1030:
1214:
1140:
136:
517:
1124:
115:
1238:
attacked its prey from below and behind, much like the modern great white shark does. The deepest bite marks on the dolphin's ribs indicate the shark aimed for the abdomen of its prey to inflict a fatal bite quickly and incapacitate its prey, and that when the dolphin was attacked a second time, it
1504:
Dana J. Ehret; Bruce J. Macfadden; Douglas S. Jones; Thomas J. Devries; David A. Foster; Rodolfo Salas-Gismondi (2012). "Origin of the white shark
Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru". Palaeontology. 55 (6): 1139–1153.
1051:
teeth being found worldwide. The species is divided into two forms based on tooth morphology, each with a unique evolutionary line. The maximum adult length is estimated between 5 and 7 m (16 and 23 ft). Smaller individuals were about 2.6–4.5 metres (8.5–14.8 ft) long.
1105:
and their roots are more uneven in thickness and shape, with more narrow builds and rounder lobes, somewhat making the appearance of a heart. But the crowns of the lower teeth of this species are described to be more straight and identical to the lower teeth of
1264:
1156:
The broad form is characterized by broad-shaped teeth often described as being identical to that of modern great whites besides the lack of serrations. Fossil evidence shows that the broad-form is the direct ancestor of the genus
933:, which was a move that was first proposed by Glikman (1964) but was long rejected beforehand. Later discoveries supports Ward & Bonavi's conclusion of its ancestry to the great white. An analysis of a newly discovered
1228:. It most likely would have been one of the major predators in its ecosystem, preying upon small whales and other mammals. Trace fossils in the form of tooth marks on the bones of a Pliocene dolphin of the species
1881:
McCormack, Jeremy; Griffiths, Michael L.; Kim, Sora L.; Shimada, Kenshu; Karnes, Molly; Maisch, Harry; Pederzani, Sarah; Bourgon, Nicolas; Jaouen, Klervia; Becker, Martin A.; Jöns, Niels (May 31, 2022).
1239:
was bitten near the dorsal fin, suggesting that the dolphin rolled over while injured. The size of the bites indicates further that the shark responsible was estimated to be 4 m (13 ft) long.
1246:
from Peru described in 2017 showcases that the shark also included fish in its diet. Stomach contents preserved with the fossil show this shark also fed on pilchards such as ancient sardines (genus
1043:
teeth can grow up to 8.9 cm (3.5 in) in length, suggesting a very large shark. Its body was probably very similar to that of modern great whites. It is also believed to have a
635:. Although this conclusion is widely accepted, some scientists disagree, with a study by Whitenack and Gottfried (2010) demonstrating geometrically morphological differences between
443:
can reach lengths up to 3.5 in (7.5 cm) and are found worldwide. It is believed to be an ancestor to the great white shark, an argument supported by the transitional species
2015:
Collareta, Alberto; Landini, Walter; Chacaltana Budiel, César
Augusto; Valdivia Vera, Waldir; Altamirano Sierra, Ali J.; Urbina Schmitt, Mario; Bianucci, Giovanni (March 2017).
1714:
Cione, Alberto Luis; Cabrera, Daniel
Alfredo; Barla, María Julia (2012). "Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic".
1006:
and the great white "have been documented from the same deposits, hence the former cannot be a chronospecific ancestor of the latter." He also criticized the fact that the "
1081:. True to its name, each tooth is "hooked", the point of which is shifted away from the middle axis, in the direction of the corners of the mouth. In this, the teeth of
2148:
2081:
1303:
554:, although some indications show that he had coined the taxon as early as 1838. Throughout the early and mid 20th century, different genera and species of other
598:). Eventually, later studies would begin to show that much of the sharks within the genus and two species are distinct from each other, discarding the genus
2017:"A well preserved skeleton of the fossil shark Cosmopolitodus hastalis from the late Miocene of Peru, featuring fish remains as fossilized stomach contents"
1525:"A Well Preserved Skeleton of the Fossil Shark Cosmopolitodus Hastalis from the Late Miocene of Peru, Featuring Fish Remains as Fossilized Stomach Contents"
2189:
2135:
2068:
1969:
Ciampaglio, C., Ricketts,D. Miocene
Madness: Fossil Finder ID and Information Guide. Bakersfield, California. Buena Vista Museum of Natural History.
1811:
2229:
2204:
1282:
621:, citing the similarities between the teeth of the two and that of modern mako sharks. In 2001, a study by Ward & Bonavi (2001) considered
2214:
602:
altogether and creating another issue on what new taxa the sharks should be placed in. As of now, the genus is still uncertain and debated.
2234:
1524:
2209:
2199:
1615:"A morphometric approach for addressing toothbased species delimitation in fossil mako sharks, Isurus (Elasmobranchii: Lamniformes)"
1667:"Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru"
1110:. The hooked teeth, which probably would be efficient for gripping prey, would suggest a diet of smaller and medium-sized animals.
1029:
1665:
Ehret, Dana J.; MacFadden, Bruce J.; Jones, Douglas S.; Devries, Thomas J.; Foster, David A.; Salas-Gismondi, Rodolfo (2012).
1188:
teeth. Unlike the broad-form, the narrow form is believed to be the ancestor of two extinct sharks, the hooked-tooth "mako" (
929:
is a direct ancestor of the great white and is more related to it than other makos. They moved the species into the genus
135:
1375:
1824:
1044:
1213:
983:. However, the study also noted that this would only be done if the putative sister species is conspecific with
963:
teeth showing basal serrations. Cione et al. (2012) also noted a possibility of moving all species in the genus
2224:
1761:"New example of Cosmopolitodus hastalis (Lamniformes, Lamnidae) from the Miocene of South Korea | Zoodiversity"
1461:
2219:
1330:"The occurrence of early Pleistocene marine fish remains from the Gulf Coast of Mobile County, Alabama, USA"
482:
1760:
2036:
1010:
1797:
1013:
has never been tested through phylogenetic analyses," and denoted that as of 2021, the argument that the
2161:
2094:
1193:
783:
588:
582:
532:
1420:
Zevallos-Valdivia, L.; Martínez-Pérez, C.; García-Flores, V.; Chávez-Valencia, A.; Botella, H. (2023).
1810:
Patrick L. Jambura; Julia Türtscher; Faviel A. López-Romero; Catalina
Pimiento; Jürgen Kriwet (2019).
1895:
1723:
1678:
1626:
935:
826:
445:
1884:"Trophic position of Otodus megalodon and great white sharks through time revealed by zinc isotopes"
1069:
576:
2001:
1937:
1863:
1696:
1644:
1443:
1402:
1351:
1230:
1182:
The narrow form has more slender teeth than its broad-form counterpart, which resemble broadened
306:
130:
2166:
2099:
1139:
2153:
2194:
2086:
2016:
1929:
1911:
1855:
1847:
922:
843:
563:
34:
1979:
1919:
1903:
1839:
1772:
1739:
1731:
1686:
1634:
1544:
1536:
1529:
Rivista
Italiana di Paleontologia e Stratigrafia (Research in Paleontology and Stratigraphy)
1433:
1394:
1341:
516:
1164:
1380:
comb. nov. (Chondrichthyes, Elasmobranchii) from the
Miocene of Pohang City, South Korea"
1174:
to reflect its difference from the narrow-form, although this is scientifically invalid.
1063:
teeth are somewhat similar to those of other mako sharks, especially the extinct species
1899:
1727:
1682:
1630:
1487:
1123:
1924:
1883:
1277:
1078:
951:, remarking that the two genera were separated solely due to the lack of serrations in
187:
177:
647:
is considered as the "traditional view" in the debate regarding the shark's taxonomy.
114:
2183:
1941:
1700:
1691:
1666:
1595:
1570:
1447:
1406:
1355:
1304:"Skeletal remains of the Miocene lamniform shark, Cosmopolitodus hastalis, from Peru"
1269:
1252:), suggesting bony fish featured prominently in the shark's diet while it was young.
1225:
1184:
547:
470:
450:
260:
72:
1867:
1648:
1506:
1263:
631:
531:, especially the status of its genus, has long been subject to debate. The initial
243:
1823:
Pimiento, C.; Cantalapiedra, J.L.; Shimada, K.; Field, D.J.; Smaers, J.B. (2019).
1735:
2059:
1159:
1098:
555:
440:
432:
394:
207:
47:
2126:
1907:
1639:
1614:
1540:
1398:
1259:
92:
57:
1915:
1851:
1777:
1248:
1074:
972:
594:
537:
390:
147:
97:
41:
1933:
1859:
1217:
Specimen CPI-7899, featuring aggregate of skeletal and dermal fish remains
546:
for the narrow and broad-form variations respectively by Swiss naturalist
2120:
2053:
1346:
1329:
386:
217:
167:
87:
82:
67:
62:
52:
30:
1744:
987:. The study concluded that there is indeed a putative sister species of
2140:
2073:
1596:"Additions to and a review of the Miocene Shark and Ray fauna of Malta"
975:
that would occur if one of them, which was traditionally identified as
102:
77:
1843:
1438:
1421:
921:
and noted a strong morphological similarity between it and the extant
157:
2030:
389:
that lived between thirty and one million years ago during the late
1212:
1028:
515:
489:
397:
382:
197:
943:'s ancestry to the great white, however, they also proposed that
422:
was reassigned to the genus, and thus became the second species
2034:
2002:"Jaws -- 4 million BC: How an extinct shark attacked its prey"
1812:
On the origin of the great white shark
Carcharodon carcharias.
1073:. Adult upper teeth are generally 2–4 cm long with an un
1825:"Evolutionary pathways toward gigantism in sharks and rays"
426:. However, some researchers still consider both species of
1170:
This form is also commonly labeled by its species synonym
1571:"Central Paratethyan shark fauna (Ipolytarnóc, Hungary)"
1328:
Ebersole, J.A.; Ebersole, S.M.; Cicimurri, D.J. (2017).
939:
by Ehret et al. (2012) further cemented the theory of
1002:
However, Yun argued that the tooth fossil remains of
609:, a review by Holec et al. (1995) placed the species
1957:
Fossil Shark Teeth of the World: A collector's guide
1613:
Whitenack †, Lisa B.; Gottfried, Michael D. (2010).
289:
276:
250:
2110:
2043:
1422:"First record of palaeozoic vertebrates from Peru"
1167:in Peru show an evolutionary mosaic between them.
1798:"Great White Sharks and Prehistoric White Sharks"
1462:"Carcharodon hastalis Agassiz 1843 (white shark)"
1017:lineage with narrow, serrated teeth evolved from
917:Ward & Bonavi (2001) reexamined the teeth of
574:included two species of extinct ancestral makos,
1507:https://doi.org/10.1111/j.1475-4983.2012.01201.x
566:. Sharks previously identified as variations of
558:would be clumped into the two taxa as the genus
8:
1792:
1790:
1788:
1519:
1517:
1515:
1021:with broad, unserrated teeth is uncertain.
477:" meaning "citizen of the world" and ὀδών "
2031:
959:and pointing out examples of Late Miocene
605:Shortly after the discarding of the genus
113:
20:
1923:
1776:
1743:
1690:
1638:
1564:
1562:
1437:
1345:
629:as conspecific and declared the latter a
500:is derived from the Ancient Greek ξίφος "
1660:
1658:
1242:A well-preserved skeleton of a juvenile
999:for it, resolving the paraphyly issue.
1590:
1588:
1294:
1376:"A tooth of the extinct lamnid shark,
1283:List of prehistoric cartilaginous fish
496:meaning "spear". The disputed species
1369:
1367:
1365:
617:as a species of mako under the genus
7:
1486:Crumpton, Nick (November 14, 2012).
955:and the lack of lateral cusplets in
552:Recherches sur les poissons fossiles
1488:"Great whites 'not from megashark'"
818:
760:
752:
745:
721:
699:
677:
655:
453:analyses have been done for proof.
2021:Repositorio Institucional Ingemmet
1619:Journal of Vertebrate Paleontology
979:, is a putative sister species of
14:
2190:Prehistoric fish of South America
1311:Erlanger Geologische Abhandlungen
1759:Yun, Chan-gyu (August 7, 2020).
1692:10.1111/j.1475-4983.2012.01201.x
1569:Kocsis, László (February 2007).
1426:Spanish Journal of Palaeontology
1262:
1138:
1122:
410:(other common names include the
134:
592:), and the false-toothed mako (
16:Extinct genus of mackerel shark
1:
2230:Fossil taxa described in 1843
2205:Fossil taxa described in 1964
1736:10.1016/j.geobios.2011.06.002
1387:Acta Palaeontologica Romaniae
2215:Pliocene species extinctions
1190:Cosmopolitodus/Isurus planus
1077:cutting edge and no lateral
504:" meaning "sword" and ὀδών "
2235:Taxa named by Louis Agassiz
925:. The study concluded that
2251:
1908:10.1038/s41467-022-30528-9
1224:was a confirmed hunter of
2210:Miocene first appearances
1640:10.1080/02724630903409055
1399:10.35463/j.apr.2022.01.02
1163:, and specimens from the
1045:cosmopolitan distribution
910:and its close relatives.
840:
823:
816:
780:
765:
758:
750:
743:
726:
719:
704:
697:
682:
675:
660:
312:
305:
273:
268:
249:
242:
131:Scientific classification
129:
121:
112:
23:
2200:Prehistoric shark genera
488:may be derived from the
1778:10.15407/zoo2020.05.433
1541:10.13130/2039-4942/8005
995:and proposed the taxon
730:Cosmopolitodus hastalis
481:" meaning "tooth". The
416:broad-tooth white shark
403:Cosmopolitodus hastalis
255:Cosmopolitodus hastalis
1302:Klug, S.; Kriwet, J.,
1218:
1037:
997:Carcharodon plicatilis
844:Carcharodon carcharias
643:. The scientific name
562:began to be used as a
524:
400:. Its type species is
2162:Paleobiology Database
2095:Paleobiology Database
1980:"Great White Lineage"
1888:Nature Communications
1378:Cosmopolitodus planus
1231:Astadelphis gastaldii
1216:
1198:Carcharomodus escheri
1032:
906:Relationship between
784:Carcharomodus escheri
768:Cosmopolitodus planus
589:Carcharomodus escheri
586:, the serrated mako (
519:
449:, but as of 2021, no
2112:Carcharodon hastalis
1575:Geologica Carpathica
1347:10.18476/pale.v10.a6
1097:teeth are much more
971:to avoid a possible
936:Carcharodon hubbelli
827:Carcharodon hubbelli
469:is derived from the
446:Carcharodon hubbelli
316:Carcharodon hastalis
1955:Cocke, Joe (2002).
1900:2022NatCo..13.2980M
1728:2012Geobi..45..167C
1683:2012Palgy..55.1139E
1631:2010JVPal..30...17W
947:should be moved to
595:Parotodus benedenii
535:was first given as
508:" meaning "tooth".
1219:
1185:Isurus oxyrhinchus
1038:
663:Isurus schoutedeni
583:Isurus retroflexus
550:in his 1843 paper
525:
412:extinct giant mako
2177:
2176:
2037:Taxon identifiers
1844:10.1111/evo.13680
1439:10.7203/sjp.25691
923:great white shark
914:
913:
900:
899:
891:
890:
882:
881:
873:
872:
864:
863:
855:
854:
804:
803:
795:
794:
756:Narrow form
685:Isurus praecursor
572:Oxyrhina xiphodon
568:Oxyrhina hastalis
564:wastebasket taxon
544:Oxyrhina xiphodon
512:Taxonomic history
374:
373:
368:
360:
352:
348:Oxyrhina xiphodon
344:
340:Oxyrhina hastalis
336:
328:
320:
299:
286:
238:
2242:
2170:
2169:
2157:
2156:
2144:
2143:
2131:
2130:
2129:
2103:
2102:
2090:
2089:
2077:
2076:
2064:
2063:
2062:
2032:
2025:
2024:
2012:
2006:
2005:
1998:
1992:
1991:
1989:
1987:
1976:
1970:
1967:
1961:
1960:
1952:
1946:
1945:
1927:
1878:
1872:
1871:
1829:
1820:
1814:
1808:
1802:
1801:
1794:
1783:
1782:
1780:
1756:
1750:
1749:
1747:
1711:
1705:
1704:
1694:
1677:(6): 1139–1153.
1662:
1653:
1652:
1642:
1610:
1604:
1603:
1592:
1583:
1582:
1566:
1557:
1556:
1554:
1552:
1543:. Archived from
1521:
1510:
1502:
1496:
1495:
1483:
1477:
1476:
1474:
1472:
1458:
1452:
1451:
1441:
1417:
1411:
1410:
1384:
1374:Yun, C. (2021).
1371:
1360:
1359:
1349:
1325:
1319:
1318:
1308:
1299:
1272:
1267:
1266:
1209:Hunting behavior
1142:
1126:
819:
814:Broad form
761:
753:
746:
722:
700:
678:
656:
651:
650:
527:The taxonomy of
520:Fossil teeth of
408:broad-tooth mako
366:
358:
350:
342:
334:
326:
318:
297:
291:
284:
278:
252:
236:
229:
139:
138:
122:Tooth series of
117:
107:
44:
29:Temporal range:
21:
2250:
2249:
2245:
2244:
2243:
2241:
2240:
2239:
2225:Pliocene sharks
2180:
2179:
2178:
2173:
2165:
2160:
2152:
2147:
2139:
2134:
2125:
2124:
2119:
2106:
2098:
2093:
2085:
2080:
2072:
2067:
2058:
2057:
2052:
2039:
2029:
2028:
2014:
2013:
2009:
2000:
1999:
1995:
1985:
1983:
1978:
1977:
1973:
1968:
1964:
1954:
1953:
1949:
1880:
1879:
1875:
1827:
1822:
1821:
1817:
1809:
1805:
1796:
1795:
1786:
1758:
1757:
1753:
1713:
1712:
1708:
1664:
1663:
1656:
1612:
1611:
1607:
1594:
1593:
1586:
1568:
1567:
1560:
1550:
1548:
1547:on June 1, 2022
1523:
1522:
1513:
1503:
1499:
1485:
1484:
1480:
1470:
1468:
1460:
1459:
1455:
1419:
1418:
1414:
1382:
1373:
1372:
1363:
1334:Palaeodiversity
1327:
1326:
1322:
1306:
1301:
1300:
1296:
1291:
1268:
1261:
1258:
1211:
1206:
1194:serrated "mako"
1180:
1165:Pisco Formation
1154:
1153:
1152:
1151:
1150:
1143:
1135:
1134:
1127:
1116:
1058:
1027:
967:into the genus
915:
901:
892:
883:
874:
865:
856:
805:
796:
645:Isurus hastalis
627:Isurus xiphodon
623:Isurus hastalis
514:
464:
459:
359:(Agassiz, 1856)
351:(Agassiz, 1843)
343:(Agassiz, 1843)
335:(Agassiz, 1843)
332:Isurus xiphodon
327:(Agassiz, 1843)
324:Isurus hastalis
319:(Agassiz, 1843)
285:(Agassiz, 1843)
264:
258:
235:
227:
133:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
27:
17:
12:
11:
5:
2248:
2246:
2238:
2237:
2232:
2227:
2222:
2220:Miocene sharks
2217:
2212:
2207:
2202:
2197:
2192:
2182:
2181:
2175:
2174:
2172:
2171:
2158:
2145:
2132:
2116:
2114:
2108:
2107:
2105:
2104:
2091:
2078:
2065:
2049:
2047:
2045:Cosmopolitodus
2041:
2040:
2035:
2027:
2026:
2007:
1993:
1971:
1962:
1947:
1873:
1838:(3): 588–599.
1815:
1803:
1784:
1771:(5): 433–438.
1751:
1722:(2): 167–172.
1706:
1654:
1605:
1584:
1558:
1511:
1497:
1478:
1453:
1412:
1361:
1320:
1293:
1292:
1290:
1287:
1286:
1285:
1280:
1278:List of sharks
1274:
1273:
1257:
1254:
1226:marine mammals
1210:
1207:
1205:
1202:
1179:
1176:
1145:A narrow form
1144:
1137:
1136:
1128:
1121:
1120:
1119:
1118:
1117:
1115:
1112:
1057:
1054:
1026:
1023:
1019:Cosmopolitodus
1008:Cosmopolitodus
1004:Cosmopolitodus
991:distinct from
965:Cosmopolitodus
945:Cosmopolitodus
931:Cosmopolitodus
912:
911:
903:
902:
898:
897:
894:
893:
889:
888:
885:
884:
880:
879:
876:
875:
871:
870:
867:
866:
862:
861:
858:
857:
853:
852:
849:
848:
839:
836:
835:
832:
831:
822:
817:
815:
811:
810:
807:
806:
802:
801:
798:
797:
793:
792:
789:
788:
779:
776:
775:
772:
771:
764:
759:
757:
751:
749:
744:
742:
739:
738:
735:
734:
725:
720:
718:
715:
714:
711:
710:
703:
698:
696:
693:
692:
689:
688:
681:
676:
674:
671:
670:
667:
666:
659:
654:
649:
513:
510:
473:κοσμοπολίτης "
467:Cosmopolitodus
463:
460:
458:
455:
430:as species of
428:Cosmopolitodus
387:mackerel shark
381:is an extinct
378:Cosmopolitodus
372:
371:
370:
369:
367:(Jordan, 1907)
361:
356:Oxyrhina plana
353:
345:
337:
329:
321:
310:
309:
303:
302:
301:
300:
287:
271:
270:
266:
265:
259:
247:
246:
240:
239:
232:Cosmopolitodus
225:
221:
220:
215:
211:
210:
205:
201:
200:
195:
191:
190:
188:Elasmobranchii
185:
181:
180:
178:Chondrichthyes
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
28:
25:Cosmopolitodus
15:
13:
10:
9:
6:
4:
3:
2:
2247:
2236:
2233:
2231:
2228:
2226:
2223:
2221:
2218:
2216:
2213:
2211:
2208:
2206:
2203:
2201:
2198:
2196:
2193:
2191:
2188:
2187:
2185:
2168:
2163:
2159:
2155:
2150:
2146:
2142:
2137:
2133:
2128:
2122:
2118:
2117:
2115:
2113:
2109:
2101:
2096:
2092:
2088:
2083:
2079:
2075:
2070:
2066:
2061:
2055:
2051:
2050:
2048:
2046:
2042:
2038:
2033:
2022:
2018:
2011:
2008:
2003:
1997:
1994:
1981:
1975:
1972:
1966:
1963:
1958:
1951:
1948:
1943:
1939:
1935:
1931:
1926:
1921:
1917:
1913:
1909:
1905:
1901:
1897:
1893:
1889:
1885:
1877:
1874:
1869:
1865:
1861:
1857:
1853:
1849:
1845:
1841:
1837:
1833:
1826:
1819:
1816:
1813:
1807:
1804:
1799:
1793:
1791:
1789:
1785:
1779:
1774:
1770:
1766:
1762:
1755:
1752:
1746:
1741:
1737:
1733:
1729:
1725:
1721:
1717:
1710:
1707:
1702:
1698:
1693:
1688:
1684:
1680:
1676:
1672:
1671:Palaeontology
1668:
1661:
1659:
1655:
1650:
1646:
1641:
1636:
1632:
1628:
1624:
1620:
1616:
1609:
1606:
1601:
1597:
1591:
1589:
1585:
1580:
1576:
1572:
1565:
1563:
1559:
1546:
1542:
1538:
1534:
1530:
1526:
1520:
1518:
1516:
1512:
1508:
1501:
1498:
1493:
1489:
1482:
1479:
1467:
1463:
1457:
1454:
1449:
1445:
1440:
1435:
1432:(1): 95–100.
1431:
1427:
1423:
1416:
1413:
1408:
1404:
1400:
1396:
1392:
1388:
1381:
1379:
1370:
1368:
1366:
1362:
1357:
1353:
1348:
1343:
1340:(1): 97–115.
1339:
1335:
1331:
1324:
1321:
1316:
1312:
1305:
1298:
1295:
1288:
1284:
1281:
1279:
1276:
1275:
1271:
1270:Sharks portal
1265:
1260:
1255:
1253:
1251:
1250:
1245:
1240:
1237:
1233:
1232:
1227:
1223:
1215:
1208:
1203:
1201:
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1191:
1187:
1186:
1177:
1175:
1173:
1168:
1166:
1162:
1161:
1148:
1141:
1132:
1129:A broad form
1125:
1113:
1111:
1109:
1104:
1100:
1096:
1092:
1088:
1084:
1080:
1076:
1072:
1071:
1070:Isurus desori
1066:
1062:
1055:
1053:
1050:
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1042:
1035:
1031:
1024:
1022:
1020:
1016:
1012:
1009:
1005:
1000:
998:
994:
990:
989:C. carcharias
986:
982:
981:C. carcharias
978:
974:
970:
966:
962:
958:
957:C. carcharias
954:
950:
946:
942:
938:
937:
932:
928:
924:
920:
909:
905:
904:
896:
895:
887:
886:
878:
877:
869:
868:
860:
859:
851:
850:
847:
846:
845:
838:
837:
834:
833:
830:
829:
828:
821:
820:
813:
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808:
800:
799:
791:
790:
787:
786:
785:
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774:
773:
770:
769:
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762:
755:
754:
748:
747:
741:
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737:
736:
733:
732:
731:
724:
723:
717:
716:
713:
712:
709:
708:
707:Isurus desori
702:
701:
695:
694:
691:
690:
687:
686:
680:
679:
673:
672:
669:
668:
665:
664:
658:
657:
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652:
648:
646:
642:
638:
634:
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624:
620:
616:
612:
608:
603:
601:
597:
596:
591:
590:
585:
584:
579:
578:
577:Isurus desori
573:
569:
565:
561:
557:
553:
549:
548:Louis Agassiz
545:
541:
539:
534:
533:binomial name
530:
523:
518:
511:
509:
507:
503:
499:
495:
491:
487:
484:
483:specific name
480:
476:
472:
471:Ancient Greek
468:
461:
456:
454:
452:
448:
447:
442:
437:
435:
434:
429:
425:
421:
420:Isurus planus
417:
413:
409:
405:
404:
399:
396:
393:to the Early
392:
388:
384:
380:
379:
365:
364:Isurus planus
362:
357:
354:
349:
346:
341:
338:
333:
330:
325:
322:
317:
314:
313:
311:
308:
304:
298:Agassiz, 1856
296:
295:
288:
283:
282:
275:
274:
272:
267:
262:
257:
256:
248:
245:
241:
237:Glikman, 1964
234:
233:
226:
223:
222:
219:
216:
213:
212:
209:
206:
203:
202:
199:
198:Selachimorpha
196:
194:Subdivision:
193:
192:
189:
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
40:30–1.07
36:
32:
26:
22:
19:
2111:
2044:
2020:
2010:
1996:
1984:. Retrieved
1982:. elasmo.com
1974:
1965:
1956:
1950:
1891:
1887:
1876:
1835:
1831:
1818:
1806:
1768:
1765:Zoodiversity
1764:
1754:
1745:11336/139163
1719:
1715:
1709:
1674:
1670:
1625:(1): 17–25.
1622:
1618:
1608:
1600:ResearchGate
1599:
1578:
1574:
1549:. Retrieved
1545:the original
1532:
1528:
1500:
1491:
1481:
1471:December 17,
1469:. Retrieved
1465:
1456:
1429:
1425:
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1377:
1337:
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1310:
1297:
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1235:
1234:reveal that
1229:
1221:
1220:
1204:Paleobiology
1197:
1189:
1183:
1181:
1171:
1169:
1158:
1155:
1146:
1130:
1107:
1102:
1099:heterodontic
1094:
1090:
1089:differ from
1086:
1082:
1068:
1064:
1060:
1059:
1048:
1040:
1039:
1033:
1018:
1014:
1007:
1003:
1001:
996:
992:
988:
984:
980:
976:
968:
964:
960:
956:
952:
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944:
940:
934:
930:
926:
918:
916:
907:
842:
841:
825:
824:
782:
781:
767:
766:
729:
728:
727:
706:
705:
684:
683:
662:
661:
644:
640:
636:
632:nomen dubium
630:
626:
622:
618:
614:
610:
606:
604:
599:
593:
587:
581:
575:
571:
567:
559:
551:
543:
536:
528:
526:
521:
505:
501:
497:
493:
485:
478:
475:kosmopolítēs
474:
466:
465:
451:phylogenetic
444:
438:
431:
427:
423:
419:
418:). In 2021,
415:
411:
407:
402:
401:
377:
376:
375:
363:
355:
347:
339:
331:
323:
315:
293:
292:
280:
279:
254:
253:
244:Type species
231:
230:
123:
24:
18:
1894:(1): 2980.
1535:(1). 2017.
1466:Fossilworks
1393:(1): 9–16.
1244:C. hastalis
1236:C. hastalis
1222:C. hastalis
1192:), and the
1178:Narrow-form
1160:Carcharodon
1147:C. hastalis
1131:C. hastalis
1108:C. hastalis
1091:C. hastalis
1065:C. hastalis
1049:C. hastalis
1041:C. hastalis
1034:C. hastalis
1025:Description
1015:Carcharodon
993:C. hastalis
985:C. hastalis
977:C. xiphodon
969:Carcharodon
961:C. hastalis
953:C. hastalis
949:Carcharodon
941:C. hastalis
927:I. hastalis
919:I. hastalis
908:C. hastalis
641:I. xiphodon
637:I. hastalis
556:lamniformes
529:C. hastalis
522:C. hastalis
433:Carcharodon
395:Pleistocene
281:C. hastalis
208:Lamniformes
124:C. hastalis
2184:Categories
1289:References
1114:Broad-form
1011:morphotype
184:Subclass:
2060:Q20686304
1986:April 20,
1942:249235478
1916:2041-1723
1852:1558-5646
1832:Evolution
1701:128666594
1551:August 8,
1448:256585278
1407:242113412
1356:134476316
1249:Sardinops
1103:C. planus
1095:I. desori
1087:I. desori
1083:C. planus
1061:C. planus
973:paraphyly
462:Etymology
424:C. planus
391:Oligocene
294:C. planus
154:Kingdom:
148:Eukaryota
35:Calabrian
2195:Lamnidae
2154:11083619
2127:Q3803933
2121:Wikidata
2054:Wikidata
1934:35641494
1868:59224442
1860:30675721
1649:85821523
1581:: 27–40.
1492:BBC News
1256:See also
1172:xiphodon
1079:cusplets
1075:serrated
615:xiphodon
611:hastalis
607:Oxyrhina
600:Oxyrhina
560:Oxyrhina
540:hastalis
538:Oxyrhina
498:xiphodon
486:hastalis
457:Taxonomy
307:Synonyms
269:Species
218:Lamnidae
214:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
31:Rupelian
2141:5816397
2087:1223316
2074:4824543
1925:9156768
1896:Bibcode
1724:Bibcode
1716:Geobios
1679:Bibcode
1627:Bibcode
1047:, with
261:Agassiz
224:Genus:
204:Order:
174:Class:
2167:265174
2100:118027
1940:
1932:
1922:
1914:
1866:
1858:
1850:
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1647:
1446:
1405:
1354:
619:Isurus
502:xíphos
406:, the
398:epochs
263:, 1843
2149:IRMNG
2082:IRMNG
1938:S2CID
1864:S2CID
1828:(PDF)
1697:S2CID
1645:S2CID
1444:S2CID
1403:S2CID
1383:(PDF)
1352:S2CID
1307:(PDF)
1149:tooth
1133:tooth
1101:than
1056:Teeth
494:hasta
492:word
490:Latin
441:teeth
383:genus
2136:GBIF
2069:GBIF
1988:2022
1930:PMID
1912:ISSN
1856:PMID
1848:ISSN
1553:2019
1473:2021
1317:: 99
1085:and
1067:and
1036:jaws
639:and
625:and
613:and
580:and
542:and
506:odṓn
479:odṓn
439:Its
414:and
48:PreꞒ
1920:PMC
1904:doi
1840:doi
1773:doi
1740:hdl
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1537:doi
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1342:doi
570:or
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1364:^
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98:Pg
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2004:.
1990:.
1959:.
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1906::
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88:J
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63:S
58:O
53:Ꞓ
33:-
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