129:
750:
3439:
891:
3445:
4063:
3995:
147:
4025:
852:
899:
1748:
1677:
4001:
4051:
1623:
609:
691:. Some of these beds extend for hundreds of meters and contain thousands of individuals of all ages and all levels of completion. Scientists have speculated that the high density and number of individuals would be explained if they had perished while trying to cross a flooded river. A discovery of thousands of
867:
bore a single large horn over their noses. These horns curved forwards or backwards depending on the specimen. Skull ornamentation was reduced as animals aged. The frill was relatively short compared to the total skull length and could grow to over half a meter (68.8 cm) long in the oldest and
1687:
The large frills and nasal horns of the ceratopsians are among the most distinctive facial adornments of all dinosaurs. Their function has been the subject of debate since the first horned dinosaurs were discovered. Common theories concerning the function of ceratopsian frills and horns include
1767:
dinosaurs in the region. Large herbivores like the ceratopsians living in North
America during the Late Cretaceous had "remarkably small geographic ranges" despite their large body size and high mobility. This restricted distribution strongly contrasts with modern
2661:
Ekhtiari, S.; Chiba, K.; Popovic, S.; Crowther, R.; Wohl, G.; Wong, A. K. O.; Tanke, D. H.; Dufault, D. M.; Geen, O. D.; Parasu, N.; Crowther, M. A.; Evans, D. C. (2020). "First case of osteosarcoma in a dinosaur: a multimodal diagnosis".
1882:"Centrosaurus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B.
128:
862:
were large dinosaurs, although not as large as some of their relatives, reaching 5–5.5 metres (16–18 ft) long and 2–2.5 metric tons (2.2–2.8 short tons) in body mass situated atop stocky limbs. Like other centrosaurines,
667:, "lizard") and refers to, the series of small hornlets placed along the margin of their frills, not the nasal horns (which were unknown when the dinosaur was named). The genus is not to be confused with the
1730:
had reached an aggressive stage. The cancer would have resulted in a severe limp that would have made the ceratopsian more vulnerable to predation. However, the fact that it was part of a herd allowed the
1735:
to survive much longer than would be expected for an animal infected with such severe disease. The individual itself is believed to have died from drowning in the flash flood that created the
710:
Because of the variation between species and even individual specimens of centrosaurines, there has been much debate over which genera and species are valid, particularly whether
749:
4190:
2709:
Lehman, T. M., 2001, Late
Cretaceous dinosaur provinciality: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 310-328.
2551:
Rogers, R. R. (1990). "Taphonomy of three dinosaur bone beds in the Upper
Cretaceous Two Medicine Formation, northwestern Montana: Evidence for drought-related mortality".
2380:
Ryan, M.J.; A.P. Russell (2005). "A new centrosaurine ceratopsid from the Oldman
Formation of Alberta and its implications for centrosaurine taxonomy and systematics".
1698:
skull lesions found that bone injuries on the skulls were more likely caused by intraspecific combat (horn-to-horn combat) rather than predatory attacks. The frills of
874:
is distinguished by having two large hornlets which hook forwards over the frill. A pair of small upwards directed horns is also found over the eyes. The frills of
926:. It so closely resembles the latter of these that some paleontologists have considered them to represent the same animal. Other members of the Centrosaurinae
4139:
2272:
Benson, Roger B. J.; Campione, Nicolás E.; Carrano, Matthew T.; Mannion, Philip D.; Sullivan, Corwin; Upchurch, Paul; Evans, David C. (2014-05-06).
4225:
4205:
4220:
4210:
3438:
2256:
1907:
1657:
in what is now
Alberta. The mass deaths may have been caused by otherwise non-herding animals gathering around a waterhole during a drought.
2760:
3444:
2471:
Kentaro Chiba; Michael J. Ryan; Federico Fanti; Mark A. Loewen; David C. Evans (2018). "New material and systematic re-evaluation of
2535:
2359:
2066:"Craniofacial ontogeny in centrosaurine dinosaurs (Ornithischia: Ceratopsidae): taphonomic and behavioral phylogenetic implications"
1965:
1932:
1891:
534:
4200:
4018:
1781:
2274:"Rates of Dinosaur Body Mass Evolution Indicate 170 Million Years of Sustained Ecological Innovation on the Avian Stem Lineage"
2730:
2007:, with a preliminary analysis of ceratopsian phylogeny. Ph.D. Dissertation. University of Pennsylvania, Philadelphia. 227 pp.
4215:
4041:
4195:
3994:
146:
1480:
1645:
in Canada suggest that they were gregarious animals and could have traveled in large herds. A bone bed composed of
1521:
1429:
1197:
1180:
1149:
890:
4185:
1504:
1301:
2851:
2753:
1016:
684:
2449:
1739:
bone bed where it was found. The specimen remains one of the few dinosaur specimens found with severe cancer.
4180:
2969:
1654:
591:
2436:
Ryan, M.J. (2007). "A new basal centrosaurine ceratopsid from the Oldman
Formation, southeastern Alberta".
4079:
1334:
1223:
4152:
4028:
2878:
1456:
1375:
1103:
999:
338:
4000:
2096:
M.J. Ryan and D.C. Evans, 2005, "Ornithischian dinosaurs". In: P.J. Currie and E.B. Koppelhus (eds.),
4126:
2615:
2560:
2484:
2389:
2215:
2030:
1840:
1240:
1059:
703:, is believed to be the largest bed of dinosaur bones ever discovered. The area is now known as the
3134:
2746:
580:
1904:
3854:
3804:
3236:
2687:
2576:
2500:
2453:
2415:
2046:
1856:
1681:
1627:
1284:
879:
704:
603:
366:
141:
2527:
2519:
1957:
1949:
4157:
4117:
2351:
3937:
3875:
3838:
3312:
3199:
2679:
2643:
2531:
2355:
2313:
2295:
2252:
2184:
1961:
1928:
1887:
1702:
were too thin to be used for defense against predators, although the thicker, solid frills of
1415:
1045:
789:
798:, there is no firm evidence for sexual dimorphism in any ceratopsid. Others have synonymized
4055:
3967:
3765:
3590:
3292:
2987:
2671:
2633:
2623:
2568:
2492:
2445:
2405:
2397:
2343:
2303:
2285:
2223:
2174:
2164:
2077:
2038:
1848:
932:
894:
A large hook from the back of a
Centrosaurus frill. These hooks are diagnostic of the genus.
490:
1641:
were adapted to shear through tough plant material. The discovery of gigantic bone beds of
3861:
3813:
3772:
3708:
3611:
3562:
3264:
3188:
3143:
3017:
3003:
2962:
2836:
2820:
1911:
1824:
950:
584:
37:
2619:
2564:
2488:
2393:
2219:
2034:
1844:
3868:
3827:
3758:
3745:
3716:
3701:
3671:
3632:
3618:
3535:
3496:
3375:
3285:
3250:
3243:
3123:
3073:
2996:
2976:
2638:
2599:
2308:
2273:
2245:
2179:
2148:
2082:
2065:
1688:
defense from predators, combat within the species, and visual display. A 2009 study of
978:
696:
625:
621:
567:
357:
314:
294:
281:
265:
223:
2675:
1710:
were most likely used "for species recognition and/or other forms of visual display".
4174:
4067:
3946:
3925:
3895:
3886:
3786:
3680:
3625:
3604:
3576:
3569:
3555:
3548:
3354:
3343:
3336:
3278:
3271:
3257:
3215:
3150:
3116:
3087:
3040:
2691:
2504:
2457:
2419:
2344:
2050:
1827:; Burns, M. E.; Sissons, R. L. (2009). "A redescription of the ankylosaurid dinosaur
1263:
1253:
956:
916:
794:
776:. His assessments have been partially followed, with other researchers not accepting
637:
79:
2722:
1860:
914:
gives its name to the
Centrosaurinae subfamily. Its closest relatives appear to be
3960:
3911:
3820:
3793:
3779:
3687:
3660:
3653:
3646:
3597:
3583:
3475:
3462:
3393:
3382:
3367:
3329:
3321:
3303:
3208:
3174:
3157:
3109:
3102:
3058:
2863:
2811:
2595:
1723:
898:
839:
719:
718:
are valid genera, undiagnosable, or possibly members of the opposite sex. In 1996,
672:
570:
252:
207:
20:
2475:(Dinosauria, Ceratopsidae) from the Judith River Formation (Campanian, Montana)".
1726:
in its right fibula. Examination of the cancerous lesions in the bone suggest the
1722:
recovered from
Dinosaur Provincial Park in 1989 was discovered to have crippling
851:
2628:
2290:
4111:
3953:
3847:
3730:
3694:
3517:
3229:
3094:
1747:
1690:
1676:
1442:
944:
938:
922:
903:
755:
427:
406:
391:
376:
54:
3723:
3639:
3489:
3222:
3010:
2944:
2931:
2890:
2769:
2042:
1772:
faunas whose large herbivores' ranges "typical ... span much of a continent."
1634:
239:
99:
64:
4102:
2299:
1981:
Dodson, P. "Quantitative aspects of relative growth and sexual dimorphism in
1794:
Lambe, L. M. (1904). "On the squamoso-parietal crest of the horned dinosaurs
2204:"A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta"
2017:
Lehman, T. M. (1998). "A gigantic skull and skeleton of the horned dinosaur
1769:
1764:
964:
927:
668:
577:
573:
158:
104:
48:
41:
2683:
2647:
2317:
2188:
2110:
Russell, L.S. (1930). "Upper
Cretaceous dinosaur faunas of North America".
2734:, from the Dinosaur Encyclopaedia at Univ. of Illinois at Urbana-Champaign
1622:
608:
4096:
3524:
3510:
3503:
3066:
2802:
2793:
1852:
191:
178:
94:
89:
74:
69:
59:
2496:
4144:
3918:
3080:
3024:
2580:
1831:
Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus".
1760:
700:
688:
629:
109:
84:
2410:
2228:
2203:
2169:
2787:
1727:
587:
168:
4073:
2572:
2401:
2524:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
2098:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
1954:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
1927:. Princeton University Press: Princeton, New Jersey, pp. 197–199.
1746:
1675:
1621:
897:
889:
850:
748:
607:
563:
2003:
Forster, C. A. (1990). The cranial morphology and systematics of
4131:
4077:
3460:
2929:
2780:
2742:
1905:
http://www.edmontonsun.com/news/canada/2010/06/18/14439211.html
1903:
Scientists find dino deathbed, signs of carnage. Edmonton Sun.
2738:
2520:"Ceratopsian bonebeds: occurrence, origins, and significance"
2202:
Ryan, M. J.; Evans, D. C.; Shepherd, K. M.; Sues, H. (2012).
2142:
2140:
2138:
662:
655:
648:
641:
1878:
1876:
1874:
1872:
1870:
967:
presented below represents a phylogenetic analysis by Chiba
520:
1804:
Proceedings and Transactions of the Royal Society of Canada
523:
511:
499:
2450:
10.1666/0022-3360(2007)81[376:ANBCCF]2.0.CO;2
1706:
might have evolved to protect their necks. The frills of
676:, the name of which is derived from the same Greek word.
2147:
Frederickson, J. A.; Tumarkin-Deratzian, A. R. (2014).
814:. It has also been suggested as the direct ancestor of
4039:
2334:
Dodson, P. (1990). "On the status of the ceratopsids
535:
526:
517:
502:
496:
493:
620:
remains were discovered and named by paleontologist
508:
505:
4086:
3936:
3894:
3885:
3837:
3803:
3744:
3670:
3534:
3473:
3392:
3353:
3311:
3302:
3198:
3172:
3133:
3038:
2986:
2942:
2889:
2862:
2835:
2559:(5). SEPM Society for Sedimentary Geology: 394–41.
1680:Restoration of a herd swimming, as hypothesised by
514:
2526:. Bloomington: Indiana University Press. pp.
2350:. Cambridge: Cambridge University Press. pp.
2244:
2100:. Indiana University Press, Bloomington pp 312-348
1956:. Bloomington: Indiana University Press. pp.
2346:Dinosaur Systematics: Perspectives and Approaches
2112:Proceedings of the American Philosophical Society
2522:. In Currie, Phillip J.; Koppelhus, Eva (eds.).
2375:
2373:
2371:
2064:Sampson, S. D.; Ryan, M.J.; Tanke, D.H. (1997).
1952:. In Currie, Phillip J.; Koppelhus, Eva (eds.).
837:, described in 2005, was moved to the new genus
792:has been proposed for a more basal ceratopsian,
1759:fossils haven't been found outside of southern
594:, dating from 76.5 to 75.5 million years ago.
2754:
1763:even though they are among the most abundant
8:
2518:Eberth, David A.; Getty, Michael A. (2005).
136:Skull ROM 767 from Dinosaur Provincial Park
2329:
2327:
1886:. Publications International, LTD. p. 135.
818:. A 2014 study of changes during growth in
547:
4191:Late Cretaceous dinosaurs of North America
4074:
3891:
3750:
3540:
3481:
3470:
3457:
3359:
3308:
3180:
3050:
3046:
2954:
2950:
2939:
2926:
2868:
2841:
2832:
2777:
2761:
2747:
2739:
2342:". In Carpenter, K.; Currie, P.J. (eds.).
1948:Ryan, Michael J.; Evans, David C. (2005).
882:and small hornlets along the outer edges.
127:
27:
2705:
2703:
2701:
2637:
2627:
2431:
2429:
2409:
2307:
2289:
2227:
2178:
2168:
2081:
2070:Zoological Journal of the Linnean Society
878:were moderately long, with fairly large
4046:
1925:The Horned Dinosaurs: A Natural History
1816:
590:. Their remains have been found in the
2130:The Princeton Field Guide to Dinosaurs
1943:
1941:
1673:as the environment changed over time.
830:, representing a middle growth stage.
1661:is found lower in the formation than
7:
734:to warrant separate genera and that
2132:, Princeton University Press p. 261
1755:Thomas M. Lehman has observed that
1496:
1472:
1448:
1421:
1350:
1326:
1276:
1269:
1259:
1172:
1165:
1141:
1051:
1041:
991:
984:
974:
742:more closely than either resembled
469:
463:
451:
445:
433:
418:
412:
397:
382:
2382:Canadian Journal of Earth Sciences
2247:Princeton Field Guide to Dinosaurs
2208:Canadian Journal of Earth Sciences
2083:10.1111/j.1096-3642.1997.tb00340.x
1833:Journal of Vertebrate Paleontology
772:, may actually have been a female
14:
1802:from the Cretaceous of Alberta".
4061:
4049:
4024:
4023:
3999:
3993:
3443:
3437:
2727:at The Dinosaur Picture Database
1782:Timeline of ceratopsian research
489:
145:
764:Dodson believed one species of
722:found enough variation between
2251:. Princeton University Press.
806:, or considered it a separate
1:
4226:Ceratopsians of North America
4206:Fossil taxa described in 1904
2676:10.1016/S1470-2045(20)30171-6
636:means "pointed lizard" (from
4221:Late Cretaceous ceratopsians
4211:Taxa named by Lawrence Lambe
2629:10.1371/journal.pone.0004252
2594:Farke, A.A.; Wolff, E.D.S.;
2291:10.1371/journal.pbio.1001853
1751:Skin impression of AMNH 5427
902:Complete skulls arranged in
656:
642:
1481:Pachyrhinosaurus canadensis
4242:
2149:"Craniofacial ontogeny in
1829:Dyoplosaurus acutosquameus
1653:remains is known from the
1522:Pachyrhinosaurus perotorum
1430:Einiosaurus procurvicornis
1198:Wendiceratops pinhornensis
1181:Sinoceratops zhuchengensis
1150:Xenoceratops foremostensis
855:Size comparison with human
663:
649:
601:
18:
16:Extinct genus of dinosaurs
4013:
3991:
3753:
3543:
3484:
3469:
3456:
3434:
3362:
3183:
3053:
3049:
2957:
2953:
2938:
2925:
2871:
2844:
2831:
2776:
2243:Paul, Gregory S. (2010).
2043:10.1017/S0022336000027220
1950:"Ornithischian Dinosaurs"
1518:
1505:Pachyrhinosaurus lakustai
1501:
1494:
1477:
1470:
1453:
1446:
1426:
1419:
1372:
1355:
1348:
1331:
1324:
1302:Styracosaurus albertensis
1298:
1281:
1274:
1267:
1257:
1237:
1220:
1194:
1177:
1170:
1163:
1146:
1139:
1117:
1100:
1087:
1074:
1056:
1049:
1039:
1013:
996:
989:
982:
816:Styracosaurus albertensis
695:fossils near the town of
372:
365:
344:
337:
142:Scientific classification
140:
135:
126:
30:
2019:Pentaceratops sternbergi
1017:Machairoceratops cronusi
788:as a valid genus. While
685:Dinosaur Provincial Park
679:Later, vast bonebeds of
19:Not to be confused with
4201:Paleontology in Alberta
2970:Micropachycephalosaurus
2477:Journal of Paleontology
2438:Journal of Paleontology
2023:Journal of Paleontology
1987:Journal of Paleontology
1655:Dinosaur Park Formation
826:is a junior synonym of
812:Centrosaurus nasicornus
592:Dinosaur Park Formation
2602:Evidence of Combat in
1800:Monoclonius canadensis
1752:
1684:
1630:
1335:Coronosaurus brinkmani
1224:Albertaceratops nesmoi
907:
895:
856:
778:Monoclonius nasicornus
761:
613:
460:Eucentrosaurus apertus
4153:Paleobiology Database
1750:
1679:
1625:
1457:Achelousaurus horneri
1376:Spinops sternbergorum
1104:Nasutoceratops titusi
1000:Diabloceratops eatoni
901:
893:
854:
752:
611:
4216:Ornithischian genera
2473:Medusaceratops lokii
2151:Centrosaurus apertus
1914:retrieved 18 06 2010
1884:The Age of Dinosaurs
1853:10.1671/039.029.0405
1796:Centrosaurus apertus
1720:Centrosaurus apertus
1359:Centrosaurus apertus
1241:Medusaceratops lokii
1060:Avaceratops lammersi
624:in strata along the
598:Discovery and naming
471:Chure & McIntosh
351:Centrosaurus apertus
4196:Dinosaur Park fauna
3135:Archaeoceratopsidae
2664:The Lancet Oncology
2620:2009PLoSO...4.4252F
2565:1990Palai...5..394R
2497:10.1017/jpa.2017.62
2489:2018JPal...92..272C
2394:2005CaJES..42.1369R
2220:2012CaJES..49.1251R
2035:1998JPal...72..894L
1923:Dodson, P. (1996).
1845:2009JVPal..29.1117A
1825:Arbour, V.M., V. M.
1626:Cast of AMNH 5427,
442:Centrosaurus flexus
3855:Crittendenceratops
3805:Albertaceratopsini
2128:Paul, G.S., 2010,
2021:from New Mexico".
1910:2010-06-22 at the
1753:
1685:
1665:, indicating that
1631:
1628:Museum of Victoria
1285:Rubeosaurus ovatus
908:
896:
857:
762:
705:Hilda mega-bonebed
614:
604:Hilda mega-bonebed
4168:
4167:
4080:Taxon identifiers
4037:
4036:
4009:
4008:
3989:
3988:
3985:
3984:
3981:
3980:
3977:
3976:
3938:Pachyrhinosaurini
3876:Yehuecauhceratops
3839:Nasutoceratopsini
3740:
3739:
3452:
3451:
3432:
3431:
3428:
3427:
3424:
3423:
3420:
3419:
3416:
3415:
3412:
3411:
3313:Protoceratopsidae
3200:Leptoceratopsidae
3168:
3167:
3034:
3033:
2921:
2920:
2917:
2916:
2913:
2912:
2909:
2908:
2258:978-0-691-13720-9
2229:10.1139/e2012-056
2170:10.7717/peerj.252
1743:Paleobiogeography
1669:was displaced by
1614:
1613:
1605:
1604:
1596:
1595:
1587:
1586:
1578:
1577:
1569:
1568:
1560:
1559:
1551:
1550:
1542:
1541:
1533:
1532:
1416:Pachyrhinosaurini
1405:
1404:
1396:
1395:
1387:
1386:
1313:
1312:
1209:
1208:
1128:
1127:
1046:Nasutoceratopsini
1028:
1027:
790:sexual dimorphism
480:
479:
474:
468:
456:
450:
438:
423:
417:
402:
387:
318:
4233:
4186:Campanian genera
4161:
4160:
4148:
4147:
4135:
4134:
4122:
4121:
4120:
4107:
4106:
4105:
4075:
4066:
4065:
4064:
4054:
4053:
4052:
4045:
4027:
4026:
4004:
4003:
3997:
3968:Pachyrhinosaurus
3892:
3766:Machairoceratops
3751:
3591:Coahuilaceratops
3541:
3482:
3471:
3458:
3447:
3442:
3441:
3360:
3309:
3293:Zhuchengceratops
3181:
3051:
3047:
2988:Chaoyangsauridae
2955:
2951:
2940:
2927:
2869:
2842:
2833:
2826:
2825:
2778:
2763:
2756:
2749:
2740:
2710:
2707:
2696:
2695:
2670:(8): 1021−1022.
2658:
2652:
2651:
2641:
2631:
2591:
2585:
2584:
2548:
2542:
2541:
2515:
2509:
2508:
2468:
2462:
2461:
2433:
2424:
2423:
2413:
2388:(7): 1369–1387.
2377:
2366:
2365:
2349:
2331:
2322:
2321:
2311:
2293:
2269:
2263:
2262:
2250:
2240:
2234:
2233:
2231:
2199:
2193:
2192:
2182:
2172:
2144:
2133:
2126:
2120:
2119:
2107:
2101:
2094:
2088:
2087:
2085:
2061:
2055:
2054:
2014:
2008:
2001:
1995:
1994:
1978:
1972:
1971:
1945:
1936:
1921:
1915:
1901:
1895:
1880:
1865:
1864:
1839:(4): 1117–1135.
1821:
1811:
1497:
1473:
1449:
1422:
1351:
1327:
1277:
1270:
1260:
1173:
1166:
1142:
1119:Malta new taxon
1052:
1042:
992:
985:
975:
933:Pachyrhinosaurus
868:largest adults.
666:
665:
659:
652:
651:
645:
612:Life restoration
561:
558:
555:
552:
549:
542:
538:
533:
532:
529:
528:
525:
522:
519:
516:
513:
510:
507:
504:
501:
498:
495:
472:
466:
454:
448:
436:
421:
415:
400:
385:
353:
349:
313:
306:
293:
280:
264:
251:
238:
222:
206:
150:
149:
131:
121:
51:
36:Temporal range:
28:
4241:
4240:
4236:
4235:
4234:
4232:
4231:
4230:
4171:
4170:
4169:
4164:
4156:
4151:
4143:
4138:
4130:
4125:
4116:
4115:
4110:
4101:
4100:
4095:
4082:
4072:
4062:
4060:
4050:
4048:
4040:
4038:
4033:
4005:
3998:
3973:
3932:
3881:
3862:Furcatoceratops
3833:
3814:Albertaceratops
3799:
3773:Menefeeceratops
3736:
3709:Tatankaceratops
3666:
3612:Mercuriceratops
3563:Arrhinoceratops
3530:
3478:
3465:
3448:
3436:
3408:
3388:
3349:
3298:
3265:Montanoceratops
3194:
3189:Graciliceratops
3177:
3164:
3144:Archaeoceratops
3129:
3043:
3030:
3018:Xuanhuaceratops
3004:Hualianceratops
2982:
2963:Albalophosaurus
2947:
2934:
2905:
2885:
2858:
2852:Avemetatarsalia
2837:Avemetatarsalia
2827:
2821:Marginocephalia
2783:
2782:
2772:
2767:
2719:
2714:
2713:
2708:
2699:
2660:
2659:
2655:
2593:
2592:
2588:
2573:10.2307/3514834
2550:
2549:
2545:
2538:
2517:
2516:
2512:
2470:
2469:
2465:
2435:
2434:
2427:
2402:10.1139/e05-029
2379:
2378:
2369:
2362:
2333:
2332:
2325:
2284:(5): e1001853.
2271:
2270:
2266:
2259:
2242:
2241:
2237:
2201:
2200:
2196:
2146:
2145:
2136:
2127:
2123:
2109:
2108:
2104:
2095:
2091:
2063:
2062:
2058:
2016:
2015:
2011:
2002:
1998:
1980:
1979:
1975:
1968:
1947:
1946:
1939:
1922:
1918:
1912:Wayback Machine
1902:
1898:
1881:
1868:
1823:
1822:
1818:
1793:
1790:
1778:
1745:
1716:
1620:
1615:
1606:
1597:
1588:
1579:
1570:
1561:
1552:
1543:
1534:
1406:
1397:
1388:
1314:
1210:
1129:
1029:
951:Albertaceratops
888:
849:
822:concluded that
606:
600:
585:Late Cretaceous
559:
556:
553:
550:
540:
536:
492:
488:
470:
464:
462:
452:
446:
444:
434:
432:
419:
413:
411:
398:
396:
383:
381:
361:
355:
347:
346:
333:
330:C. apertus
312:
304:
291:
278:
262:
249:
236:
220:
204:
144:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
47:76.5–75.5
46:
45:
38:Late Cretaceous
34:
24:
17:
12:
11:
5:
4239:
4237:
4229:
4228:
4223:
4218:
4213:
4208:
4203:
4198:
4193:
4188:
4183:
4181:Centrosaurines
4173:
4172:
4166:
4165:
4163:
4162:
4149:
4136:
4123:
4108:
4092:
4090:
4084:
4083:
4078:
4071:
4070:
4058:
4035:
4034:
4032:
4031:
4021:
4014:
4011:
4010:
4007:
4006:
3992:
3990:
3987:
3986:
3983:
3982:
3979:
3978:
3975:
3974:
3972:
3971:
3964:
3957:
3950:
3942:
3940:
3934:
3933:
3931:
3930:
3922:
3915:
3908:
3900:
3898:
3889:
3883:
3882:
3880:
3879:
3872:
3869:Nasutoceratops
3865:
3858:
3851:
3843:
3841:
3835:
3834:
3832:
3831:
3828:Medusaceratops
3824:
3817:
3809:
3807:
3801:
3800:
3798:
3797:
3790:
3783:
3776:
3769:
3762:
3759:Diabloceratops
3754:
3748:
3746:Centrosaurinae
3742:
3741:
3738:
3737:
3735:
3734:
3727:
3720:
3717:Titanoceratops
3713:
3705:
3702:Regaliceratops
3698:
3691:
3684:
3676:
3674:
3672:Triceratopsini
3668:
3667:
3665:
3664:
3657:
3650:
3643:
3636:
3633:Sierraceratops
3629:
3622:
3619:Navajoceratops
3615:
3608:
3601:
3594:
3587:
3580:
3573:
3566:
3559:
3552:
3544:
3538:
3536:Chasmosaurinae
3532:
3531:
3529:
3528:
3521:
3514:
3507:
3500:
3497:Brachyceratops
3493:
3485:
3479:
3474:
3467:
3466:
3461:
3454:
3453:
3450:
3449:
3435:
3433:
3430:
3429:
3426:
3425:
3422:
3421:
3418:
3417:
3414:
3413:
3410:
3409:
3407:
3406:
3405:
3404:
3398:
3396:
3390:
3389:
3387:
3386:
3379:
3376:Turanoceratops
3372:
3363:
3357:
3351:
3350:
3348:
3347:
3340:
3333:
3326:
3317:
3315:
3306:
3300:
3299:
3297:
3296:
3289:
3286:Unescoceratops
3282:
3275:
3268:
3261:
3254:
3251:Ischioceratops
3247:
3244:Gryphoceratops
3240:
3233:
3226:
3219:
3212:
3204:
3202:
3196:
3195:
3193:
3192:
3184:
3178:
3173:
3170:
3169:
3166:
3165:
3163:
3162:
3154:
3147:
3139:
3137:
3131:
3130:
3128:
3127:
3124:Sasayamagnomus
3120:
3113:
3106:
3099:
3091:
3084:
3077:
3074:Auroraceratops
3070:
3063:
3054:
3044:
3039:
3036:
3035:
3032:
3031:
3029:
3028:
3021:
3014:
3007:
3000:
2997:Chaoyangsaurus
2992:
2990:
2984:
2983:
2981:
2980:
2977:Psittacosaurus
2973:
2966:
2958:
2948:
2943:
2936:
2935:
2930:
2923:
2922:
2919:
2918:
2915:
2914:
2911:
2910:
2907:
2906:
2904:
2903:
2902:
2901:
2895:
2893:
2887:
2886:
2884:
2883:
2882:
2881:
2872:
2866:
2860:
2859:
2857:
2856:
2855:
2854:
2845:
2839:
2829:
2828:
2824:
2823:
2814:
2805:
2796:
2790:
2781:
2774:
2773:
2768:
2766:
2765:
2758:
2751:
2743:
2737:
2736:
2728:
2718:
2717:External links
2715:
2712:
2711:
2697:
2653:
2586:
2543:
2536:
2510:
2483:(2): 272–288.
2463:
2444:(2): 376–396.
2425:
2367:
2360:
2323:
2264:
2257:
2235:
2194:
2134:
2121:
2102:
2089:
2076:(3): 293–337.
2056:
2029:(5): 894–906.
2009:
1996:
1973:
1966:
1937:
1916:
1896:
1866:
1815:
1814:
1813:
1812:
1789:
1786:
1785:
1784:
1777:
1774:
1744:
1741:
1718:A specimen of
1715:
1714:Paleopathology
1712:
1637:, the jaws of
1619:
1616:
1612:
1611:
1608:
1607:
1603:
1602:
1599:
1598:
1594:
1593:
1590:
1589:
1585:
1584:
1581:
1580:
1576:
1575:
1572:
1571:
1567:
1566:
1563:
1562:
1558:
1557:
1554:
1553:
1549:
1548:
1545:
1544:
1540:
1539:
1536:
1535:
1531:
1530:
1527:
1526:
1517:
1514:
1513:
1510:
1509:
1500:
1495:
1493:
1490:
1489:
1486:
1485:
1476:
1471:
1469:
1466:
1465:
1462:
1461:
1452:
1447:
1445:
1439:
1438:
1435:
1434:
1425:
1420:
1418:
1412:
1411:
1408:
1407:
1403:
1402:
1399:
1398:
1394:
1393:
1390:
1389:
1385:
1384:
1381:
1380:
1371:
1368:
1367:
1364:
1363:
1354:
1349:
1347:
1344:
1343:
1340:
1339:
1330:
1325:
1323:
1320:
1319:
1316:
1315:
1311:
1310:
1307:
1306:
1297:
1294:
1293:
1290:
1289:
1280:
1275:
1273:
1268:
1266:
1258:
1256:
1250:
1249:
1246:
1245:
1236:
1233:
1232:
1229:
1228:
1219:
1216:
1215:
1212:
1211:
1207:
1206:
1203:
1202:
1193:
1190:
1189:
1186:
1185:
1176:
1171:
1169:
1164:
1162:
1159:
1158:
1155:
1154:
1145:
1140:
1138:
1135:
1134:
1131:
1130:
1126:
1125:
1122:
1121:
1116:
1113:
1112:
1109:
1108:
1099:
1096:
1095:
1092:
1091:
1086:
1083:
1082:
1079:
1078:
1073:
1070:
1069:
1066:
1065:
1055:
1050:
1048:
1040:
1038:
1035:
1034:
1031:
1030:
1026:
1025:
1022:
1021:
1012:
1009:
1008:
1005:
1004:
995:
990:
988:
983:
981:
979:Centrosaurinae
973:
887:
886:Classification
884:
848:
845:
683:were found in
653:, "point" and
626:Red Deer River
622:Lawrence Lambe
599:
596:
557:pointed lizard
478:
477:
476:
475:
457:
439:
424:
403:
388:
370:
369:
363:
362:
356:
342:
341:
335:
334:
326:
324:
320:
319:
302:
298:
297:
289:
285:
284:
276:
269:
268:
266:Centrosaurinae
260:
256:
255:
247:
243:
242:
234:
227:
226:
224:Neornithischia
218:
211:
210:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
35:
15:
13:
10:
9:
6:
4:
3:
2:
4238:
4227:
4224:
4222:
4219:
4217:
4214:
4212:
4209:
4207:
4204:
4202:
4199:
4197:
4194:
4192:
4189:
4187:
4184:
4182:
4179:
4178:
4176:
4159:
4154:
4150:
4146:
4141:
4137:
4133:
4128:
4124:
4119:
4113:
4109:
4104:
4098:
4094:
4093:
4091:
4089:
4085:
4081:
4076:
4069:
4059:
4057:
4047:
4043:
4030:
4022:
4020:
4016:
4015:
4012:
4002:
3996:
3970:
3969:
3965:
3963:
3962:
3958:
3956:
3955:
3951:
3949:
3948:
3947:Achelousaurus
3944:
3943:
3941:
3939:
3935:
3928:
3927:
3926:Styracosaurus
3923:
3921:
3920:
3916:
3914:
3913:
3909:
3907:
3906:
3902:
3901:
3899:
3897:
3896:Centrosaurini
3893:
3890:
3888:
3887:Eucentrosaura
3884:
3878:
3877:
3873:
3871:
3870:
3866:
3864:
3863:
3859:
3857:
3856:
3852:
3850:
3849:
3845:
3844:
3842:
3840:
3836:
3830:
3829:
3825:
3823:
3822:
3818:
3816:
3815:
3811:
3810:
3808:
3806:
3802:
3796:
3795:
3791:
3789:
3788:
3787:Wendiceratops
3784:
3782:
3781:
3777:
3775:
3774:
3770:
3768:
3767:
3763:
3761:
3760:
3756:
3755:
3752:
3749:
3747:
3743:
3733:
3732:
3728:
3726:
3725:
3721:
3719:
3718:
3714:
3711:
3710:
3706:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3685:
3683:
3682:
3681:Eotriceratops
3678:
3677:
3675:
3673:
3669:
3663:
3662:
3658:
3656:
3655:
3651:
3649:
3648:
3644:
3642:
3641:
3637:
3635:
3634:
3630:
3628:
3627:
3626:Pentaceratops
3623:
3621:
3620:
3616:
3614:
3613:
3609:
3607:
3606:
3605:Kosmoceratops
3602:
3600:
3599:
3595:
3593:
3592:
3588:
3586:
3585:
3581:
3579:
3578:
3577:Bravoceratops
3574:
3572:
3571:
3570:Bisticeratops
3567:
3565:
3564:
3560:
3558:
3557:
3556:Anchiceratops
3553:
3551:
3550:
3549:Agujaceratops
3546:
3545:
3542:
3539:
3537:
3533:
3527:
3526:
3522:
3520:
3519:
3515:
3513:
3512:
3508:
3506:
3505:
3501:
3499:
3498:
3494:
3492:
3491:
3487:
3486:
3483:
3480:
3477:
3472:
3468:
3464:
3459:
3455:
3446:
3440:
3402:
3401:
3400:
3399:
3397:
3395:
3391:
3385:
3384:
3380:
3378:
3377:
3373:
3370:
3369:
3365:
3364:
3361:
3358:
3356:
3355:Ceratopsoidea
3352:
3346:
3345:
3344:Protoceratops
3341:
3339:
3338:
3337:Breviceratops
3334:
3332:
3331:
3327:
3324:
3323:
3319:
3318:
3316:
3314:
3310:
3307:
3305:
3301:
3295:
3294:
3290:
3288:
3287:
3283:
3281:
3280:
3279:Udanoceratops
3276:
3274:
3273:
3272:Prenoceratops
3269:
3267:
3266:
3262:
3260:
3259:
3258:Leptoceratops
3255:
3253:
3252:
3248:
3246:
3245:
3241:
3239:
3238:
3234:
3232:
3231:
3227:
3225:
3224:
3220:
3218:
3217:
3216:Bainoceratops
3213:
3211:
3210:
3206:
3205:
3203:
3201:
3197:
3191:
3190:
3186:
3185:
3182:
3179:
3176:
3171:
3160:
3159:
3155:
3153:
3152:
3151:Helioceratops
3148:
3146:
3145:
3141:
3140:
3138:
3136:
3132:
3126:
3125:
3121:
3119:
3118:
3117:Mosaiceratops
3114:
3112:
3111:
3107:
3105:
3104:
3100:
3097:
3096:
3092:
3090:
3089:
3088:Koreaceratops
3085:
3083:
3082:
3078:
3076:
3075:
3071:
3069:
3068:
3064:
3061:
3060:
3056:
3055:
3052:
3048:
3045:
3042:
3041:Neoceratopsia
3037:
3027:
3026:
3022:
3020:
3019:
3015:
3013:
3012:
3008:
3006:
3005:
3001:
2999:
2998:
2994:
2993:
2991:
2989:
2985:
2979:
2978:
2974:
2972:
2971:
2967:
2965:
2964:
2960:
2959:
2956:
2952:
2949:
2946:
2941:
2937:
2933:
2928:
2924:
2899:
2898:
2897:
2896:
2894:
2892:
2888:
2880:
2876:
2875:
2874:
2873:
2870:
2867:
2865:
2861:
2853:
2849:
2848:
2847:
2846:
2843:
2840:
2838:
2834:
2830:
2822:
2818:
2815:
2813:
2809:
2806:
2804:
2800:
2797:
2795:
2791:
2789:
2785:
2784:
2779:
2775:
2771:
2764:
2759:
2757:
2752:
2750:
2745:
2744:
2741:
2735:
2733:
2729:
2726:
2725:
2721:
2720:
2716:
2706:
2704:
2702:
2698:
2693:
2689:
2685:
2681:
2677:
2673:
2669:
2665:
2657:
2654:
2649:
2645:
2640:
2635:
2630:
2625:
2621:
2617:
2613:
2609:
2605:
2603:
2597:
2590:
2587:
2582:
2578:
2574:
2570:
2566:
2562:
2558:
2554:
2547:
2544:
2539:
2537:0-253-34595-2
2533:
2529:
2525:
2521:
2514:
2511:
2506:
2502:
2498:
2494:
2490:
2486:
2482:
2478:
2474:
2467:
2464:
2459:
2455:
2451:
2447:
2443:
2439:
2432:
2430:
2426:
2421:
2417:
2412:
2407:
2403:
2399:
2395:
2391:
2387:
2383:
2376:
2374:
2372:
2368:
2363:
2361:0-521-36672-0
2357:
2353:
2348:
2347:
2341:
2337:
2330:
2328:
2324:
2319:
2315:
2310:
2305:
2301:
2297:
2292:
2287:
2283:
2279:
2275:
2268:
2265:
2260:
2254:
2249:
2248:
2239:
2236:
2230:
2225:
2221:
2217:
2213:
2209:
2205:
2198:
2195:
2190:
2186:
2181:
2176:
2171:
2166:
2162:
2158:
2154:
2152:
2143:
2141:
2139:
2135:
2131:
2125:
2122:
2118:(4): 133–159.
2117:
2113:
2106:
2103:
2099:
2093:
2090:
2084:
2079:
2075:
2071:
2067:
2060:
2057:
2052:
2048:
2044:
2040:
2036:
2032:
2028:
2024:
2020:
2013:
2010:
2006:
2000:
1997:
1992:
1988:
1984:
1983:Protoceratops
1977:
1974:
1969:
1967:0-253-34595-2
1963:
1959:
1955:
1951:
1944:
1942:
1938:
1934:
1933:0-691-02882-6
1930:
1926:
1920:
1917:
1913:
1909:
1906:
1900:
1897:
1893:
1892:0-7853-0443-6
1889:
1885:
1879:
1877:
1875:
1873:
1871:
1867:
1862:
1858:
1854:
1850:
1846:
1842:
1838:
1834:
1830:
1826:
1820:
1817:
1809:
1805:
1801:
1797:
1792:
1791:
1787:
1783:
1780:
1779:
1775:
1773:
1771:
1766:
1762:
1758:
1749:
1742:
1740:
1738:
1734:
1729:
1725:
1721:
1713:
1711:
1709:
1705:
1701:
1697:
1693:
1692:
1683:
1682:mega-bonebeds
1678:
1674:
1672:
1671:Styracosaurus
1668:
1664:
1663:Styracosaurus
1660:
1656:
1652:
1651:Styracosaurus
1648:
1644:
1640:
1636:
1629:
1624:
1617:
1610:
1609:
1601:
1600:
1592:
1591:
1583:
1582:
1574:
1573:
1565:
1564:
1556:
1555:
1547:
1546:
1538:
1537:
1529:
1528:
1525:
1524:
1523:
1516:
1515:
1512:
1511:
1508:
1507:
1506:
1499:
1498:
1492:
1491:
1488:
1487:
1484:
1483:
1482:
1475:
1474:
1468:
1467:
1464:
1463:
1460:
1459:
1458:
1451:
1450:
1444:
1441:
1440:
1437:
1436:
1433:
1432:
1431:
1424:
1423:
1417:
1414:
1413:
1410:
1409:
1401:
1400:
1392:
1391:
1383:
1382:
1379:
1378:
1377:
1370:
1369:
1366:
1365:
1362:
1361:
1360:
1353:
1352:
1346:
1345:
1342:
1341:
1338:
1337:
1336:
1329:
1328:
1322:
1321:
1318:
1317:
1309:
1308:
1305:
1304:
1303:
1296:
1295:
1292:
1291:
1288:
1287:
1286:
1279:
1278:
1272:
1271:
1265:
1264:Centrosaurini
1262:
1261:
1255:
1254:Eucentrosaura
1252:
1251:
1248:
1247:
1244:
1243:
1242:
1235:
1234:
1231:
1230:
1227:
1226:
1225:
1218:
1217:
1214:
1213:
1205:
1204:
1201:
1200:
1199:
1192:
1191:
1188:
1187:
1184:
1183:
1182:
1175:
1174:
1168:
1167:
1161:
1160:
1157:
1156:
1153:
1152:
1151:
1144:
1143:
1137:
1136:
1133:
1132:
1124:
1123:
1120:
1115:
1114:
1111:
1110:
1107:
1106:
1105:
1098:
1097:
1094:
1093:
1090:
1085:
1084:
1081:
1080:
1077:
1072:
1071:
1068:
1067:
1064:
1063:(ANSP 15800)
1062:
1061:
1054:
1053:
1047:
1044:
1043:
1037:
1036:
1033:
1032:
1024:
1023:
1020:
1019:
1018:
1011:
1010:
1007:
1006:
1003:
1002:
1001:
994:
993:
987:
986:
980:
977:
976:
972:
970:
966:
961:
959:
958:
957:Achelousaurus
953:
952:
947:
946:
941:
940:
935:
934:
929:
925:
924:
919:
918:
917:Styracosaurus
913:
905:
900:
892:
885:
883:
881:
877:
873:
869:
866:
861:
853:
846:
844:
842:
841:
836:
831:
829:
825:
824:C. nasicornus
821:
817:
813:
809:
805:
801:
800:C. nasicornus
797:
796:
795:Protoceratops
791:
787:
783:
782:Styracosaurus
779:
775:
774:Styracosaurus
771:
770:M. nasicornus
767:
759:
757:
751:
747:
745:
741:
737:
736:Styracosaurus
733:
729:
728:Styracosaurus
725:
721:
717:
713:
708:
706:
702:
698:
694:
690:
686:
682:
677:
675:
674:
670:
660:
658:
646:
644:
639:
635:
631:
627:
623:
619:
610:
605:
597:
595:
593:
589:
586:
582:
579:
575:
572:
569:
568:centrosaurine
565:
545:
544:
531:
486:
485:
461:
458:
443:
440:
431:
429:
425:
410:
408:
404:
395:
393:
389:
380:
378:
374:
373:
371:
368:
364:
359:
354:
352:
343:
340:
339:Binomial name
336:
332:
331:
325:
322:
321:
316:
311:
310:
303:
300:
299:
296:
295:Centrosaurini
290:
287:
286:
283:
282:Eucentrosaura
277:
274:
271:
270:
267:
261:
258:
257:
254:
248:
245:
244:
241:
235:
232:
229:
228:
225:
219:
216:
213:
212:
209:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
39:
33:
29:
26:
22:
4118:Centrosaurus
4088:Centrosaurus
4087:
3966:
3961:Stellasaurus
3959:
3952:
3945:
3924:
3917:
3912:Coronosaurus
3910:
3905:Centrosaurus
3904:
3903:
3874:
3867:
3860:
3853:
3846:
3826:
3821:Lokiceratops
3819:
3812:
3794:Xenoceratops
3792:
3785:
3780:Sinoceratops
3778:
3771:
3764:
3757:
3729:
3722:
3715:
3707:
3700:
3693:
3688:Nedoceratops
3686:
3679:
3661:Vagaceratops
3659:
3654:Utahceratops
3652:
3647:Terminocavus
3645:
3638:
3631:
3624:
3617:
3610:
3603:
3598:Judiceratops
3596:
3589:
3584:Chasmosaurus
3582:
3575:
3568:
3561:
3554:
3547:
3523:
3516:
3509:
3502:
3495:
3488:
3476:Ceratopsidae
3463:Ceratopsidae
3394:Ceratopsidae
3383:Zuniceratops
3381:
3374:
3368:Ajkaceratops
3366:
3342:
3335:
3330:Bagaceratops
3328:
3322:Ajkaceratops
3320:
3304:Coronosauria
3291:
3284:
3277:
3270:
3263:
3256:
3249:
3242:
3235:
3228:
3221:
3214:
3209:Asiaceratops
3207:
3187:
3175:Euceratopsia
3158:Yamaceratops
3156:
3149:
3142:
3122:
3115:
3110:Microceratus
3108:
3103:Liaoceratops
3101:
3093:
3086:
3079:
3072:
3065:
3059:Ajkaceratops
3057:
3023:
3016:
3009:
3002:
2995:
2975:
2968:
2961:
2879:Ornithischia
2864:Ornithischia
2816:
2812:Ornithischia
2807:
2798:
2732:Centrosaurus
2731:
2724:Centrosaurus
2723:
2667:
2663:
2656:
2614:(1): e4252.
2611:
2607:
2604:Triceratops"
2601:
2589:
2556:
2552:
2546:
2523:
2513:
2480:
2476:
2472:
2466:
2441:
2437:
2385:
2381:
2345:
2340:Centrosaurus
2339:
2335:
2281:
2278:PLOS Biology
2277:
2267:
2246:
2238:
2214:(10): 1251.
2211:
2207:
2197:
2160:
2156:
2150:
2129:
2124:
2115:
2111:
2105:
2097:
2092:
2073:
2069:
2059:
2026:
2022:
2018:
2012:
2004:
1999:
1990:
1986:
1982:
1976:
1953:
1924:
1919:
1899:
1883:
1836:
1832:
1828:
1819:
1807:
1803:
1799:
1795:
1757:Centrosaurus
1756:
1754:
1737:Centrosaurus
1736:
1733:Centrosaurus
1732:
1724:osteosarcoma
1719:
1717:
1708:Centrosaurus
1707:
1703:
1700:Centrosaurus
1699:
1696:Centrosaurus
1695:
1689:
1686:
1670:
1667:Centrosaurus
1666:
1662:
1659:Centrosaurus
1658:
1650:
1647:Centrosaurus
1646:
1643:Centrosaurus
1642:
1639:Centrosaurus
1638:
1632:
1618:Paleobiology
1520:
1519:
1503:
1502:
1479:
1478:
1455:
1454:
1428:
1427:
1374:
1373:
1358:
1357:
1356:
1333:
1332:
1300:
1299:
1283:
1282:
1239:
1238:
1222:
1221:
1196:
1195:
1179:
1178:
1148:
1147:
1118:
1102:
1101:
1088:
1075:
1058:
1057:
1015:
1014:
998:
997:
968:
962:
955:
949:
943:
937:
931:
921:
915:
912:Centrosaurus
911:
909:
876:Centrosaurus
875:
872:Centrosaurus
871:
870:
865:Centrosaurus
864:
860:Centrosaurus
859:
858:
840:Coronosaurus
838:
835:C. brinkmani
834:
833:The species
832:
827:
823:
820:Centrosaurus
819:
815:
811:
808:Centrosaurus
807:
803:
799:
793:
785:
781:
780:as a female
777:
773:
769:
765:
763:
754:
743:
740:Centrosaurus
739:
735:
731:
727:
724:Centrosaurus
723:
720:Peter Dodson
715:
712:Centrosaurus
711:
709:
693:Centrosaurus
692:
681:Centrosaurus
680:
678:
673:Kentrosaurus
671:
654:
640:
634:Centrosaurus
633:
618:Centrosaurus
617:
615:
484:Centrosaurus
483:
482:
481:
467:, 1904)
459:
449:, 1914)
441:
426:
416:, 1940)
409:longirostris
405:
390:
375:
350:
345:
329:
328:
309:Centrosaurus
308:
307:
272:
253:Ceratopsidae
230:
214:
208:Ornithischia
198:
185:
32:Centrosaurus
31:
25:
21:Kentrosaurus
4112:Wikispecies
3954:Einiosaurus
3848:Avaceratops
3731:Triceratops
3695:Ojoceratops
3518:Monoclonius
3230:Ferrisaurus
3095:Kulceratops
2596:Tanke, D.H.
2336:Monoclonius
2005:Triceratops
1704:Triceratops
1691:Triceratops
1635:ceratopsids
1633:Like other
1443:Pachyrostra
945:Einiosaurus
939:Avaceratops
923:Monoclonius
904:ontogenetic
847:Description
786:Monoclonius
766:Monoclonius
756:Monoclonius
744:Monoclonius
732:Monoclonius
716:Monoclonius
632:. The name
571:ceratopsian
473:, 1989
455:, 1915
437:, 1917
428:Monoclonius
422:, 1964
407:Monoclonius
401:, 1902
392:Monoclonius
386:, 1917
377:Monoclonius
259:Subfamily:
4175:Categories
4017:See also:
3724:Torosaurus
3640:Spiclypeus
3490:Agathaumas
3403:see below↓
3223:Cerasinops
3011:Stenopelix
2945:Ceratopsia
2932:Ceratopsia
2900:see below↓
2891:Ceratopsia
2803:Dinosauria
2770:Ceratopsia
2411:1880/47001
1993:: 929–940.
1788:References
910:The genus
828:C. apertus
804:C. apertus
760:" skeleton
758:nasicornus
738:resembled
687:, also in
616:The first
602:See also:
414:(Sternberg
379:nasicornus
240:Ceratopsia
192:Dinosauria
4056:Dinosaurs
2786:Kingdom:
2692:225473251
2505:134031275
2458:130607301
2420:128478038
2300:1545-7885
2051:132807103
1810:(4): 1–9.
1770:mammalian
1765:Judithian
1089:CMN 8804
965:cladogram
928:subfamily
880:fenestrae
843:in 2012.
810:species:
669:stegosaur
578:Campanian
323:Species:
165:Kingdom:
159:Eukaryota
42:Campanian
4097:Wikidata
4029:Category
4019:Timeline
3525:Polyonax
3511:Dysganus
3504:Ceratops
3067:Aquilops
2794:Chordata
2792:Phylum:
2788:Animalia
2684:32758461
2648:19172995
2608:PLOS ONE
2598:(2009).
2318:24802911
2189:24688836
2163:: e252.
1908:Archived
1861:85665879
1776:See also
1076:MOR 692
971:(2017):
930:include
574:dinosaur
367:Synonyms
246:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
4145:7236956
4132:4531582
4103:Q131589
4042:Portals
3919:Spinops
3237:Gremlin
3025:Yinlong
2639:2617760
2616:Bibcode
2581:3514834
2561:Bibcode
2553:PALAIOS
2528:501–536
2485:Bibcode
2390:Bibcode
2309:4011683
2216:Bibcode
2180:3933270
2031:Bibcode
1958:312–348
1841:Bibcode
1761:Alberta
714:and/or
701:Alberta
689:Alberta
650:κέντρον
643:kentron
630:Alberta
562:) is a
551:
430:cutleri
394:dawsoni
348:†
327:†
301:Genus:
288:Tribe:
116:↓
4068:Canada
2690:
2682:
2646:
2636:
2579:
2534:
2503:
2456:
2418:
2358:
2354:–243.
2316:
2306:
2298:
2255:
2187:
2177:
2049:
1964:
1931:
1890:
1859:
1728:cancer
969:et al.
954:, and
730:, and
664:σαῦρος
657:sauros
588:Canada
539:-troh-
465:(Lambe
447:(Brown
4158:38854
2817:Clade
2808:Clade
2799:Clade
2688:S2CID
2577:JSTOR
2501:S2CID
2454:S2CID
2416:S2CID
2157:PeerJ
2047:S2CID
1857:S2CID
1806:. 2.
906:order
802:with
753:The "
697:Hilda
638:Greek
576:from
564:genus
453:Lambe
435:Brown
399:Lambe
384:Brown
358:Lambe
315:Lambe
273:Clade
231:Clade
215:Clade
199:Clade
186:Clade
4140:GBIF
2877:see
2850:see
2680:PMID
2644:PMID
2532:ISBN
2356:ISBN
2338:and
2314:PMID
2296:ISSN
2253:ISBN
2185:PMID
1962:ISBN
1929:ISBN
1888:ISBN
1798:and
1694:and
1649:and
963:The
920:and
548:lit.
420:Kuhn
360:1904
317:1904
55:PreꞒ
4127:EoL
3081:Beg
2672:doi
2634:PMC
2624:doi
2569:doi
2493:doi
2446:doi
2406:hdl
2398:doi
2352:231
2304:PMC
2286:doi
2224:doi
2175:PMC
2165:doi
2078:doi
2074:121
2039:doi
1985:".
1849:doi
784:or
628:in
583:of
581:age
566:of
543:-əs
541:SOR
537:SEN
521:ɔːr
44:),
4177::
4155::
4142::
4129::
4114::
4099::
2819::
2810::
2801::
2700:^
2686:.
2678:.
2668:21
2666:.
2642:.
2632:.
2622:.
2610:.
2606:.
2575:.
2567:.
2555:.
2530:.
2499:.
2491:.
2481:92
2479:.
2452:.
2442:81
2440:.
2428:^
2414:.
2404:.
2396:.
2386:42
2384:.
2370:^
2326:^
2312:.
2302:.
2294:.
2282:12
2280:.
2276:.
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