316:
296:
135:
113:
232:
273:
in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the
Permian to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous
689:
Huang, Di-Ying; Bechly, GĂĽnter; Nel, Patricia; Engel, Michael S.; Prokop, Jakub; Azar, Dany; Cai, Chen-Yang; van de Kamp, Thomas; Staniczek, Arnold H.; Garrouste, Romain; Krogmann, Lars (March 2016).
269:
it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late
Carboniferous. The group was first named as a suborder by
286:
position of the group. The morphology of the mouthparts suggests that they were capable of suction feeding and chewing, with preserved angiosperm pollen grains in the gut of
691:"New fossil insect order Permopsocida elucidates major radiation and evolution of suction feeding in hemimetabolous insects (Hexapoda: Acercaria)"
871:
Liang, F.; Zhang, W.; Liu, X. (2016). "A new genus and species of the paraneopteran family
Archipsyllidae in mid-Cretaceous amber of Myanmar".
791:
278:). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in
953:
Prokop, J.; Rosová, K.; Pecharová, M.; Sroka, P.; Leipner, A.; Nel, A. (2024). "The structure of wing in the earliest
Permopsocida".
756:"A new genus and species of the family Archipsyllidae (Insecta: Paraneoptera: Permopsocida) from the Late Triassic of Queensland"
633:
1078:
134:
242:
315:
295:
524:
121:
814:"Bridging the gap between chewing and sucking in the hemipteroid insects:new insights from Cretaceous amber"
397:
990:
378:
1083:
1037:
417:
1055:
702:
579:
552:
429:
370:
270:
560:
473:
441:
657:
591:
497:
343:
288:
645:
409:
626:
614:
536:
485:
461:
355:
129:
1042:
390:
1028:
970:
935:
896:
888:
853:
845:
783:
775:
736:
718:
517:
962:
927:
880:
835:
825:
767:
755:
726:
710:
362:
706:
112:
731:
690:
602:
567:
1072:
598:
401:
283:
279:
70:
304:
266:
246:
186:
1013:
1022:
918:—a new bizarre genus of †Permopsocida (Insecta) from Burmese Cretaceous amber".
45:
966:
931:
884:
830:
813:
771:
606:
374:
275:
90:
55:
17:
892:
849:
779:
722:
548:
382:
308:
258:
250:
166:
146:
95:
974:
939:
900:
857:
787:
740:
231:
1007:
544:
421:
366:
300:
85:
80:
65:
60:
50:
292:
suggesting that at least some members of the group were pollenivorous.
528:
100:
75:
840:
714:
637:
571:
262:
176:
156:
984:
299:
Comparison of the head morphology of
Permopsocida (b) with that of
1050:
314:
294:
230:
254:
988:
241:
914:
Beutel, R. G.; Prokop, J.; MĂĽller, P.; Pohl, H. (2019). "â€
350:
OsnabrĂĽck
Formation, Germany, Carboniferous (Moscovian)
997:
448:Croudace Bay Formation, Australia, Changhsingian
436:Croudace Bay Formation, Australia, Changhsingian
331:From Huang et al 2016 and subsequent literature
282:, which has helped clarify the morphology and
8:
985:
111:
31:
839:
829:
730:
504:Wellington Formation, Kansas, Artinskian
492:Wellington Formation, Kansas, Artinskian
480:Wellington Formation, Kansas, Artinskian
468:Wellington Formation, Kansas, Artinskian
676:
245:) to the Mid-Cretaceous. It is part of
955:Arthropod Structure & Development
7:
1056:03E0821E-FFAD-FFD6-FF70-59CB83812620
812:Yoshizawa, K.; Lienhard, C. (2016).
684:
682:
680:
373:, Kansas, Oklahoma, Early Permian (
601:, Myanmar, mid Cretaceous (latest
543:"Green Series", Germany, Toarcian
25:
395:Prokop, Garrouste & Nel, 2016
754:Lambkin, Kevin J. (2018-02-20).
133:
794:from the original on 2020-02-25
650:Yoshizawa & Lienhard, 2016
586:Zaza Formation, Russia, Aptian
1:
365:, New Mexico, Early Permian (
636:, Australia, Late Triassic (
619:Liang, Zhang & Liu, 2016
570:, Russia, Early Cretaceous (
420:", Germany, Early Jurassic (
555:, Kazakhstan, Late Jurassic
400:, Australia, Late Permian (
1100:
547:, China, Middle Jurassic (
527:, Russia, Middle Permian (
967:10.1016/j.asd.2024.101358
932:10.11646/zootaxa.4576.2.9
885:10.11646/zootaxa.4105.5.4
831:10.11646/zootaxa.4079.2.5
772:10.11646/zootaxa.4382.1.9
381:, Russia, Early Permian (
214:
209:
130:Scientific classification
128:
119:
110:
34:
27:Extinct genus of insects
525:Iva-Gora Beds Formation
122:Psocorrhyncha burmitica
664:Burmese amber, Myanmar
652:Burmese amber, Myanmar
621:Burmese amber, Myanmar
398:Croudace Bay Formation
323:
312:
235:
1079:Extinct insect orders
1038:Paleobiology Database
318:
298:
234:
634:Blackstone Formation
553:Karabastau Formation
379:Koshelevka Formation
371:Wellington Formation
271:Robert John Tillyard
41:Moscovian–Cenomanian
707:2016NatSR...623004H
348:Prokop et al., 2024
695:Scientific Reports
596:Beutel et al. 2019
324:
313:
236:
1066:
1065:
991:Taxon identifiers
715:10.1038/srep23004
663:
662:Huang et al. 2016
651:
632:
620:
597:
585:
584:Vishniakova, 1976
566:
565:Vishniakova, 1976
542:
523:
512:
503:
491:
479:
467:
456:
447:
435:
415:
396:
361:
349:
338:
229:
228:
205:
16:(Redirected from
1091:
1059:
1058:
1046:
1045:
1033:
1032:
1031:
1018:
1017:
1016:
986:
979:
978:
950:
944:
943:
911:
905:
904:
868:
862:
861:
843:
833:
809:
803:
802:
800:
799:
751:
745:
744:
734:
686:
661:
649:
630:
618:
595:
583:
564:
541:Handlirsch, 1908
540:
522:Vishnyakova 1976
521:
511:Handlirsch, 1906
510:
509:†Archipsyllidae
501:
489:
477:
465:
454:
445:
433:
413:
394:
363:Bursum Formation
359:
347:
336:
203:
198:
138:
137:
115:
105:
42:
38:Temporal range:
32:
21:
1099:
1098:
1094:
1093:
1092:
1090:
1089:
1088:
1069:
1068:
1067:
1062:
1054:
1049:
1041:
1036:
1027:
1026:
1021:
1012:
1011:
1006:
993:
983:
982:
952:
951:
947:
913:
912:
908:
870:
869:
865:
811:
810:
806:
797:
795:
753:
752:
748:
688:
687:
678:
673:
580:Archipsyllopsis
453:†Permopsocidae
430:Austropsocidium
329:
223:†Archipsyllidae
202:
196:
132:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
1097:
1095:
1087:
1086:
1081:
1071:
1070:
1064:
1063:
1061:
1060:
1047:
1034:
1019:
1003:
1001:
995:
994:
989:
981:
980:
945:
926:(2): 357–366.
906:
863:
804:
766:(1): 192–194.
746:
675:
674:
672:
669:
668:
667:
666:
665:
653:
641:
622:
610:
587:
575:
568:Zaza Formation
561:Archipsyllodes
556:
532:
507:
506:
505:
493:
490:Carpenter 1932
481:
478:Carpenter 1932
474:Lithopsocidium
469:
466:Tillyard, 1926
455:Tillyard, 1926
451:
450:
449:
442:Stenopsocidium
437:
425:
405:
386:
360:Tillyard, 1926
351:
337:Tillyard, 1926
328:
325:
227:
226:
225:
224:
221:
220:†Permopsocidae
218:
212:
211:
207:
206:
204:Tillyard, 1926
194:
190:
189:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
37:
26:
24:
18:Archipsyllidae
14:
13:
10:
9:
6:
4:
3:
2:
1096:
1085:
1082:
1080:
1077:
1076:
1074:
1057:
1052:
1048:
1044:
1039:
1035:
1030:
1024:
1020:
1015:
1009:
1005:
1004:
1002:
1000:
996:
992:
987:
976:
972:
968:
964:
960:
956:
949:
946:
941:
937:
933:
929:
925:
921:
917:
916:Bittacopsocus
910:
907:
902:
898:
894:
890:
886:
882:
879:(5): 483–90.
878:
874:
867:
864:
859:
855:
851:
847:
842:
837:
832:
827:
824:(2): 229–45.
823:
819:
815:
808:
805:
793:
789:
785:
781:
777:
773:
769:
765:
761:
757:
750:
747:
742:
738:
733:
728:
724:
720:
716:
712:
708:
704:
700:
696:
692:
685:
683:
681:
677:
670:
660:
659:
658:Psocorrhyncha
654:
648:
647:
642:
639:
635:
631:Lambkin, 2018
629:
628:
623:
617:
616:
611:
608:
604:
600:
599:Burmese amber
594:
593:
592:Bittacopsocus
588:
582:
581:
576:
573:
569:
563:
562:
557:
554:
550:
546:
539:
538:
533:
530:
526:
520:
519:
514:
513:
508:
502:Tillyard 1926
500:
499:
498:Progonopsocus
494:
488:
487:
482:
476:
475:
470:
464:
463:
458:
457:
452:
446:Tillyard 1935
444:
443:
438:
434:Tillyard 1935
432:
431:
426:
423:
419:
414:Ansorge, 1996
412:
411:
406:
403:
402:Changhsingian
399:
393:
392:
387:
384:
380:
376:
372:
368:
364:
358:
357:
352:
346:
345:
344:Carbonopsocus
340:
339:
335:†Psocidiidae
334:
333:
332:
326:
322:
321:Psocorrhyncha
317:
310:
306:
302:
297:
293:
291:
290:
289:Psocorrhyncha
285:
281:
280:Burmese amber
277:
272:
268:
264:
260:
256:
252:
248:
244:
240:
233:
222:
219:
216:
215:
213:
208:
201:
195:
192:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
124:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
33:
30:
19:
1084:Paraneoptera
1029:Permopsocida
999:Permopsocida
998:
958:
954:
948:
923:
919:
915:
909:
876:
872:
866:
821:
817:
807:
796:. Retrieved
763:
759:
749:
701:(1): 23004.
698:
694:
656:
646:Mydiognathus
644:
625:
613:
590:
578:
559:
535:
516:
496:
484:
472:
460:
440:
428:
418:Green Series
410:Liassopsocus
408:
389:
354:
342:
330:
320:
305:Thysanoptera
287:
284:phylogenetic
267:Paraneoptera
249:, alongside
247:Paraneoptera
239:Permopsocida
238:
237:
217:†Psocidiidae
200:Permopsocida
199:
187:Paraneoptera
183:(unranked):
120:
35:Permopsocida
29:
1023:Wikispecies
627:Dinmopsylla
615:Burmopsylla
537:Archipsylla
486:Orthopsocus
462:Permopsocus
356:Dichentomum
253:(including
1073:Categories
961:. 101358.
841:2115/61234
798:2020-05-01
671:References
607:Cenomanian
605:-earliest
391:Hypopsylla
375:Artinskian
276:Cenomanian
167:Arthropoda
1014:Q27096119
893:1175-5334
850:1175-5334
780:1175-5334
723:2045-2322
549:Callovian
383:Kungurian
309:Hemiptera
265:. Within
251:bark lice
243:Moscovian
210:Families
153:Kingdom:
147:Eukaryota
1008:Wikidata
975:38704966
940:31715767
901:27394792
858:27396002
792:Archived
788:29689945
741:26961785
545:Daohugou
518:Eopsylla
422:Toarcian
367:Asselian
327:Taxonomy
319:Head of
307:(c) and
301:Psocodea
163:Phylum:
157:Animalia
143:Domain:
920:Zootaxa
873:Zootaxa
818:Zootaxa
760:Zootaxa
732:4785345
703:Bibcode
529:Roadian
193:Order:
177:Insecta
173:Class:
1043:337717
973:
938:
899:
891:
856:
848:
786:
778:
739:
729:
721:
638:Norian
603:Albian
572:Aptian
263:thrips
1051:Plazi
971:PMID
936:PMID
924:4576
897:PMID
889:ISSN
877:4105
854:PMID
846:ISSN
822:4079
784:PMID
776:ISSN
764:4382
737:PMID
719:ISSN
303:(a)
261:and
259:bugs
255:lice
46:Preęž’
963:doi
928:doi
881:doi
836:hdl
826:doi
768:doi
727:PMC
711:doi
551:),
369:)
311:(d)
257:),
1075::
1053::
1040::
1025::
1010::
969:.
959:80
957:.
934:.
922:.
895:.
887:.
875:.
852:.
844:.
834:.
820:.
816:.
790:.
782:.
774:.
762:.
758:.
735:.
725:.
717:.
709:.
697:.
693:.
679:^
385:),
377:)
96:Pg
977:.
965::
942:.
930::
903:.
883::
860:.
838::
828::
801:.
770::
743:.
713::
705::
699:6
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407:â€
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