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Gracillariidae

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820: 1699: 204: 31: 56: 354:, which can provide a useful means of identification, especially if the hostplant is known. The subfamilies differ by the adult moth resting posture (Davis and Robinson, 1999). Most Gracillariinae rest with the front of the body steeply raised; Lithocolletinae and Phyllocnistinae rest with the body parallel to the surface; in Lithocolletinae often with the head lowered. 393:. Later instars are cylindrical with round heads, and have chewing mouthparts, legs and functional spinnerets; these feed on plant tissue. Some species additionally have a spinning instar that does not feed, and others have a non-feeding instar before this spinning instar. 1377:
Kawahara, Akito Y.; Plotkin, David; Ohshima, Issei; Lopez-Vaamonde, Carlos; Houlihan, Peter R.; Breinholt, Jesse W.; Kawakita, Atsushi; Xiao, Lei; Regier, Jerome C.; Davis, Donald R.; Kumata, Tosio; Sohn, Jae-Cheon; De Prins, Jurate; Mitter, Charles (2017).
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Li, Xuankun; St Laurent, Ryan; Earl, Chandra; Doorenweerd, Camiel; van Nieukerken, Erik J.; Davis, Donald R.; Johns, Chris A.; Kawakita, Atsushi; Kobayashi, Shigeki; Zwick, Andreas; Lopez-Vaamonde, Carlos; Ohshima, Issei; Kawahara, Akito Y. (2022).
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Lopez-Vaamonde C; Wikström N; Labandeira C; Godfray HC; Simon J. Goodman; James M Cook (1 July 2006). "Fossil-calibrated molecular phylogenies reveal that leaf-mining moths radiated millions of years after their host plants".
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of Gracillariidae (see below). A complete checklist is available of all currently recognised species. There are many undescribed species in the tropics but there is also an online catalogue of
1482:"A new Brazilian Passiflora leafminer: Spinivalva gaucha, gen. n., sp. n. (Lepidoptera, Gracillariidae, Gracillariinae), the first gracillariid without a sap-feeding instar" 1632:
Kawahara, A. Y.; Plotkin, D.; Ohshima, I.; Lopez-Vaamonde, C.; Houlihan, P. R.; Breinholt, J. W.; Davis, D. R.; Kumata, T.; Sohn, J.-C.; De Prins, J.; Mitter, C. (2017).
3825: 819: 1380:"A molecular phylogeny and revised higher-level classification for the leaf-mining moth family G racillariidae and its implications for larval host-use evolution" 1634:"A molecular phylogeny and revised higher-level classification for the leaf-mining moth family Gracillariidae and its implications for larval host-use evolution" 3966: 3864: 365:: a basal row with large scales covering an apical row with smaller scales. The male genitalia has only four pair of muscles and lacks a gnathos. The female 1766: 1749: 169: 3915: 3799: 3838: 1794: 1161: 1118: 298:(Winteraceae), and has leaf mines structurally similar in structure to fossils (see "Fossils"). While there have been some recent 1351: 1193:"New leaf-mining moths from Chile, with remarks on the history and composition of Phyllocnistinae (Lepidoptera: Gracillariidae)" 1244: 1721: 3611: 1729: 1188: 3843: 3851: 203: 55: 1559:"Ninety-seven million years of angiosperm-insect association: paleobiological insights into the meaning of coevolution" 1435:"Phylogeny of gracillariid leaf-mining moths: evolution of larval behaviour inferred from phylogenomic and Sanger data" 3708: 3675: 3606: 3869: 3778: 3791: 1787: 952: 240: 208: 1167: 276:; the South African fauna is quite well known. Although Japanese and Russian authors have recognised additional 1108:"Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness - Lepidoptera" 958: 929: 3951: 1738: 3956: 3628: 919: 862: 160: 151: 132: 1707: 269: 3900: 3183: 1762: 1745: 1017: 281: 1755: 3928: 3713: 1671:
Davis, D.R, and Robinson, G.S. (1999). The Tineoidea and Gracillarioidea. In: Kristensen, N.P. (ed.),
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The family is an old one, with fossil Phyllocnistinae mines known from 97-million-year-old rocks in
3666: 3199: 2974: 2966: 1304: 1302:; James M. Cook (2003). "EVOLUTIONARY DYNAMICS OF HOST-PLANT USE IN A GENUS OF LEAF-MINING MOTHS". 825: 256: 950:
species. The present taxonomic status is unknown, but is probably a junior subjective synonym of
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while others instead roll leaves, and there are also larvae which mine other plant parts or form
378: 50: 3905: 3887: 3734: 3690: 2544: 1606: 1588: 1525: 1507: 1462: 1454: 1407: 1321: 1269: 1261: 1212: 1157: 874: 635: 398: 358: 38: 30: 3830: 1078:, this species was recorded from southern France, where it was said to have been reared on a 280:, there are three currently recognised subfamilies, Phyllocnistinae of which is likely to be 3892: 3525: 3016: 2575: 1653: 1614: 1596: 1580: 1515: 1497: 1446: 1399: 1329: 1313: 1277: 1253: 1220: 1204: 1023: 220: 3094: 2770: 2647: 2523: 2515: 2481: 2455: 2447: 2114: 2085: 2079: 2070: 1881: 1875: 1863: 1733: 1711: 1299: 1063:, this species was described from Switzerland. Larvae were probably bred from a mine on a 892: 859: 847: 745: 582: 567: 273: 166: 157: 117: 1649: 1576: 1493: 1434: 1395: 1107: 3786: 3773: 3315: 3102: 2780: 2705: 2671: 2629: 2554: 2182: 1520: 1192: 973: 853: 841: 708: 650: 503: 497: 441: 426: 148: 3680: 3945: 3543: 3468: 3376: 3323: 3302: 3294: 3165: 3045: 3037: 2757: 2639: 2634: 2624: 2403: 2395: 2382: 2374: 2345: 2306: 2174: 2119: 2106: 2097: 2022: 2014: 2005: 1967: 1961: 1949: 1601: 1558: 1347: 1257: 886: 730: 455:
occurred long after that of the larval hostplants, rather than demonstrating a tight
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Brito, Rosangela; Gonçalves, Gislene; Vargas, Hector; Moreira, Gilson (2013-04-17).
3739: 3562: 3530: 3512: 3460: 3392: 3352: 3152: 3144: 3110: 2995: 2987: 2808: 2798: 2785: 2739: 2731: 2718: 2710: 2697: 2655: 2616: 2606: 2583: 2562: 2531: 2494: 2486: 2473: 2353: 2301: 1847: 488: 449: 434: 362: 351: 307: 299: 263: 190: 1726: 1039:, this species was described from Norway. The present taxonomic status is unknown. 310:
framework for the subfamilies of Gracillariidae. Some genera are very large, e.g.
985:, this species was described from Italy. The present taxonomic status is unknown. 3856: 3812: 3660: 3520: 3478: 3402: 3397: 3368: 3336: 3231: 3172: 3008: 2932: 2924: 2911: 2903: 2874: 2824: 2749: 2663: 2598: 2593: 2588: 2570: 2361: 2296: 2224: 2203: 2195: 2145: 1924: 1918: 1906: 1804: 1544:"HOSTS - the Hostplants and Caterpillars Database at the Natural History Museum" 1086: 1011: 996: 880: 787: 456: 445: 422: 338:
Gracillariidae occur in all terrestrial regions of the world except Antarctica.
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Lepidoptera, Moths and Butterflies, 1: Evolution, Systematics, and Biogeography
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Proceedings of the National Academy of Sciences of the United States of America
909:, this species was described from Switzerland. Larvae were recorded feeding on 3586: 3494: 3486: 3439: 3423: 3410: 3384: 3360: 3344: 3265: 3252: 3244: 3223: 3215: 3131: 3123: 3066: 3058: 3024: 2882: 2816: 2803: 2775: 2762: 2684: 2311: 2253: 2158: 1772: 944:, this species was described from Finland. The larval hostplant is probably a 868: 833:
Gracillariidae phylogeny has been revised in 2017 and is now containing eight
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trees or shrubs. Patterns of hostplant shifting have been inferred for many
386: 347: 87: 67: 1610: 1529: 1466: 1273: 1009:, this species was described from Germany. Larvae were recorded feeding on 1618: 1333: 1281: 3721: 3700: 3645: 3431: 3207: 2953: 2945: 2890: 2845: 2502: 2420: 2288: 2153: 2136: 1989: 1841: 1048:, this species was described from Italy. Larvae were recorded feeding on 472: 385:
have a flattened head and body, modified mandibles, and lack functional
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that includes several economic, horticultural or recently invasive pest
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C C Labandeira; D L Dilcher; D R Davis; D L Wagner (6 December 1994).
1450: 3817: 3502: 1829: 1817: 492: 476: 468: 382: 294: 259: 97: 77: 3804: 3622: 1021:. The present taxonomic status is unknown. It might be a synonym of 3765: 1689: 1317: 3923: 946: 911: 818: 357:
Some additional features can be seen under close examination. The
289: 202: 1716: 381:, meaning there is a major change within the larval stage. Early 410: 3626: 1986: 1946: 1903: 1860: 1776: 1704: 390: 267: 3752: 402:
is an exception to this trend, lacking a sap-feeding stage.
3226:(gossamer-winged butterflies: blues, coppers and relatives) 3447:(daggers, sallows, owlet moths, quakers, cutworms, darts) 1717:
Global Taxonomic Database of Gracillariidae (Lepidoptera)
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These generally small (wingspan 5–20 mm) moths are
3434:(underwing, tiger, tussock, litter, snout, owlet moths) 306:
species there is need for a satisfactory modern global
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Gracillariidae have various lifestyles. Some larvae
3635: 3555: 3511: 3477: 3422: 3335: 3314: 3293: 3264: 3243: 3164: 3143: 3122: 3093: 3075: 3057: 3036: 3007: 2986: 2965: 2944: 2923: 2902: 2865: 2836: 2748: 2730: 2615: 2553: 2514: 2464: 2446: 2419: 2394: 2373: 2344: 2323: 2279: 2252: 2223: 2194: 2144: 2135: 2105: 2096: 2078: 2069: 2044: 2035: 2013: 2004: 1960: 1917: 1874: 475:. There are other fossil mines known from rocks of 2885:(castniid moths: giant butterfly-moths, sun moths) 1679:(35): 91-117. Walter de Gruyter. Berlin, New York. 1700:Natural History Museum Lepidoptera genus database 1069:species. The present taxonomic status is unknown. 1150:Prins, Willy De; Prins, Jurate De (1998-01-01). 483:age. There are also two adult moths known from 1788: 8: 1293: 1291: 1236: 1234: 181: 3623: 3234:(brush-footed, or four-footed butterflies) 2443: 2439: 2276: 2249: 2141: 2102: 2075: 2041: 2010: 2001: 1997: 1983: 1957: 1943: 1914: 1900: 1871: 1857: 1795: 1781: 1773: 915:. The present taxonomic status is unknown. 29: 20: 1657: 1600: 1519: 1501: 361:flagellomeres of adults have two rows of 284:. In this subfamily, the primitive genus 3413:(hawk moths, sphinx moths and hornworms) 3210:(whites, yellows, orangetips, sulphurs) 1099: 3326:(eggars, snout moths, or lappet moths) 421:Many host plants are known, generally 1695:Natural History Museum hosts database 7: 3967:Taxa named by Henry Tibbats Stainton 3929:DE22D21F-C50D-FFF0-FF13-FE16FC69FA18 3880:6528288e-031a-410e-8024-1583ec01d9e1 3792:8eb27281-8f44-4f20-8a84-894fa5172965 1427: 1425: 1423: 1421: 1372: 1370: 1368: 1145: 1143: 1141: 1139: 448:species has shown that the burst of 723: 701: 628: 606: 560: 553: 546: 539: 532: 524: 377:Most larval Gracillariidae undergo 197:About 101 genera and 1,866 species 2848:(carpenter millers, or goat moths) 2765:(burnet, forester, or smoky moths) 883:Kuznetzov & Baryshnikova, 2001 369:is short and laterally flattened. 14: 2169:(Amazonian primitive ghost moths) 1124:from the original on 10 June 2014 1258:10.1111/J.1420-9101.2005.01070.X 1073:"Phyllonorycter" sessilifoliella 850:Li, Ohshima & Kawahara, 2022 54: 2185:(African primitive ghost moths) 1354:from the original on 2007-03-24 1245:Journal of Evolutionary Biology 1170:from the original on 2023-09-30 2489:(Old World spiny-winged moths) 1: 3387:(Kentish glory and relatives) 2206:(New Zealand primitive moths) 1727:Fauna Europaea : Experts 1722:Fauna Europaea Gracillariidae 829:) tree damage in Parma, Italy 3533:(oriental swallowtail moths) 2811:(planthopper parasite moths) 1034:"Lithocolletis" norvegicella 966:"Lithocolletis" graeseriella 889:Kawahara & Ohshima, 2016 877:RĂ©al & Balachowsky, 1966 871:Kawahara & Ohshima, 2016 844:Kawahara & Ohshima, 2016 518:Phylogeny of Gracillariidae 231:and the principal family of 3607:Taxonomy of the Lepidoptera 3194:(American moth-butterflies) 3155:(Old World butterfly-moths) 3134:(picture-winged leaf moths) 1884:(mandibulate archaic moths) 1769:Featured Creatures Web site 1752:Featured Creatures Web site 1004:"Lithocolletis" lativitella 939:"Phyllonorycter" fennicella 823:Horse-chestnut leaf miner ( 3983: 3105:(tropical fruitworm moths) 2642:(case-bearers, case moths) 2406:(trumpet leaf miner moths) 2161:(swift moths, ghost moths) 1153:World Catalogue of Insects 930:"Gracilaria" delicatulella 498:Gracillariites lithuanicus 3602: 3583: 3355:(American silkworm moths) 3186:(swallowtail butterflies) 2856:(dudgeon carpenter moths) 2565:(false diamondback moths) 2476:(burrowing webworm moths) 2442: 2000: 1996: 1982: 1956: 1942: 1913: 1899: 1870: 1856: 1812: 1043:"Gracillaria" pistaciella 953:Lithocolletis viminetorum 785: 743: 728: 721: 706: 699: 648: 633: 626: 611: 604: 580: 565: 558: 551: 544: 537: 530: 292:feeds on the plant genus 241:horse-chestnut leaf miner 209:Phyllocnistis magnoliella 196: 189: 145: 140: 51:Scientific classification 49: 37: 28: 23: 3347:(Australian lappet moth) 2356:(pigmy, or midget moths) 1675:. Handbuch der Zoologie 1585:10.1073/PNAS.91.25.12278 1503:10.3897/zookeys.291.4910 959:Lithocolletis salictella 920:"Gracilaria" confectella 251:Taxonomy and systematics 2177:(miniature ghost moths) 1298:Carlos Lopez-Vaamonde; 980:"Lithocolletis" italica 389:or legs; these feed on 3556:Superfamily unassigned 2977:(bristle-legged moths) 2526:(ribbed cocoon makers) 2335:(Andean endemic moths) 2291:(fairy longhorn moths) 1705:Afrotropical Checklist 983:Herrich-Schäffer, 1855 895:Herrich-Schäffer, 1857 830: 511:Subfamilies and genera 213: 182:Subfamilies and genera 43:Gracillaria syringella 3901:Paleobiology Database 3463:(prominents, kittens) 2998:(fringe-tufted moths) 2054:Acanthopteroctetidae 2046:Acanthopteroctetoidea 1638:Systematic Entomology 1384:Systematic Entomology 1300:H. Charles J. Godfray 989:"Ornix" jyngipennella 822: 429:species in the genus 206: 3787:Fauna Europaea (new) 2975:Schreckensteiniidae 2967:Schreckensteinioidea 2956:(false burnet moths) 2827:(slug, or cup moths) 2385:(Gondwanaland moths) 2364:(white eyecap moths) 2122:(archaic bell moths) 1740:Caloptilia azaleella 1197:Tropical Lepidoptera 1051:Pistacia terebinthus 813:from Li et al. 2022 3305:(sack bearer moths) 3027:(false plume moths) 3019:(many-plumed moths) 2877:(little bear moths) 2819:(slug caterpillars) 2674:(long-horned moths) 2658:(grass-miner moths) 2056:(archaic sun moths) 1650:2017SysEn..42...60K 1577:1994PNAS...9112278L 1494:2013ZooK..291....1B 1396:2017SysEn..42...60K 826:Cameraria ohridella 245:Cameraria ohridella 1759:, citrus peelminer 1742:, azalea leafminer 1732:2012-02-05 at the 1710:2007-02-05 at the 1659:10.1111/syen.12210 1404:10.1111/syen.12210 1018:Pyrus scandinavica 831: 453:adaptive radiation 379:hypermetamorphosis 302:-based studies of 272:2007-02-05 at the 266:described species 214: 45:(Fabricius, 1794) 3939: 3938: 3888:Open Tree of Life 3629:Taxon identifiers 3620: 3619: 3598: 3597: 3594: 3593: 3579: 3578: 3575: 3574: 3571: 3570: 3547: 3534: 3498: 3490: 3464: 3456: 3448: 3435: 3414: 3406: 3388: 3380: 3372: 3364: 3356: 3348: 3327: 3306: 3285: 3277: 3256: 3235: 3227: 3219: 3211: 3203: 3195: 3187: 3177: 3156: 3135: 3114: 3113:(fruitworm moths) 3106: 3049: 3028: 3020: 2999: 2978: 2957: 2936: 2915: 2914:(metalmark moths) 2894: 2893:(clearwing moths) 2886: 2878: 2857: 2849: 2828: 2820: 2812: 2789: 2766: 2722: 2714: 2701: 2700:(concealer moths) 2693: 2675: 2667: 2659: 2651: 2643: 2602: 2579: 2576:Glyphipterigidae 2566: 2545:Roeslerstammiidae 2535: 2527: 2506: 2498: 2490: 2477: 2415: 2414: 2407: 2386: 2365: 2357: 2336: 2315: 2292: 2219: 2218: 2215: 2214: 2207: 2186: 2178: 2170: 2162: 2131: 2130: 2123: 2092: 2091: 2065: 2064: 2057: 2031: 2030: 1978: 1977: 1974: 1973: 1962:Heterobathmioidea 1938: 1937: 1934: 1933: 1928: 1895: 1894: 1891: 1890: 1885: 1451:10.1111/cla.12490 1163:978-87-88757-64-4 1077: 1062: 1061:MĂĽller-Rutz, 1934 1058:"Ornix" quercella 1047: 1038: 1029: 1008: 993: 992:Heydenreich, 1851 984: 970: 943: 935: 925: 908: 907:MĂĽller-Rutz, 1920 904:"Ornix" blandella 899:Unplaced species 875:Oecophyllembiinae 817: 816: 807: 806: 798: 797: 774: 773: 765: 764: 756: 755: 688: 687: 679: 678: 670: 669: 661: 660: 636:Oecophyllembiinae 593: 592: 501:Kozlov, 1987 and 417:Larval hostplants 399:Spinivalva gaucha 212:in magnolia leaf. 201: 200: 136: 39:Privet leaf miner 3974: 3932: 3931: 3919: 3918: 3909: 3908: 3896: 3895: 3883: 3882: 3873: 3872: 3860: 3859: 3857:NBNSYS0000160082 3847: 3846: 3834: 3833: 3821: 3820: 3808: 3807: 3795: 3794: 3782: 3781: 3769: 3768: 3756: 3755: 3743: 3742: 3730: 3729: 3717: 3716: 3704: 3703: 3694: 3693: 3684: 3683: 3671: 3670: 3669: 3656: 3655: 3654: 3624: 3546:(geometer moths) 3545: 3532: 3526:Pseudobistonidae 3496: 3488: 3462: 3454: 3446: 3433: 3412: 3404: 3386: 3378: 3370: 3362: 3354: 3346: 3325: 3304: 3283: 3275: 3254: 3233: 3225: 3217: 3209: 3201: 3193: 3185: 3176: 3169: 3154: 3133: 3112: 3104: 3047: 3026: 3018: 2997: 2976: 2955: 2934: 2913: 2892: 2884: 2876: 2855: 2847: 2826: 2818: 2810: 2787: 2764: 2720: 2712: 2699: 2691: 2673: 2665: 2657: 2649: 2648:Cosmopterigidae 2641: 2600: 2577: 2564: 2533: 2525: 2524:Bucculatricidae 2504: 2496: 2488: 2475: 2444: 2440: 2405: 2384: 2363: 2355: 2334: 2313: 2290: 2277: 2250: 2205: 2184: 2176: 2168: 2160: 2142: 2121: 2103: 2076: 2055: 2042: 2011: 2002: 1998: 1984: 1968:Heterobathmiidae 1958: 1944: 1926: 1915: 1901: 1883: 1882:Micropterigidae 1876:Micropterigoidea 1872: 1858: 1797: 1790: 1783: 1774: 1664: 1663: 1661: 1629: 1623: 1622: 1604: 1571:(25): 12278–82. 1554: 1548: 1547: 1540: 1534: 1533: 1523: 1505: 1477: 1471: 1470: 1429: 1416: 1415: 1374: 1363: 1362: 1360: 1359: 1348:"Gracillariidae" 1344: 1338: 1337: 1312:(8): 1804–1821. 1295: 1286: 1285: 1252:(4): 1314–1326. 1238: 1229: 1228: 1185: 1179: 1178: 1176: 1175: 1147: 1134: 1133: 1131: 1129: 1123: 1112: 1104: 1075: 1060: 1045: 1036: 1027: 1024:Tinea lantanella 1006: 991: 982: 968: 941: 933: 923: 906: 724: 702: 629: 607: 561: 554: 547: 540: 533: 525: 515: 514: 444:study mainly of 286:Prophyllocnistis 219:is an important 170:Herrich-Schäffer 131: 59: 58: 33: 21: 3982: 3981: 3977: 3976: 3975: 3973: 3972: 3971: 3942: 3941: 3940: 3935: 3927: 3922: 3914: 3912: 3904: 3899: 3891: 3886: 3878: 3876: 3868: 3863: 3855: 3850: 3842: 3837: 3829: 3824: 3816: 3811: 3803: 3798: 3790: 3785: 3777: 3772: 3764: 3759: 3751: 3746: 3738: 3733: 3725: 3720: 3712: 3707: 3699: 3697: 3689: 3687: 3679: 3674: 3665: 3664: 3659: 3650: 3649: 3644: 3631: 3621: 3616: 3612:Lists by region 3590: 3589:is not a clade. 3585:Note: division 3567: 3551: 3507: 3473: 3418: 3379:(Dryandra moth) 3371:(Brahmin moths) 3331: 3310: 3289: 3260: 3239: 3170: 3168: 3160: 3139: 3118: 3103:Copromorphidae 3095:Copromorphoidea 3089: 3071: 3053: 3032: 3003: 2982: 2961: 2940: 2935:(tortrix moths) 2919: 2898: 2861: 2832: 2788:(flannel moths) 2771:Himantopteridae 2744: 2726: 2672:Lecithoceridae 2666:(twirler moths) 2611: 2549: 2534:(Douglas moths) 2516:Gracillarioidea 2510: 2497:(bagworm moths) 2482:Arrhenophanidae 2460: 2448:Simaethistoidea 2434: 2432: 2430: 2428: 2426: 2424: 2422: 2411: 2390: 2369: 2340: 2319: 2271: 2269: 2267: 2265: 2263: 2261: 2259: 2257: 2255: 2244: 2242: 2240: 2238: 2236: 2234: 2232: 2230: 2228: 2226: 2211: 2204:Mnesarchaeidae 2196:Mnesarchaeoidea 2190: 2183:Prototheoridae 2127: 2115:Aenigmatineidae 2088: 2080:Lophocoronoidea 2061: 2027: 1992: 1970: 1952: 1950:Heterobathmiina 1930: 1925:Agathiphagidae 1919:Agathiphagoidea 1909: 1887: 1866: 1852: 1808: 1801: 1734:Wayback Machine 1712:Wayback Machine 1686: 1668: 1667: 1631: 1630: 1626: 1556: 1555: 1551: 1542: 1541: 1537: 1479: 1478: 1474: 1431: 1430: 1419: 1376: 1375: 1366: 1357: 1355: 1346: 1345: 1341: 1297: 1296: 1289: 1240: 1239: 1232: 1189:Donald R. Davis 1187: 1186: 1182: 1173: 1171: 1164: 1156:. Vol. 6. 1149: 1148: 1137: 1127: 1125: 1121: 1110: 1106: 1105: 1101: 1096: 893:Phyllocnistinae 860:Lithocolletinae 848:Callicercopinae 808: 799: 775: 766: 757: 746:Callicercopinae 689: 680: 671: 662: 594: 583:Phyllocnistinae 568:Lithocolletinae 528:Gracillariidae 513: 465: 419: 375: 344: 336: 274:Wayback Machine 253: 167:Phyllocnistinae 158:Lithocolletinae 130: 118:Gracillarioidea 53: 24:Gracillariidae 17: 16:Family of moths 12: 11: 5: 3980: 3978: 3970: 3969: 3964: 3959: 3954: 3952:Gracillariidae 3944: 3943: 3937: 3936: 3934: 3933: 3920: 3916:gracillariidae 3910: 3897: 3884: 3874: 3861: 3848: 3835: 3822: 3809: 3796: 3783: 3774:Fauna Europaea 3770: 3757: 3744: 3731: 3718: 3705: 3695: 3691:Gracillariidae 3685: 3681:Gracillariidae 3672: 3667:Gracillariidae 3657: 3641: 3639: 3637:Gracillariidae 3633: 3632: 3627: 3618: 3617: 3615: 3614: 3609: 3603: 3600: 3599: 3596: 3595: 3592: 3591: 3584: 3581: 3580: 3577: 3576: 3573: 3572: 3569: 3568: 3566: 3565: 3559: 3557: 3553: 3552: 3550: 3549: 3541: 3536: 3528: 3523: 3517: 3515: 3509: 3508: 3506: 3505: 3500: 3492: 3483: 3481: 3475: 3474: 3472: 3471: 3466: 3458: 3450: 3442: 3437: 3428: 3426: 3420: 3419: 3417: 3416: 3408: 3400: 3395: 3390: 3382: 3374: 3366: 3358: 3350: 3341: 3339: 3333: 3332: 3330: 3329: 3324:Lasiocampidae 3320: 3318: 3316:Lasiocampoidea 3312: 3311: 3309: 3308: 3299: 3297: 3291: 3290: 3288: 3287: 3279: 3270: 3268: 3262: 3261: 3259: 3258: 3249: 3247: 3241: 3240: 3238: 3237: 3229: 3221: 3213: 3205: 3197: 3189: 3180: 3178: 3162: 3161: 3159: 3158: 3149: 3147: 3141: 3140: 3138: 3137: 3128: 3126: 3120: 3119: 3117: 3116: 3108: 3099: 3097: 3091: 3090: 3088: 3087: 3081: 3079: 3073: 3072: 3070: 3069: 3063: 3061: 3059:Whalleyanoidea 3055: 3054: 3052: 3051: 3046:Pterophoridae 3042: 3040: 3038:Pterophoroidea 3034: 3033: 3031: 3030: 3022: 3013: 3011: 3005: 3004: 3002: 3001: 2992: 2990: 2984: 2983: 2981: 2980: 2971: 2969: 2963: 2962: 2960: 2959: 2950: 2948: 2942: 2941: 2939: 2938: 2929: 2927: 2921: 2920: 2918: 2917: 2908: 2906: 2900: 2899: 2897: 2896: 2888: 2880: 2871: 2869: 2863: 2862: 2860: 2859: 2851: 2842: 2840: 2834: 2833: 2831: 2830: 2822: 2814: 2806: 2801: 2796: 2791: 2786:Megalopygidae 2783: 2781:Somabrachyidae 2778: 2773: 2768: 2760: 2754: 2752: 2746: 2745: 2743: 2742: 2736: 2734: 2728: 2727: 2725: 2724: 2721:(timber moths) 2716: 2713:(flower moths) 2708: 2706:Pterolonchidae 2703: 2695: 2692:(mompha moths) 2687: 2682: 2677: 2669: 2661: 2653: 2650:(cosmet moths) 2645: 2640:Coleophoridae 2637: 2632: 2630:Batrachedridae 2627: 2621: 2619: 2613: 2612: 2610: 2609: 2604: 2601:(ermine moths) 2599:Yponomeutidae 2596: 2591: 2586: 2581: 2573: 2568: 2559: 2557: 2555:Yponomeutoidea 2551: 2550: 2548: 2547: 2542: 2540:Gracillariidae 2537: 2529: 2520: 2518: 2512: 2511: 2509: 2508: 2505:(fungus moths) 2500: 2492: 2484: 2479: 2470: 2468: 2462: 2461: 2459: 2458: 2456:Simaethistidae 2452: 2450: 2437: 2417: 2416: 2413: 2412: 2410: 2409: 2400: 2398: 2392: 2391: 2389: 2388: 2383:Palaephatidae 2379: 2377: 2375:Palaephatoidea 2371: 2370: 2368: 2367: 2359: 2350: 2348: 2342: 2341: 2339: 2338: 2329: 2327: 2321: 2320: 2318: 2317: 2309: 2304: 2299: 2294: 2285: 2283: 2274: 2247: 2221: 2220: 2217: 2216: 2213: 2212: 2210: 2209: 2200: 2198: 2192: 2191: 2189: 2188: 2180: 2175:Palaeosetidae 2172: 2164: 2156: 2150: 2148: 2139: 2133: 2132: 2129: 2128: 2126: 2125: 2120:Neopseustidae 2117: 2111: 2109: 2107:Neopseustoidea 2100: 2094: 2093: 2090: 2089: 2086:Lophocoronidae 2084: 2082: 2073: 2067: 2066: 2063: 2062: 2060: 2059: 2050: 2048: 2039: 2033: 2032: 2029: 2028: 2026: 2025: 2019: 2017: 2015:Eriocranioidea 2008: 1994: 1993: 1987: 1980: 1979: 1976: 1975: 1972: 1971: 1966: 1964: 1954: 1953: 1947: 1940: 1939: 1936: 1935: 1932: 1931: 1923: 1921: 1911: 1910: 1904: 1897: 1896: 1893: 1892: 1889: 1888: 1880: 1878: 1868: 1867: 1861: 1854: 1853: 1851: 1850: 1844: 1838: 1832: 1826: 1820: 1813: 1810: 1809: 1802: 1800: 1799: 1792: 1785: 1777: 1771: 1770: 1757:Marmara gulosa 1753: 1736: 1724: 1719: 1714: 1702: 1697: 1692: 1685: 1684:External links 1682: 1681: 1680: 1666: 1665: 1624: 1549: 1535: 1472: 1445:(3): 277–300. 1417: 1364: 1339: 1318:10.1554/02-470 1287: 1230: 1191:(1 May 1994). 1180: 1162: 1135: 1098: 1097: 1095: 1092: 1091: 1090: 1070: 1055: 1040: 1031: 1007:Sorhagen, 1900 1001: 986: 977: 974:Phyllonorycter 969:Sorhagen, 1900 963: 936: 926: 916: 897: 896: 890: 884: 878: 872: 866: 857: 856:Stainton, 1854 854:Gracillariinae 851: 845: 842:Acrocercopinae 815: 814: 810: 809: 805: 804: 801: 800: 796: 795: 792: 791: 784: 781: 780: 777: 776: 772: 771: 768: 767: 763: 762: 759: 758: 754: 753: 750: 749: 742: 739: 738: 735: 734: 727: 722: 720: 717: 716: 713: 712: 709:Gracillariinae 705: 700: 698: 695: 694: 691: 690: 686: 685: 682: 681: 677: 676: 673: 672: 668: 667: 664: 663: 659: 658: 655: 654: 651:Acrocercopinae 647: 644: 643: 640: 639: 632: 627: 625: 622: 621: 618: 617: 610: 605: 603: 600: 599: 596: 595: 591: 590: 587: 586: 579: 576: 575: 572: 571: 564: 559: 557: 552: 550: 545: 543: 538: 536: 531: 529: 523: 520: 519: 512: 509: 507:Kozlov, 1987. 464: 461: 457:coevolutionary 442:DNA sequencing 431:Phyllonorycter 427:United Kingdom 423:dicotyledonous 418: 415: 396:The Brazilian 374: 371: 343: 342:Identification 340: 335: 332: 328:Phyllonorycter 252: 249: 217:Gracillariidae 199: 198: 194: 193: 187: 186: 174: 173: 164: 155: 149:Gracillariinae 143: 142: 138: 137: 128:Gracillariidae 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 65: 61: 60: 47: 46: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 3979: 3968: 3965: 3963: 3960: 3958: 3957:Moth families 3955: 3953: 3950: 3949: 3947: 3930: 3925: 3921: 3917: 3911: 3907: 3902: 3898: 3894: 3889: 3885: 3881: 3875: 3871: 3866: 3862: 3858: 3853: 3849: 3845: 3840: 3836: 3832: 3827: 3823: 3819: 3814: 3810: 3806: 3801: 3797: 3793: 3788: 3784: 3780: 3775: 3771: 3767: 3762: 3758: 3754: 3749: 3745: 3741: 3736: 3732: 3728: 3723: 3719: 3715: 3710: 3706: 3702: 3696: 3692: 3686: 3682: 3677: 3673: 3668: 3662: 3658: 3653: 3647: 3643: 3642: 3640: 3638: 3634: 3630: 3625: 3613: 3610: 3608: 3605: 3604: 3601: 3588: 3582: 3564: 3561: 3560: 3558: 3554: 3548: 3542: 3540: 3537: 3535: 3531:Epicopeiidae 3529: 3527: 3524: 3522: 3519: 3518: 3516: 3514: 3510: 3504: 3501: 3499: 3493: 3491: 3485: 3484: 3482: 3480: 3476: 3470: 3469:Oenosandridae 3467: 3465: 3461:Notodontidae 3459: 3457: 3451: 3449: 3443: 3441: 3438: 3436: 3430: 3429: 3427: 3425: 3421: 3415: 3409: 3407: 3401: 3399: 3396: 3394: 3391: 3389: 3383: 3381: 3375: 3373: 3367: 3365: 3359: 3357: 3353:Apatelodidae 3351: 3349: 3343: 3342: 3340: 3338: 3334: 3328: 3322: 3321: 3319: 3317: 3313: 3307: 3303:Mimallonidae 3301: 3300: 3298: 3296: 3295:Mimallonoidea 3292: 3286: 3280: 3278: 3276:(snout moths) 3272: 3271: 3269: 3267: 3263: 3257: 3251: 3250: 3248: 3246: 3242: 3236: 3230: 3228: 3222: 3220: 3214: 3212: 3206: 3204: 3198: 3196: 3190: 3188: 3184:Papilionidae 3182: 3181: 3179: 3174: 3167: 3166:Papilionoidea 3163: 3157: 3153:Callidulidae 3151: 3150: 3148: 3146: 3145:Calliduloidea 3142: 3136: 3130: 3129: 3127: 3125: 3121: 3115: 3111:Carposinidae 3109: 3107: 3101: 3100: 3098: 3096: 3092: 3086: 3083: 3082: 3080: 3078: 3074: 3068: 3067:Whalleyanidae 3065: 3064: 3062: 3060: 3056: 3050: 3048:(plume moths) 3044: 3043: 3041: 3039: 3035: 3029: 3023: 3021: 3015: 3014: 3012: 3010: 3006: 3000: 2996:Epermeniidae 2994: 2993: 2991: 2989: 2988:Epermenioidea 2985: 2979: 2973: 2972: 2970: 2968: 2964: 2958: 2952: 2951: 2949: 2947: 2943: 2937: 2931: 2930: 2928: 2926: 2922: 2916: 2910: 2909: 2907: 2905: 2901: 2895: 2889: 2887: 2881: 2879: 2873: 2872: 2870: 2868: 2864: 2858: 2854:Dudgeoneidae 2852: 2850: 2844: 2843: 2841: 2839: 2835: 2829: 2823: 2821: 2815: 2813: 2809:Epipyropidae 2807: 2805: 2804:Cyclotornidae 2802: 2800: 2797: 2795: 2792: 2790: 2784: 2782: 2779: 2777: 2774: 2772: 2769: 2767: 2761: 2759: 2758:Heterogynidae 2756: 2755: 2753: 2751: 2747: 2741: 2738: 2737: 2735: 2733: 2732:Galacticoidea 2729: 2723: 2719:Xyloryctidae 2717: 2715: 2711:Scythrididae 2709: 2707: 2704: 2702: 2698:Oecophoridae 2696: 2694: 2688: 2686: 2685:Metachandidae 2683: 2681: 2678: 2676: 2670: 2668: 2662: 2660: 2656:Elachistidae 2654: 2652: 2646: 2644: 2638: 2636: 2635:Blastobasidae 2633: 2631: 2628: 2626: 2625:Autostichidae 2623: 2622: 2620: 2618: 2614: 2608: 2605: 2603: 2597: 2595: 2592: 2590: 2587: 2585: 2582: 2580: 2578:(sedge moths) 2574: 2572: 2569: 2567: 2563:Acrolepiidae 2561: 2560: 2558: 2556: 2552: 2546: 2543: 2541: 2538: 2536: 2532:Douglasiidae 2530: 2528: 2522: 2521: 2519: 2517: 2513: 2507: 2501: 2499: 2493: 2491: 2487:Eriocottidae 2485: 2483: 2480: 2478: 2474:Acrolophidae 2472: 2471: 2469: 2467: 2463: 2457: 2454: 2453: 2451: 2449: 2445: 2441: 2438: 2436: 2418: 2408: 2404:Tischeriidae 2402: 2401: 2399: 2397: 2396:Tischerioidea 2393: 2387: 2381: 2380: 2378: 2376: 2372: 2366: 2360: 2358: 2354:Nepticulidae 2352: 2351: 2349: 2347: 2346:Nepticuloidea 2343: 2337: 2333:Andesianidae 2331: 2330: 2328: 2326: 2325:Andesianoidea 2322: 2316: 2314:(yucca moths) 2310: 2308: 2307:Incurvariidae 2305: 2303: 2300: 2298: 2295: 2293: 2287: 2286: 2284: 2282: 2278: 2275: 2273: 2251: 2248: 2246: 2222: 2208: 2202: 2201: 2199: 2197: 2193: 2187: 2181: 2179: 2173: 2171: 2167:Neotheoridae 2165: 2163: 2157: 2155: 2152: 2151: 2149: 2147: 2143: 2140: 2138: 2134: 2124: 2118: 2116: 2113: 2112: 2110: 2108: 2104: 2101: 2099: 2095: 2087: 2083: 2081: 2077: 2074: 2072: 2071:Lophocoronina 2068: 2058: 2052: 2051: 2049: 2047: 2043: 2040: 2038: 2037:Acanthoctesia 2034: 2024: 2023:Eriocraniidae 2021: 2020: 2018: 2016: 2012: 2009: 2007: 2003: 1999: 1995: 1991: 1985: 1981: 1969: 1965: 1963: 1959: 1955: 1951: 1945: 1941: 1929: 1927:(kauri moths) 1922: 1920: 1916: 1912: 1908: 1902: 1898: 1886: 1879: 1877: 1873: 1869: 1865: 1859: 1855: 1849: 1845: 1843: 1839: 1837: 1833: 1831: 1827: 1825: 1821: 1819: 1815: 1814: 1811: 1806: 1798: 1793: 1791: 1786: 1784: 1779: 1778: 1775: 1768: 1764: 1760: 1758: 1754: 1751: 1747: 1743: 1741: 1737: 1735: 1731: 1728: 1725: 1723: 1720: 1718: 1715: 1713: 1709: 1706: 1703: 1701: 1698: 1696: 1693: 1691: 1688: 1687: 1683: 1678: 1674: 1670: 1669: 1660: 1655: 1651: 1647: 1643: 1639: 1635: 1628: 1625: 1620: 1616: 1612: 1608: 1603: 1598: 1594: 1590: 1586: 1582: 1578: 1574: 1570: 1566: 1565: 1560: 1553: 1550: 1545: 1539: 1536: 1531: 1527: 1522: 1517: 1513: 1509: 1504: 1499: 1495: 1491: 1488:(291): 1–26. 1487: 1483: 1476: 1473: 1468: 1464: 1460: 1456: 1452: 1448: 1444: 1440: 1436: 1428: 1426: 1424: 1422: 1418: 1413: 1409: 1405: 1401: 1397: 1393: 1389: 1385: 1381: 1373: 1371: 1369: 1365: 1353: 1349: 1343: 1340: 1335: 1331: 1327: 1323: 1319: 1315: 1311: 1307: 1306: 1301: 1294: 1292: 1288: 1283: 1279: 1275: 1271: 1267: 1263: 1259: 1255: 1251: 1247: 1246: 1237: 1235: 1231: 1226: 1222: 1218: 1214: 1210: 1206: 1202: 1198: 1194: 1190: 1184: 1181: 1169: 1165: 1159: 1155: 1154: 1146: 1144: 1142: 1140: 1136: 1120: 1116: 1109: 1103: 1100: 1093: 1089: 1088: 1083: 1082: 1074: 1071: 1068: 1067: 1059: 1056: 1053: 1052: 1046:Rondani, 1876 1044: 1041: 1035: 1032: 1028:Schrank, 1802 1026: 1025: 1020: 1019: 1014: 1013: 1005: 1002: 999: 998: 990: 987: 981: 978: 976: 975: 967: 964: 961: 960: 955: 954: 949: 948: 940: 937: 932: 931: 927: 922: 921: 917: 914: 913: 905: 902: 901: 900: 894: 891: 888: 887:Parornichinae 885: 882: 879: 876: 873: 870: 867: 864: 861: 858: 855: 852: 849: 846: 843: 840: 839: 838: 836: 828: 827: 821: 812: 811: 803: 802: 794: 793: 790: 789: 783: 782: 779: 778: 770: 769: 761: 760: 752: 751: 748: 747: 741: 740: 737: 736: 733: 732: 731:Parornichinae 726: 725: 719: 718: 715: 714: 711: 710: 704: 703: 697: 696: 693: 692: 684: 683: 675: 674: 666: 665: 657: 656: 653: 652: 646: 645: 642: 641: 638: 637: 631: 630: 624: 623: 620: 619: 616: 615: 609: 608: 602: 601: 598: 597: 589: 588: 585: 584: 578: 577: 574: 573: 570: 569: 563: 562: 556: 555: 549: 548: 542: 541: 535: 534: 527: 526: 522: 521: 517: 516: 510: 508: 506: 505: 500: 499: 494: 490: 486: 482: 478: 474: 470: 462: 460: 458: 454: 451: 447: 443: 439: 436: 432: 428: 424: 416: 414: 412: 408: 403: 401: 400: 394: 392: 388: 384: 380: 372: 370: 368: 364: 360: 355: 353: 349: 341: 339: 333: 331: 329: 325: 321: 317: 313: 309: 305: 301: 297: 296: 291: 287: 283: 279: 275: 271: 268: 265: 261: 258: 255:There are 98 250: 248: 246: 242: 238: 234: 230: 227:in the order 226: 222: 218: 211: 210: 205: 195: 192: 188: 185: 183: 179: 171: 168: 165: 162: 159: 156: 153: 150: 147: 146: 144: 139: 134: 129: 126: 123: 122: 119: 116: 114:Superfamily: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 89: 86: 83: 82: 79: 76: 73: 72: 69: 66: 63: 62: 57: 52: 48: 44: 40: 36: 32: 27: 22: 19: 3636: 3563:Millieriidae 3544:Geometridae 3513:Geometroidea 3497:(gold moths) 3455:(tuft moths) 3405:(saturniids) 3403:Saturniidae 3393:Eupterotidae 3377:Carthaeidae 3369:Brahmaeidae 3363:(silk moths) 3284:(grass moth) 3255:(teak moths) 3232:Nymphalidae 3218:(metalmarks) 3200:Hesperiidae 2933:Tortricidae 2925:Tortricoidea 2912:Choreutidae 2904:Choreutoidea 2875:Brachodidae 2825:Limacodidae 2799:Anomoeotidae 2740:Galacticidae 2664:Gelechiidae 2617:Gelechioidea 2607:Ypsolophidae 2584:Heliodinidae 2539: 2362:Opostegidae 2302:Heliozelidae 2154:Anomosetidae 2098:Neopseustina 1848:Holometabola 1846:Superorder: 1840:Infraclass: 1756: 1739: 1690:Tree of Life 1676: 1672: 1644:(1): 60–81. 1641: 1637: 1627: 1568: 1562: 1552: 1538: 1485: 1475: 1442: 1438: 1390:(1): 60–81. 1387: 1383: 1356:. 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Retrieved 1114: 1102: 1085: 1079: 1076:Hering, 1957 1072: 1064: 1057: 1049: 1042: 1037:Strand, 1919 1033: 1022: 1016: 1010: 1003: 995: 988: 979: 972: 965: 957: 951: 945: 942:Hering, 1924 938: 934:Walker, 1864 928: 924:Walker, 1864 918: 910: 903: 898: 832: 824: 786: 744: 729: 707: 649: 634: 612: 581: 566: 502: 496: 489:Baltic amber 466: 450:evolutionary 437: 435:sister genus 430: 420: 404: 397: 395: 376: 373:Life history 356: 352:caterpillars 345: 337: 334:Distribution 327: 323: 319: 315: 311: 308:phylogenetic 300:DNA sequence 293: 285: 264:Afrotropical 254: 244: 239:such as the 216: 215: 207: 177: 175: 141:Subfamilies 127: 42: 18: 3962:Leaf miners 3813:iNaturalist 3661:Wikispecies 3521:Sematuridae 3495:Cimeliidae 3489:(hook-tips) 3487:Drepanidae 3479:Drepanoidea 3411:Sphingidae 3398:Phiditiidae 3385:Endromidae 3361:Bombycidae 3345:Anthelidae 3337:Bombycoidea 3253:Hyblaeidae 3245:Hyblaeoidea 3224:Lycaenidae 3216:Riodinidae 3173:butterflies 3132:Thyrididae 3124:Thyridoidea 3025:Tineodidae 3017:Alucitidae 3009:Alucitoidea 2883:Castniidae 2817:Dalceridae 2763:Zygaenidae 2750:Zygaenoidea 2594:Plutellidae 2589:Lyonetiidae 2571:Bedelliidae 2312:Prodoxidae 2297:Cecidosidae 2159:Hepialidae 2146:Hepialoidea 1864:Zeugloptera 1805:Lepidoptera 1209:10088/20928 1115:mapress.com 1087:nomen nudum 1012:Sorbus aria 997:nomen nudum 881:Ornixolinae 835:subfamilies 788:Ornixolinae 446:Palaearctic 440:. 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Index

Privet leaf miner, "Gracillaria syringella" (Fabricius, 1794)
Privet leaf miner
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Lepidoptera
Gracillarioidea
Gracillariidae
Stainton
Gracillariinae
Stainton
Lithocolletinae
Stainton
Phyllocnistinae
Herrich-Schäffer
Subfamilies and genera
Diversity

Phyllocnistis magnoliella
family
insects
Lepidoptera
leaf miners
species
horse-chestnut leaf miner
described
genera

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