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Plant density

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65: 161: 1830: 100: 223: 189:(LAI, leaf area per ground area). The higher the Leaf Area Index, the higher the fraction of intercepted sunlight will be, but the gain in light interception and photosynthesis will not match the increase in LAI, and this is the reason that total biomass per ground area saturates at high plant densities. 56:
to die off, in a process that has been named 'self-thinning'. The remaining plants perform better as fewer plants will now compete for resources. A key factor in agronomy and forestry is plant population density, which provides an experimental approach for better understanding plant-plant competition.
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for light, nutrients and water, individual plants will not be able to take up all resources that are required for optimal growth. This indicates that plant density not only depends on the space available to grow but it is also determined by the amount of resources available. Especially in the case of
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of a monostand increases with increasing density, up to the point where the biomass saturates. This is what has been dubbed 'constant final yield', and refers to the total plant biomass per unit ground area. Seed production per ground area is not constant, but often declines with density after total
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than bigger plants, even less than would be expected on the basis of their size differences. As plant density increases it will affect the structure of the plant as well as the developmental patterns of the plant. This is called 'asymmetric competition'  and will cause some subordinate plants
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Contrary to the total biomass per unit ground area, which increases with density until reaching saturation, the average biomass of individual plants in a monostand strongly declines with plant density, such that for every doubling in density individual plants will become ~30-40% smaller. Plants in
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Experiments with herbaceous plants have been carried out with extremely high densities (up to 80,000 plants per square meter). At such high densities, these plants will start to compete soon after germination, and eventually a large number of those individuals (up to 95%) will die. In agriculture,
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is the number of individual plants present per unit of ground area. It is most easily interpreted in the case of monospecific stands, where all plants belong to the same species and have germinated at the same time. However, it could also indicate the number of individual plants found at a given
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Root growth in environments with high plant density show that there will be fewer roots per plant and but the length and general density of the individual root remain somewhat the same, this is expected to still cause issues for the plant in future growth.
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of even-aged individuals that are sown or planted at the same time. These can be referred to as 'monostands' and are often studied in the context of agricultural, horticultural or silvicultural questions. However, they are also highly relevant in
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Average plant height or vegetation height often remains remarkably similar, but a very consistent difference is that the stems of high-density plants have a much smaller diameter. They also have fewer side shoots
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farmers avoid these very high densities as they do not contribute to seed yield. Normal densities in modern agriculture depend on final plant size and vary between 5-10 plants per square meter for
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Effect of plant density on (a) total shoot mass and (b) seed mass per unit ground area. Schematised figure, inspired a.o. by experiments with maize by Li et al. (2015).
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Because densely-grown plants are smaller, they will also produce fewer seeds per individual. But also the seed production as a fraction of total plant biomass (
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Plant density is defined as the number of plants present per unit area of ground. In nature, plant densities can be especially high when seeds present in a
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Li J, Xie RZ, Wang KR, Ming B, Guo YQ, Zhang GQ, Li SK (2015). "Variations in Maize dry matter, harvest index, and grain yield with plant density".
331:"Consequences of plant population size and density for plant-pollinator interactions and plant performance: Plant-pollinator interactions" 272:
In dense stands, there is a strong gradient of light from top to bottom. Lower leaves in high-density stands will therefore have a lower
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than similar leaves of plants in open stands. There are indications that also the well-illuminated top leaves may have a lower
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Leaf size of the largest full-grown leaf of Maize plants grown at a low (L), intermediate (I), and high (H) plant density.
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Postma, Johannes A.; Hecht, Vera L.; Hikosaka, Kouki; Nord, Eric A.; Pons, Thijs L.; Poorter, Hendrik (2021).
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and they are often smaller and more narrow (see photo). Leaves of high-density plants are thinner (higher
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germinate after winter, or in a forest understory after a tree fall opens a gap in the canopy. Due to
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higher density stands invest relatively more of their biomass in stems (higher
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Postma JA, Hecht VL, Hikosaka K, Nord EA, Pons TL, Poorter H (December 2020).
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Mustajärvi, Kaisa; Siikamäki, Pirkko; Rytkönen, Saara; Lammi, Antti (2001).
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Effect of a low (L), intermediate (I) and high (H) plant density on Maize.
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Many of the processes related to plant density can well be studied in
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Number of individual plants present per unit of ground area
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Weiner J, Freckleton RP (2010). "Constant final yield".
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in many other ways and at different integration levels:
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Annual Review of Ecology, Evolution, and Systematics
1772: 1736: 1626: 1499: 1415: 1311: 1261: 986: 916: 859: 821: 795: 731: 651: 86:biomass per ground area reached its maximum value. 210:Apart from their weight, plants will change their 611: 230:Individual plants in dense stands have fewer 8: 1717:International Association for Plant Taxonomy 553: 551: 128:. Unsourced material may be challenged and 818: 618: 604: 596: 575: 438: 397: 387: 346: 173:till 200-300 plants per square meter for 148:Learn how and when to remove this message 51:light, smaller plants will take up fewer 1661:International Code of Nomenclature (ICN) 362:Dhaliwal DS, Williams MM (2020-02-07). 321: 507:10.1146/annurev-ecolsys-102209-144642 81:. In general, the total above-ground 7: 126:adding citations to reliable sources 193:The individual plant in a monostand 25: 1666:ICN for Cultivated Plants (ICNCP) 207:), and less in leaves and roots. 1829: 1828: 348:10.1046/j.1365-2745.2001.00521.x 255:in the case of herbs and trees. 98: 90:Plant density and self-thinning 1: 564:Plant, Cell & Environment 427:Plant, Cell & Environment 251:) in the case of grasses, or 1707:History of plant systematics 1294:Thorns, spines, and prickles 480:Population biology of plants 389:10.1371/journal.pone.0228809 1881: 1523:Alternation of generations 1824: 1673:Cultivated plant taxonomy 1636:Biological classification 633: 482:. London: Academic Press. 310:Intraspecific competition 284:in densely-grown plants. 1533:Evolutionary development 1184:Hypanthium (Floral cup) 282:photosynthetic capacity 38:Definition and concepts 1799:by author abbreviation 1723:Plant taxonomy systems 1641:Botanical nomenclature 227: 165: 69: 1806:Botanical expeditions 542:10.2134/agronj14.0522 225: 163: 67: 1538:Evolutionary history 1528:Double fertilization 1380:Cellular respiration 122:improve this section 757:Non-vascular plants 534:2015AgrJ..107..829L 380:2020PLoSO..1528809D 274:photosynthetic rate 1262:Surface structures 1057:Flower development 478:Harper JL (1977). 335:Journal of Ecology 278:transpiration rate 228: 205:Stem Mass Fraction 166: 70: 1842: 1841: 1481:Herbaceous plants 1307: 1306: 577:10.1111/pce.13968 440:10.1111/pce.13968 158: 157: 150: 16:(Redirected from 1872: 1832: 1831: 1811:Individual trees 1486:Secondary growth 1457:Succulent plants 1445:Prostrate shrubs 1328:Apical dominance 1313:Plant physiology 1274:Epicuticular wax 819: 812: 803:Plant morphology 620: 613: 606: 597: 590: 589: 579: 570:(4): 1072–1094. 555: 546: 545: 522:Agronomy Journal 517: 511: 510: 490: 484: 483: 475: 469: 468: 442: 433:(4): 1072–1094. 418: 412: 411: 401: 391: 359: 353: 352: 350: 326: 153: 146: 142: 139: 133: 102: 94: 21: 18:Planting density 1880: 1879: 1875: 1874: 1873: 1871: 1870: 1869: 1845: 1844: 1843: 1838: 1820: 1789:Botanical terms 1782: 1768: 1732: 1678:Citrus taxonomy 1656:Author citation 1622: 1516: 1495: 1417: 1411: 1407:Turgor pressure 1315: 1303: 1257: 1072:Floral symmetry 990: 982: 912: 901:Vascular bundle 896:Vascular tissue 855: 815: 806: 805: 791: 762:Vascular plants 727: 723:Plant pathology 647: 629: 624: 594: 593: 557: 556: 549: 519: 518: 514: 492: 491: 487: 477: 476: 472: 420: 419: 415: 374:(2): e0228809. 361: 360: 356: 328: 327: 323: 318: 306: 290: 288:Seed production 270: 261: 244: 220: 200: 195: 187:Leaf Area Index 154: 143: 137: 134: 119: 103: 92: 62: 40: 28: 23: 22: 15: 12: 11: 5: 1878: 1876: 1868: 1867: 1862: 1857: 1847: 1846: 1840: 1839: 1837: 1836: 1825: 1822: 1821: 1819: 1818: 1813: 1808: 1803: 1802: 1801: 1791: 1785: 1783: 1781: 1780: 1779:Related topics 1777: 1773: 1770: 1769: 1767: 1766: 1761: 1756: 1751: 1746: 1740: 1738: 1734: 1733: 1731: 1730: 1728:Taxonomic rank 1725: 1720: 1714: 1709: 1704: 1703: 1702: 1701: 1700: 1695: 1690: 1680: 1670: 1669: 1668: 1663: 1658: 1653: 1648: 1646:Botanical name 1638: 1632: 1630: 1628:Plant taxonomy 1624: 1623: 1621: 1620: 1619: 1618: 1613: 1612: 1611: 1604:Megasporangium 1601: 1600: 1599: 1592:Microsporangia 1584: 1583: 1582: 1577: 1572: 1567: 1557: 1552: 1547: 1546: 1545: 1535: 1530: 1525: 1519: 1517: 1515: 1514: 1509: 1503: 1497: 1496: 1494: 1493: 1488: 1483: 1478: 1477: 1476: 1475: 1474: 1464: 1459: 1454: 1453: 1452: 1447: 1437: 1432: 1430:Cushion plants 1421: 1419: 1413: 1412: 1410: 1409: 1404: 1399: 1394: 1389: 1384: 1383: 1382: 1377: 1367: 1365:Plant hormones 1362: 1357: 1356: 1355: 1348:Photosynthesis 1345: 1340: 1335: 1330: 1325: 1319: 1317: 1309: 1308: 1305: 1304: 1302: 1301: 1296: 1291: 1286: 1281: 1276: 1271: 1265: 1263: 1259: 1258: 1256: 1255: 1250: 1245: 1240: 1235: 1234: 1233: 1228: 1223: 1213: 1212: 1211: 1206: 1201: 1196: 1186: 1181: 1180: 1179: 1178: 1177: 1172: 1167: 1166: 1165: 1160: 1140: 1135: 1130: 1129: 1128: 1127: 1126: 1121: 1111: 1106: 1101: 1096: 1091: 1081: 1080: 1079: 1074: 1069: 1067:Floral formula 1064: 1062:Floral diagram 1059: 1054: 1044: 1043: 1042: 1037: 1032: 1031: 1030: 1025: 1015: 1005: 1000: 994: 992: 991:(incl. Flower) 984: 983: 981: 980: 979: 978: 973: 968: 967: 966: 961: 951: 941: 936: 931: 926: 920: 918: 914: 913: 911: 910: 905: 904: 903: 893: 891:Storage organs 888: 883: 882: 881: 871: 865: 863: 857: 856: 854: 853: 848: 843: 838: 833: 827: 825: 816: 814: 813: 799: 793: 792: 790: 789: 784: 779: 777:Spermatophytes 774: 769: 764: 759: 754: 749: 747:Archaeplastida 744: 738: 736: 729: 728: 726: 725: 720: 715: 710: 709: 708: 701:Phytogeography 698: 696:Phytochemistry 693: 688: 683: 678: 673: 668: 663: 657: 655: 653:Subdisciplines 649: 648: 646: 645: 640: 634: 631: 630: 625: 623: 622: 615: 608: 600: 592: 591: 547: 528:(3): 829–834. 512: 485: 470: 413: 354: 320: 319: 317: 314: 313: 312: 305: 302: 289: 286: 269: 266: 260: 257: 243: 240: 219: 216: 199: 196: 194: 191: 156: 155: 106: 104: 97: 91: 88: 61: 58: 39: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 1877: 1866: 1863: 1861: 1860:Plant ecology 1858: 1856: 1853: 1852: 1850: 1835: 1827: 1826: 1823: 1817: 1814: 1812: 1809: 1807: 1804: 1800: 1797: 1796: 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1003:Antheridium 998:Archegonium 846:Plasmodesma 823:Plant cells 686:Paleobotany 681:Ethnobotany 666:Astrobotany 501:: 173–192. 48:competition 1849:Categories 1597:Microspore 1587:Sporangium 1565:Artificial 1253:Sporophyte 1248:Sporophyll 1243:Receptacle 1138:Gynandrium 1008:Androecium 917:Vegetative 787:Angiosperm 782:Gymnosperm 676:Dendrology 316:References 268:Physiology 138:March 2022 60:Monostands 34:location. 1794:Botanists 1712:Herbarium 1609:Megaspore 1507:Evolution 1450:Subshrubs 1418:and habit 1343:Nutrition 1338:Cellulose 1333:Bulk flow 1316:Materials 1279:Epidermis 1143:Gynoecium 1124:Endosperm 1119:Dispersal 1035:Staminode 971:Sessility 959:Cataphyll 879:Mesophyll 831:Cell wall 772:Lycophyte 752:Bryophyte 706:Geobotany 691:Phycology 465:227523495 449:0140-7791 212:phenotype 109:does not 53:resources 44:seed bank 1834:Category 1754:Forestry 1744:Agronomy 1737:Practice 1688:Cultivar 1683:Cultigen 1543:timeline 1435:Rosettes 1323:Aleurone 1299:Trichome 1216:Perianth 1028:Filament 886:Meristem 809:glossary 671:Bryology 586:33280135 457:33280135 408:32032371 368:PLOS ONE 304:See also 253:branches 183:forestry 1512:Ecology 1269:Cuticle 1099:Capsule 1089:Anatomy 1040:Tapetum 964:Petiole 939:Rhizome 934:Rhizoid 861:Tissues 851:Vacuole 841:Plastid 643:Outline 638:History 530:Bibcode 399:7006923 376:Bibcode 249:tillers 198:Biomass 130:removed 115:sources 83:biomass 79:ecology 1865:Botany 1816:Plants 1719:(IAPT) 1472:Lianas 1440:Shrubs 1392:Starch 1284:Nectar 1204:Raceme 1170:Stigma 1158:Locule 1148:Carpel 1109:Pyrena 1047:Flower 1023:Anther 1018:Stamen 1013:Pollen 735:groups 627:Botany 584:  463:  455:  447:  406:  396:  232:leaves 218:Leaves 179:Barley 1776:Lists 1693:Group 1616:Spore 1550:Flora 1467:Vines 1462:Trees 1425:Habit 1397:Sugar 1289:Stoma 1231:Sepal 1226:Petal 1221:Tepal 1209:Umbel 1194:Bract 1175:Style 1163:Ovule 1153:Ovary 1094:Berry 1084:Fruit 1077:Whorl 944:Shoot 742:Algae 733:Plant 461:S2CID 259:Roots 242:Stems 181:. In 171:Maize 1698:Grex 1580:Self 1114:Seed 976:Stem 954:Leaf 929:Root 924:Bulb 908:Wood 869:Cork 767:Fern 582:PMID 453:PMID 445:ISSN 404:PMID 298:seed 175:Rice 113:any 111:cite 1387:Sap 1104:Nut 949:Bud 572:doi 538:doi 526:107 503:doi 435:doi 394:PMC 384:doi 343:doi 236:SLA 177:or 124:by 1851:: 580:. 568:44 566:. 562:. 550:^ 536:. 524:. 499:41 497:. 459:. 451:. 443:. 431:44 429:. 425:. 402:. 392:. 382:. 372:15 370:. 366:. 339:89 337:. 333:. 300:. 811:) 807:( 619:e 612:t 605:v 588:. 574:: 544:. 540:: 532:: 509:. 505:: 467:. 437:: 410:. 386:: 378:: 351:. 345:: 247:( 151:) 145:( 140:) 136:( 132:. 118:. 20:)

Index

Planting density
seed bank
competition
resources

monocultures
ecology
biomass

cite
sources
improve this section
adding citations to reliable sources
removed
Learn how and when to remove this message

Maize
Rice
Barley
forestry
Leaf Area Index
Stem Mass Fraction
phenotype

leaves
SLA
tillers
branches
photosynthetic rate
transpiration rate

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