305:
285:
124:
102:
221:
262:
in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the
Permian to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous
678:
Huang, Di-Ying; Bechly, GĂĽnter; Nel, Patricia; Engel, Michael S.; Prokop, Jakub; Azar, Dany; Cai, Chen-Yang; van de Kamp, Thomas; Staniczek, Arnold H.; Garrouste, Romain; Krogmann, Lars (March 2016).
258:
it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late
Carboniferous. The group was first named as a suborder by
275:
position of the group. The morphology of the mouthparts suggests that they were capable of suction feeding and chewing, with preserved angiosperm pollen grains in the gut of
680:"New fossil insect order Permopsocida elucidates major radiation and evolution of suction feeding in hemimetabolous insects (Hexapoda: Acercaria)"
860:
Liang, F.; Zhang, W.; Liu, X. (2016). "A new genus and species of the paraneopteran family
Archipsyllidae in mid-Cretaceous amber of Myanmar".
780:
267:). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in
942:
Prokop, J.; Rosová, K.; Pecharová, M.; Sroka, P.; Leipner, A.; Nel, A. (2024). "The structure of wing in the earliest
Permopsocida".
745:"A new genus and species of the family Archipsyllidae (Insecta: Paraneoptera: Permopsocida) from the Late Triassic of Queensland"
622:
1067:
123:
231:
304:
284:
513:
110:
803:"Bridging the gap between chewing and sucking in the hemipteroid insects:new insights from Cretaceous amber"
386:
979:
367:
1072:
1026:
406:
1044:
691:
568:
541:
418:
359:
259:
549:
462:
430:
646:
580:
486:
332:
277:
634:
398:
615:
603:
525:
474:
450:
344:
118:
1031:
379:
1017:
959:
924:
885:
877:
842:
834:
772:
764:
725:
707:
506:
951:
916:
869:
824:
814:
756:
744:
715:
699:
351:
695:
101:
720:
679:
591:
556:
1061:
587:
390:
272:
268:
59:
293:
255:
235:
175:
1002:
1011:
907:—a new bizarre genus of †Permopsocida (Insecta) from Burmese Cretaceous amber".
34:
955:
920:
873:
819:
802:
760:
595:
363:
264:
79:
44:
881:
838:
768:
711:
537:
371:
297:
247:
239:
155:
135:
84:
963:
928:
889:
846:
776:
729:
220:
996:
533:
410:
355:
289:
74:
69:
54:
49:
39:
281:
suggesting that at least some members of the group were pollenivorous.
517:
89:
64:
829:
703:
626:
560:
251:
165:
145:
973:
288:
Comparison of the head morphology of
Permopsocida (b) with that of
1039:
303:
283:
219:
243:
977:
230:
903:
Beutel, R. G.; Prokop, J.; MĂĽller, P.; Pohl, H. (2019). "â€
339:
OsnabrĂĽck
Formation, Germany, Carboniferous (Moscovian)
986:
437:Croudace Bay Formation, Australia, Changhsingian
425:Croudace Bay Formation, Australia, Changhsingian
320:From Huang et al 2016 and subsequent literature
271:, which has helped clarify the morphology and
8:
974:
100:
20:
828:
818:
719:
493:Wellington Formation, Kansas, Artinskian
481:Wellington Formation, Kansas, Artinskian
469:Wellington Formation, Kansas, Artinskian
457:Wellington Formation, Kansas, Artinskian
665:
234:) to the Mid-Cretaceous. It is part of
944:Arthropod Structure & Development
7:
1045:03E0821E-FFAD-FFD6-FF70-59CB83812620
801:Yoshizawa, K.; Lienhard, C. (2016).
673:
671:
669:
362:, Kansas, Oklahoma, Early Permian (
590:, Myanmar, mid Cretaceous (latest
532:"Green Series", Germany, Toarcian
14:
384:Prokop, Garrouste & Nel, 2016
743:Lambkin, Kevin J. (2018-02-20).
122:
783:from the original on 2020-02-25
639:Yoshizawa & Lienhard, 2016
575:Zaza Formation, Russia, Aptian
1:
354:, New Mexico, Early Permian (
625:, Australia, Late Triassic (
608:Liang, Zhang & Liu, 2016
559:, Russia, Early Cretaceous (
409:", Germany, Early Jurassic (
544:, Kazakhstan, Late Jurassic
389:, Australia, Late Permian (
1089:
536:, China, Middle Jurassic (
516:, Russia, Middle Permian (
956:10.1016/j.asd.2024.101358
921:10.11646/zootaxa.4576.2.9
874:10.11646/zootaxa.4105.5.4
820:10.11646/zootaxa.4079.2.5
761:10.11646/zootaxa.4382.1.9
370:, Russia, Early Permian (
203:
198:
119:Scientific classification
117:
108:
99:
23:
16:Extinct genus of insects
514:Iva-Gora Beds Formation
111:Psocorrhyncha burmitica
653:Burmese amber, Myanmar
641:Burmese amber, Myanmar
610:Burmese amber, Myanmar
387:Croudace Bay Formation
312:
301:
224:
1068:Extinct insect orders
1027:Paleobiology Database
307:
287:
223:
623:Blackstone Formation
542:Karabastau Formation
368:Koshelevka Formation
360:Wellington Formation
260:Robert John Tillyard
30:Moscovian–Cenomanian
696:2016NatSR...623004H
337:Prokop et al., 2024
684:Scientific Reports
585:Beutel et al. 2019
313:
302:
225:
1055:
1054:
980:Taxon identifiers
704:10.1038/srep23004
652:
651:Huang et al. 2016
640:
621:
609:
586:
574:
573:Vishniakova, 1976
555:
554:Vishniakova, 1976
531:
512:
501:
492:
480:
468:
456:
445:
436:
424:
404:
385:
350:
338:
327:
218:
217:
194:
1080:
1048:
1047:
1035:
1034:
1022:
1021:
1020:
1007:
1006:
1005:
975:
968:
967:
939:
933:
932:
900:
894:
893:
857:
851:
850:
832:
822:
798:
792:
791:
789:
788:
740:
734:
733:
723:
675:
650:
638:
619:
607:
584:
572:
553:
530:Handlirsch, 1908
529:
511:Vishnyakova 1976
510:
500:Handlirsch, 1906
499:
498:†Archipsyllidae
490:
478:
466:
454:
443:
434:
422:
402:
383:
352:Bursum Formation
348:
336:
325:
192:
187:
127:
126:
104:
94:
31:
27:Temporal range:
21:
1088:
1087:
1083:
1082:
1081:
1079:
1078:
1077:
1058:
1057:
1056:
1051:
1043:
1038:
1030:
1025:
1016:
1015:
1010:
1001:
1000:
995:
982:
972:
971:
941:
940:
936:
902:
901:
897:
859:
858:
854:
800:
799:
795:
786:
784:
742:
741:
737:
677:
676:
667:
662:
569:Archipsyllopsis
442:†Permopsocidae
419:Austropsocidium
318:
212:†Archipsyllidae
191:
185:
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
1086:
1084:
1076:
1075:
1070:
1060:
1059:
1053:
1052:
1050:
1049:
1036:
1023:
1008:
992:
990:
984:
983:
978:
970:
969:
934:
915:(2): 357–366.
895:
852:
793:
755:(1): 192–194.
735:
664:
663:
661:
658:
657:
656:
655:
654:
642:
630:
611:
599:
576:
564:
557:Zaza Formation
550:Archipsyllodes
545:
521:
496:
495:
494:
482:
479:Carpenter 1932
470:
467:Carpenter 1932
463:Lithopsocidium
458:
455:Tillyard, 1926
444:Tillyard, 1926
440:
439:
438:
431:Stenopsocidium
426:
414:
394:
375:
349:Tillyard, 1926
340:
326:Tillyard, 1926
317:
314:
216:
215:
214:
213:
210:
209:†Permopsocidae
207:
201:
200:
196:
195:
193:Tillyard, 1926
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
115:
114:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
32:
26:
15:
13:
10:
9:
6:
4:
3:
2:
1085:
1074:
1071:
1069:
1066:
1065:
1063:
1046:
1041:
1037:
1033:
1028:
1024:
1019:
1013:
1009:
1004:
998:
994:
993:
991:
989:
985:
981:
976:
965:
961:
957:
953:
949:
945:
938:
935:
930:
926:
922:
918:
914:
910:
906:
905:Bittacopsocus
899:
896:
891:
887:
883:
879:
875:
871:
868:(5): 483–90.
867:
863:
856:
853:
848:
844:
840:
836:
831:
826:
821:
816:
813:(2): 229–45.
812:
808:
804:
797:
794:
782:
778:
774:
770:
766:
762:
758:
754:
750:
746:
739:
736:
731:
727:
722:
717:
713:
709:
705:
701:
697:
693:
689:
685:
681:
674:
672:
670:
666:
659:
649:
648:
647:Psocorrhyncha
643:
637:
636:
631:
628:
624:
620:Lambkin, 2018
618:
617:
612:
606:
605:
600:
597:
593:
589:
588:Burmese amber
583:
582:
581:Bittacopsocus
577:
571:
570:
565:
562:
558:
552:
551:
546:
543:
539:
535:
528:
527:
522:
519:
515:
509:
508:
503:
502:
497:
491:Tillyard 1926
489:
488:
487:Progonopsocus
483:
477:
476:
471:
465:
464:
459:
453:
452:
447:
446:
441:
435:Tillyard 1935
433:
432:
427:
423:Tillyard 1935
421:
420:
415:
412:
408:
403:Ansorge, 1996
401:
400:
395:
392:
391:Changhsingian
388:
382:
381:
376:
373:
369:
365:
361:
357:
353:
347:
346:
341:
335:
334:
333:Carbonopsocus
329:
328:
324:†Psocidiidae
323:
322:
321:
315:
311:
310:Psocorrhyncha
306:
299:
295:
291:
286:
282:
280:
279:
278:Psocorrhyncha
274:
270:
269:Burmese amber
266:
261:
257:
253:
249:
245:
241:
237:
233:
229:
222:
211:
208:
205:
204:
202:
197:
190:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
113:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
1073:Paraneoptera
1018:Permopsocida
988:Permopsocida
987:
947:
943:
937:
912:
908:
904:
898:
865:
861:
855:
810:
806:
796:
785:. Retrieved
752:
748:
738:
690:(1): 23004.
687:
683:
645:
635:Mydiognathus
633:
614:
602:
579:
567:
548:
524:
505:
485:
473:
461:
449:
429:
417:
407:Green Series
399:Liassopsocus
397:
378:
343:
331:
319:
309:
294:Thysanoptera
276:
273:phylogenetic
256:Paraneoptera
238:, alongside
236:Paraneoptera
228:Permopsocida
227:
226:
206:†Psocidiidae
189:Permopsocida
188:
176:Paraneoptera
172:(unranked):
109:
24:Permopsocida
18:
1012:Wikispecies
616:Dinmopsylla
604:Burmopsylla
526:Archipsylla
475:Orthopsocus
451:Permopsocus
345:Dichentomum
242:(including
1062:Categories
950:. 101358.
830:2115/61234
787:2020-05-01
660:References
596:Cenomanian
594:-earliest
380:Hypopsylla
364:Artinskian
265:Cenomanian
156:Arthropoda
1003:Q27096119
882:1175-5334
839:1175-5334
769:1175-5334
712:2045-2322
538:Callovian
372:Kungurian
298:Hemiptera
254:. Within
240:bark lice
232:Moscovian
199:Families
142:Kingdom:
136:Eukaryota
997:Wikidata
964:38704966
929:31715767
890:27394792
847:27396002
781:Archived
777:29689945
730:26961785
534:Daohugou
507:Eopsylla
411:Toarcian
356:Asselian
316:Taxonomy
308:Head of
296:(c) and
290:Psocodea
152:Phylum:
146:Animalia
132:Domain:
909:Zootaxa
862:Zootaxa
807:Zootaxa
749:Zootaxa
721:4785345
692:Bibcode
518:Roadian
182:Order:
166:Insecta
162:Class:
1032:337717
962:
927:
888:
880:
845:
837:
775:
767:
728:
718:
710:
627:Norian
592:Albian
561:Aptian
252:thrips
1040:Plazi
960:PMID
925:PMID
913:4576
886:PMID
878:ISSN
866:4105
843:PMID
835:ISSN
811:4079
773:PMID
765:ISSN
753:4382
726:PMID
708:ISSN
292:(a)
250:and
248:bugs
244:lice
35:Preęž’
952:doi
917:doi
870:doi
825:hdl
815:doi
757:doi
716:PMC
700:doi
540:),
358:)
300:(d)
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1064::
1042::
1029::
1014::
999::
958:.
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946:.
923:.
911:.
884:.
876:.
864:.
841:.
833:.
823:.
809:.
805:.
779:.
771:.
763:.
751:.
747:.
724:.
714:.
706:.
698:.
686:.
682:.
668:^
374:),
366:)
85:Pg
966:.
954::
931:.
919::
892:.
872::
849:.
827::
817::
790:.
759::
732:.
702::
694::
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