 2569: 2563: 2562: 2560: 2558: 2539: 2533: 2532: 2530: 2528: 2517: 2511: 2510: 2508: 2506: 2501:on 19 April 2007 2497:. Archived from 2487: 2481: 2480: 2478: 2477: 2462: 2456: 2455: 2453: 2451: 2436: 2430: 2429: 2427: 2425: 2414: 2408: 2407: 2390:(3–4): 109–122. 2379: 2373: 2372: 2336: 2330: 2329: 2301: 2295: 2294: 2292: 2291: 2277: 2271: 2270: 2268: 2266: 2252: 2246: 2245: 2243: 2241: 2229:Spielman, Bari. 2226: 2220: 2219: 2217: 2215: 2192: 2186: 2185: 2183: 2181: 2162: 2156: 2155: 2153: 2151: 2142:. Archived from 2136: 2130: 2129: 2127: 2126: 2111: 2105: 2104: 2089: 2083: 2082: 2061: 2055: 2054: 2042: 2032: 2026: 2025: 2017: 2011: 2010: 2008: 1974: 1968: 1967: 1965: 1963: 1954:. Archived from 1938: 1932: 1931: 1924: 1918: 1917: 1915: 1913: 1904:. Archived from 1898: 1892: 1891: 1883: 1877: 1876: 1874: 1872: 1866:Alley Cat Rescue 1858: 1852: 1851: 1849: 1847: 1833: 1827: 1826: 1824: 1822: 1804: 1798: 1797: 1795: 1794: 1780: 1774: 1773: 1761: 1755: 1754: 1726: 1720: 1719: 1717: 1715: 1700: 1694: 1693: 1691: 1689: 1674: 1665: 1664: 1636: 1630: 1629: 1618:. pp. 5–6. 1617: 1610: 1599: 1590: 1589: 1569: 1560: 1559: 1525: 1519: 1518: 1508: 1476: 1470: 1469: 1449: 1443: 1442: 1440: 1438: 1423: 1414: 1413: 1407: 1402: 1400: 1392: 1376: 1370: 1369: 1367: 1365: 1349: 1324: 1323: 1313: 1273: 1267: 1266: 1226: 1220: 1219: 1179: 1173: 1172: 1162: 1122: 1116: 1115: 1113: 1112: 1098: 1092: 1091: 1071: 1007: 990: 897: 889: 882: 881: 750:, and many cats 723:is likely these 685: 497: 479: 461: 447: 433: 364:An article from 355: 311: 310: 307: 306: 303: 300: 297: 199: 118: 117: 89: 4241: 4240: 4234: 4233: 4232: 4230: 4229: 4228: 4209: 4208: 4207: 4202: 4184: 4158: 4121:Cats by country 4115: 3982: 3975: 3933: 3897: 3773:Neva Masquerade 3600: 3587: 3581: 3575: 3502: 3375: 3368: 3359:Righting reflex 3291: 3246: 3214: 3171: 3166: 3136: 3135: 3128: 3115: 3114: 3097: 3090: 3069: 3068: 3061: 3031: 3030: 3026: 2988: 2987: 2980: 2973: 2960: 2959: 2938: 2916: 2915: 2904: 2897: 2884: 2883: 2879: 2872: 2859: 2858: 2845: 2838: 2825: 2824: 2815: 2808: 2787: 2786: 2777: 2736:(8): e0183455. 2719: 2718: 2714: 2707: 2694: 2693: 2689: 2653: 2652: 2648: 2641: 2626: 2625: 2621: 2611: 2609: 2607: 2592: 2591: 2587: 2577: 2575: 2571: 2570: 2566: 2556: 2554: 2553:on 20 June 2018 2541: 2540: 2536: 2526: 2524: 2519: 2518: 2514: 2504: 2502: 2489: 2488: 2484: 2475: 2473: 2464: 2463: 2459: 2449: 2447: 2438: 2437: 2433: 2423: 2421: 2416: 2415: 2411: 2381: 2380: 2376: 2347:(11): 1526–32. 2338: 2337: 2333: 2303: 2302: 2298: 2289: 2287: 2279: 2278: 2274: 2264: 2262: 2254: 2253: 2249: 2239: 2237: 2228: 2227: 2223: 2213: 2211: 2209: 2194: 2193: 2189: 2179: 2177: 2164: 2163: 2159: 2149: 2147: 2138: 2137: 2133: 2124: 2122: 2113: 2112: 2108: 2091: 2090: 2086: 2079: 2065:Morris, Desmond 2063: 2062: 2058: 2051: 2034: 2033: 2029: 2019: 2018: 2014: 1997: 1976: 1975: 1971: 1961: 1959: 1940: 1939: 1935: 1926: 1925: 1921: 1911: 1909: 1908:on 20 June 2018 1900: 1899: 1895: 1885: 1884: 1880: 1870: 1868: 1860: 1859: 1855: 1845: 1843: 1835: 1834: 1830: 1820: 1818: 1806: 1805: 1801: 1792: 1790: 1788:The Spruce Pets 1782: 1781: 1777: 1763: 1762: 1758: 1728: 1727: 1723: 1713: 1711: 1709:animalwised.com 1702: 1701: 1697: 1687: 1685: 1676: 1675: 1668: 1638: 1637: 1633: 1626: 1615: 1608: 1601: 1600: 1593: 1571: 1570: 1563: 1527: 1526: 1522: 1485:Current Biology 1478: 1477: 1473: 1466: 1451: 1450: 1446: 1436: 1434: 1425: 1424: 1417: 1403: 1393: 1378: 1377: 1373: 1363: 1361: 1351: 1350: 1327: 1282:Current Zoology 1275: 1274: 1270: 1228: 1227: 1223: 1200:10.2307/1416947 1181: 1180: 1176: 1124: 1123: 1119: 1110: 1108: 1100: 1099: 1095: 1088: 1073: 1072: 1068: 1063: 1041: 1028: 996: 981: 971: 947: 945:Social learning 927: 914: 792: 779: 761: 716:kneading action 712: 710:Kneading (cats) 706: 693:social grooming 674: 672: 667: 623: 606: 593: 584: 514: 505: 504: 503: 502: 501: 498: 489: 488: 487: 480: 471: 470: 469: 462: 453: 452: 451: 448: 439: 438: 437: 434: 423: 411: 402: 386: 366:The Spruce Pets 346: 344: 335: 294: 290: 279: 273: 188: 186: 180: 133: 132: 124: 122: 121: 120: 119: 103: 100: 93: 87: 80: 35:play. As such, 17: 12: 11: 5: 4239: 4238: 4235: 4227: 4226: 4221: 4211: 4210: 4204: 4203: 4201: 4200: 4189: 4186: 4185: 4183: 4182: 4177: 4172: 4166: 4164: 4160: 4159: 4157: 4156: 4151: 4149:United Kingdom 4146: 4141: 4136: 4131: 4125: 4123: 4117: 4116: 4114: 4113: 4108: 4103: 4098: 4096:Skin disorders 4093: 4088: 4083: 4078: 4073: 4068: 4063: 4058: 4053: 4051:Leukemia virus 4048: 4043: 4038: 4033: 4028: 4023: 4018: 4013: 4008: 4003: 3998: 3993: 3987: 3985: 3977: 3976: 3974: 3973: 3968: 3963: 3958: 3953: 3947: 3945: 3939: 3938: 3935: 3934: 3932: 3931: 3926: 3921: 3916: 3911: 3905: 3903: 3899: 3898: 3896: 3895: 3890: 3888:Turkish Angora 3885: 3880: 3875: 3870: 3865: 3860: 3855: 3850: 3845: 3840: 3835: 3830: 3825: 3820: 3815: 3810: 3805: 3800: 3795: 3790: 3785: 3780: 3775: 3770: 3765: 3760: 3755: 3750: 3745: 3740: 3735: 3730: 3725: 3720: 3715: 3710: 3705: 3700: 3695: 3690: 3685: 3680: 3675: 3670: 3668:Chinese Li Hua 3665: 3660: 3655: 3650: 3645: 3640: 3635: 3630: 3625: 3620: 3615: 3610: 3604: 3602: 3593: 3577: 3576: 3574: 3573: 3568: 3563: 3558: 3553: 3548: 3543: 3538: 3533: 3528: 3523: 3518: 3512: 3510: 3504: 3503: 3501: 3500: 3495: 3494: 3493: 3488: 3483: 3478: 3468: 3463: 3458: 3453: 3451:Cats and Islam 3448: 3443: 3438: 3437: 3436: 3431: 3421: 3416: 3411: 3406: 3401: 3396: 3391: 3386: 3380: 3378: 3370: 3369: 3367: 3366: 3361: 3356: 3351: 3346: 3341: 3340: 3339: 3334: 3324: 3323: 3322: 3312: 3307: 3301: 3299: 3293: 3292: 3290: 3289: 3284: 3283: 3282: 3277: 3267: 3262: 3256: 3254: 3248: 3247: 3245: 3244: 3239: 3234: 3229: 3224: 3219: 3212: 3207: 3202: 3197: 3192: 3187: 3181: 3179: 3173: 3172: 3167: 3165: 3164: 3157: 3150: 3142: 3134: 3133: 3126: 3095: 3088: 3059: 3024: 2997:(1): 159–177. 2978: 2971: 2936: 2902: 2895: 2877: 2870: 2843: 2836: 2813: 2806: 2775: 2712: 2705: 2687: 2666:(10): 1071–9. 2646: 2639: 2619: 2605: 2585: 2564: 2534: 2512: 2482: 2457: 2431: 2409: 2374: 2331: 2312:(2): 141–150. 2296: 2272: 2247: 2221: 2207: 2187: 2157: 2131: 2106: 2084: 2077: 2056: 2049: 2027: 2012: 1995: 1969: 1933: 1919: 1893: 1878: 1853: 1841:Dictionary.com 1828: 1799: 1775: 1756: 1737:(5): 568–577. 1721: 1695: 1666: 1647:(6): 559–570. 1631: 1624: 1591: 1561: 1520: 1471: 1464: 1444: 1415: 1406:|journal= 1371: 1325: 1288:(3): 331–339. 1268: 1241:(6): 559–570. 1221: 1194:(2): 184–205. 1174: 1117: 1093: 1086: 1065: 1064: 1062: 1059: 1058: 1057: 1052: 1047: 1040: 1037: 1027: 1024: 970: 967: 946: 943: 926: 923: 913: 910: 907: 906: 903: 899: 898: 890: 827:Urine spraying 791: 788: 778: 775: 760: 757: 708:Main article: 705: 702: 671: 668: 666: 663: 645:disease, head 622: 619: 605: 602: 592: 589: 583: 580: 579: 578: 572: 566: 560: 554: 548: 542: 536: 530: 524: 513: 510: 499: 492: 491: 490: 481: 474: 473: 472: 463: 456: 455: 454: 449: 442: 441: 440: 435: 428: 427: 426: 425: 424: 422: 419: 410: 407: 401: 398: 385: 382: 343: 340: 334: 331: 275:Main article: 272: 269: 182:Main article: 179: 176: 152: 151: 148: 145: 123: 101: 96: 95: 94: 85: 84: 83: 79: 76: 49:social ladders 15: 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Manee 3718:German Rex 3608:Abyssinian 3508:Registries 3491:Ship's cat 3476:Bodega cat 3232:Calico cat 3177:Felinology 3019:1297110751 2871:085198715X 2527:5 November 2476:2024-03-18 2290:2024-03-18 2125:2024-03-18 1793:2024-03-18 1714:5 November 1688:5 November 1364:5 November 1137:(1): e18. 1111:2024-03-05 1061:References 854:amino acid 829:is also a 816:pheromones 744:stretching 725:pheromones 649:, or drug 569:Stretching 539:Aggressive 466:Cyprus cat 318:littermate 128:media help 45:feral cats 4129:Australia 4091:Roundworm 4026:Heartworm 3983:disorders 3943:Landraces 3929:Serengeti 3878:Tonkinese 3848:Singapura 3808:Pixie-bob 3803:Peterbald 3723:Himalayan 3688:Devon Rex 3663:Chartreux 3584:full list 3446:Feral cat 3429:fictional 3389:Cat cafés 3374:Human–cat 3265:Declawing 3237:Tabby cat 3227:Black cat 3195:Dwarf cat 2547:PetMD.com 2240:7 January 2097:ASPCA.org 1556:0425-1644 1302:1674-5507 1255:0012-1630 1208:0002-9556 1151:1229-845X 969:Dominance 949:Cats are 866:peptidase 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Index

Two cats crouch, facing each other, on either side of a door. Both cats ears are angled back.
body language
Cats
domestication
feral cats
social ladders
tactile
olfactory communication
body language
grooming
marking their territory
A cat meowing
media help
purring
prophylactic
conspecific
Purr
felines
positive emotional state
University of Vienna
larynxes
vocal folds
harmonics
electromyographic
vocal folds
larynx
glottis
genus
Felis
Panthera

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