3237:
152:
1429:
1239:
788:
3226:
3232:
176:
3243:
1572:
1048:
1181:
766:
1456:
1149:
1026:
1651:
1312:
877:
1485:
1290:
826:
540:
1550:
1070:
3266:
1622:
1521:
1268:
937:
612:
25:
744:
1728:
1210:
959:
578:
and discovered small bumps on the surface, known as quill knobs. The same feature is present in some bird bones, and represents the attachment point for strong secondary wing feathers. This finding provided the first direct evidence that velociraptorines, like all other maniraptorans, had feathers.
587:
According to Currie (1995), Velociraptorinae can be distinguished based on the following characteristics: dromaeosaurids with maxillary and dentary teeth possessing denticles on the anterior carinae that are significantly smaller than the posterior denticles, and which have a second premaxillary
664:(2012) also supported a traditional, monophyletic of Velociraptorinae. However, some studies found a very different group of dromaeosaurids in velociraptorinae, such as Longrich and Currie (2009), which found
2291:
Godefroit, Pascal; Cau, Andrea; Hu, Dong-Yu; Escuillié, François; Wu, Wenhao; Dyke, Gareth (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds".
623:
and supposed closely related species. It was not until 1998 that this group was defined as a clade by Paul Sereno. Sereno defined the group as all dromaeosaurids more closely related to
595:(2012), Velociraptorinae can be distinguished based on the following unambiguous characteristics: the posterior opening of the basisphenoid recess is divided into two small, circular
2113:
Currie, P.J. 1995. New information on the anatomy and relationships of
Dromaeosaurus albertensis (Dinosauria: Theropoda). Journal of Vertebrate Paleontology 15: 576–591.
588:
tooth that is significantly larger than the third and fourth premaxillary teeth; dromaeosaurids with nasals that appear depressed, when observed in lateral view.
2419:
1970:
Jasinski, Steven E.; Sullivan, Robert M.; Carter, Aja M.; Johnson, Erynn H.; Dalman, Sebastian G.; Zariwala, Juned; Currie, Philip J. (2022-11-07).
547:
While most velociraptorines were generally small animals, at least one species may have achieved gigantic sizes comparable to those found among the
43:
674:
to be a member of a more basal group they named
Saurornitholestinae. A larger analysis in 2013 found some traditional velociraptorines, such as
2344:"New Dromaeosaurid Dinosaur (Theropoda, Dromaeosauridae) from New Mexico and Biodiversity of Dromaeosaurids at the end of the Cretaceous"
3259:
1846:
2210:
Sereno, P. C. (1998). "A rationale for phylogenetic definitions, with application to the higher-level taxonomy of
Dinosauria".
3236:
3225:
175:
2412:
3242:
565:, but they appear to belong to a velociraptorine, judging by the shape of the teeth and the anatomy of their serrations.
3231:
1124:
below follows a 2012 analysis by Turner, Makovicky and Norell, using a dataset of 474 characters scored for 111 taxa.
635:, they vary widely regarding which species are actually velociraptorines and which are either more basal or closer to
2047:
Naish, D. Hutt, and
Martill, D.M. (2001). "Saurischian dinosaurs: theropods." in Martill, D.M. and Naish, D. (eds).
2181:
1977:, a southern eudromaeosaur (Theropoda: Dromaeosauridae: Eudromaeosauria) from the latest Cretaceous of New Mexico"
1645:
310:
161:
2516:
2405:
1868:"A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight"
599:
by a thin bar of bone; the dorsal tympanic recess is present as a deep, posterolaterally directed concavity;
3374:
3286:
1674:
1428:
1238:
787:
642:
Novas and Pol (2005) found a distinct velociraptorine clade close to the traditional view, which included
316:
3346:
3269:
515:
are previously assigned to the velociraptorines, they are now reclassified within different lineages of
151:
3324:
2570:
2300:
2078:
1934:
33:
2987:
1424:
724:
2724:
2543:
2324:
2159:
2009:
1952:
170:
2027:
3351:
519:. Several teeth that may belong to indeterminate velociraptorines have also been discovered in
3184:
3075:
3011:
2907:
2700:
2383:
2365:
2316:
2273:
2096:
2001:
1993:
1899:
1866:
Hartman, S.; Mortimer, M.; Wahl, W. R.; Lomax, D. R.; Lippincott, J.; Lovelace, D. M. (2019).
1571:
1328:
1233:
1047:
782:
3054:
2914:
2892:
2373:
2355:
2308:
2263:
2253:
2219:
2177:
2149:
2141:
2086:
1985:
1942:
1889:
1879:
1180:
765:
704:
600:
551:. So far, this unnamed giant velociraptorine is known only from isolated teeth found on the
140:
1455:
1148:
1025:
3168:
3022:
2925:
2638:
2616:
2582:
2501:
2428:
1650:
1451:
1410:
1311:
1144:
1131:
1006:
876:
714:
631:. While several studies have since recovered a group of dromaeosaurids closely related to
569:
548:
516:
498:
494:
412:
275:
259:
38:
2242:"A microraptorine (Dinosauria–Dromaeosauridae) from the Late Cretaceous of North America"
2304:
2082:
1938:
1484:
1289:
825:
2826:
2378:
2343:
2268:
2241:
1972:
1894:
1867:
1549:
1069:
619:
When erected by
Barsbold in 1983, Velociraptorinae was conceived as a group containing
539:
1621:
1520:
1267:
936:
3368:
3147:
3061:
2953:
2938:
2877:
2870:
2863:
2853:
2788:
2781:
2710:
2013:
1956:
1698:
1398:
1020:
552:
528:
512:
507:
392:
381:
359:
136:
98:
2223:
2163:
2028:
Velociraptorine dromaeosaurid teeth from the
Kimmeridgian (Late Jurassic) of Germany
3206:
3117:
3038:
3031:
2996:
2975:
2960:
2899:
2766:
2759:
2752:
2742:
2689:
2674:
2328:
1616:
1566:
1306:
1042:
871:
738:
680:
644:
574:
524:
475:
338:
299:
1947:
1918:
3333:
3318:
3161:
3139:
3090:
3046:
3003:
2968:
2946:
2885:
2796:
2731:
2555:
1744:
1479:
1284:
1175:
975:
820:
760:
666:
650:
444:
422:
73:
3309:
2360:
611:
3154:
3131:
3083:
2834:
2803:
2681:
2659:
2467:
2458:
2198:
1544:
1064:
561:
402:
370:
233:
118:
83:
2369:
1997:
497:. Definitive fossils attributed to the subfamily have only been found in the
3192:
3124:
2819:
2811:
2717:
2528:
2476:
2258:
2091:
2062:
1515:
1389:
1262:
1121:
931:
700:
455:
349:
246:
187:
123:
2387:
2320:
2277:
2100:
2005:
1903:
743:
3303:
3176:
3097:
3068:
2842:
2652:
2452:
2051:. The Palaeontological Association, Field Guides to Fossils. 10, 242–309.
893:
596:
559:. The teeth belong to an animal the size of dromaeosaurines of the genus
490:
433:
220:
207:
113:
108:
93:
88:
78:
2312:
3199:
2485:
1884:
1722:
1204:
953:
703:
below follows a 2009 analysis by paleontologists
Nicholas Longrich and
693:
656:
556:
520:
465:
128:
103:
3338:
692:, previously found to be a velociraptorine by most analyses, to be an
2774:
2666:
2446:
2190:
2154:
1989:
197:
3280:
2129:
1923:, a dromaeosaurid theropod from the Upper Cretaceous of Tajikistan"
1727:
1209:
958:
670:
to be a non-velociraptorine, non-dromaeosaurine eudromaeosaur, and
2397:
2145:
610:
538:
325:
2184:[Carnivorous dinosaurs from the Cretaceous of Mongolia]
502:
3284:
2439:
2401:
2130:"A Review of Dromaeosaurid Systematics and Paravian Phylogeny"
1392:
below follows the phylogenetic analysis performed by
Jasinski
18:
2212:
Neues
Jahrbuch für Geologie und Paläontologie - Abhandlungen
2026:
van der Lubbe, T., Richter, U. and Knotschke, N. (2009). "
707:, using a dataset of 114 characters scored for 23 taxa.
2128:
Turner, A. H.; Makovicky, P. J.; Norell, M. A. (2012).
2342:
Jasinski, S. E.; Sullivan, R. M.; Dodson, P. (2020).
2061:
Turner, A.H.; Makovicky, P.J.; Norell, M.A. (2007).
471:
461:
451:
440:
429:
418:
408:
398:
388:
377:
366:
355:
345:
334:
306:
3293:
3107:
3021:
2986:
2924:
2852:
2741:
2699:
2636:
2581:
2554:
2527:
2500:
2134:Bulletin of the American Museum of Natural History
1971:
511:as the basalmost member. While numerous taxa from
688:, making them dromaeosaurines. This study found
2246:Proceedings of the National Academy of Sciences
2235:
2233:
2413:
684:, while others to be more closely related to
8:
603:are present in all of the dorsal vertebrae.
660:. A cladistic analysis conducted by Turner
32:It has been suggested that this article be
3281:
2930:
2644:
2633:
2560:
2533:
2506:
2497:
2436:
2420:
2406:
2398:
150:
59:
2377:
2359:
2267:
2257:
2153:
2123:
2121:
2119:
2090:
1946:
1917:Averianov, A. O.; Lopatin, A. V. (2023).
1893:
1883:
2240:Longrich, N. R.; Currie, P. J. (2009).
1858:
2063:"Feather quill knobs in the dinosaur
7:
654:, and material that was later named
1714:
1690:
1666:
1637:
1608:
1536:
1507:
1500:
1471:
1444:
1417:
1405:
1254:
1225:
1196:
1167:
1137:
1127:
1012:
923:
863:
851:
841:
812:
730:
720:
710:
1847:Timeline of dromaeosaurid research
615:Velociraptorine skeletons to scale
583:Distinguishing anatomical features
14:
3265:
3264:
3241:
3235:
3230:
3224:
2182:"Хищные динозавры мела Монголии"
1726:
1649:
1620:
1570:
1548:
1519:
1483:
1454:
1427:
1310:
1288:
1266:
1237:
1208:
1179:
1147:
1068:
1046:
1024:
957:
935:
875:
824:
786:
764:
742:
174:
23:
1973:"Osteology and reassessment of
1396:2020 during the description of
2049:Dinosaurs of the Isle of Wight
543:Velociraptorine skull to scale
16:Extinct subfamily of dinosaurs
1:
2032:Acta Palaeontologica Polonica
1948:10.1016/j.cretres.2023.105524
572:studied front limb bones of
3391:
2361:10.1038/s41598-020-61480-7
1975:Dineobellator notohesperus
49:Proposed since April 2024.
3254:
3222:
2933:
2647:
2632:
2613:
2563:
2536:
2509:
2496:
2435:
2224:10.1127/njgpa/210/1998/41
1741:
1719:
1712:
1695:
1688:
1671:
1664:
1646:Velociraptor mongoliensis
1642:
1635:
1613:
1606:
1563:
1541:
1534:
1512:
1505:
1498:
1476:
1469:
1449:
1442:
1422:
1415:
1325:
1303:
1281:
1259:
1252:
1230:
1223:
1201:
1194:
1172:
1165:
1142:
1135:
1061:
1039:
1017:
1010:
972:
950:
928:
921:
890:
868:
861:
849:
839:
817:
810:
779:
757:
735:
728:
718:
331:
324:
311:Velociraptor mongoliensis
305:
298:
171:Scientific classification
169:
162:Velociraptor mongoliensis
158:
149:
62:
678:, to be more basal than
165:(specimen MPC-D 100/54)
2259:10.1073/pnas.0811664106
2092:10.1126/science.1145076
1921:Kansaignathus sogdianus
69:Santonian–Maastrichtian
1675:Velociraptor osmolskae
616:
544:
489:is a subfamily of the
3347:Paleobiology Database
1982:The Anatomical Record
614:
542:
591:According to Turner
2988:Saurornitholestinae
2313:10.1038/nature12168
2305:2013Natur.498..359G
2083:2007Sci...317.1721T
1939:2023CrRes.14705524A
1927:Cretaceous Research
1425:Saurornitholestinae
725:Saurornitholestinae
2348:Scientific Reports
1885:10.7717/peerj.7247
617:
545:
3362:
3361:
3287:Taxon identifiers
3278:
3277:
3250:
3249:
3220:
3219:
3216:
3215:
3185:Saurornitholestes
3076:Saurornitholestes
3012:Saurornitholestes
2701:Halszkaraptorinae
2628:
2627:
2624:
2623:
2609:
2608:
2605:
2604:
2601:
2600:
2299:(7454): 359–362.
2252:(13): 5002−5007.
1837:
1836:
1828:
1827:
1819:
1818:
1810:
1809:
1801:
1800:
1792:
1791:
1783:
1782:
1774:
1773:
1765:
1764:
1756:
1755:
1592:
1591:
1583:
1582:
1385:
1384:
1376:
1375:
1367:
1366:
1358:
1357:
1349:
1348:
1340:
1339:
1234:Saurornitholestes
1117:
1116:
1108:
1107:
1099:
1098:
1090:
1089:
1081:
1080:
996:
995:
987:
986:
905:
904:
799:
798:
783:Saurornitholestes
672:Saurornitholestes
484:
483:
294:
56:
55:
51:
3382:
3355:
3354:
3342:
3341:
3329:
3328:
3327:
3325:Velociraptorinae
3314:
3313:
3312:
3295:Velociraptorinae
3282:
3268:
3267:
3245:
3239:
3234:
3228:
3109:Velociraptorinae
3055:Dromaeosauroides
2931:
2915:Zhongjianosaurus
2893:Sinornithosaurus
2645:
2634:
2561:
2534:
2507:
2498:
2491:
2490:
2437:
2422:
2415:
2408:
2399:
2392:
2391:
2381:
2363:
2339:
2333:
2332:
2288:
2282:
2281:
2271:
2261:
2237:
2228:
2227:
2207:
2201:
2199:Translated paper
2197:
2187:
2174:
2168:
2167:
2157:
2125:
2114:
2111:
2105:
2104:
2094:
2058:
2052:
2045:
2039:
2024:
2018:
2017:
1990:10.1002/ar.25103
1979:
1967:
1961:
1960:
1950:
1914:
1908:
1907:
1897:
1887:
1863:
1730:
1715:
1691:
1667:
1653:
1638:
1624:
1609:
1603:Velociraptorinae
1574:
1552:
1537:
1523:
1508:
1501:
1487:
1472:
1458:
1445:
1431:
1418:
1406:
1314:
1292:
1270:
1255:
1241:
1226:
1212:
1197:
1183:
1168:
1162:Velociraptorinae
1151:
1138:
1128:
1072:
1050:
1028:
1013:
961:
939:
924:
918:
879:
864:
858:
852:
846:Velociraptorinae
842:
828:
813:
790:
768:
746:
731:
721:
711:
705:Philip J. Currie
523:, dating to the
487:Velociraptorinae
473:
463:
453:
442:
431:
420:
410:
400:
390:
379:
368:
357:
347:
336:
308:
292:
289:Velociraptorinae
287:
274:
258:
179:
178:
154:
144:
141:Early Cretaceous
133:
70:
66:Temporal range:
63:Velociraptorines
60:
47:
27:
26:
19:
3390:
3389:
3385:
3384:
3383:
3381:
3380:
3379:
3365:
3364:
3363:
3358:
3350:
3345:
3337:
3332:
3323:
3322:
3317:
3308:
3307:
3302:
3289:
3279:
3274:
3246:
3212:
3169:Luanchuanraptor
3103:
3023:Dromaeosaurinae
3017:
2982:
2926:Eudromaeosauria
2920:
2848:
2737:
2695:
2641:
2639:Dromaeosauridae
2620:
2617:Dromaeosauridae
2597:
2583:Dromaeosauridae
2577:
2550:
2523:
2517:Avemetatarsalia
2502:Avemetatarsalia
2492:
2442:
2441:
2431:
2429:Dromaeosauridae
2426:
2396:
2395:
2341:
2340:
2336:
2290:
2289:
2285:
2239:
2238:
2231:
2209:
2208:
2204:
2185:
2176:
2175:
2171:
2127:
2126:
2117:
2112:
2108:
2060:
2059:
2055:
2046:
2042:
2025:
2021:
1969:
1968:
1964:
1916:
1915:
1911:
1865:
1864:
1860:
1855:
1843:
1838:
1829:
1820:
1811:
1802:
1793:
1784:
1775:
1766:
1757:
1593:
1584:
1452:Dromaeosaurinae
1411:Eudromaeosauria
1386:
1377:
1368:
1359:
1350:
1341:
1145:Dromaeosaurinae
1132:Eudromaeosauria
1118:
1109:
1100:
1091:
1082:
1007:Dromaeosaurinae
997:
988:
919:
916:
906:
859:
856:
800:
715:Eudromaeosauria
609:
585:
570:paleontologists
549:dromaeosaurines
537:
517:Eudromaeosauria
499:Late Cretaceous
495:Dromaeosauridae
413:Luanchuanraptor
320:
314:
291:
285:
276:Eudromaeosauria
272:
260:Dromaeosauridae
256:
173:
145:
134:
132:
131:
126:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
68:
67:
64:
52:
39:Eudromaeosauria
28:
24:
17:
12:
11:
5:
3388:
3386:
3378:
3377:
3375:Eudromaeosaurs
3367:
3366:
3360:
3359:
3357:
3356:
3343:
3330:
3315:
3299:
3297:
3291:
3290:
3285:
3276:
3275:
3273:
3272:
3262:
3255:
3252:
3251:
3248:
3247:
3223:
3221:
3218:
3217:
3214:
3213:
3211:
3210:
3203:
3196:
3189:
3181:
3173:
3165:
3158:
3151:
3144:
3136:
3128:
3121:
3113:
3111:
3105:
3104:
3102:
3101:
3094:
3087:
3080:
3072:
3065:
3058:
3051:
3043:
3035:
3027:
3025:
3019:
3018:
3016:
3015:
3008:
3000:
2992:
2990:
2984:
2983:
2981:
2980:
2972:
2965:
2957:
2950:
2943:
2934:
2928:
2922:
2921:
2919:
2918:
2911:
2904:
2896:
2889:
2882:
2874:
2867:
2859:
2857:
2850:
2849:
2847:
2846:
2839:
2831:
2827:Unquillosaurus
2823:
2816:
2808:
2800:
2793:
2785:
2778:
2771:
2763:
2756:
2748:
2746:
2739:
2738:
2736:
2735:
2728:
2721:
2714:
2706:
2704:
2697:
2696:
2694:
2693:
2686:
2678:
2671:
2663:
2656:
2648:
2642:
2637:
2630:
2629:
2626:
2625:
2622:
2621:
2614:
2611:
2610:
2607:
2606:
2603:
2602:
2599:
2598:
2596:
2595:
2594:
2593:
2587:
2585:
2579:
2578:
2576:
2575:
2574:
2573:
2564:
2558:
2552:
2551:
2549:
2548:
2547:
2546:
2537:
2531:
2525:
2524:
2522:
2521:
2520:
2519:
2510:
2504:
2494:
2493:
2489:
2488:
2479:
2470:
2461:
2455:
2449:
2440:
2433:
2432:
2427:
2425:
2424:
2417:
2410:
2402:
2394:
2393:
2334:
2283:
2229:
2202:
2192:(in Russian).
2169:
2115:
2106:
2077:(5845): 1721.
2053:
2040:
2019:
1962:
1909:
1857:
1856:
1854:
1851:
1850:
1849:
1842:
1839:
1835:
1834:
1831:
1830:
1826:
1825:
1822:
1821:
1817:
1816:
1813:
1812:
1808:
1807:
1804:
1803:
1799:
1798:
1795:
1794:
1790:
1789:
1786:
1785:
1781:
1780:
1777:
1776:
1772:
1771:
1768:
1767:
1763:
1762:
1759:
1758:
1754:
1753:
1750:
1749:
1740:
1737:
1736:
1733:
1732:
1718:
1713:
1711:
1708:
1707:
1704:
1703:
1694:
1689:
1687:
1684:
1683:
1680:
1679:
1670:
1665:
1663:
1660:
1659:
1656:
1655:
1641:
1636:
1634:
1631:
1630:
1627:
1626:
1612:
1607:
1605:
1599:
1598:
1595:
1594:
1590:
1589:
1586:
1585:
1581:
1580:
1577:
1576:
1562:
1559:
1558:
1555:
1554:
1540:
1535:
1533:
1530:
1529:
1526:
1525:
1511:
1506:
1504:
1499:
1497:
1494:
1493:
1490:
1489:
1475:
1470:
1468:
1465:
1464:
1461:
1460:
1448:
1443:
1441:
1438:
1437:
1434:
1433:
1421:
1416:
1414:
1404:
1383:
1382:
1379:
1378:
1374:
1373:
1370:
1369:
1365:
1364:
1361:
1360:
1356:
1355:
1352:
1351:
1347:
1346:
1343:
1342:
1338:
1337:
1334:
1333:
1324:
1321:
1320:
1317:
1316:
1302:
1299:
1298:
1295:
1294:
1280:
1277:
1276:
1273:
1272:
1258:
1253:
1251:
1248:
1247:
1244:
1243:
1229:
1224:
1222:
1219:
1218:
1215:
1214:
1200:
1195:
1193:
1190:
1189:
1186:
1185:
1171:
1166:
1164:
1158:
1157:
1154:
1153:
1141:
1136:
1134:
1126:
1115:
1114:
1111:
1110:
1106:
1105:
1102:
1101:
1097:
1096:
1093:
1092:
1088:
1087:
1084:
1083:
1079:
1078:
1075:
1074:
1060:
1057:
1056:
1053:
1052:
1038:
1035:
1034:
1031:
1030:
1016:
1011:
1009:
1003:
1002:
999:
998:
994:
993:
990:
989:
985:
984:
981:
980:
971:
968:
967:
964:
963:
949:
946:
945:
942:
941:
927:
922:
920:
915:
912:
911:
908:
907:
903:
902:
899:
898:
889:
886:
885:
882:
881:
867:
862:
860:
855:
850:
848:
840:
838:
835:
834:
831:
830:
816:
811:
809:
806:
805:
802:
801:
797:
796:
793:
792:
778:
775:
774:
771:
770:
756:
753:
752:
749:
748:
734:
729:
727:
719:
717:
709:
608:
607:Classification
605:
584:
581:
536:
533:
482:
481:
480:
479:
469:
459:
449:
438:
427:
416:
406:
396:
386:
375:
364:
353:
343:
329:
328:
322:
321:
315:
303:
302:
296:
295:
293:Barsbold, 1983
283:
279:
278:
270:
263:
262:
254:
250:
249:
244:
237:
236:
231:
224:
223:
218:
211:
210:
205:
201:
200:
195:
191:
190:
185:
181:
180:
167:
166:
156:
155:
147:
146:
127:
122:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
71:
65:
54:
53:
31:
29:
22:
15:
13:
10:
9:
6:
4:
3:
2:
3387:
3376:
3373:
3372:
3370:
3353:
3348:
3344:
3340:
3335:
3331:
3326:
3320:
3316:
3311:
3305:
3301:
3300:
3298:
3296:
3292:
3288:
3283:
3271:
3263:
3261:
3257:
3256:
3253:
3244:
3240:
3238:
3233:
3227:
3209:
3208:
3204:
3202:
3201:
3197:
3195:
3194:
3190:
3187:
3186:
3182:
3179:
3178:
3174:
3171:
3170:
3166:
3164:
3163:
3159:
3157:
3156:
3152:
3150:
3149:
3148:Kansaignathus
3145:
3142:
3141:
3137:
3134:
3133:
3129:
3127:
3126:
3122:
3120:
3119:
3115:
3114:
3112:
3110:
3106:
3100:
3099:
3095:
3093:
3092:
3088:
3086:
3085:
3081:
3078:
3077:
3073:
3071:
3070:
3066:
3064:
3063:
3062:Dromaeosaurus
3059:
3057:
3056:
3052:
3049:
3048:
3044:
3041:
3040:
3036:
3034:
3033:
3029:
3028:
3026:
3024:
3020:
3014:
3013:
3009:
3006:
3005:
3001:
2999:
2998:
2994:
2993:
2991:
2989:
2985:
2978:
2977:
2973:
2971:
2970:
2966:
2963:
2962:
2958:
2956:
2955:
2954:Dineobellator
2951:
2949:
2948:
2944:
2941:
2940:
2936:
2935:
2932:
2929:
2927:
2923:
2917:
2916:
2912:
2910:
2909:
2905:
2902:
2901:
2897:
2895:
2894:
2890:
2888:
2887:
2883:
2880:
2879:
2878:Hesperonychus
2875:
2873:
2872:
2871:Graciliraptor
2868:
2866:
2865:
2864:Changyuraptor
2861:
2860:
2858:
2855:
2854:Microraptoria
2851:
2845:
2844:
2840:
2837:
2836:
2832:
2829:
2828:
2824:
2822:
2821:
2817:
2814:
2813:
2809:
2806:
2805:
2801:
2799:
2798:
2794:
2791:
2790:
2789:Ornithodesmus
2786:
2784:
2783:
2782:Neuquenraptor
2779:
2777:
2776:
2772:
2769:
2768:
2764:
2762:
2761:
2757:
2755:
2754:
2750:
2749:
2747:
2744:
2740:
2734:
2733:
2729:
2727:
2726:
2722:
2720:
2719:
2715:
2713:
2712:
2711:Halszkaraptor
2708:
2707:
2705:
2702:
2698:
2692:
2691:
2687:
2684:
2683:
2679:
2677:
2676:
2672:
2669:
2668:
2664:
2662:
2661:
2657:
2655:
2654:
2650:
2649:
2646:
2643:
2640:
2635:
2631:
2619:
2618:
2612:
2591:
2590:
2589:
2588:
2586:
2584:
2580:
2572:
2568:
2567:
2566:
2565:
2562:
2559:
2557:
2553:
2545:
2541:
2540:
2539:
2538:
2535:
2532:
2530:
2526:
2518:
2514:
2513:
2512:
2511:
2508:
2505:
2503:
2499:
2495:
2487:
2483:
2480:
2478:
2474:
2471:
2469:
2465:
2462:
2460:
2456:
2454:
2450:
2448:
2444:
2443:
2438:
2434:
2430:
2423:
2418:
2416:
2411:
2409:
2404:
2403:
2400:
2389:
2385:
2380:
2375:
2371:
2367:
2362:
2357:
2353:
2349:
2345:
2338:
2335:
2330:
2326:
2322:
2318:
2314:
2310:
2306:
2302:
2298:
2294:
2287:
2284:
2279:
2275:
2270:
2265:
2260:
2255:
2251:
2247:
2243:
2236:
2234:
2230:
2225:
2221:
2217:
2213:
2206:
2203:
2200:
2195:
2191:
2183:
2179:
2173:
2170:
2165:
2161:
2156:
2151:
2147:
2146:10.1206/748.1
2143:
2139:
2135:
2131:
2124:
2122:
2120:
2116:
2110:
2107:
2102:
2098:
2093:
2088:
2084:
2080:
2076:
2072:
2068:
2066:
2057:
2054:
2050:
2044:
2041:
2038:(3): 401-408.
2037:
2033:
2029:
2023:
2020:
2015:
2011:
2007:
2003:
1999:
1995:
1991:
1987:
1983:
1978:
1976:
1966:
1963:
1958:
1954:
1949:
1944:
1940:
1936:
1932:
1928:
1924:
1922:
1919:"New data on
1913:
1910:
1905:
1901:
1896:
1891:
1886:
1881:
1877:
1873:
1869:
1862:
1859:
1852:
1848:
1845:
1844:
1840:
1833:
1832:
1824:
1823:
1815:
1814:
1806:
1805:
1797:
1796:
1788:
1787:
1779:
1778:
1770:
1769:
1761:
1760:
1752:
1751:
1748:
1747:
1746:
1739:
1738:
1735:
1734:
1731:
1729:
1725:
1724:
1717:
1716:
1710:
1709:
1706:
1705:
1702:
1701:
1700:
1699:Dineobellator
1693:
1692:
1686:
1685:
1682:
1681:
1678:
1677:
1676:
1669:
1668:
1662:
1661:
1658:
1657:
1654:
1652:
1648:
1647:
1640:
1639:
1633:
1632:
1629:
1628:
1625:
1623:
1619:
1618:
1611:
1610:
1604:
1601:
1600:
1597:
1596:
1588:
1587:
1579:
1578:
1575:
1573:
1569:
1568:
1561:
1560:
1557:
1556:
1553:
1551:
1547:
1546:
1539:
1538:
1532:
1531:
1528:
1527:
1524:
1522:
1518:
1517:
1510:
1509:
1503:
1502:
1496:
1495:
1492:
1491:
1488:
1486:
1482:
1481:
1474:
1473:
1467:
1466:
1463:
1462:
1459:
1457:
1453:
1447:
1446:
1440:
1439:
1436:
1435:
1432:
1430:
1426:
1420:
1419:
1413:
1412:
1408:
1407:
1403:
1401:
1400:
1399:Dineobellator
1395:
1391:
1381:
1380:
1372:
1371:
1363:
1362:
1354:
1353:
1345:
1344:
1336:
1335:
1332:
1331:
1330:
1323:
1322:
1319:
1318:
1315:
1313:
1309:
1308:
1301:
1300:
1297:
1296:
1293:
1291:
1287:
1286:
1279:
1278:
1275:
1274:
1271:
1269:
1265:
1264:
1257:
1256:
1250:
1249:
1246:
1245:
1242:
1240:
1236:
1235:
1228:
1227:
1221:
1220:
1217:
1216:
1213:
1211:
1207:
1206:
1199:
1198:
1192:
1191:
1188:
1187:
1184:
1182:
1178:
1177:
1170:
1169:
1163:
1160:
1159:
1156:
1155:
1152:
1150:
1146:
1140:
1139:
1133:
1130:
1129:
1125:
1123:
1113:
1112:
1104:
1103:
1095:
1094:
1086:
1085:
1077:
1076:
1073:
1071:
1067:
1066:
1059:
1058:
1055:
1054:
1051:
1049:
1045:
1044:
1037:
1036:
1033:
1032:
1029:
1027:
1023:
1022:
1021:Dromaeosaurus
1015:
1014:
1008:
1005:
1004:
1001:
1000:
992:
991:
983:
982:
979:
978:
977:
970:
969:
966:
965:
962:
960:
956:
955:
948:
947:
944:
943:
940:
938:
934:
933:
926:
925:
914:
913:
910:
909:
901:
900:
897:
896:
895:
888:
887:
884:
883:
880:
878:
874:
873:
866:
865:
854:
853:
847:
844:
843:
837:
836:
833:
832:
829:
827:
823:
822:
815:
814:
808:
807:
804:
803:
795:
794:
791:
789:
785:
784:
777:
776:
773:
772:
769:
767:
763:
762:
755:
754:
751:
750:
747:
745:
741:
740:
733:
732:
726:
723:
722:
716:
713:
712:
708:
706:
702:
697:
695:
691:
687:
686:Dromaeosaurus
683:
682:
677:
673:
669:
668:
663:
659:
658:
653:
652:
647:
646:
640:
638:
637:Dromaeosaurus
634:
630:
629:Dromaeosaurus
626:
622:
613:
606:
604:
602:
598:
594:
589:
582:
580:
577:
576:
571:
566:
564:
563:
558:
554:
553:Isle of Wight
550:
541:
534:
532:
530:
529:Late Jurassic
527:stage of the
526:
522:
518:
514:
513:North America
510:
509:
508:Kansaignathus
504:
500:
496:
492:
488:
478:
477:
470:
468:
467:
460:
458:
457:
450:
447:
446:
439:
436:
435:
428:
425:
424:
417:
415:
414:
407:
405:
404:
397:
395:
394:
393:Kansaignathus
387:
384:
383:
382:Dineobellator
376:
373:
372:
365:
362:
361:
354:
352:
351:
344:
341:
340:
333:
332:
330:
327:
323:
318:
313:
312:
304:
301:
297:
290:
284:
281:
280:
277:
271:
268:
265:
264:
261:
255:
252:
251:
248:
245:
242:
239:
238:
235:
232:
229:
226:
225:
222:
219:
216:
213:
212:
209:
206:
203:
202:
199:
196:
193:
192:
189:
186:
183:
182:
177:
172:
168:
164:
163:
157:
153:
148:
142:
138:
137:Late Jurassic
130:
125:
120:
115:
110:
105:
100:
95:
90:
85:
80:
75:
61:
58:
50:
45:
41:
40:
35:
30:
21:
20:
3294:
3229:
3207:Velociraptor
3205:
3198:
3191:
3183:
3175:
3167:
3160:
3153:
3146:
3138:
3130:
3123:
3118:Acheroraptor
3116:
3108:
3096:
3089:
3082:
3074:
3067:
3060:
3053:
3045:
3039:Dakotaraptor
3037:
3032:Achillobator
3030:
3010:
3002:
2997:Atrociraptor
2995:
2976:Zhenyuanlong
2974:
2967:
2961:Tianyuraptor
2959:
2952:
2945:
2937:
2913:
2906:
2900:Tianyuraptor
2898:
2891:
2884:
2876:
2869:
2862:
2841:
2833:
2825:
2818:
2810:
2802:
2795:
2787:
2780:
2773:
2767:Dakotaraptor
2765:
2760:Buitreraptor
2758:
2753:Austroraptor
2751:
2743:Unenlagiinae
2730:
2723:
2716:
2709:
2690:Zhenyuanlong
2688:
2680:
2675:Tianyuraptor
2673:
2665:
2658:
2651:
2615:
2481:
2472:
2463:
2351:
2347:
2337:
2296:
2292:
2286:
2249:
2245:
2218:(1): 41–83.
2215:
2211:
2205:
2193:
2189:
2178:Barsbold, R.
2172:
2137:
2133:
2109:
2074:
2070:
2065:Velociraptor
2064:
2056:
2048:
2043:
2035:
2031:
2022:
1984:: ar.25103.
1981:
1974:
1965:
1930:
1926:
1920:
1912:
1875:
1871:
1861:
1743:
1742:
1721:
1720:
1697:
1696:
1673:
1672:
1644:
1643:
1617:Acheroraptor
1615:
1614:
1602:
1567:Achillobator
1565:
1564:
1543:
1542:
1514:
1513:
1478:
1477:
1450:
1423:
1409:
1397:
1393:
1387:
1327:
1326:
1307:Velociraptor
1305:
1304:
1283:
1282:
1261:
1260:
1232:
1231:
1203:
1202:
1174:
1173:
1161:
1143:
1119:
1063:
1062:
1043:Achillobator
1041:
1040:
1019:
1018:
974:
973:
952:
951:
930:
929:
892:
891:
872:Velociraptor
870:
869:
845:
819:
818:
781:
780:
759:
758:
739:Atrociraptor
737:
736:
698:
689:
685:
681:Velociraptor
679:
675:
671:
665:
661:
655:
649:
645:Velociraptor
643:
641:
636:
633:Velociraptor
632:
628:
625:Velociraptor
624:
621:Velociraptor
620:
618:
592:
590:
586:
575:Velociraptor
573:
567:
560:
546:
525:Kimmeridgian
506:
501:deposits of
486:
485:
476:Velociraptor
474:
464:
454:
443:
432:
421:
411:
401:
391:
380:
369:
358:
348:
339:Acheroraptor
337:
309:
300:Type species
288:
266:
240:
227:
214:
160:
57:
48:
37:
3334:iNaturalist
3319:Wikispecies
3162:Linheraptor
3140:Deinonychus
3091:Yurgovuchia
3047:Deinonychus
3004:Bambiraptor
2969:Vectiraptor
2947:Bambiraptor
2886:Microraptor
2797:Pamparaptor
2732:Natovenator
2571:Maniraptora
2556:Maniraptora
2354:(1): 5105.
1745:Linheraptor
1480:Deinonychus
1285:Deinonychus
1176:Bambiraptor
976:Linheraptor
821:Deinonychus
761:Bambiraptor
667:Deinonychus
651:Deinonychus
601:pleurocoels
535:Description
445:Deinonychus
423:Linheraptor
282:Subfamily:
3258:See also:
3132:Boreonykus
3084:Utahraptor
2835:Variraptor
2804:Pyroraptor
2682:Variraptor
2660:Pyroraptor
2592:see below↓
2468:Dinosauria
2459:Sauropsida
1933:. 105524.
1853:References
1545:Utahraptor
1065:Utahraptor
562:Utahraptor
371:Boreonykus
234:Saurischia
221:Dinosauria
135:Uncertain
3125:Adasaurus
2820:Unenlagia
2812:Rahonavis
2718:Hulsanpes
2544:Theropoda
2529:Theropoda
2477:Theropoda
2445:Kingdom:
2370:2045-2322
2155:2246/6352
2140:: 1–206.
2014:253382718
1998:1932-8486
1957:257453407
1878:: e7247.
1516:Adasaurus
1390:cladogram
1263:Adasaurus
1122:cladogram
932:Adasaurus
701:cladogram
696:instead.
350:Adasaurus
247:Theropoda
194:Kingdom:
188:Eukaryota
159:Skull of
3369:Category
3310:Q1066137
3304:Wikidata
3270:Category
3260:Timeline
3177:Nuthetes
3098:Zapsalis
3069:Itemirus
2843:Ypupiara
2725:Mahakala
2653:Daurlong
2453:Chordata
2451:Phylum:
2447:Animalia
2388:32218481
2321:23719374
2278:19289829
2180:(1983).
2164:83572446
2101:17885130
2006:36342817
1904:31333906
1841:See also
894:Itemirus
627:than to
597:foramina
568:In 2007
491:theropod
434:Nuthetes
253:Family:
208:Chordata
204:Phylum:
198:Animalia
184:Domain:
3200:Tsaagan
2486:Paraves
2457:Class:
2379:7099077
2329:4364892
2301:Bibcode
2269:2664043
2079:Bibcode
2071:Science
1935:Bibcode
1895:6626525
1723:Tsaagan
1205:Tsaagan
954:Tsaagan
917:unnamed
857:unnamed
694:avialan
676:Tsaagan
657:Tsaagan
557:England
521:Germany
505:, with
466:Tsaagan
143:records
44:Discuss
3339:123143
2939:Balaur
2908:Wulong
2775:Diuqin
2667:Shanag
2386:
2376:
2368:
2327:
2319:
2293:Nature
2276:
2266:
2162:
2099:
2012:
2004:
1996:
1955:
1902:
1892:
1394:et al.
1329:Balaur
690:Balaur
662:et al.
593:et al.
493:group
360:Balaur
326:Genera
319:, 1924
317:Osborn
34:merged
3352:54553
2482:Clade
2473:Clade
2464:Clade
2325:S2CID
2196:: 89.
2186:(PDF)
2160:S2CID
2010:S2CID
1953:S2CID
1872:PeerJ
267:Clade
241:Clade
228:Clade
215:Clade
36:into
3193:Shri
3155:Kuru
2569:see
2542:see
2515:see
2384:PMID
2366:ISSN
2317:PMID
2274:PMID
2097:PMID
2002:PMID
1994:ISSN
1900:PMID
1388:The
1120:The
699:The
503:Asia
456:Shri
403:Kuru
74:PreꞒ
2374:PMC
2356:doi
2309:doi
2297:498
2264:PMC
2254:doi
2250:106
2220:doi
2216:210
2150:hdl
2142:doi
2138:371
2087:doi
2075:317
2030:."
1986:doi
1943:doi
1931:147
1890:PMC
1880:doi
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2118:^
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2008:.
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1992:.
1980:.
1951:.
1941:.
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1925:.
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124:Pg
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2331:.
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448:?
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419:†
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389:†
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378:†
374:?
367:†
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346:†
342:?
335:†
307:†
286:†
273:†
257:†
139:-
129:N
119:K
114:J
109:T
104:P
99:C
94:D
89:S
84:O
79:Ꞓ
46:)
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