244:
308:
61:
1535:
1076:
1664:
1069:
1063:
1529:
40:
1783:
255:
had a very wide skull due to a widened temporal region. Viewed from above, it looks roughly triangular. The skull is about 20 centimetres (7.9 in) long, with nearly half its length in the postorbital region behind the eye sockets. It has large
260:
and smaller buccal, or cheek teeth, along the thick upper and lower jaws. The lower jaw is robust and curved upward, with a well-developed symphyseal region where the two halves of the jaw meet.
1729:
243:
1824:
1716:
385:
278:
by
Ivakhnenko 2011 as they shared a wide temporal region, Liu and Abdala 2020 found little relation between the two genera and instead classifies
1858:
1848:
1843:
307:
1863:
1534:
1817:
378:
1853:
60:
1868:
1075:
1068:
1810:
1684:
1667:
1062:
371:
225:
1755:
221:
1177:
1528:
1205:
168:
832:
1261:
994:
733:
1630:
1595:
899:
860:
1602:
1408:
912:
675:
668:
654:
640:
476:
349:
55:
1760:
618:
1734:
332:
Ivakhnenko, M.F. (2011). "Permian and
Triassic therocephals (Eutherapsida) of Eastern Europe".
1742:
1505:
874:
1747:
1581:
1433:
1184:
341:
1268:
1226:
1156:
1121:
825:
809:
782:
712:
689:
632:
609:
563:
1794:
1498:
1491:
1457:
1401:
1247:
1001:
888:
881:
867:
748:
625:
445:
275:
147:
1837:
1567:
1442:
1422:
1254:
1240:
1163:
1149:
1008:
980:
932:
818:
791:
762:
696:
538:
525:
488:
394:
353:
311:
206:
134:
1644:
1637:
1616:
1609:
1554:
1477:
1449:
1381:
1354:
1343:
1329:
1320:
1275:
1170:
1128:
1015:
987:
923:
853:
839:
598:
582:
257:
229:
213:
176:
30:
1588:
1574:
1293:
1282:
1219:
1212:
1142:
964:
953:
946:
798:
741:
682:
661:
647:
270:
1707:
1415:
1363:
1233:
1191:
1107:
1094:
1030:
973:
770:
591:
570:
556:
436:
345:
287:
118:
39:
1790:
1623:
1468:
1336:
1302:
1135:
939:
846:
461:
427:
283:
209:
105:
72:
1782:
274:
were considered closely related and classified as the only two members of
1701:
1394:
1370:
726:
418:
300:
295:
92:
1721:
1309:
1484:
1198:
412:
217:
82:
1678:
306:
242:
363:
1682:
1551:
1092:
523:
405:
367:
16:
Extinct genus of therapsids from the Late
Permian of Russia
1798:
183:
1691:
1467:
1432:
1380:
1353:
1319:
1292:
1105:
1029:
963:
922:
898:
808:
781:
710:
608:
581:
536:
487:
460:
1818:
379:
8:
314:major consensus tree (Liu and Abdala, 2020)
1825:
1811:
1679:
1548:
1386:
1113:
1102:
1089:
904:
718:
548:
544:
533:
520:
466:
457:
402:
386:
372:
364:
38:
20:
251:In comparison to other therocephalians,
324:
7:
1779:
1777:
220:. Fossils have been found from the
205:is an extinct genus of herbivorous
268:Although it and the South African
14:
1781:
1663:
1662:
1533:
1527:
1074:
1067:
1061:
59:
1:
1859:Lopingian synapsids of Europe
1849:Fossil taxa described in 2011
1797:. You can help Knowledge by
1885:
1776:
1844:Extinct animals of Russia
1658:
1562:
1547:
1525:
1389:
1116:
1101:
1088:
1059:
907:
721:
551:
547:
532:
519:
469:
456:
401:
346:10.1134/S0031030111090012
182:
175:
56:Scientific classification
54:
46:
37:
23:
334:Paleontological Journal
1793:-related article is a
315:
248:
226:Nizhny Novgorod Oblast
1756:Paleobiology Database
310:
294:forms an exclusively
246:
240:, was named in 2011.
222:Tonshayevsky District
1864:Therocephalia genera
304:from China instead.
316:
249:
1854:Eutherocephalians
1806:
1805:
1771:
1770:
1743:Open Tree of Life
1685:Taxon identifiers
1676:
1675:
1654:
1653:
1543:
1542:
1523:
1522:
1519:
1518:
1515:
1514:
1506:Traversodontoides
1178:Jiucaiyuangnathus
1084:
1083:
1057:
1056:
1053:
1052:
1049:
1048:
1025:
1024:
875:Promoschorhynchus
706:
705:
515:
514:
511:
510:
507:
506:
198:
197:
171:
1876:
1827:
1820:
1813:
1785:
1778:
1764:
1763:
1751:
1750:
1738:
1737:
1725:
1724:
1712:
1711:
1710:
1680:
1666:
1665:
1582:Blattoidealestes
1549:
1538:
1537:
1531:
1434:Ericiolacertidae
1387:
1185:Macroscelesaurus
1114:
1103:
1090:
1079:
1078:
1071:
1065:
905:
719:
549:
545:
534:
521:
467:
458:
451:
450:
403:
388:
381:
374:
365:
358:
357:
329:
193:Ivakhnenko, 2011
185:
166:
159:
146:
133:
64:
63:
42:
29:Temporal range:
21:
1884:
1883:
1879:
1878:
1877:
1875:
1874:
1873:
1869:Therapsid stubs
1834:
1833:
1832:
1831:
1774:
1772:
1767:
1759:
1754:
1746:
1741:
1733:
1728:
1720:
1715:
1706:
1705:
1700:
1687:
1677:
1672:
1650:
1558:
1539:
1532:
1511:
1463:
1428:
1376:
1349:
1315:
1288:
1269:Silphictidoides
1227:Scalopocephalus
1206:Nanictocephalus
1157:Ictidosuchoides
1122:Homodontosaurus
1110:
1097:
1080:
1073:
1045:
1021:
959:
918:
894:
826:Annatherapsidus
810:Akidnognathidae
804:
783:Chthonosauridae
777:
715:
713:Eutherocephalia
702:
690:Pristerognathus
633:Biarmosuchoides
610:Scylacosauridae
604:
577:
564:Porosteognathus
541:
528:
503:
483:
452:
408:
407:
397:
392:
362:
361:
340:(9): 981–1144.
331:
330:
326:
321:
266:
194:
191:
165:
157:
144:
131:
58:
49:Purlovia maxima
47:Restoration of
33:
27:
17:
12:
11:
5:
1882:
1880:
1872:
1871:
1866:
1861:
1856:
1851:
1846:
1836:
1835:
1830:
1829:
1822:
1815:
1807:
1804:
1803:
1786:
1769:
1768:
1766:
1765:
1752:
1739:
1726:
1713:
1697:
1695:
1689:
1688:
1683:
1674:
1673:
1671:
1670:
1659:
1656:
1655:
1652:
1651:
1649:
1648:
1641:
1634:
1627:
1620:
1613:
1606:
1599:
1592:
1585:
1578:
1571:
1563:
1560:
1559:
1552:
1545:
1544:
1541:
1540:
1526:
1524:
1521:
1520:
1517:
1516:
1513:
1512:
1510:
1509:
1502:
1499:Scalopognathus
1495:
1492:Microgomphodon
1488:
1481:
1473:
1471:
1465:
1464:
1462:
1461:
1458:Silphedosuchus
1454:
1446:
1438:
1436:
1430:
1429:
1427:
1426:
1419:
1412:
1405:
1402:Nothogomphodon
1398:
1390:
1384:
1378:
1377:
1375:
1374:
1367:
1359:
1357:
1351:
1350:
1348:
1347:
1340:
1333:
1325:
1323:
1317:
1316:
1314:
1313:
1306:
1298:
1296:
1290:
1289:
1287:
1286:
1279:
1272:
1265:
1258:
1251:
1248:Scalopolacerta
1244:
1237:
1230:
1223:
1216:
1209:
1202:
1195:
1188:
1181:
1174:
1167:
1160:
1153:
1146:
1139:
1132:
1125:
1117:
1111:
1106:
1099:
1098:
1093:
1086:
1085:
1082:
1081:
1060:
1058:
1055:
1054:
1051:
1050:
1047:
1046:
1044:
1043:
1042:
1041:
1035:
1033:
1027:
1026:
1023:
1022:
1020:
1019:
1012:
1005:
1002:Moschowhaitsia
998:
991:
984:
977:
969:
967:
961:
960:
958:
957:
950:
943:
936:
928:
926:
920:
919:
917:
916:
908:
902:
896:
895:
893:
892:
889:Zorillodontops
885:
882:Shiguaignathus
878:
871:
868:Olivierosuchus
864:
857:
850:
843:
836:
833:Cerdosuchoides
829:
822:
814:
812:
806:
805:
803:
802:
795:
787:
785:
779:
778:
776:
775:
767:
759:
752:
749:Perplexisaurus
745:
738:
730:
722:
716:
711:
708:
707:
704:
703:
701:
700:
693:
686:
679:
672:
665:
658:
651:
644:
637:
629:
626:Alopecognathus
622:
614:
612:
606:
605:
603:
602:
595:
587:
585:
579:
578:
576:
575:
567:
560:
552:
542:
537:
530:
529:
524:
517:
516:
513:
512:
509:
508:
505:
504:
502:
501:
500:
499:
493:
491:
485:
484:
482:
481:
480:
479:
470:
464:
454:
453:
449:
448:
446:Eutheriodontia
439:
430:
421:
415:
406:
399:
398:
393:
391:
390:
383:
376:
368:
360:
359:
323:
322:
320:
317:
276:Nanictidopidae
265:
264:Classification
262:
247:Holotype skull
207:therocephalian
196:
195:
192:
180:
179:
173:
172:
155:
151:
150:
148:Nanictidopidae
142:
138:
137:
129:
122:
121:
116:
109:
108:
103:
96:
95:
90:
86:
85:
80:
76:
75:
70:
66:
65:
52:
51:
44:
43:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1881:
1870:
1867:
1865:
1862:
1860:
1857:
1855:
1852:
1850:
1847:
1845:
1842:
1841:
1839:
1828:
1823:
1821:
1816:
1814:
1809:
1808:
1802:
1800:
1796:
1792:
1787:
1784:
1780:
1775:
1762:
1757:
1753:
1749:
1744:
1740:
1736:
1731:
1727:
1723:
1718:
1714:
1709:
1703:
1699:
1698:
1696:
1694:
1690:
1686:
1681:
1669:
1661:
1660:
1657:
1647:
1646:
1642:
1640:
1639:
1635:
1633:
1632:
1628:
1626:
1625:
1621:
1619:
1618:
1614:
1612:
1611:
1607:
1605:
1604:
1600:
1598:
1597:
1593:
1591:
1590:
1586:
1584:
1583:
1579:
1577:
1576:
1572:
1570:
1569:
1568:Alopecorhinus
1565:
1564:
1561:
1557:
1556:
1550:
1546:
1536:
1530:
1508:
1507:
1503:
1501:
1500:
1496:
1494:
1493:
1489:
1487:
1486:
1482:
1480:
1479:
1475:
1474:
1472:
1470:
1466:
1460:
1459:
1455:
1452:
1451:
1447:
1445:
1444:
1443:Ericiolacerta
1440:
1439:
1437:
1435:
1431:
1425:
1424:
1423:Scaloposaurus
1420:
1418:
1417:
1413:
1411:
1410:
1406:
1404:
1403:
1399:
1397:
1396:
1392:
1391:
1388:
1385:
1383:
1379:
1373:
1372:
1368:
1366:
1365:
1361:
1360:
1358:
1356:
1352:
1346:
1345:
1341:
1339:
1338:
1334:
1332:
1331:
1327:
1326:
1324:
1322:
1318:
1312:
1311:
1307:
1305:
1304:
1300:
1299:
1297:
1295:
1291:
1285:
1284:
1280:
1278:
1277:
1273:
1271:
1270:
1266:
1264:
1263:
1262:Scaloposuchus
1259:
1257:
1256:
1255:Scaloporhinus
1252:
1250:
1249:
1245:
1243:
1242:
1241:Scalopodontes
1238:
1236:
1235:
1231:
1229:
1228:
1224:
1222:
1221:
1217:
1215:
1214:
1210:
1208:
1207:
1203:
1201:
1200:
1196:
1194:
1193:
1189:
1187:
1186:
1182:
1180:
1179:
1175:
1173:
1172:
1168:
1166:
1165:
1164:Ictidosuchops
1161:
1159:
1158:
1154:
1152:
1151:
1150:Ictidognathus
1147:
1145:
1144:
1140:
1138:
1137:
1133:
1131:
1130:
1126:
1124:
1123:
1119:
1118:
1115:
1112:
1109:
1104:
1100:
1096:
1091:
1087:
1077:
1072:
1070:
1064:
1039:
1038:
1037:
1036:
1034:
1032:
1028:
1018:
1017:
1013:
1011:
1010:
1009:Theriognathus
1006:
1004:
1003:
999:
997:
996:
995:Microwhaitsia
992:
990:
989:
985:
983:
982:
981:Ictidochampsa
978:
976:
975:
971:
970:
968:
966:
962:
956:
955:
951:
949:
948:
944:
942:
941:
937:
935:
934:
933:Gorochovetzia
930:
929:
927:
925:
921:
915:
914:
910:
909:
906:
903:
901:
897:
891:
890:
886:
884:
883:
879:
877:
876:
872:
870:
869:
865:
863:
862:
858:
856:
855:
851:
849:
848:
844:
842:
841:
837:
835:
834:
830:
828:
827:
823:
821:
820:
819:Akidnognathus
816:
815:
813:
811:
807:
801:
800:
796:
794:
793:
792:Chthonosaurus
789:
788:
786:
784:
780:
773:
772:
768:
765:
764:
763:Scylacosuchus
760:
758:
757:
753:
751:
750:
746:
744:
743:
739:
736:
735:
734:Kotelcephalon
731:
729:
728:
724:
723:
720:
717:
714:
709:
699:
698:
697:Scylacosaurus
694:
692:
691:
687:
685:
684:
680:
678:
677:
673:
671:
670:
666:
664:
663:
659:
657:
656:
652:
650:
649:
645:
643:
642:
638:
635:
634:
630:
628:
627:
623:
621:
620:
616:
615:
613:
611:
607:
601:
600:
596:
594:
593:
589:
588:
586:
584:
580:
573:
572:
568:
566:
565:
561:
559:
558:
554:
553:
550:
546:
543:
540:
539:Therocephalia
535:
531:
527:
526:Therocephalia
522:
518:
497:
496:
495:
494:
492:
490:
489:Therocephalia
486:
478:
474:
473:
472:
471:
468:
465:
463:
459:
455:
447:
443:
440:
438:
434:
431:
429:
425:
422:
420:
416:
414:
410:
409:
404:
400:
396:
395:Therocephalia
389:
384:
382:
377:
375:
370:
369:
366:
355:
351:
347:
343:
339:
335:
328:
325:
318:
313:
312:Therocephalia
309:
305:
303:
302:
297:
293:
289:
285:
281:
277:
273:
272:
263:
261:
259:
254:
245:
241:
239:
235:
231:
227:
223:
219:
215:
211:
208:
204:
203:
190:
189:
181:
178:
174:
170:
164:
163:
156:
153:
152:
149:
143:
140:
139:
136:
135:Therocephalia
130:
127:
124:
123:
120:
117:
114:
111:
110:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
62:
57:
53:
50:
45:
41:
36:
32:
26:
22:
19:
1799:expanding it
1788:
1773:
1692:
1645:Trochosuchus
1643:
1638:Trochosaurus
1636:
1631:Trochorhinus
1629:
1622:
1617:Scymnosaurus
1615:
1610:Ictidosaurus
1608:
1601:
1596:Hyaenasuchus
1594:
1587:
1580:
1573:
1566:
1555:Nomina dubia
1553:
1504:
1497:
1490:
1483:
1478:Antecosuchus
1476:
1456:
1450:Pedaeosaurus
1448:
1441:
1421:
1414:
1407:
1400:
1393:
1382:Bauriamorpha
1369:
1362:
1355:Regisauridae
1344:Tetracynodon
1342:
1335:
1330:Choerosaurus
1328:
1321:Lycideopidae
1308:
1301:
1281:
1276:Silpholestes
1274:
1267:
1260:
1253:
1246:
1239:
1232:
1225:
1218:
1211:
1204:
1197:
1190:
1183:
1176:
1171:Ictidosuchus
1169:
1162:
1155:
1148:
1141:
1134:
1129:Icticephalus
1127:
1120:
1066:
1016:Viatkosuchus
1014:
1007:
1000:
993:
988:Megawhaitsia
986:
979:
972:
952:
945:
938:
931:
924:Hofmeyriidae
911:
900:Whaitsioidea
887:
880:
873:
866:
861:Notaelurodon
859:
854:Moschorhinus
852:
845:
840:Euchambersia
838:
831:
824:
817:
797:
790:
769:
761:
755:
754:
747:
740:
732:
725:
695:
688:
681:
674:
667:
660:
653:
646:
639:
631:
624:
617:
599:Simorhinella
597:
590:
583:Lycosuchidae
569:
562:
555:
441:
432:
423:
337:
333:
327:
299:
291:
279:
269:
267:
258:canine teeth
252:
250:
237:
233:
230:type species
214:Late Permian
201:
200:
199:
187:
186:
177:Type species
167:Ivakhnenko,
161:
160:
125:
112:
99:
48:
31:Late Permian
24:
18:
1603:Ictidoparia
1589:Dongusaurus
1575:Arnognathus
1409:Notictoides
1294:Karenitidae
1283:Yikezhaogia
1220:Rhigosaurus
1213:Polycynodon
1143:Ictidodraco
965:Whaitsiidae
954:Mirotenthes
947:Ictidostoma
913:Ophidostoma
799:Ichibengops
742:Nanictidops
683:Pardosuchus
676:Maraisaurus
669:Koksharovia
662:Julognathus
655:Hyorhynchus
648:Glanosuchus
641:Eutheriodon
298:clade with
280:Nanictidops
271:Nanictidops
1838:Categories
1416:Ordosiodon
1364:Regisaurus
1234:Scalopodon
1192:Malasaurus
1108:Baurioidea
1095:Baurioidea
1040:see below↓
1031:Baurioidea
974:Dalongkoua
771:Cynodontia
619:Alopecodon
592:Lycosuchus
571:Cynodontia
557:Gorynychus
498:see below↓
437:Therapsida
319:References
288:Baurioidea
286:member of
210:therapsids
119:Therapsida
1791:therapsid
1624:Tamboeria
1469:Bauriidae
1337:Lycideops
1303:Karenites
1136:Ictidodon
940:Hofmeyria
847:Jiufengia
477:Synapsida
462:Synapsida
428:Synapsida
411:Kingdom:
354:128958135
296:Laurasian
284:basalmost
238:P. maxima
212:from the
188:P. maxima
106:Synapsida
79:Kingdom:
73:Eukaryota
1735:11913268
1708:Q4383895
1702:Wikidata
1693:Purlovia
1668:Category
1395:Hazhenia
1371:Urumchia
756:Purlovia
727:Caodeyao
419:Chordata
417:Phylum:
413:Animalia
301:Caodeyao
292:Purlovia
290:, while
253:Purlovia
234:Purlovia
202:Purlovia
162:Purlovia
141:Family:
93:Chordata
89:Phylum:
83:Animalia
69:Domain:
25:Purlovia
1748:7667721
1722:9315845
1310:Mupashi
282:as the
154:Genus:
1761:362040
1485:Bauria
1199:Muchia
352:
228:. The
218:Russia
1789:This
1730:IRMNG
442:Clade
433:Clade
424:Clade
350:S2CID
126:Clade
113:Clade
100:Clade
1795:stub
1717:GBIF
475:see
169:2011
342:doi
232:of
224:of
216:of
1840::
1758::
1745::
1732::
1719::
1704::
444::
435::
426::
348:.
338:45
336:.
236:,
128::
115::
102::
1826:e
1819:t
1812:v
1801:.
1453:?
774:?
766:?
737:?
636:?
574:?
387:e
380:t
373:v
356:.
344::
184:†
158:†
145:†
132:†
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