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Docodonta

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5641: 5646: 927: 799:(fit together), the pseudotalonid acts as a receptacle for cusp Y of the upper molar. Cusp Y is often termed the "pseudoprotocone" in this relationship. At the same time, cusp b of the lower molar shears into an area labial to cusp Y. Occlusion is completed when the rest of the upper molar slides between adjacent lower molar teeth, letting the rear edge of the preceding lower molar scrape against cusp X. This shearing-and-grinding process is more specialized than in any other early mammaliaform. 906: 148: 7251: 670: 659: 983:, and recurved molar cusps, which suggests a possible diet of fish or aquatic invertebrates. It was thought possible that docodonts had tendencies towards semi-aquatic habits, given their presence in wetland environments, although this could also be explained by the ease with which these environments preserve fossils compared with more terrestrial ones. Recent discoveries of other complete docodontans such as the specialised digging species 879: 774:
wear facet is found on the labial edge of cusp X, extending along the crest leading to cusp A. Cusp Y, a unique feature of docodonts, is positioned directly behind cusp X. Many docodonts have one or two additional cusps (cusps B and E) in front of cusp A. Cusp B is almost always present and is usually shifted slightly labial relative to cusp A. Cusp E, which may be absent in later docodonts, is positioned lingual to cusp B.
7456: 6357: 6351: 727: 6345: 7246: 120: 837:, an unusual collection of Jurassic mammals with tall pointed cusps. Relative to docodonts, shuotheriids have pseudotalonids which are positioned further forwards in their lower molars. This is potentially another case of convergent evolution, as shuotheriid are often considered true mammals related to modern 1523:" mammaliaforms, which had fairly simple lower molars with a straight row of large cusps. However, re-evaluations of mammaliaform tooth homology in the late 1990s established that docodonts were not closely related to either morganucodonts or therians. Instead, they were found to be similar to certain early " 773:
Cusps A and C lie in a row along the labial edge of the tooth (i.e., on the outer side, facing the cheek). Cusp A is located in front of cusp C and is typically the largest cusp in the upper molars. Cusp X lies lingual to cusp A (i.e., positioned inwards, towards the midline of the skull). A distinct
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was much smaller than either, at fewer than 156 g in adult body mass. The authors propose that the standard condition in modern small mammals (very high metabolism, rapid growth, and short lifespan) would not be adopted until true mammals in the Jurassic mammals. In addition, docodonts contradict
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The lower molars are longer than wide. On average, they have seven cusps arranged in two rows. The labial/outer row has the largest cusp, cusp a, which lies between two more cusps. The other major labial cusps are cusp b (a slightly smaller cusp in front of cusp a) and cusp d (a much smaller cusp
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Two additional lingual cusps may be present: cusp e and cusp df. Cusp e lies in front of cusp g and is roughly lingual to cusp b. Cusp df (“docodont cuspule f”) lies behind cusp c and is lingual to cusp d. There is some variation in the relative sizes, position, or even presence of some of these
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When seen from below, the upper molars have an overall subtriangular or figure-eight shape, wider (from side to side) than they are long (from front to back). The bulk of the tooth makes up four major cusps: cusps A, C, X, and Y. This overall structure is similar to the
1551:). These "symmetrodonts" have three major cusps (c, a, and b) set in a triangular arrangement on their lower molars. These cusps would be homologous to cusps c, a, and g in docodonts, which have a similar size and position. The lingual cusp (cusp X) is prominent in 714:. Most mammaliaforms have fairly simple molars primarily suited for shearing and slicing food. Docodonts, on the other hand, have developed specialized molars with crushing surfaces. The shape of each molar is defined by a characteristic pattern of conical 625:, plate-like ossified cartilage caps which terminate bone growth in adulthood. This suggests that docodont bones continued growing throughout their lifetime, like some other mammaliaforms and early mammals. The ankle is distinctive, with a downturned 344:, another group of Jurassic mammaliaforms, also shared some dental characteristics with docodonts. One study has suggested that shuotheriids are closely related to docodonts, though others consider shuotheriids to be true mammals, perhaps related to 5059:
Averianov, A. O.; Martin, T.; Lopatin, A. V.; Skutschas, P. P.; Vitenko, D. D.; Schellhorn, R.; Kolosov, P. N. (2024). "Docodontans from the Lower Cretaceous of Yakutia, Russia: new insights into diversity, morphology, and phylogeny of Docodonta".
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specimens found that docodonts had a slower metabolism and lower growth rates relative to modern mammals of the same size. The juvenile, which was 7 to 24 months old at the time of its death, was only 49% through the process of replacing its
388:-like species, hinting at impressive ecological diversity within the group. Many docodonts have muscular limbs and broad tail vertebrae, adaptations for burrowing or swimming. Like true mammals, docodonts have hair, a saddle-shaped 4730:
Alexander Averianov; Thomas Martin; Alexey Lopatin; Pavel Skutschas; Rico Schellhorn; Petr Kolosov; Dmitry Vitenko (2018). "A high-latitude fauna of mid-Mesozoic mammals from Yakutia, Russia". PLoS ONE. 13 (7): e0199983.
465:) is still relatively large and rests against the petrosal bone of the braincase, a remnant of a pre-mammalian style jaw joint. In true mammals, the postdentary elements detach fully and shrink further, becoming the 1527:", a broad and polyphyletic grouping of mammaliaforms with triangular upper molars. In particular, the closest relatives of Docodonta have been identified as certain Late Triassic "symmetrodonts", such as 441:) in the rear part of the dentary's inner edge. Nevertheless, they are very slender, hosting hooked prongs which start to converge towards an oval-shaped area immediately behind the dentary. The 437:, a connection which is strengthened relative to earlier mammaliaforms. The other bones in the jaw, known as postdentary elements, are still connected to the dentary and lie within a groove (the 4306:
Krusat, G. 1991 Functional morphology of Haldanodon exspectatus (Mammalia, Docodonta) from the Upper Jurassic of Portugal. Fifth Symposium on Mesozoic Terrestrial Ecosystems and Biota.
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Zhou, Chang-Fu; Bhullar, Bhart-Anjan; Neander, April; Martin, Thomas; Luo, Zhe-Xi (19 Jul 2019). "New Jurassic mammaliaform sheds light on early evolution of mammal-like hyoid bones".
859:, even proposed that shuotheriids and docodonts were most closely related to each other among mammaliaforms. The study named a new clade, Docodontiformes, to encompass the two groups. 795:
A distinct concavity or basin is apparent in the front half of each lower molar, between cusps a, g, and b. This basin has been named the pseudotalonid. When the upper and lower teeth
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teeth. Docodont teeth have been described as "pseudotribosphenic": a cusp on the inner half of the upper molar grinds into a basin on the front half of the lower molar, like a
3846:"Cranio-mandibular anatomy of Haldanodon exspectatus (Docodonta; Mammalia) from the late Jurassic of Portugal and its implications to the evolution of mammalian characters" 830:, as therian talonids lie at the back of the lower molar rather than the front. The opposite is true for docodont teeth, which have been described as "pseudotribosphenic". 324:
teeth of therian mammals. There is much uncertainty for how docodont teeth developed from their simpler ancestors. Their closest relatives may have been certain Triassic "
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behind cusp a). The lingual/inner row is shifted backwards (relative to the labial row) and has two large cusps: cusp g (at the front) and cusp c (at the back).
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like the docodonts. Nevertheless, docodonts are still closely related to crown-Mammalia, to a greater extent than many other early mammaliaform groups such as
4841:"Reigitherium (Meridiolestida, Mesungulatoidea) an Enigmatic Late Cretaceous Mammal from Patagonia, Argentina: Morphology, Affinities, and Dental Evolution" 396:. On the other hand, the postdentary bones are still attached to the jaw and skull, the nares (bony nostril rims) have yet to fuse, and in most species the 3989:
Panciroli, Elsa; Benson, Roger B. J.; Fernandez, Vincent; Humpage, Matthew; Martín-Serra, Alberto; Walsh, Stig; Luo, Zhe-Xi; Fraser, Nicholas C. (2021).
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which connects to two pairs of bony structures: the anterior hyoid cornu (a jointed series of rods which snake up to the braincase), and the posterior
7597: 3845: 1066: 1024:. Based on jaw length, the juvenile was 51-59% the weight of the 7-year-old adult. The closest comparisons among modern mammals were monotremes and 351:
For much of their history of study, docodont fossils were represented by isolated teeth and jaws. The first docodont known from decent remains was
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Kühne W. G. and Krusat, G. 1972. Legalisierung des Taxon Haldanodon (Mammalia, Docodonta). Neues Jahrbuch für Geologie Monatshefte 1972:300-302
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Martin, T.; Averianov, A. O.; Lang, A. J.; Schultz, J. A.; Wings, O. (2024). "Docodontans (Mammaliaformes) from the Late Jurassic of Germany".
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L. P. Tatarinov (1994). "On an unusual mammalian tooth from the Mongolian Jurassic". Journal of the Geological Society of London. 128: 119–125.
592:(rib pedestals), supporting powerful tail musculature. Most docodonts have gradually shrinking ribs, forming a subdued transition between the 4668: 4325:
Panciroli, Elsa; Benson, Roger B. J.; Fernandez, Vincent; Fraser, Nicholas C.; Humpage, Matt; Luo, Zhe-Xi; Newham, Elis; Walsh, Stig (2024).
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Prasad GVR, and Manhas BK. 2007. A new docodont mammal from the Jurassic Kota Formation of India. Palaeontologia electronica, 10.2: 1-11.
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earlier assumptions that high metabolism evolved in sync with ecological diversity, since their diversity far outpaces their metabolism.
493:(bony nostril holes) are small and paired, rather than fused into a single opening, and the rear edge of each naris is formed by a large 5645: 878: 1655:
of these groups (in their widest form) are not found in any other analyses, and therefore not accepted by all mammal palaeontologists.
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Docodont fossils have been recognized since the 1880s, but their relationships and diversity have only recently been well-established.
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Panciroli, Elsa; Benson, Roger B J; Fernandez, Vincent; Butler, Richard J; Fraser, Nicholas C; Luo, Zhe-Xi; Walsh, Stig (2021-07-30).
1087:, though most others consider haramiyidans to be closer to mammals than docodonts are. Docodonts may lie crownward of haramiyidans in 905: 4366:"Mandibular and dental characteristics of Late Triassic mammaliaform Haramiyavia and their ramifications for basal mammal evolution" 1092: 5018:
Waldman, M and Savage, R.J.G 1972 The first Jurassic mammal from Scotland. Journal of the Geological Society of London 128:119-125
743:* Left side: right maxilla molar and left dentary molar in occlusal view (looking onto the teeth). Cusp nomenclature is labelled. 3529:
Martin, Thomas; Jäger, Kai R. K.; Plogschties, Thorsten; Schwermann, Achim H.; Brinkkötter, Janka J.; Schultz, Julia A. (2020).
308:(modern-day North America, Europe, and Asia). They are distinguished from other early mammaliaforms by their relatively complex 7582: 1096: 550:. It allows for a narrower and more muscular throat and tongue, which are correlated with uniquely mammalian behaviors such as 5103: 4749:
Lopatin, A. V.; Averianov, A. O.; Kuzmin, I. T.; Boitsova, E. A.; Saburov, P. G.; Ivantsov, S. V.; Skutschas, P. P. (2020).
4128:(Tegotheriidae, Mammalia), a docodont from the Middle Jurassic of Western Siberia and phylogenetic analysis of Docodonta". 1513:
instead argued that docodont teeth were impossible to homologize with modern mammals. He drew comparisons to the teeth of
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mammal. Some authors have suggested splitting Docodonta into two families (Simpsonodontidae and Tegotheriidae), but the
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Luo, Zhe-Xi; Meng, Qing-Jin; Grossnickle, David M.; Liu, Di; Neander, April I.; Zhang, Yu-Guang; Ji, Qiang (2017).
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lacks lumbar ribs, so it has an abrupt transition from the thoracic to lumbar vertebrae like many modern mammals.
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Mao, Fangyuan; Li, Zhiyu; Wang, Zhili; Zhang, Chi; Rich, Thomas; Vickers-Rich, Patricia; Meng, Jin (2024-04-03).
3171: 3043: 4818:"A highly derived docodont from the Patagonian Late Cretaceous: evolutionary implications for Gondwanan mammals" 4049:(Mammalia, Docodonta) from the Late Jurassic (Kimmeridgian) of Portugal and its bearing for mammalian evolution" 5139: 3030: 3019: 457:) sends down a particularly well-developed prong known as the manubrium, which is sensitive to vibrations. The 129: 3530: 1509:", part of a broad group ancestral to true therian mammals according to their complex molars. A 1956 paper by 481:
Docodont skulls are generally fairly low, and in general form are similar to other early mammaliforms such as
4988: 3458: 6356: 4092: 2997: 745:* Right side: left maxilla and dentary molars in lingual view (from the perspective of the tongue, right). 7322: 4909: 7476: 6344: 4186:"Reconsideration of the systematic position of the Middle Jurassic mammaliaforms Itatodon and Paritatodon" 1502: 732: 546:). This hyoid system affords greater strength and flexibility than the simple, U-shaped hyoids of earlier 6350: 5106:. In Proceedings of the Zoological Society of London, 109:329-356). Oxford, UK: Blackwell Publishing Ltd. 7549: 4212: 3572:"A Swimming Mammaliaform from the Middle Jurassic and Ecomorphological Diversification of Early Mammals" 1088: 796: 449:, fits into a deep slot on the dentary which opens backwards, a characteristic unique to docodonts. The 7245: 4364:
Luo, Zhe-Xi; Gatesy, Stephen M.; Jenkins, Farish A.; Amaral, William W.; Shubin, Neil H. (2015-12-22).
5952: 4751:"A New Finding of a Docodontan (Mammaliaformes, Docodonta) in the Lower Cretaceous of Western Siberia" 3716:
Meng, Qing-Jin; Ji, Qiang; Zhang, Yu-Guang; Liu, Di; Grossnickle, David M.; Luo, Zhe-Xi (2015-02-13).
7523: 6980: 6414: 5592: 5069: 4967: 4951: 4910:"New docodontians from the Middle Jurassic of Siberia and reanalysis of docodonta interrelationships" 4762: 4605: 4444: 4377: 4245: 4137: 3938: 3799: 3729: 3657: 3583: 3400: 3281: 3070: 1058: 827: 589: 369: 317: 7382: 6604: 7010: 6564: 5470: 5132: 5119: 4816:
Pascual, Rosendo; Goin, Francisco J.; Gonzalez, Pablo; Ardolino, Alberto; Puerta, Pablo F. (2000).
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based on a phylogenetic analysis of Zhou et al. (2019) focusing on a wide range of mammaliamorphs:
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Luo, Zhe-Xi; Meng, Qing-Jin; Ji, Qiang; Liu, Di; Zhang, Yu-Guang; Neander, April I. (2015-02-13).
1069:. However, modern authors usually limit the term "Mammalia" to the crown group, excluding earlier 494: 7577: 7459: 7392: 7144: 7116: 7049: 6844: 6742: 6705: 6544: 6514: 5480: 5330: 5237: 5085: 5041: 5001: 4932: 4878: 4786: 4629: 4593: 4476: 4277: 4161: 4022: 3962: 3883: 3823: 3753: 3681: 3607: 3470: 3182: 2709: 1815: 855: 811: 767: 751: 593: 585: 438: 400: 321: 142: 7352: 7072: 7064: 6045: 5792: 741:
Upper (maxilla) molars are pink and lower (dentary) molars are blue. Anterior is to the right.
7554: 7187: 4560: 539: 7302: 7225: 7025: 6851: 6712: 6006: 5806: 5364: 4993: 4870: 4778: 4710: 4664: 4621: 4468: 4460: 4413: 4395: 4346: 4269: 4261: 4153: 4096: 4070: 4014: 3954: 3907: 3857: 3815: 3745: 3673: 3599: 3508: 3462: 3193: 1592: 610: 543: 535: 313: 249: 3990: 853:, have been considered members of either group. A 2024 study, describing the new shuotheriid 7541: 7312: 7109: 6689: 6125: 6036: 5972: 5761: 5077: 5033: 4983: 4975: 4924: 4860: 4852: 4770: 4732: 4702: 4656: 4613: 4452: 4433:"New evidence for mammaliaform ear evolution and feeding adaptation in a Jurassic ecosystem" 4403: 4385: 4338: 4253: 4200: 4145: 4060: 4006: 3995:(Mammaliformes, Docodonta) and the emergence of ecomorphological diversity in early mammals" 3946: 3899: 3807: 3737: 3665: 3591: 3500: 3454: 1608: 597: 577: 404: 397: 301: 43: 669: 658: 633:
which connects to the tibia via a trochlea (pulley-like joint). The only known specimen of
7372: 6283: 6169: 6052: 5875: 5822: 5676: 5663: 5604: 5490: 5401: 5380: 3645: 3571: 1640: 1588: 1565:, was originally considered to have been a very early mammaliaform which lived during the 1510: 1021: 1017: 630: 581: 518: 389: 297: 275: 39: 7514: 4496:"A Catalogue of the Mesozoic Mammalia in the Geological Department of the British Museum" 5073: 4971: 4766: 4609: 4448: 4381: 4249: 4141: 3942: 3803: 3733: 3661: 3587: 7423: 7418: 7279: 7158: 7057: 6995: 6719: 6504: 6387: 6374: 6308: 6220: 6185: 6099: 5982: 5859: 5206: 4928: 4408: 4365: 2371: 1686: 1672: 1648: 1130: 1113: 1074: 1070: 715: 638: 600:
regions of the spine. However, this developmental trait is not universal. For example,
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is variable between docodonts, as with many other mammaliaforms. The components of the
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Docodonts also see the first occurrence of a mammalian-style saddle-shaped complex of
7571: 7428: 7170: 7130: 6938: 6924: 6917: 6873: 6816: 6733: 6574: 6534: 6444: 6269: 6115: 6022: 5892: 5707: 5540: 5433: 5410: 5089: 5045: 5005: 4817: 4790: 4649:"The Late Triassic Record of Cynodonts: Time of Innovations in the Mammalian Lineage" 4326: 4165: 4088: 4065: 4044: 4026: 3966: 3926: 3827: 3757: 3717: 3685: 3492: 3442: 3373: 3327: 3314: 3303: 3292: 3226: 3054: 2668: 2534: 2503: 2347: 2102: 2078: 1791: 1597: 1524: 834: 711: 510: 502: 486: 462: 381: 358: 341: 325: 309: 77: 4936: 4882: 3718:"An arboreal docodont from the Jurassic and mammaliaform ecological diversification" 3611: 3474: 975:
from the Upper Jurassic of Portugal, were specialised for a semi-aquatic lifestyle.
296:). They were among the most common mammaliaforms of their time, persisting from the 7433: 7409: 7404: 7094: 7036: 6902: 6888: 6809: 6795: 6765: 6584: 6474: 6424: 6290: 6206: 6178: 6073: 6066: 6059: 6015: 5836: 5737: 5697: 5567: 5551: 5524: 4655:, Topics in Geobiology, Cham: Springer International Publishing, pp. 407–445, 4617: 4480: 4281: 3270: 3248: 3062: 2692: 2591: 2323: 2299: 2126: 2028: 1760: 1635: 1603: 1515: 1420: 1404: 1257: 1240: 1216: 991: 965: 933: 885: 718:, with sharp, concave crests connecting the center of each cusp to adjacent cusps. 703: 680: 602: 568: 530: 506: 446: 442: 368:. Recently, exceptionally preserved skeletons have been discovered in the Jurassic 5081: 5037: 4979: 1095:, but shift stemward relative to haramiyidans when the same data is put through a 4736: 4660: 4185: 3903: 1665:
Topology of Zhou et al. (2019), based on tooth, cranial, and postcranial traits:
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falls outside the docodont family tree, albeit as a close relative to the group.
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of living mammals. Previously, docodonts were sometimes regarded as belonging to
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Averianov, A. A.; Lopatin, A. V.; Krasnolutskii, S. A.; Ivantsov, S. V. (2010).
3384: 3360: 3349: 3338: 3204: 3008: 2766: 2634: 2551: 2476: 2452: 2275: 2143: 1997: 1839: 1733: 1709: 1617: 1557: 1520: 1506: 1377: 1320: 1079: 1054: 1000: 960: 945: 917: 894: 849: 823: 763: 726: 290: 52: 4706: 4648: 4342: 3504: 963:, but recent more complete fossils from China have shown this is not the case. 7002: 6972: 6959: 6895: 6866: 6837: 6823: 6698: 6645: 6624: 6434: 6297: 6276: 6255: 6135: 6080: 5902: 5852: 5845: 5784: 5717: 5533: 5460: 5450: 5387: 5340: 5299: 5286: 5249: 5197: 5155: 4856: 4774: 4432: 4315:
Paleontology and Geology of the Upper Jurassic Morrison Formation: Bulletin 36
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Cladograms based on phylogenetic analyses focusing on docodont relationships:
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mammals were traditionally thought to have been primarily ground dwelling and
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The forelimbs and hindlimbs generally have strong muscle attachments, and the
525: 498: 353: 218: 205: 97: 62: 7499: 4997: 4874: 4782: 4714: 4690: 4625: 4464: 4399: 4350: 4265: 4157: 4074: 4018: 3911: 3861: 3646:"Evolutionary development in basal mammaliaforms as revealed by a docodontan" 3512: 3493:"Jurassic shuotheriids show earliest dental diversification of mammaliaforms" 3466: 1647:, has previously been described as a docodont, though it is now considered a 7101: 6675: 6655: 6594: 5992: 5942: 5510: 5373: 5222: 5188: 4840: 4390: 3950: 3811: 3741: 3669: 3595: 2419: 1652: 1644: 1535: 1498: 1102: 1050: 1046: 1042: 995:
suggest Docodonta were more ecologically diverse than previously suspected.
897: 838: 819: 807: 759: 642: 626: 622: 361: 345: 330: 192: 159: 102: 4472: 4417: 4273: 4204: 3958: 3819: 3749: 3677: 3603: 1065:, owing to the complexity of their molars and the fact that they possess a 4327:"Jurassic fossil juvenile reveals prolonged life history in early mammals" 787:
cusps, though docodonts in general have a fairly consistent cusp pattern.
7493: 7362: 7332: 7194: 7151: 7123: 6880: 6749: 6454: 6227: 6155: 5179: 4647:
Abdala, Fernando; Gaetano, Leandro C. (2018), Tanner, Lawrence H. (ed.),
3888:(Mammaliformes: Docodonta) from the Middle Jurassic of the British Isles" 2651: 2402: 1973: 1626: 1580:
teeth, completely unrelated to docodonts or any Mesozoic mammaliaforms.
1544: 1529: 1428: 1359: 1062: 956: 942: 843: 841:. Docodont and shuotheriid teeth are so similar that some genera, namely 707: 573: 551: 547: 466: 426: 385: 365: 336: 305: 286: 279: 179: 92: 87: 72: 67: 57: 4456: 4257: 2258:
Topology of Panciroli et al. (2021), based on dentary and tooth traits:
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The lineage of Docodonta evolved prior to the origin of living mammals:
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of the braincase. Vertebrae at the base of the tail often have expanded
392:, and reduced postdentary jaw bones which are beginning to develop into 119: 17: 7342: 6830: 5770: 5727: 5417: 4865: 4633: 4010: 3158: 3147: 3136: 3125: 3114: 3103: 3092: 3081: 2814: 1887: 1629:. However, this identification is not certain, and in recent analyses, 1612: 1574: 1566: 803: 699: 689: 685: 470: 450: 430: 107: 82: 3570:
Ji, Qiang; Luo, Zhe-Xi; Yuan, Chong-Xi; Tabrum, Alan R. (2006-02-24).
7214: 5320: 5173: 4538:"Early Cretaceous mammals and the evolution of mammalian molar teeth" 1548: 1540: 1083:. Some authors also consider docodonts to lie crownward of the order 980: 815: 755: 293: 169: 7470: 4691:"Putative Triassic stem mammal Tikitherium copei is a Neogene shrew" 802:"Pseudotalonid" and "pseudoprotocone" are names which reference the 1621:, has been described from India, which was previously part of the 1577: 1562: 1025: 890: 618: 490: 458: 377: 373: 4594:"An alternative hypothesis on the origin of docodont molar teeth" 3540:. München, Germany: Verlag Dr. Friedrich Pfeil. pp. 187–214. 7528: 4950:
Panciroli, Elsa; Benson, Roger B. J.; Luo, Zhe-Xi (2019-05-04).
3531:"Molar diversity and functional adaptations in Mesozoic mammals" 614: 563: 7474: 7266: 6372: 5661: 5284: 5166: 5128: 4839:
Harper, Tony; Parras, Ana; Rougier, Guillermo W. (2019-12-01).
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on its ankle, similar to defensive structures observed in male
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bone. The dentary connects to the cranium via a joint with the
4234:"Transformation and diversification in early mammal evolution" 3443:"Analysis of Molar Structure and Phylogeny of Docodont Genera" 5124: 1661: 513:
bones of the skull roof are flat and broad, and there is no
1003:, such as wide, shortened digits in the hands for digging. 698:
Like other mammaliaforms, docodont teeth include peg-like
1053:. In other words, docodonts are outside of the mammalian 999:
shows many of the same physical traits as the modern day
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Averianov, Alexander O.; Voyta, Leonid L. (2024-02-28).
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10.2992/0145-9058(2007)39[27:AOMSAP]2.0.CO;2
4956:(Docodonta) from the Middle Jurassic of Skye, Scotland" 4561:"Reassessment of the late Triassic symmetrodont mammal 4653:
The Late Triassic World: Earth in a Time of Transition
3536:. In Martin, Thomas; von Koenigswald, Wighart (eds.). 1569:
stage of the Triassic. Later investigation found that
814:. Tribosphenic teeth show up in the oldest fossils of 566:
is found in a well-preserved specimen of the docodont
7388: 7378: 7368: 7358: 7348: 7338: 7328: 7318: 7308: 7298: 7288: 6956: 6783: 6730: 6686: 6671: 6661: 6651: 6641: 6630: 6620: 6610: 6600: 6590: 6580: 6570: 6560: 6550: 6540: 6530: 6520: 6510: 6500: 6490: 6480: 6470: 6460: 6450: 6440: 6430: 6420: 6410: 6400: 6166: 6151: 6141: 6131: 6121: 6111: 6033: 6003: 5988: 5978: 5968: 5958: 5948: 5938: 5928: 5918: 5908: 5898: 5888: 5833: 5803: 5758: 5743: 5733: 5723: 5713: 5703: 5693: 5578: 5563: 5521: 5506: 5496: 5486: 5476: 5466: 5456: 5446: 5398: 5361: 5346: 5336: 5326: 5316: 1400: 1343: 1316: 1150: 1127: 497:, a bone which is no longer present in mammals. The 7483: 7403: 7277: 7212: 7169: 7035: 6955: 6782: 6729: 6685: 6385: 6307: 6165: 6098: 6032: 6002: 5873: 5832: 5802: 5757: 5674: 5603: 5577: 5550: 5520: 5431: 5397: 5360: 5297: 5248: 5221: 4802: 4800: 4124:
Averianov, A. O.; Lopatin, A. V. (December 2006). "
3844:Lillegraven, Jason A.; Krusat, Georg (1991-10-01). 3220:
Kühne & Krusat 1972 sensu Sigoneau-Russell 2003
429:(lower jaw), formed primarily by the tooth-bearing 4293: 4291: 3075:Sigogneau-Russell 2003 sensu Panciroli et al. 2021 4989:20.500.11820/75714386-2baa-4512-b4c8-add5719f129b 4217:: CS1 maint: DOI inactive as of September 2024 ( 3925:Hoffmann, Simone; Krause, David W. (2019-07-19). 1587:, abruptly appearing in the fossil record in the 1057:, which only includes animals descended from the 4903: 4901: 4519:Memoirs of the Peabody Museum of Yale University 3884:"New species of mammaliaform and the cranium of 1505:in the 1920s argued that they were specialized " 969:from the Middle Jurassic of China, and possibly 4559:Sigogneu-Russell, Denise; Hahn, Renate (1995). 4370:Proceedings of the National Academy of Sciences 584:are unfused, attaching to the large and porous 3447:Bulletin of Carnegie Museum of Natural History 1595:; the youngest known members of the group are 5140: 770:docodont cusps with those of modern mammals. 684:, from the Early Cretaceous of Russia. Right 617:is flexed inwards. All limb bones except the 572:, though hair likely evolved much earlier in 8: 1583:Unambiguous docodonts are restricted to the 4917:Proceedings of the Zoological Institute RAS 979:had a flattened tail, similar to that of a 833:Pseudotribosphenic teeth are also found in 645:and several other early-branching mammals. 7471: 7274: 7263: 7041: 6964: 6393: 6382: 6369: 6104: 5881: 5686: 5682: 5671: 5658: 5556: 5439: 5309: 5305: 5294: 5281: 5227: 5218: 5163: 5147: 5133: 5125: 562:The oldest unambiguous fossil evidence of 118: 31: 4987: 4864: 4407: 4389: 4064: 4053:Zoological Journal of the Linnean Society 3892:Zoological Journal of the Linnean Society 3332:Maschenko, Lopatin & Voronkevich 2002 3059:Freeman 1979 sensu Sigogneau-Russell 2001 766:. However, there is little consensus for 678:Toothed jaw fragments and molar teeth of 725: 4184:Wang, Yuan-Qing; Li, Chuan-Kui (2016). 3412: 1591:. Very few docodonts survived into the 874: 4500:Trustees of the British Museum, London 4210: 4038: 4036: 3984: 3982: 3980: 3978: 3976: 1555:. Another proposed docodont relative, 501:expand at the back and overlook thick 4726: 4724: 4587: 4585: 4179: 4177: 4175: 4119: 4117: 3839: 3837: 3785: 3783: 3781: 3779: 3777: 3775: 3773: 3771: 3769: 3767: 3524: 3522: 3486: 3484: 3436: 7: 3877: 3875: 3873: 3871: 3711: 3709: 3707: 3705: 3703: 3701: 3699: 3697: 3695: 3639: 3637: 3635: 3633: 3631: 3629: 3627: 3625: 3623: 3621: 3565: 3563: 3561: 3559: 3557: 3555: 3553: 3551: 3549: 3547: 3441:Luo, Zhe-Xi; Martin, Thomas (2007). 3434: 3432: 3430: 3428: 3426: 3424: 3422: 3420: 3418: 3416: 818:, the mammalian subgroup containing 810:crushing complex which characterize 485:. The snout is long and has several 4097:"Were mammals originally venomous?" 3538:Mammalian Teeth - Form and Function 2806: 2782: 2758: 2734: 2684: 2583: 2576: 2526: 2519: 2495: 2468: 2444: 2394: 2387: 2363: 2339: 2315: 2291: 2267: 2260: 2118: 2094: 2070: 2020: 2013: 1989: 1965: 1879: 1855: 1831: 1807: 1783: 1776: 1752: 1725: 1701: 1679: 1667: 1393: 1366: 1336: 1309: 1232: 1208: 1184: 1160: 1145: 1120: 1108: 1011:A 2024 study on adult and juvenile 4960:Journal of Vertebrate Paleontology 4598:Journal of Vertebrate Paleontology 1573:was likely a misidentification of 737:from the Late Jurassic of Portugal 534:has a straight, sideways-oriented 25: 5104:The teeth of the Jurassic mammals 7455: 7454: 7249: 7244: 6355: 6349: 6343: 5644: 5639: 4929:10.31610/trudyzin/2010.314.2.121 4513:Simpson, George Gaylord (1929). 4494:Simpson, George Gaylord (1928). 4066:10.1111/j.1096-3642.2005.00187.x 925: 904: 877: 668: 657: 146: 4952:"The mandible and dentition of 989:, and specialised tree-dweller 4845:Journal of Mammalian Evolution 4695:Journal of Mammalian Evolution 4618:10.1080/02724634.1997.10010988 425:Docodonts have a long and low 407:transition is rather subdued. 27:Extinct order of mammaliaforms 1: 5082:10.1016/j.cretres.2024.105836 5038:10.1080/08912963.2023.2300635 4980:10.1080/02724634.2019.1621884 4572:Acta Palaeontologica Polonica 4104:Acta Palaeontologica Polonica 4043:Martin, Thomas (2005-10-01). 869:Docodont ecological variation 863:Paleoecology and paleobiology 7598:Taxa named by Miklós Kretzoi 4737:10.1371/journal.pone.0199983 4661:10.1007/978-3-319-68009-5_11 4592:Butler, P. M. (1997-06-19). 4515:"American Mesozoic Mammalia" 3242:Lopatin & Averianov 2005 706:, and numerous interlocking 517:forming the rear rim of the 376:. Chinese docodonts include 7603:Prehistoric tetrapod orders 7588:Bathonian first appearances 3378:Martin & Averianov 2004 3260:Krusatodon kirtlingtonensis 1067:dentary-squamosal jaw joint 7619: 4707:10.1007/s10914-024-09703-w 4343:10.1038/s41586-024-07733-1 3904:10.1093/zoolinnean/zlaa144 3505:10.1038/s41586-024-07258-7 688:(upper two rows) and left 467:ossicles of the middle ear 137:(blue) Scale bars = 10 mm 7450: 7273: 7262: 7242: 7044: 6967: 6396: 6381: 6368: 6341: 6107: 5884: 5689: 5685: 5670: 5657: 5637: 5559: 5442: 5312: 5308: 5293: 5280: 5230: 5217: 5162: 4857:10.1007/s10914-018-9437-x 4775:10.1134/S1028334X20090123 4536:Patterson, Bryan (1956). 4150:10.1134/s0031030106060098 3374:Tashkumyrodon desideratus 3286:(Sigogneau-Russell, 1998) 3172:Docofossor brachydactylus 3044:Castorocauda lutrasimilis 3035:Waldman & Savage 1972 3013:Waldman & Savage 1972 2828: 2811: 2804: 2787: 2780: 2763: 2756: 2739: 2732: 2706: 2689: 2682: 2665: 2648: 2631: 2605: 2588: 2581: 2574: 2548: 2531: 2524: 2517: 2500: 2493: 2473: 2466: 2449: 2442: 2416: 2399: 2392: 2385: 2368: 2361: 2344: 2337: 2320: 2313: 2296: 2289: 2272: 2265: 2140: 2123: 2116: 2099: 2092: 2075: 2068: 2042: 2025: 2018: 2011: 1994: 1987: 1970: 1963: 1901: 1884: 1877: 1860: 1853: 1836: 1829: 1812: 1805: 1788: 1781: 1774: 1757: 1750: 1730: 1723: 1706: 1699: 1684: 1677: 1615:. One disputed docodont, 1418: 1398: 1391: 1371: 1364: 1341: 1334: 1314: 1307: 1254: 1237: 1230: 1213: 1206: 1189: 1182: 1165: 1158: 1143: 1125: 1118: 469:and embracing a circular 262: 257: 143:Scientific classification 141: 126: 117: 34: 4954:Borealestes serendipitus 4045:"Postcranial anatomy of 3209:Prasad & Manhas 2007 3187:Pfretzschner et al. 2005 445:bone, also known as the 304:across the continent of 289:closely related to true 130:Borealestes serendipitus 4391:10.1073/pnas.1519387112 4193:Palaeontologia Polonica 4130:Paleontological Journal 3951:10.1126/science.aay2061 3812:10.1126/science.aau9345 3742:10.1126/science.1260879 3670:10.1126/science.1260880 3596:10.1126/science.1123026 3249:Khorotherium yakutensis 2998:Agilodocodon scansorius 1022:permanent (adult) teeth 1007:Metabolism and lifespan 681:Khorotherium yakutensis 576:. The structure of the 7583:Taxa described in 1946 5120:Docodonta from Palaeos 4755:Doklady Earth Sciences 4207:(inactive 2024-09-12). 4205:10.4202/pp.2016.67_249 4047:Haldanodon exspectatus 3850:Rocky Mountain Geology 3328:Sibirotherium rossicus 3319:Sigogneau-Russell 2003 3264:Sigogneau-Russell 2003 3227:Hutegotherium yaomingi 3216:Haldanodon exspectatus 1503:George Gaylord Simpson 746: 7550:Paleobiology Database 4093:Kielan-Jaworowska, Z. 3365:Averianov et al. 2010 3271:Microdocodon gracilis 3253:Averianov et al. 2018 3231:Averianov et al. 2010 3198:Averianov et al. 2024 3024:Panciroli et al. 2021 1089:phylogenetic analyses 758:mammals, like modern 733:Haldanodon expectatus 729: 127:Skeletal diagrams of 4232:Luo, Zhe-Xi (2007). 3401:Evolution of mammals 3282:Paritatodon kermacki 3071:Dobunnodon mussettae 1059:last common ancestor 955:Docodonts and other 828:convergent evolution 826:. This is a case of 754:teeth found in true 590:transverse processes 558:Postcranial skeleton 370:Tiaojishan Formation 318:convergent evolution 316:. This is a case of 5074:2024CrRes.15805836A 5062:Cretaceous Research 4972:2019JVPal..39E1884P 4767:2020DokES.494..667L 4610:1997JVPal..17..435B 4457:10.1038/nature23483 4449:2017Natur.548..326L 4382:2015PNAS..112E7101L 4376:(51): E7101–E7109. 4258:10.1038/nature06277 4250:2007Natur.450.1011L 4244:(7172): 1011–1019. 4142:2006PalJ...40..668A 4126:Itatodon tatarinovi 3943:2019Sci...365..222H 3804:2019Sci...365..276Z 3734:2015Sci...347..764M 3662:2015Sci...347..760L 3588:2006Sci...311.1123J 3582:(5764): 1123–1127. 3343:Kermack et al. 1987 3238:Itatodon tatarinovi 3097:Rougier et al. 2014 3055:Cyrtlatherium canei 1623:Southern Hemisphere 1585:Northern Hemisphere 542:(which link to the 515:postorbital process 461:(also known as the 453:(also known as the 394:middle ear ossicles 7593:Aptian extinctions 5030:Historical Biology 4011:10.1111/pala.12577 3385:Tegotherium gubini 3205:Gondtherium dattai 3108:Martin et al. 2024 812:tribosphenic teeth 747: 586:occipital condyles 477:Cranium and throat 439:postdentary trough 7565: 7564: 7477:Taxon identifiers 7468: 7467: 7446: 7445: 7442: 7441: 7258: 7257: 7240: 7239: 7236: 7235: 7208: 7207: 7204: 7203: 6364: 6363: 6339: 6338: 6335: 6334: 6331: 6330: 6327: 6326: 6007:Prozostrodontidae 5869: 5868: 5807:Probainognathidae 5653: 5652: 5635: 5634: 5631: 5630: 5627: 5626: 5623: 5622: 5427: 5426: 5365:Charassognathidae 5276: 5275: 5272: 5271: 5268: 5267: 5102:Butler PM. 1939. 4670:978-3-319-68009-5 4443:(7667): 326–329. 4337:(8026): 815–822. 3937:(6450): 222–223. 3798:(6450): 276–279. 3728:(6223): 764–768. 3656:(6223): 760–764. 3390: 3379: 3366: 3355: 3344: 3333: 3320: 3309: 3298: 3287: 3276: 3265: 3254: 3243: 3232: 3221: 3210: 3199: 3188: 3177: 3164: 3153: 3142: 3131: 3120: 3109: 3098: 3087: 3076: 3060: 3049: 3036: 3025: 3014: 3003: 2986: 2985: 2978: 2977: 2969: 2968: 2960: 2959: 2951: 2950: 2942: 2941: 2933: 2932: 2924: 2923: 2915: 2914: 2906: 2905: 2897: 2896: 2888: 2887: 2879: 2878: 2870: 2869: 2861: 2860: 2852: 2851: 2843: 2842: 2721: 2720: 2620: 2619: 2563: 2562: 2431: 2430: 2254: 2253: 2245: 2244: 2236: 2235: 2227: 2226: 2218: 2217: 2209: 2208: 2200: 2199: 2191: 2190: 2182: 2181: 2173: 2172: 2164: 2163: 2155: 2154: 2057: 2056: 1952: 1951: 1943: 1942: 1934: 1933: 1925: 1924: 1916: 1915: 1593:Cretaceous Period 1494: 1493: 1485: 1484: 1476: 1475: 1467: 1466: 1458: 1457: 1449: 1448: 1440: 1439: 1296: 1295: 1287: 1286: 1278: 1277: 1269: 1268: 1097:Bayesian analysis 1093:maximum parsimony 692:(lower four rows) 611:olecranon process 544:thyroid cartilage 314:mortar-and-pestle 269: 268: 253: 16:(Redirected from 7610: 7558: 7557: 7545: 7544: 7532: 7531: 7519: 7518: 7517: 7504: 7503: 7502: 7472: 7458: 7457: 7390: 7380: 7370: 7360: 7350: 7340: 7330: 7320: 7310: 7300: 7290: 7275: 7264: 7253: 7248: 7042: 6965: 6958: 6785: 6732: 6690:Kuehneotheriidae 6688: 6673: 6663: 6653: 6643: 6632: 6622: 6612: 6602: 6592: 6582: 6572: 6562: 6552: 6542: 6532: 6522: 6512: 6502: 6492: 6482: 6472: 6462: 6452: 6442: 6432: 6422: 6412: 6402: 6394: 6383: 6370: 6359: 6353: 6347: 6168: 6153: 6143: 6133: 6126:Botucaraitherium 6123: 6113: 6105: 6037:Tritheledontidae 6035: 6005: 5990: 5980: 5973:Santacruzgnathus 5970: 5960: 5953:Pseudotriconodon 5950: 5940: 5930: 5920: 5910: 5900: 5890: 5882: 5835: 5805: 5762:Chiniquodontidae 5760: 5745: 5735: 5725: 5715: 5705: 5695: 5687: 5683: 5672: 5659: 5648: 5643: 5580: 5565: 5557: 5523: 5508: 5498: 5488: 5478: 5468: 5458: 5448: 5440: 5400: 5363: 5348: 5338: 5328: 5318: 5310: 5306: 5295: 5282: 5228: 5219: 5212: 5211: 5164: 5149: 5142: 5135: 5126: 5107: 5100: 5094: 5093: 5056: 5050: 5049: 5025: 5019: 5016: 5010: 5009: 4991: 4947: 4941: 4940: 4914: 4905: 4896: 4893: 4887: 4886: 4868: 4836: 4830: 4829: 4813: 4807: 4804: 4795: 4794: 4746: 4740: 4728: 4719: 4718: 4686: 4680: 4679: 4678: 4677: 4644: 4638: 4637: 4589: 4580: 4579: 4569: 4556: 4550: 4549: 4533: 4527: 4526: 4510: 4504: 4503: 4491: 4485: 4484: 4428: 4422: 4421: 4411: 4393: 4361: 4355: 4354: 4322: 4316: 4313: 4307: 4304: 4298: 4295: 4286: 4285: 4229: 4223: 4222: 4216: 4208: 4190: 4181: 4170: 4169: 4121: 4112: 4111: 4101: 4085: 4079: 4078: 4068: 4040: 4031: 4030: 3986: 3971: 3970: 3927:"Tongues untied" 3922: 3916: 3915: 3898:(4): 1323–1362. 3879: 3866: 3865: 3841: 3832: 3831: 3787: 3762: 3761: 3713: 3690: 3689: 3641: 3616: 3615: 3567: 3542: 3541: 3535: 3526: 3517: 3516: 3488: 3479: 3478: 3438: 3388: 3377: 3364: 3353: 3342: 3331: 3318: 3307: 3296: 3285: 3275:Zhou et al. 2019 3274: 3263: 3252: 3241: 3230: 3219: 3208: 3197: 3186: 3183:Dsungarodon zuoi 3175: 3162: 3151: 3140: 3129: 3118: 3107: 3096: 3085: 3074: 3058: 3047: 3034: 3023: 3012: 3002:Meng et al. 2015 3001: 2807: 2783: 2759: 2735: 2685: 2584: 2577: 2527: 2520: 2496: 2469: 2445: 2395: 2388: 2364: 2340: 2316: 2292: 2268: 2261: 2119: 2095: 2071: 2021: 2014: 1990: 1966: 1880: 1856: 1832: 1808: 1784: 1777: 1753: 1726: 1702: 1680: 1668: 1662: 1609:Early Cretaceous 1402: 1394: 1367: 1345: 1337: 1318: 1310: 1233: 1209: 1185: 1161: 1152: 1146: 1129: 1121: 1109: 929: 908: 881: 672: 661: 578:vertebral column 528:(throat bones). 302:Early Cretaceous 248: 243: 151: 150: 122: 112: 49: 48:Bathonian–Aptian 44:Early Cretaceous 38:Temporal range: 32: 21: 7618: 7617: 7613: 7612: 7611: 7609: 7608: 7607: 7568: 7567: 7566: 7561: 7553: 7548: 7540: 7535: 7527: 7522: 7513: 7512: 7507: 7498: 7497: 7492: 7479: 7469: 7464: 7438: 7408: 7399: 7373:Panchetocynodon 7269: 7254: 7232: 7200: 7165: 7031: 6981:Hypsiprymnopsis 6951: 6778: 6725: 6681: 6415:Brachyzostrodon 6390: 6377: 6360: 6323: 6303: 6284:Xenocretosuchus 6170:Tritylodontidae 6161: 6094: 6053:Diarthrognathus 6028: 5998: 5878: 5876:Prozostrodontia 5865: 5828: 5823:Probainognathus 5798: 5753: 5679: 5677:Probainognathia 5666: 5664:Probainognathia 5649: 5619: 5605:Probainognathia 5599: 5573: 5546: 5516: 5491:Platycraniellus 5436: 5423: 5402:Procynosuchidae 5393: 5381:Charassognathus 5356: 5302: 5289: 5264: 5244: 5213: 5169: 5168: 5158: 5153: 5116: 5111: 5110: 5101: 5097: 5058: 5057: 5053: 5027: 5026: 5022: 5017: 5013: 4966:(3): e1621884. 4949: 4948: 4944: 4912: 4907: 4906: 4899: 4894: 4890: 4838: 4837: 4833: 4815: 4814: 4810: 4805: 4798: 4748: 4747: 4743: 4729: 4722: 4688: 4687: 4683: 4675: 4673: 4671: 4646: 4645: 4641: 4591: 4590: 4583: 4567: 4558: 4557: 4553: 4535: 4534: 4530: 4512: 4511: 4507: 4493: 4492: 4488: 4430: 4429: 4425: 4363: 4362: 4358: 4324: 4323: 4319: 4314: 4310: 4305: 4301: 4296: 4289: 4231: 4230: 4226: 4209: 4188: 4183: 4182: 4173: 4123: 4122: 4115: 4099: 4087: 4086: 4082: 4042: 4041: 4034: 3991:"Postcrania of 3988: 3987: 3974: 3924: 3923: 3919: 3881: 3880: 3869: 3843: 3842: 3835: 3789: 3788: 3765: 3715: 3714: 3693: 3643: 3642: 3619: 3569: 3568: 3545: 3533: 3528: 3527: 3520: 3490: 3489: 3482: 3440: 3439: 3414: 3409: 3397: 3304:P. inexpectatus 3194:Ergetiis ichchi 3176:Luo et al. 2015 3031:B. serendipitus 3020:B. cuillinensis 2992: 2987: 2979: 2970: 2961: 2952: 2943: 2934: 2925: 2916: 2907: 2898: 2889: 2880: 2871: 2862: 2853: 2844: 2722: 2621: 2564: 2432: 2255: 2246: 2237: 2228: 2219: 2210: 2201: 2192: 2183: 2174: 2165: 2156: 2058: 1953: 1944: 1935: 1926: 1917: 1649:meridiolestidan 1641:Late Cretaceous 1589:Middle Jurassic 1511:Bryan Patterson 1495: 1486: 1477: 1468: 1459: 1450: 1441: 1297: 1288: 1279: 1270: 1156: 1039: 1018:deciduous teeth 1009: 953: 952: 951: 948: 930: 921: 909: 900: 882: 871: 870: 865: 793: 791:Tooth occlusion 780: 744: 742: 730:Molar teeth of 724: 696: 695: 694: 693: 675: 674: 673: 664: 663: 662: 651: 560: 479: 423: 418: 416:Skeletal traits 413: 390:hyoid apparatus 298:Middle Jurassic 247: 241: 145: 135:B. cuillinensis 113: 111: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 47: 46: 40:Middle Jurassic 36: 28: 23: 22: 15: 12: 11: 5: 7616: 7614: 7606: 7605: 7600: 7595: 7590: 7585: 7580: 7570: 7569: 7563: 7562: 7560: 7559: 7546: 7533: 7520: 7505: 7489: 7487: 7481: 7480: 7475: 7466: 7465: 7463: 7462: 7451: 7448: 7447: 7444: 7443: 7440: 7439: 7437: 7436: 7431: 7426: 7424:Morganucodonta 7421: 7419:Dromatheriidae 7415: 7413: 7401: 7400: 7398: 7397: 7386: 7383:Redondagnathus 7376: 7366: 7356: 7346: 7336: 7326: 7316: 7306: 7296: 7285: 7283: 7280:Incertae sedis 7271: 7270: 7267: 7260: 7259: 7256: 7255: 7243: 7241: 7238: 7237: 7234: 7233: 7231: 7230: 7229: 7228: 7219: 7217: 7210: 7209: 7206: 7205: 7202: 7201: 7199: 7198: 7191: 7184: 7176: 7174: 7167: 7166: 7164: 7163: 7159:Gondwanatheria 7155: 7148: 7141: 7134: 7127: 7120: 7113: 7106: 7098: 7091: 7083: 7076: 7069: 7061: 7058:Arboroharamiya 7054: 7045: 7039: 7033: 7032: 7030: 7029: 7022: 7015: 7007: 6999: 6996:Kalaallitkigun 6992: 6985: 6977: 6968: 6962: 6953: 6952: 6950: 6949: 6942: 6935: 6928: 6921: 6914: 6906: 6899: 6892: 6885: 6877: 6870: 6863: 6855: 6848: 6841: 6834: 6827: 6820: 6813: 6806: 6799: 6791: 6789: 6780: 6779: 6777: 6776: 6769: 6762: 6754: 6746: 6738: 6736: 6727: 6726: 6724: 6723: 6720:Kuehneotherium 6716: 6709: 6702: 6694: 6692: 6683: 6682: 6680: 6679: 6669: 6659: 6649: 6639: 6628: 6618: 6608: 6605:Paikasigudodon 6598: 6588: 6578: 6568: 6558: 6548: 6538: 6528: 6518: 6508: 6505:Erythrotherium 6498: 6488: 6478: 6468: 6458: 6448: 6438: 6428: 6418: 6408: 6397: 6391: 6388:Mammaliaformes 6386: 6379: 6378: 6375:Mammaliaformes 6373: 6366: 6365: 6362: 6361: 6342: 6340: 6337: 6336: 6333: 6332: 6329: 6328: 6325: 6324: 6322: 6321: 6320: 6319: 6313: 6311: 6309:Mammaliaformes 6305: 6304: 6302: 6301: 6294: 6287: 6280: 6273: 6266: 6259: 6252: 6245: 6238: 6231: 6224: 6221:Kayentatherium 6217: 6210: 6203: 6196: 6189: 6186:Bienotheroides 6182: 6174: 6172: 6163: 6162: 6160: 6159: 6149: 6139: 6129: 6119: 6108: 6102: 6100:Mammaliamorpha 6096: 6095: 6093: 6092: 6085: 6077: 6070: 6063: 6056: 6049: 6041: 6039: 6030: 6029: 6027: 6026: 6019: 6011: 6009: 6000: 5999: 5997: 5996: 5986: 5983:Therioherpeton 5976: 5966: 5956: 5946: 5936: 5926: 5916: 5906: 5896: 5885: 5879: 5874: 5871: 5870: 5867: 5866: 5864: 5863: 5860:Trucidocynodon 5856: 5849: 5841: 5839: 5830: 5829: 5827: 5826: 5819: 5811: 5809: 5800: 5799: 5797: 5796: 5789: 5781: 5774: 5766: 5764: 5755: 5754: 5752: 5751: 5741: 5731: 5721: 5711: 5701: 5690: 5680: 5675: 5668: 5667: 5662: 5655: 5654: 5651: 5650: 5638: 5636: 5633: 5632: 5629: 5628: 5625: 5624: 5621: 5620: 5618: 5617: 5616: 5615: 5609: 5607: 5601: 5600: 5598: 5597: 5596: 5595: 5586: 5584: 5575: 5574: 5572: 5571: 5560: 5554: 5548: 5547: 5545: 5544: 5537: 5529: 5527: 5518: 5517: 5515: 5514: 5504: 5494: 5484: 5474: 5464: 5454: 5443: 5437: 5432: 5429: 5428: 5425: 5424: 5422: 5421: 5414: 5406: 5404: 5395: 5394: 5392: 5391: 5384: 5377: 5369: 5367: 5358: 5357: 5355: 5354: 5344: 5334: 5324: 5313: 5303: 5298: 5291: 5290: 5285: 5278: 5277: 5274: 5273: 5270: 5269: 5266: 5265: 5263: 5262: 5261: 5260: 5254: 5252: 5246: 5245: 5243: 5242: 5241: 5240: 5231: 5225: 5215: 5214: 5210: 5209: 5207:Eutheriodontia 5200: 5191: 5182: 5176: 5167: 5160: 5159: 5154: 5152: 5151: 5144: 5137: 5129: 5123: 5122: 5115: 5114:External links 5112: 5109: 5108: 5095: 5051: 5020: 5011: 4942: 4923:(2): 121–148. 4897: 4888: 4851:(4): 447–478. 4831: 4808: 4796: 4761:(1): 667–669. 4741: 4720: 4681: 4669: 4639: 4604:(2): 435–439. 4581: 4551: 4528: 4505: 4486: 4423: 4356: 4317: 4308: 4299: 4287: 4224: 4171: 4136:(6): 668–677. 4113: 4080: 4059:(2): 219–248. 4032: 3972: 3917: 3867: 3833: 3763: 3691: 3617: 3543: 3518: 3480: 3411: 3410: 3408: 3405: 3404: 3403: 3396: 3393: 3392: 3391: 3389:Tatarinov 1994 3380: 3369: 3368: 3367: 3356: 3334: 3323: 3322: 3321: 3310: 3288: 3277: 3266: 3255: 3244: 3233: 3222: 3211: 3200: 3189: 3178: 3167: 3166: 3165: 3154: 3143: 3132: 3121: 3110: 3099: 3077: 3066: 3050: 3048:Ji et al. 2006 3039: 3038: 3037: 3026: 3004: 2991: 2988: 2984: 2983: 2976: 2975: 2972: 2971: 2967: 2966: 2963: 2962: 2958: 2957: 2954: 2953: 2949: 2948: 2945: 2944: 2940: 2939: 2936: 2935: 2931: 2930: 2927: 2926: 2922: 2921: 2918: 2917: 2913: 2912: 2909: 2908: 2904: 2903: 2900: 2899: 2895: 2894: 2891: 2890: 2886: 2885: 2882: 2881: 2877: 2876: 2873: 2872: 2868: 2867: 2864: 2863: 2859: 2858: 2855: 2854: 2850: 2849: 2846: 2845: 2841: 2840: 2837: 2836: 2827: 2824: 2823: 2820: 2819: 2810: 2805: 2803: 2800: 2799: 2796: 2795: 2786: 2781: 2779: 2776: 2775: 2772: 2771: 2762: 2757: 2755: 2752: 2751: 2748: 2747: 2738: 2733: 2731: 2728: 2727: 2724: 2723: 2719: 2718: 2715: 2714: 2705: 2702: 2701: 2698: 2697: 2688: 2683: 2681: 2678: 2677: 2674: 2673: 2664: 2661: 2660: 2657: 2656: 2647: 2644: 2643: 2640: 2639: 2630: 2627: 2626: 2623: 2622: 2618: 2617: 2614: 2613: 2604: 2601: 2600: 2597: 2596: 2587: 2582: 2580: 2575: 2573: 2570: 2569: 2566: 2565: 2561: 2560: 2557: 2556: 2547: 2544: 2543: 2540: 2539: 2530: 2525: 2523: 2518: 2516: 2513: 2512: 2509: 2508: 2499: 2494: 2492: 2486: 2485: 2482: 2481: 2472: 2467: 2465: 2462: 2461: 2458: 2457: 2448: 2443: 2441: 2438: 2437: 2434: 2433: 2429: 2428: 2425: 2424: 2415: 2412: 2411: 2408: 2407: 2398: 2393: 2391: 2386: 2384: 2381: 2380: 2377: 2376: 2372:Kuehneotherium 2367: 2362: 2360: 2357: 2356: 2353: 2352: 2343: 2338: 2336: 2333: 2332: 2329: 2328: 2319: 2314: 2312: 2309: 2308: 2305: 2304: 2295: 2290: 2288: 2285: 2284: 2281: 2280: 2271: 2266: 2264: 2259: 2256: 2252: 2251: 2248: 2247: 2243: 2242: 2239: 2238: 2234: 2233: 2230: 2229: 2225: 2224: 2221: 2220: 2216: 2215: 2212: 2211: 2207: 2206: 2203: 2202: 2198: 2197: 2194: 2193: 2189: 2188: 2185: 2184: 2180: 2179: 2176: 2175: 2171: 2170: 2167: 2166: 2162: 2161: 2158: 2157: 2153: 2152: 2149: 2148: 2139: 2136: 2135: 2132: 2131: 2122: 2117: 2115: 2112: 2111: 2108: 2107: 2098: 2093: 2091: 2088: 2087: 2084: 2083: 2074: 2069: 2067: 2064: 2063: 2060: 2059: 2055: 2054: 2051: 2050: 2041: 2038: 2037: 2034: 2033: 2024: 2019: 2017: 2012: 2010: 2007: 2006: 2003: 2002: 1993: 1988: 1986: 1983: 1982: 1979: 1978: 1969: 1964: 1962: 1959: 1958: 1955: 1954: 1950: 1949: 1946: 1945: 1941: 1940: 1937: 1936: 1932: 1931: 1928: 1927: 1923: 1922: 1919: 1918: 1914: 1913: 1910: 1909: 1900: 1897: 1896: 1893: 1892: 1883: 1878: 1876: 1873: 1872: 1869: 1868: 1859: 1854: 1852: 1849: 1848: 1845: 1844: 1835: 1830: 1828: 1825: 1824: 1821: 1820: 1811: 1806: 1804: 1801: 1800: 1797: 1796: 1787: 1782: 1780: 1775: 1773: 1770: 1769: 1766: 1765: 1756: 1751: 1749: 1743: 1742: 1739: 1738: 1729: 1724: 1722: 1719: 1718: 1715: 1714: 1705: 1700: 1698: 1695: 1694: 1691: 1690: 1687:Morganucodonta 1683: 1678: 1676: 1673:Mammaliaformes 1666: 1660: 1492: 1491: 1488: 1487: 1483: 1482: 1479: 1478: 1474: 1473: 1470: 1469: 1465: 1464: 1461: 1460: 1456: 1455: 1452: 1451: 1447: 1446: 1443: 1442: 1438: 1437: 1434: 1433: 1417: 1414: 1413: 1410: 1409: 1397: 1392: 1390: 1387: 1386: 1383: 1382: 1370: 1365: 1363: 1355: 1354: 1351: 1350: 1340: 1335: 1333: 1330: 1329: 1326: 1325: 1313: 1308: 1306: 1303: 1302: 1299: 1298: 1294: 1293: 1290: 1289: 1285: 1284: 1281: 1280: 1276: 1275: 1272: 1271: 1267: 1266: 1263: 1262: 1253: 1250: 1249: 1246: 1245: 1236: 1231: 1229: 1226: 1225: 1222: 1221: 1212: 1207: 1205: 1202: 1201: 1198: 1197: 1188: 1183: 1181: 1178: 1177: 1174: 1173: 1164: 1159: 1157: 1149: 1144: 1142: 1139: 1138: 1135: 1134: 1131:Morganucodonta 1124: 1119: 1117: 1114:Mammaliaformes 1107: 1075:Morganucodonta 1038: 1037:Classification 1035: 1008: 1005: 950: 949: 931: 924: 922: 920:-like burrower 910: 903: 901: 883: 876: 873: 872: 868: 867: 866: 864: 861: 792: 789: 779: 776: 723: 720: 677: 676: 667: 666: 665: 656: 655: 654: 653: 652: 650: 647: 637:has a pointed 559: 556: 521:(eye socket). 483:morganucodonts 478: 475: 422: 419: 417: 414: 412: 409: 267: 266: 260: 259: 255: 254: 239: 235: 234: 232:Mammaliaformes 229: 222: 221: 216: 209: 208: 203: 196: 195: 190: 183: 182: 177: 173: 172: 167: 163: 162: 157: 153: 152: 139: 138: 124: 123: 115: 114: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 50: 37: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 7615: 7604: 7601: 7599: 7596: 7594: 7591: 7589: 7586: 7584: 7581: 7579: 7576: 7575: 7573: 7556: 7551: 7547: 7543: 7538: 7534: 7530: 7525: 7521: 7516: 7510: 7506: 7501: 7495: 7491: 7490: 7488: 7486: 7482: 7478: 7473: 7461: 7453: 7452: 7449: 7435: 7432: 7430: 7429:Symmetrodonta 7427: 7425: 7422: 7420: 7417: 7416: 7414: 7411: 7406: 7402: 7395: 7394: 7387: 7385: 7384: 7377: 7375: 7374: 7367: 7365: 7364: 7357: 7355: 7354: 7347: 7345: 7344: 7337: 7335: 7334: 7327: 7325: 7324: 7317: 7315: 7314: 7307: 7305: 7304: 7297: 7295: 7294: 7287: 7286: 7284: 7282: 7281: 7276: 7272: 7265: 7261: 7252: 7247: 7227: 7223: 7222: 7221: 7220: 7218: 7216: 7211: 7197: 7196: 7192: 7190: 7189: 7185: 7183: 7182: 7178: 7177: 7175: 7172: 7171:Hahnodontidae 7168: 7161: 7160: 7156: 7154: 7153: 7149: 7147: 7146: 7142: 7140: 7139: 7135: 7133: 7132: 7131:Sineleutherus 7128: 7126: 7125: 7121: 7119: 7118: 7114: 7112: 7111: 7107: 7104: 7103: 7099: 7097: 7096: 7092: 7089: 7088: 7084: 7082: 7081: 7077: 7075: 7074: 7070: 7067: 7066: 7062: 7060: 7059: 7055: 7052: 7051: 7047: 7046: 7043: 7040: 7038: 7034: 7028: 7027: 7023: 7021: 7020: 7016: 7013: 7012: 7008: 7005: 7004: 7000: 6998: 6997: 6993: 6991: 6990: 6986: 6983: 6982: 6978: 6975: 6974: 6970: 6969: 6966: 6963: 6961: 6954: 6948: 6947: 6943: 6941: 6940: 6939:Tashkumyrodon 6936: 6934: 6933: 6929: 6927: 6926: 6925:Sibirotherium 6922: 6920: 6919: 6918:Peraiocynodon 6915: 6912: 6911: 6907: 6905: 6904: 6900: 6898: 6897: 6893: 6891: 6890: 6886: 6883: 6882: 6878: 6876: 6875: 6874:Hutegotherium 6871: 6869: 6868: 6864: 6861: 6860: 6856: 6854: 6853: 6849: 6847: 6846: 6842: 6840: 6839: 6835: 6833: 6832: 6828: 6826: 6825: 6821: 6819: 6818: 6817:Cyrtlatherium 6814: 6812: 6811: 6807: 6805: 6804: 6800: 6798: 6797: 6793: 6792: 6790: 6788: 6781: 6775: 6774: 6770: 6768: 6767: 6763: 6760: 6759: 6755: 6752: 6751: 6747: 6745: 6744: 6740: 6739: 6737: 6735: 6734:Shuotheriidae 6728: 6722: 6721: 6717: 6715: 6714: 6710: 6708: 6707: 6703: 6701: 6700: 6696: 6695: 6693: 6691: 6684: 6678: 6677: 6670: 6668: 6667: 6660: 6658: 6657: 6650: 6648: 6647: 6640: 6637: 6636: 6629: 6627: 6626: 6619: 6617: 6616: 6609: 6607: 6606: 6599: 6597: 6596: 6589: 6587: 6586: 6579: 6577: 6576: 6575:Megazostrodon 6569: 6567: 6566: 6565:Indozostrodon 6559: 6557: 6556: 6549: 6547: 6546: 6539: 6537: 6536: 6535:Hallautherium 6529: 6527: 6526: 6519: 6517: 6516: 6509: 6507: 6506: 6499: 6497: 6496: 6489: 6487: 6486: 6479: 6477: 6476: 6469: 6467: 6466: 6459: 6457: 6456: 6449: 6447: 6446: 6445:Cifellilestes 6439: 6437: 6436: 6429: 6427: 6426: 6419: 6417: 6416: 6409: 6407: 6406: 6399: 6398: 6395: 6392: 6389: 6384: 6380: 6376: 6371: 6367: 6358: 6354: 6352: 6346: 6317: 6316: 6315: 6314: 6312: 6310: 6306: 6300: 6299: 6295: 6293: 6292: 6288: 6286: 6285: 6281: 6279: 6278: 6274: 6272: 6271: 6270:Stereognathus 6267: 6265: 6264: 6260: 6258: 6257: 6253: 6251: 6250: 6246: 6244: 6243: 6239: 6237: 6236: 6232: 6230: 6229: 6225: 6223: 6222: 6218: 6216: 6215: 6211: 6209: 6208: 6204: 6202: 6201: 6197: 6195: 6194: 6190: 6188: 6187: 6183: 6181: 6180: 6176: 6175: 6173: 6171: 6164: 6158: 6157: 6150: 6148: 6147: 6140: 6138: 6137: 6130: 6128: 6127: 6120: 6118: 6117: 6116:Adelobasileus 6110: 6109: 6106: 6103: 6101: 6097: 6091: 6090: 6086: 6083: 6082: 6078: 6076: 6075: 6071: 6069: 6068: 6064: 6062: 6061: 6057: 6055: 6054: 6050: 6048: 6047: 6043: 6042: 6040: 6038: 6031: 6025: 6024: 6023:Pseudotherium 6020: 6018: 6017: 6013: 6012: 6010: 6008: 6001: 5995: 5994: 5987: 5985: 5984: 5977: 5975: 5974: 5967: 5965: 5964: 5957: 5955: 5954: 5947: 5945: 5944: 5937: 5935: 5934: 5927: 5925: 5924: 5917: 5915: 5914: 5907: 5905: 5904: 5897: 5895: 5894: 5893:Alemoatherium 5887: 5886: 5883: 5880: 5877: 5872: 5862: 5861: 5857: 5855: 5854: 5850: 5848: 5847: 5843: 5842: 5840: 5838: 5831: 5825: 5824: 5820: 5818: 5817: 5813: 5812: 5810: 5808: 5801: 5795: 5794: 5790: 5787: 5786: 5782: 5780: 5779: 5775: 5773: 5772: 5768: 5767: 5765: 5763: 5756: 5750: 5749: 5742: 5740: 5739: 5732: 5730: 5729: 5722: 5720: 5719: 5712: 5710: 5709: 5708:Candelariodon 5702: 5700: 5699: 5692: 5691: 5688: 5684: 5681: 5678: 5673: 5669: 5665: 5660: 5656: 5647: 5642: 5613: 5612: 5611: 5610: 5608: 5606: 5602: 5594: 5590: 5589: 5588: 5587: 5585: 5583: 5576: 5570: 5569: 5562: 5561: 5558: 5555: 5553: 5549: 5543: 5542: 5541:Progalesaurus 5538: 5536: 5535: 5531: 5530: 5528: 5526: 5519: 5513: 5512: 5505: 5503: 5502: 5495: 5493: 5492: 5485: 5483: 5482: 5475: 5473: 5472: 5471:Nanictosaurus 5465: 5463: 5462: 5455: 5453: 5452: 5445: 5444: 5441: 5438: 5435: 5434:Epicynodontia 5430: 5420: 5419: 5415: 5413: 5412: 5411:Procynosuchus 5408: 5407: 5405: 5403: 5396: 5390: 5389: 5385: 5383: 5382: 5378: 5376: 5375: 5371: 5370: 5368: 5366: 5359: 5353: 5352: 5345: 5343: 5342: 5335: 5333: 5332: 5325: 5323: 5322: 5315: 5314: 5311: 5307: 5304: 5301: 5296: 5292: 5288: 5283: 5279: 5258: 5257: 5256: 5255: 5253: 5251: 5247: 5239: 5235: 5234: 5233: 5232: 5229: 5226: 5224: 5220: 5216: 5208: 5204: 5201: 5199: 5195: 5192: 5190: 5186: 5183: 5181: 5177: 5175: 5171: 5170: 5165: 5161: 5157: 5150: 5145: 5143: 5138: 5136: 5131: 5130: 5127: 5121: 5118: 5117: 5113: 5105: 5099: 5096: 5091: 5087: 5083: 5079: 5075: 5071: 5067: 5063: 5055: 5052: 5047: 5043: 5039: 5035: 5031: 5024: 5021: 5015: 5012: 5007: 5003: 4999: 4995: 4990: 4985: 4981: 4977: 4973: 4969: 4965: 4961: 4957: 4955: 4946: 4943: 4938: 4934: 4930: 4926: 4922: 4918: 4911: 4904: 4902: 4898: 4892: 4889: 4884: 4880: 4876: 4872: 4867: 4862: 4858: 4854: 4850: 4846: 4842: 4835: 4832: 4828:(3): 395–414. 4827: 4823: 4822:Geodiversitas 4819: 4812: 4809: 4803: 4801: 4797: 4792: 4788: 4784: 4780: 4776: 4772: 4768: 4764: 4760: 4756: 4752: 4745: 4742: 4738: 4734: 4727: 4725: 4721: 4716: 4712: 4708: 4704: 4700: 4696: 4692: 4685: 4682: 4672: 4666: 4662: 4658: 4654: 4650: 4643: 4640: 4635: 4631: 4627: 4623: 4619: 4615: 4611: 4607: 4603: 4599: 4595: 4588: 4586: 4582: 4578:(3): 245–260. 4577: 4573: 4566: 4564: 4555: 4552: 4547: 4543: 4539: 4532: 4529: 4524: 4520: 4516: 4509: 4506: 4501: 4497: 4490: 4487: 4482: 4478: 4474: 4470: 4466: 4462: 4458: 4454: 4450: 4446: 4442: 4438: 4434: 4427: 4424: 4419: 4415: 4410: 4405: 4401: 4397: 4392: 4387: 4383: 4379: 4375: 4371: 4367: 4360: 4357: 4352: 4348: 4344: 4340: 4336: 4332: 4328: 4321: 4318: 4312: 4309: 4303: 4300: 4294: 4292: 4288: 4283: 4279: 4275: 4271: 4267: 4263: 4259: 4255: 4251: 4247: 4243: 4239: 4235: 4228: 4225: 4220: 4214: 4206: 4202: 4198: 4194: 4187: 4180: 4178: 4176: 4172: 4167: 4163: 4159: 4155: 4151: 4147: 4143: 4139: 4135: 4131: 4127: 4120: 4118: 4114: 4109: 4105: 4098: 4094: 4090: 4084: 4081: 4076: 4072: 4067: 4062: 4058: 4054: 4050: 4048: 4039: 4037: 4033: 4028: 4024: 4020: 4016: 4012: 4008: 4004: 4000: 3999:Palaeontology 3996: 3994: 3985: 3983: 3981: 3979: 3977: 3973: 3968: 3964: 3960: 3956: 3952: 3948: 3944: 3940: 3936: 3932: 3928: 3921: 3918: 3913: 3909: 3905: 3901: 3897: 3893: 3889: 3887: 3878: 3876: 3874: 3872: 3868: 3863: 3859: 3856:(2): 39–138. 3855: 3851: 3847: 3840: 3838: 3834: 3829: 3825: 3821: 3817: 3813: 3809: 3805: 3801: 3797: 3793: 3786: 3784: 3782: 3780: 3778: 3776: 3774: 3772: 3770: 3768: 3764: 3759: 3755: 3751: 3747: 3743: 3739: 3735: 3731: 3727: 3723: 3719: 3712: 3710: 3708: 3706: 3704: 3702: 3700: 3698: 3696: 3692: 3687: 3683: 3679: 3675: 3671: 3667: 3663: 3659: 3655: 3651: 3647: 3640: 3638: 3636: 3634: 3632: 3630: 3628: 3626: 3624: 3622: 3618: 3613: 3609: 3605: 3601: 3597: 3593: 3589: 3585: 3581: 3577: 3573: 3566: 3564: 3562: 3560: 3558: 3556: 3554: 3552: 3550: 3548: 3544: 3539: 3532: 3525: 3523: 3519: 3514: 3510: 3506: 3502: 3498: 3494: 3487: 3485: 3481: 3476: 3472: 3468: 3464: 3460: 3456: 3453:(39): 27–47. 3452: 3448: 3444: 3437: 3435: 3433: 3431: 3429: 3427: 3425: 3423: 3421: 3419: 3417: 3413: 3406: 3402: 3399: 3398: 3394: 3387: 3386: 3381: 3376: 3375: 3370: 3363: 3362: 3357: 3352: 3351: 3346: 3345: 3341: 3340: 3335: 3330: 3329: 3324: 3317: 3316: 3311: 3306: 3305: 3300: 3299: 3295: 3294: 3293:Peraiocynodon 3289: 3284: 3283: 3278: 3273: 3272: 3267: 3262: 3261: 3256: 3251: 3250: 3245: 3240: 3239: 3234: 3229: 3228: 3223: 3218: 3217: 3212: 3207: 3206: 3201: 3196: 3195: 3190: 3185: 3184: 3179: 3174: 3173: 3168: 3161: 3160: 3155: 3150: 3149: 3144: 3139: 3138: 3133: 3128: 3127: 3122: 3117: 3116: 3111: 3106: 3105: 3104:D. hercynicus 3100: 3095: 3094: 3089: 3088: 3084: 3083: 3078: 3073: 3072: 3067: 3064: 3057: 3056: 3051: 3046: 3045: 3040: 3033: 3032: 3027: 3022: 3021: 3016: 3015: 3011: 3010: 3005: 3000: 2999: 2994: 2993: 2989: 2982: 2974: 2973: 2965: 2964: 2956: 2955: 2947: 2946: 2938: 2937: 2929: 2928: 2920: 2919: 2911: 2910: 2902: 2901: 2893: 2892: 2884: 2883: 2875: 2874: 2866: 2865: 2857: 2856: 2848: 2847: 2839: 2838: 2835: 2834: 2833: 2826: 2825: 2822: 2821: 2818: 2817: 2816: 2809: 2808: 2802: 2801: 2798: 2797: 2794: 2793: 2792: 2785: 2784: 2778: 2777: 2774: 2773: 2770: 2769: 2768: 2761: 2760: 2754: 2753: 2750: 2749: 2746: 2745: 2744: 2737: 2736: 2730: 2729: 2726: 2725: 2717: 2716: 2713: 2712: 2711: 2704: 2703: 2700: 2699: 2696: 2695: 2694: 2687: 2686: 2680: 2679: 2676: 2675: 2672: 2671: 2670: 2669:Tashkumyrodon 2663: 2662: 2659: 2658: 2655: 2654: 2653: 2646: 2645: 2642: 2641: 2638: 2637: 2636: 2629: 2628: 2625: 2624: 2616: 2615: 2612: 2611: 2610: 2603: 2602: 2599: 2598: 2595: 2594: 2593: 2586: 2585: 2579: 2578: 2572: 2571: 2568: 2567: 2559: 2558: 2555: 2554: 2553: 2546: 2545: 2542: 2541: 2538: 2537: 2536: 2535:Sibirotherium 2529: 2528: 2522: 2521: 2515: 2514: 2511: 2510: 2507: 2506: 2505: 2504:Hutegotherium 2498: 2497: 2491: 2488: 2487: 2484: 2483: 2480: 2479: 2478: 2471: 2470: 2464: 2463: 2460: 2459: 2456: 2455: 2454: 2447: 2446: 2440: 2439: 2436: 2435: 2427: 2426: 2423: 2422: 2421: 2414: 2413: 2410: 2409: 2406: 2405: 2404: 2397: 2396: 2390: 2389: 2383: 2382: 2379: 2378: 2375: 2374: 2373: 2366: 2365: 2359: 2358: 2355: 2354: 2351: 2350: 2349: 2348:Megazostrodon 2342: 2341: 2335: 2334: 2331: 2330: 2327: 2326: 2325: 2318: 2317: 2311: 2310: 2307: 2306: 2303: 2302: 2301: 2294: 2293: 2287: 2286: 2283: 2282: 2279: 2278: 2277: 2270: 2269: 2263: 2262: 2257: 2250: 2249: 2241: 2240: 2232: 2231: 2223: 2222: 2214: 2213: 2205: 2204: 2196: 2195: 2187: 2186: 2178: 2177: 2169: 2168: 2160: 2159: 2151: 2150: 2147: 2146: 2145: 2138: 2137: 2134: 2133: 2130: 2129: 2128: 2121: 2120: 2114: 2113: 2110: 2109: 2106: 2105: 2104: 2103:Hutegotherium 2097: 2096: 2090: 2089: 2086: 2085: 2082: 2081: 2080: 2079:Sibirotherium 2073: 2072: 2066: 2065: 2062: 2061: 2053: 2052: 2049: 2048: 2047: 2040: 2039: 2036: 2035: 2032: 2031: 2030: 2023: 2022: 2016: 2015: 2009: 2008: 2005: 2004: 2001: 2000: 1999: 1992: 1991: 1985: 1984: 1981: 1980: 1977: 1976: 1975: 1968: 1967: 1961: 1960: 1957: 1956: 1948: 1947: 1939: 1938: 1930: 1929: 1921: 1920: 1912: 1911: 1908: 1907: 1906: 1899: 1898: 1895: 1894: 1891: 1890: 1889: 1882: 1881: 1875: 1874: 1871: 1870: 1867: 1866: 1865: 1858: 1857: 1851: 1850: 1847: 1846: 1843: 1842: 1841: 1834: 1833: 1827: 1826: 1823: 1822: 1819: 1818: 1817: 1810: 1809: 1803: 1802: 1799: 1798: 1795: 1794: 1793: 1792:Tashkumyrodon 1786: 1785: 1779: 1778: 1772: 1771: 1768: 1767: 1764: 1763: 1762: 1755: 1754: 1748: 1745: 1744: 1741: 1740: 1737: 1736: 1735: 1728: 1727: 1721: 1720: 1717: 1716: 1713: 1712: 1711: 1704: 1703: 1697: 1696: 1693: 1692: 1689: 1688: 1682: 1681: 1675: 1674: 1670: 1669: 1664: 1663: 1659: 1656: 1654: 1650: 1646: 1642: 1638: 1637: 1632: 1628: 1625:continent of 1624: 1620: 1619: 1614: 1610: 1606: 1605: 1600: 1599: 1598:Sibirotherium 1594: 1590: 1586: 1581: 1579: 1576: 1572: 1568: 1564: 1560: 1559: 1554: 1550: 1546: 1542: 1538: 1537: 1532: 1531: 1526: 1525:symmetrodonts 1522: 1518: 1517: 1512: 1508: 1504: 1500: 1490: 1489: 1481: 1480: 1472: 1471: 1463: 1462: 1454: 1453: 1445: 1444: 1436: 1435: 1432: 1430: 1426: 1422: 1416: 1415: 1412: 1411: 1408: 1407: 1406: 1396: 1395: 1389: 1388: 1385: 1384: 1381: 1379: 1375: 1369: 1368: 1362: 1361: 1357: 1356: 1353: 1352: 1349: 1348: 1339: 1338: 1332: 1331: 1328: 1327: 1324: 1323: 1322: 1312: 1311: 1305: 1304: 1301: 1300: 1292: 1291: 1283: 1282: 1274: 1273: 1265: 1264: 1261: 1260: 1259: 1252: 1251: 1248: 1247: 1244: 1243: 1242: 1235: 1234: 1228: 1227: 1224: 1223: 1220: 1219: 1218: 1211: 1210: 1204: 1203: 1200: 1199: 1196: 1195: 1194: 1187: 1186: 1180: 1179: 1176: 1175: 1172: 1171: 1170: 1163: 1162: 1155: 1148: 1147: 1141: 1140: 1137: 1136: 1133: 1132: 1123: 1122: 1116: 1115: 1111: 1110: 1106: 1104: 1100: 1098: 1094: 1090: 1086: 1082: 1081: 1076: 1072: 1071:mammaliaforms 1068: 1064: 1060: 1056: 1052: 1048: 1044: 1036: 1034: 1031: 1027: 1023: 1019: 1014: 1006: 1004: 1002: 998: 994: 993: 988: 987: 982: 978: 974: 973: 968: 967: 962: 961:insectivorous 958: 947: 944: 940: 936: 935: 928: 923: 919: 915: 914: 907: 902: 899: 896: 892: 888: 887: 880: 875: 862: 860: 858: 857: 852: 851: 846: 845: 840: 836: 831: 829: 825: 821: 817: 813: 809: 805: 800: 798: 790: 788: 784: 777: 775: 771: 769: 765: 761: 757: 753: 740: 736: 734: 728: 721: 719: 717: 713: 709: 705: 701: 691: 687: 683: 682: 671: 660: 648: 646: 644: 640: 636: 632: 628: 624: 620: 616: 612: 607: 605: 604: 599: 595: 591: 587: 583: 579: 575: 571: 570: 565: 557: 555: 553: 549: 545: 541: 537: 533: 532: 527: 522: 520: 516: 512: 508: 504: 500: 496: 492: 488: 487:plesiomorphic 484: 476: 474: 472: 468: 464: 460: 456: 452: 448: 444: 440: 436: 432: 428: 420: 415: 410: 408: 406: 402: 399: 395: 391: 387: 383: 379: 375: 371: 367: 363: 360: 359:Late Jurassic 356: 355: 349: 347: 343: 339: 338: 333: 332: 327: 326:symmetrodonts 323: 319: 315: 311: 307: 303: 299: 295: 292: 288: 284: 283:mammaliaforms 281: 277: 273: 265: 261: 256: 251: 246: 240: 237: 236: 233: 230: 227: 224: 223: 220: 217: 214: 211: 210: 207: 204: 201: 198: 197: 194: 191: 188: 185: 184: 181: 178: 175: 174: 171: 168: 165: 164: 161: 158: 155: 154: 149: 144: 140: 136: 132: 131: 125: 121: 116: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 45: 41: 33: 30: 19: 7484: 7434:Triconodonta 7410:Polyphyletic 7405:Paraphyletic 7391: 7381: 7371: 7361: 7351: 7341: 7331: 7321: 7311: 7301: 7293:Cistecynodon 7291: 7278: 7193: 7186: 7179: 7157: 7150: 7143: 7136: 7129: 7122: 7115: 7108: 7100: 7095:Maiopatagium 7093: 7085: 7080:Cryoharamiya 7078: 7071: 7063: 7056: 7048: 7037:Euharamiyida 7024: 7017: 7009: 7001: 6994: 6987: 6979: 6971: 6944: 6937: 6930: 6923: 6916: 6908: 6903:Microdocodon 6901: 6894: 6889:Khorotherium 6887: 6879: 6872: 6865: 6857: 6850: 6843: 6836: 6829: 6822: 6815: 6810:Castorocauda 6808: 6801: 6796:Agilodocodon 6794: 6786: 6771: 6766:Pseudotribos 6764: 6756: 6748: 6741: 6718: 6711: 6704: 6697: 6674: 6664: 6654: 6644: 6633: 6623: 6613: 6603: 6593: 6585:Morganucodon 6583: 6573: 6563: 6553: 6543: 6533: 6523: 6513: 6503: 6493: 6483: 6475:Dinnetherium 6473: 6465:Dianoconodon 6463: 6453: 6443: 6433: 6425:Bridetherium 6423: 6413: 6403: 6348: 6296: 6291:Yuanotherium 6289: 6282: 6275: 6268: 6261: 6254: 6247: 6240: 6233: 6226: 6219: 6212: 6207:Dinnebitodon 6205: 6198: 6191: 6184: 6179:Bienotherium 6177: 6154: 6144: 6134: 6124: 6114: 6087: 6079: 6074:Pachygenelus 6072: 6067:Irajatherium 6065: 6060:Elliotherium 6058: 6051: 6044: 6021: 6016:Prozostrodon 6014: 5991: 5981: 5971: 5961: 5951: 5941: 5933:Microconodon 5931: 5921: 5913:Dromatherium 5911: 5901: 5891: 5858: 5851: 5844: 5837:Ecteniniidae 5821: 5814: 5791: 5783: 5776: 5769: 5746: 5738:Protheriodon 5736: 5726: 5716: 5706: 5698:Agudotherium 5696: 5568:Kataigidodon 5566: 5552:Eucynodontia 5539: 5532: 5525:Galesauridae 5509: 5499: 5489: 5479: 5469: 5459: 5449: 5416: 5409: 5386: 5379: 5372: 5351:Uralocynodon 5349: 5339: 5329: 5319: 5202: 5193: 5184: 5098: 5065: 5061: 5054: 5029: 5023: 5014: 4963: 4959: 4953: 4945: 4920: 4916: 4891: 4848: 4844: 4834: 4825: 4821: 4811: 4758: 4754: 4744: 4698: 4694: 4684: 4674:, retrieved 4652: 4642: 4601: 4597: 4575: 4571: 4562: 4554: 4545: 4541: 4531: 4522: 4518: 4508: 4499: 4489: 4440: 4436: 4426: 4373: 4369: 4359: 4334: 4330: 4320: 4311: 4302: 4241: 4237: 4227: 4213:cite journal 4196: 4192: 4133: 4129: 4125: 4107: 4103: 4091:; Luo, Z-X; 4083: 4056: 4052: 4046: 4002: 3998: 3992: 3934: 3930: 3920: 3895: 3891: 3885: 3853: 3849: 3795: 3791: 3725: 3721: 3653: 3649: 3579: 3575: 3537: 3496: 3450: 3446: 3383: 3372: 3361:S. sibiricus 3359: 3354:(Kühne 1969) 3350:S. splendens 3348: 3337: 3326: 3313: 3308:Simpson 1928 3302: 3297:Simpson 1928 3291: 3280: 3269: 3258: 3247: 3236: 3225: 3214: 3203: 3192: 3181: 3170: 3163:Simpson 1929 3157: 3152:(Marsh 1887) 3146: 3141:(Marsh 1887) 3135: 3124: 3119:(Marsh 1880) 3113: 3102: 3091: 3080: 3069: 3053: 3042: 3029: 3018: 3007: 2996: 2980: 2830: 2829: 2813: 2812: 2789: 2788: 2765: 2764: 2741: 2740: 2708: 2707: 2693:Castorocauda 2691: 2690: 2667: 2666: 2650: 2649: 2633: 2632: 2607: 2606: 2592:Agilodocodon 2590: 2589: 2550: 2549: 2533: 2532: 2502: 2501: 2489: 2475: 2474: 2451: 2450: 2418: 2417: 2401: 2400: 2370: 2369: 2346: 2345: 2324:Dinnetherium 2322: 2321: 2300:Morganucodon 2298: 2297: 2274: 2273: 2142: 2141: 2127:Microdocodon 2125: 2124: 2101: 2100: 2077: 2076: 2044: 2043: 2029:Agilodocodon 2027: 2026: 1996: 1995: 1972: 1971: 1903: 1902: 1886: 1885: 1862: 1861: 1838: 1837: 1814: 1813: 1790: 1789: 1761:Castorocauda 1759: 1758: 1746: 1732: 1731: 1708: 1707: 1685: 1671: 1657: 1636:Reigitherium 1634: 1630: 1616: 1604:Khorotherium 1602: 1596: 1582: 1570: 1556: 1552: 1534: 1528: 1516:Morganucodon 1514: 1496: 1421:Theriimorpha 1419: 1405:Fruitafossor 1403: 1399: 1372: 1358: 1342: 1319: 1315: 1258:Microdocodon 1256: 1255: 1241:Agilodocodon 1239: 1238: 1217:Castorocauda 1215: 1214: 1191: 1190: 1167: 1166: 1153: 1126: 1112: 1101: 1078: 1040: 1029: 1012: 1010: 996: 992:Agilodocodon 990: 984: 977:Castorocauda 976: 970: 966:Castorocauda 964: 954: 934:Agilodocodon 932: 911: 886:Castorocauda 884: 854: 848: 842: 835:shuotheriids 832: 801: 794: 785: 781: 778:Lower molars 772: 768:homologizing 752:tribosphenic 748: 738: 731: 722:Upper molars 702:, fang-like 697: 679: 635:Castorocauda 634: 629:and a stout 608: 603:Agilodocodon 601: 569:Castorocauda 567: 561: 531:Microdocodon 529: 523: 495:septomaxilla 489:traits: the 480: 443:ectotympanic 424: 352: 350: 342:shuotheriids 335: 329: 322:tribosphenic 271: 270: 263: 244: 225: 212: 199: 186: 134: 133:(green) and 128: 29: 7509:Wikispecies 7393:Saurodesmus 7181:Cifelliodon 7145:Woodeatonia 7138:Vilevolodon 7117:Sharypovoia 7087:Kermackodon 7050:Allostaffia 7019:Theroteinus 6989:Haramiyavia 6946:Tegotherium 6932:Simpsonodon 6910:Paritatodon 6859:Gondtherium 6845:Dsungarodon 6803:Borealestes 6773:Shuotherium 6758:Paritatodon 6743:Feredocodon 6706:Kotatherium 6666:Wareolestes 6635:Sinoconodon 6615:Purbeckodon 6555:Indotherium 6545:Helvetiodon 6525:Hadrocodium 6515:Gondwanadon 6495:Eozostrodon 6485:Dyskritodon 6405:Bocaconodon 6263:Shartegodon 6249:Oligokyphus 6242:Nuurtherium 6235:Montirictus 6214:Fossiomanus 6200:Dianzhongia 6193:Bocatherium 6146:Sinoconodon 6089:Tritheledon 5963:Rewaconodon 5923:Inditherium 5816:Bonacynodon 5778:Chiniquodon 5748:Tessellatia 5593:Cynognathia 5582:Cynognathia 5501:Thrinaxodon 5481:Nythosaurus 5331:Nanocynodon 4866:11336/81478 4548:(1): 1–105. 4525:(1): 1–235. 4199:: 249–256. 4089:Hurum, J.H. 3993:Borealestes 3886:Borealestes 3339:Simpsonodon 3126:D. striatus 3009:Borealestes 2767:Borealestes 2710:Dsungarodon 2635:Simpsonodon 2552:Tegotherium 2477:Tikitherium 2453:Gondtherium 2276:Sinoconodon 2144:Tegotherium 1998:Simpsonodon 1840:Borealestes 1816:Dsungarodon 1734:Tikitherium 1710:Gondtherium 1639:, from the 1631:Gondtherium 1618:Gondtherium 1607:, from the 1571:Tikitherium 1558:Tikitherium 1521:triconodont 1519:and other " 1507:pantotheres 1423:(including 1378:Monotremata 1376:(including 1321:Hadrocodium 1080:Sinoconodon 1055:crown group 1001:golden mole 946:insectivore 918:golden mole 895:semiaquatic 856:Feredocodon 850:Paritatodon 499:nasal bones 421:Jaw and ear 411:Description 384:-like, and 357:, from the 291:crown-group 278:of extinct 7572:Categories 7353:Meurthodon 7073:Butlerodon 7065:Baidabatyr 7003:Kollikodon 6973:Avashishta 6960:Haramiyida 6896:Krusatodon 6867:Haldanodon 6838:Docofossor 6824:Dobunnodon 6699:Fluctuodon 6646:Storchodon 6625:Rosierodon 6435:Cherwellia 6318:see below↓ 6298:Yunnanodon 6277:Tritylodon 6256:Polistodon 6136:Brasilodon 6081:Riograndia 6046:Chaliminia 5903:Deccanodon 5853:Ecteninion 5846:Diegocanis 5793:Riojanodon 5785:Cromptodon 5718:Charruodon 5614:see below↓ 5534:Galesaurus 5461:Cynosaurus 5451:Bolotridon 5388:Nshimbodon 5341:Madysaurus 5300:Cynodontia 5287:Cynodontia 5259:see below↓ 5250:Cynodontia 5198:Therapsida 5156:Cynodontia 5068:: 105836. 4676:2022-01-29 4110:(1): 1–11. 3407:References 3159:D. superus 3148:D. crassus 3137:D. affinis 3130:Marsh 1881 3086:Marsh 1881 2832:Haldanodon 2791:Docofossor 2743:Dobunnodon 2609:Krusatodon 2046:Krusatodon 1905:Docofossor 1864:Haldanodon 1539:(from the 1499:Monographs 1425:Metatheria 1374:Yinotheria 1347:Haramiyida 1193:Docofossor 1169:Haldanodon 1085:Haramiyida 1051:placentals 1047:marsupials 1043:monotremes 1030:Krusatodon 1013:Krusatodon 997:Docofossor 986:Docofossor 972:Haldanodon 939:tree shrew 913:Docofossor 839:monotremes 824:placentals 820:marsupials 764:placentals 760:marsupials 643:monotremes 631:astragalus 540:thyrohyals 354:Haldanodon 346:monotremes 328:", namely 285:(advanced 264:See text. 219:Cynodontia 206:Therapsida 7578:Docodonts 7515:Docodonta 7485:Docodonta 7188:Denisodon 7102:Megaconus 6787:Docodonta 6676:Woutersia 6656:Stylidens 6595:Paceyodon 5993:Tricuspes 5943:Polonodon 5511:Vetusodon 5374:Abdalodon 5238:Synapsida 5223:Synapsida 5189:Synapsida 5172:Kingdom: 5090:267164151 5046:267167016 5006:199637122 4998:0272-4634 4875:1573-7055 4791:224811982 4783:1028-334X 4715:1573-7055 4701:(1): 10. 4626:0272-4634 4563:Woutersia 4542:Fieldiana 4465:1476-4687 4400:0027-8424 4351:1476-4687 4266:1476-4687 4166:129438779 4158:0031-0301 4075:0024-4082 4027:244822141 4019:1475-4983 3967:197663377 3912:0024-4082 3862:1555-7332 3828:197663503 3758:206562565 3686:206562572 3513:0028-0836 3467:0145-9058 3115:D. victor 3093:D. apoxys 2490:Docodonta 2420:Woutersia 1747:Docodonta 1653:monophyly 1645:Argentina 1553:Woutersia 1536:Woutersia 1154:Docodonta 1103:Cladogram 1091:based on 1028:, though 898:piscivore 808:protocone 708:premolars 627:calcaneum 623:epiphyses 574:synapsids 548:cynodonts 503:lacrimals 455:articular 435:squamosal 362:Guimarota 331:Woutersia 320:with the 287:cynodonts 272:Docodonta 245:Docodonta 193:Synapsida 166:Kingdom: 160:Eukaryota 35:Docodonts 7494:Wikidata 7460:Category 7363:Mitredon 7333:Kunminia 7323:"Hahnia" 7303:Eoraetia 7268:See also 7226:Mammalia 7215:Mammalia 7195:Hahnodon 7152:Xianshou 7124:Shenshou 7026:Thomasia 6881:Itatodon 6852:Ergetiis 6750:Itatodon 6713:Kuehneon 6455:Delsatia 6228:Lufengia 6156:Tikiodon 5180:Chordata 5178:Phylum: 5174:Animalia 4937:35820076 4883:21654055 4502:: 1–215. 4473:28792934 4418:26630008 4274:18075580 4095:(2006). 3959:31320523 3820:31320539 3750:25678661 3678:25678660 3612:46067702 3604:16497926 3475:29846648 3395:See also 3315:P. major 2652:Itatodon 2403:Delsatia 1974:Itatodon 1627:Gondwana 1545:Rhaetian 1530:Delsatia 1429:Eutheria 1360:Mammalia 1063:Mammalia 957:Mesozoic 943:arboreal 844:Itatodon 816:therians 700:incisors 594:thoracic 552:suckling 536:basihyal 511:parietal 463:quadrate 427:mandible 401:thoracic 386:squirrel 366:Portugal 364:site of 337:Delsatia 306:Laurasia 280:Mesozoic 180:Chordata 176:Phylum: 170:Animalia 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Index

Docodont
Middle Jurassic
Early Cretaceous
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Borealestes serendipitus
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Synapsida
Therapsida
Cynodontia
Mammaliaformes
Docodonta
Kretzoi
order
Mesozoic

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