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Charadriiformes

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system. The shorebird ancestor specifically evolved from a bi-parental care system, yet the species within the clade Scolopacidae evolved from a male parental care system. These transitions might have occurred for several reasons. Brooding density is correlated with male parental care. Male care systems in birds are shown to have a very low breeding density while female care systems in birds have a high breeding density. (Owens 2005). Certain rates of male and female mortality, male and female egg maturation rate, and egg death rate have been associated with particular systems as well. It has also been shown that sex role reversal is motivated by the male-biased adult sex ratio. The reason for such diversity in shorebirds, compared to other birds, has yet to be understood.
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Shorebirds pursue a larger diversity of parental care strategies than do most other avian orders. They therefore present an attractive set of examples to support the understanding of the evolution of parental care in avians generally. The ancestral avian most likely had a female parental care
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technique used by Sibley & Ahlquist was not sufficient to properly resolve the relationships in this group, and indeed it appears as if the Charadriiformes constitute a single large and very distinctive lineage of modern birds of their own.
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Owens, I.P. 2002. Male–only care and classical polyandry in birds: phylogeny, ecology and sex differences in remating opportunities. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 357:
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Tullberg, B. S., M. Ah–King and H. Temrin. 2002. Phylogenetic reconstruction of parental–care systems in the ancestors of birds. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 357:
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Tullberg, B. S., M. Ah–King and H. Temrin. 2002. Phylogenetic reconstruction of parental–care systems in the ancestors of birds. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 357:
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Bourdon, Estelle (2006): L'avifaune du PaléogÚne des phosphates du Maroc et du Togo: diversité, systématique et apports à la connaissance de la diversification des oiseaux modernes (Neornithes) . Doctoral thesis,
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Klug, H., M. B. Bonsall, and S.H Alonzo. 2013. Sex differences in life history drive evolutionary transitions among maternal, paternal, and bi-parental care. Ecology and Evolution. 3: 792–806.
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Klug, H., M. B. Bonsall, and S.H Alonzo. 2013. Sex differences in life history drive evolutionary transitions among maternal, paternal, and bi-parental care. Ecology and Evolution. 3: 792–806.
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Paton, Tara A. & Baker, Allan J. (2006): Sequences from 14 mitochondrial genes provide a well-supported phylogeny of the Charadriiform birds congruent with the nuclear RAG-1 tree.
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GĂĄl, Erika; HĂ­r, JĂĄnos; Kessler, EugĂ©n & KĂłkay, JĂłzsef (1998–99): KözĂ©psĂ”-miocĂ©n Ă”smaradvĂĄnyok, a MĂĄtraszĂ”lĂ”s, RĂĄkĂłczi-kĂĄpolna alatti ĂștbevĂĄgĂĄsbĂłl. I. A MĂĄtraszĂ”lĂ”s 1. lelĂ”hely .
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SzĂ©kely, T and J.D. Reynolds. 1995. Evolutionary transitions in parental care in shorebirds. Proceedings of the Royal Society of London. Series B: Biological Sciences. 262: 57–64.
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Ericson, Per G.P.; Envall, I.; Irestedt, M. & Norman, J.A. (2003): Inter-familial relationships of the shorebirds (Aves: Charadriiformes) based on nuclear DNA sequence data.
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van Tuinen, Marcel; Waterhouse, David & Dyke, Gareth J. (2004): Avian molecular systematics on the rebound: a fresh look at modern shorebird phylogenetic relationships.
1285:, the Charadriiformes are the only other order of modern bird to have an established fossil record within the late Cretaceous, alongside the other dinosaurs. Much of the 1200:
Phylogeny of the Charadriiformes based a study by Heiner Kuhl and collaborators published in 2020. The families and the number of species are from the list maintained by
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The auks, usually considered distinct because of their peculiar morphology, are more likely related to gulls, the "distinctness" being a result of adaptation for diving.
3045: 4391: 2504: 4430: 1835:ČernĂœ, David; Natale, Rossy (2021-07-16). "Comprehensive taxon sampling and vetted fossils help clarify the time tree of shorebirds (Aves, Charadriiformes)". 1965:"Phylogenetic relationships and divergence times of Charadriiformes genera: multigene evidence for the Cretaceous origin of at least 14 clades of shorebirds" 364:(or "Charadrii"): typical shorebirds, most of which feed by probing in the mud or picking items off the surface in both coastal and freshwater environments. 4540: 2266: 1290: 131: 130: 2382:
Paton, T.A.; Baker, A.J.; Groth, J.G. & Barrowclough, G.F. (2003): RAG-1 sequences resolve phylogenetic relationships within charadriiform birds.
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Liker, A., R. P. Freckleton, and T. Székely. 2013. The evolution of sex roles in birds is related to adult sex ratio. Nature Communications. 4: 1587.
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Liker, A., R. P. Freckleton, and T. Székely. 2013. The evolution of sex roles in birds is related to adult sex ratio. Nature Communications. 4: 1587.
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Thomas, Gavin H.; Wills, Matthew A. & Székely, Tamås (2004a): Phylogeny of shorebirds, gulls, and alcids (Aves: Charadrii) from the cytochrome-
4404: 3040: 1209: 1439:. They are now assumed to be mostly basal taxa of the charadriiforms and/or "higher waterbirds", which probably were two distinct lineages 65 4565: 2540:; Tennyson, A.J.D.; Jones, C.; McNamara, J.A. & Douglas, B.J. (2007): Miocene waterfowl and other birds from central Otago, New Zealand. 2455: 2384: 2355: 1947: 1765: 1616: 2338: 4550: 2542: 342:(seabirds), others frequent deserts, and a few are found in dense forest. Members of this group can also collectively be referred to as 2325: 127: 4570: 4560: 2019:
Case, J. A. and C. P. Tambussi. 1999. Maastrichtian record of neornithine birds in Antarctica: comments on a Late Cretaceous radiation
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Thomas, Gavin H.; Székely, Tamås; Reynolds, John D. (2007). "Sexual Conflict and the Evolution of Breeding Systems in Shorebirds".
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Thomas, Gavin H.; Székely, Tamås; Reynolds, John D. (2007). "Sexual Conflict and the Evolution of Breeding Systems in Shorebirds".
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already, and few if any are still believed to be related to the well-distinct waterfowl. Taxa formerly considered graculavids are:
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Bertelli, S.; Lindow, B. E. K.; Dyke, G. J.; Mayr, G. (2013). "Another charadriiform-like bird from the lower Eocene of Denmark".
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Kuhl., H.; Frankl-Vilches, C.; Bakker, A.; Mayr, G.; Nikolaus, G.; Boerno, S. T.; Klages, S.; Timmermann, B.; Gahr, M. (2020).
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will also take food items from beaches, or rob smaller species, and some have become adapted to inland environments.
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Charadriiformes gen. et sp. indet. (Bathans Early/Middle Miocene of Otago, New Zealand) - charadriid? scolopacid?
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Charadriiformes gen. et sp. indet. (Bathans Early/Middle Miocene of Otago, New Zealand) - charadriid? scolopacid?
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Thomas, Gavin H.; Wills, Matthew A. & Székely, Tamås (2004): A supertree approach to shorebird phylogeny.
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That the Charadriiformes are an ancient group is also borne out by the fossil record. Alongside the
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is made up of bits and pieces of birds which resemble this order. In many, this is probably due to
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Fain, Matthew G. & Houde, Peter (2004): Parallel radiations in the primary clades of birds.
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Charadriiformes gen. et sp. indet. (Bathans Early/Middle Miocene of Otago, New Zealand) - larid?
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and has members in all parts of the world. Most charadriiform birds live near water and eat
259: 4027: 3325: 2847: 2832: 2770: 2669: 2597: 2125: 2040: 1786:"An unbiased molecular approach using 3'UTRs resolves the avian family-level tree of life" 1321:. However, more complete remains of undisputed charadriiforms are known only from the mid- 1306: 772: 480: 108: 4357: 4243: 1714: 1649: 4342: 4329: 4256: 4137: 4067: 4019: 3953: 3892: 3847: 3834: 3708: 3671: 3522: 3296: 3133: 2887: 2882: 2877: 2857: 2842: 2737: 2732: 2712: 2707: 2689: 2637: 2570: 1997: 1964: 1812: 1785: 1112: 788: 646: 573: 487: 445: 377:"): these are generally larger species which take fish from the sea. Several gulls and 319: 2468: 2424: 2397: 2201: 4514: 4075: 4059: 3982: 3974: 3808: 3606: 3456: 3343: 3301: 3101: 3093: 3015: 2979: 2897: 2821: 2528: 2149: 1410:
Charadriiformes gen. et sp. indet. (Sajóvölgyi Middle Miocene of MåtraszÔlÔs, Hungary
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Avian evolution: the fossil record of birds and its paleobiological significance
1699:"Erratum to: "Another charadriiform-like bird from the Lower Eocene of Denmark"" 1554: 1491: 1350: 1310: 1298: 1228: 1057: 998: 703: 559: 539: 459: 374: 45: 2126:"A New Species of Graculavus from the Cretaceous of Wyoming (Aves: Neornithes)" 4083: 3945: 3884: 3863: 3695: 3679: 3614: 3559: 3531: 3463: 3428: 3306: 3154: 2989: 2964: 2722: 2555: 2453:
gene: parsimony, Bayesian inference, minimum evolution, and quartet puzzling.
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formerly believed to constitute the common ancestors of charadriiforms,
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The Howard & Moore Complete Checklist of the Birds of the World
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Baker, Allan J; Pereira, SĂ©rgio L; Paton, Tara A (2007-04-22).
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Bertelli, S.; Lindow, B. E. K.; Dyke, G. J.; Mayr, G. (2014).
1222: 386: 301: 1212:. The suborders are those defined by Joel Cracraft in 2013. 295: 118: 1416:(Late Miocene of Los Mansuetos, Spain) - scolopacid? larid? 283: 265: 1585:
Neornithes incerta sedis (Late Paleocene/Early Eocene of
1521:"Graculavidae" gen. et sp. indet. (Gloucester County, US) 289: 274: 1760:. Topics in Paleobiology. Wiley-Blackwell. p. 306. 1362:(Huadian Middle Eocene of Huadian, China) - charadriid? 2060:
Several wing and thorax bones of a bird the size of a
1495:(Navesink Late Cretaceous?) - charadriiform? gruiform? 1348:(Late Paleocene/Early Eocene of Jutland, Denmark) - a 1398:(John Day Early Miocene of Willow Creek, US) - larid? 354:
The order was formerly divided into three suborders:
304: 292: 262: 1370:(Early Oligocene of Boutersem, Belgium) - glareolid? 280: 277: 271: 4198: 4010: 3965: 3904: 3875: 3846: 3825: 3786: 3749: 3740: 3707: 3670: 3661: 3652: 3597: 3521: 3480: 3418: 3390: 3366: 3342: 3315: 3236: 3223: 3184: 3153: 3092: 3008: 2911: 2811: 2761: 2688: 2618: 298: 268: 1392:(Early Miocene of SE France) - larid? stercorarid? 398:lumps all the Charadriiformes together with other 2089:(MNZ S41058) of a bird apparently similar to the 1531:, which may or may not be Charadriiformes, are: 1487:(Hornerstown Late Cretaceous?) - charadriiform? 1325:onwards. Present-day orders emerged around the 122: 1779: 1777: 326:) is a diverse order of small to medium-large 2578: 2043:(MNZ S42415, S42435) of a bird the size of a 1602:Evolution of parental care in Charadriiformes 8: 2419:. Vol. 37. Elsevier. pp. 279–342. 2196:. Vol. 37. Elsevier. pp. 279–342. 1386:(Early Miocene Saint-GĂ©rand-le-Puy, France) 286: 4186: 3746: 3667: 3658: 3649: 3387: 3233: 3220: 3181: 3177: 3150: 3089: 3085: 3074: 3070: 2585: 2571: 2563: 2332:Folia Historico Naturalia Musei Matraensis 1925: 1923: 1709:(13). Pleiades Publishing Ltd: 1441–1448. 1644:(11). Pleiades Publishing Ltd: 1282–1301. 1337:. Basal or unresolved charadriiforms are: 626:– stone-curlews, thick-knees (10 species) 338:or other small animals; however, some are 20: 2130:Smithsonian Contributions to Paleobiology 1996: 1840: 1811: 1801: 1732: 1722: 1665: 1269:Learn how and when to remove this message 441:– stone-curlews, thick-knees (10 species) 1317:charadriiform somewhat reminiscent of a 1904:. International Ornithologists' Union. 1628: 1597:(Early Eocene of Silveirinha, Portugal) 1092:– gulls, terns, skimmers (103 species) 1574:(Early Paleocene of Volgograd, Russia) 569:– gulls, terns, skimmers (103 species) 1888: 1886: 1617:List of Charadriiformes by population 1291:Cretaceous–Paleogene extinction event 1060:– coursers, pratincoles (17 species) 426:The order Charadriiformes contains 3 7: 4485:AB0787CD-0A40-FF99-FDAB-F958E5CD22B4 4446:e36be5fc-7f3a-4ad4-9e35-2480e435a5ed 4348:49ff3c6a-a714-4caf-87ed-504fd4cbe627 4144: 2267:MusĂ©um national d'histoire naturelle 1251:adding citations to reliable sources 562:– coursers, pratincoles (17 species) 4168: 1908:from the original on 4 October 2020 1210:International Ornithologists' Union 1208:and David Donsker on behalf of the 1105: 1082: 1034: 1027: 1004: 935: 887: 880: 857: 849: 765: 742: 719: 696: 639: 616: 609: 601: 593: 350:Taxonomy, systematics and evolution 1902:"IOC World Bird List Version 11.2" 867:– sandpipers, snipes (98 species) 107:Possible an early origin based on 14: 4541:Extant Lutetian first appearances 3046:Late Quaternary prehistoric birds 2417:Advances in the Study of Behavior 2194:Advances in the Study of Behavior 533:– sandpipers, snipes (98 species) 410:. However, the resolution of the 4167: 4155: 4143: 4132: 4131: 4022:(New World vultures and condors) 2529:10.1111/j.0908-8857.2004.03362.x 1880:(2004), Paton & Baker (2006) 1527:Other wader- or gull-like birds 1420:The "transitional shorebirds" (" 1227: 258: 150: 2156:from the original on 8 May 2024 1790:Molecular Biology and Evolution 1238:needs additional citations for 752:– stilts, avocets (10 species) 430:, 19 families and 391 species. 791:– oystercatchers (12 species) 476:– stilts, avocets (10 species) 406:into a greatly enlarged order 1: 2469:10.1016/S1055-7903(03)00222-7 2425:10.1016/s0065-3454(07)37006-x 2398:10.1016/S1055-7903(03)00098-8 2202:10.1016/s0065-3454(07)37006-x 1558:(Lance Creek Late Cretaceous) 1550:(Lance Creek Late Cretaceous) 1540:(Lance Creek Late Cretaceous) 1378:(Early Oligocene) - turnicid? 897:– painted-snipes (3 species) 490:– oystercatchers (12 species) 141:Several members of the order 4566:Pleistocene taxonomic orders 2174:A wading bird the size of a 2124:Hope, Sylvia (4 June 1996). 1313:, Antarctica) is probably a 1303:LĂłpez de Bertodano Formation 1014:– buttonquails (18 species) 505:– painted-snipes (3 species) 4086:(woodpeckers and relatives) 2369:10.1016/j.ympev.2006.01.011 1452:- charadriiform? gruiform? 1289:' fossil record around the 548:– buttonquails (18 species) 4587: 4551:Oligocene taxonomic orders 3948:(cormorants and relatives) 3041:Extinct species since 1500 2039:S42416) and proximal left 1301:habits. Specimen VI 9901 ( 665:– sheathbills (2 species) 4571:Holocene taxonomic orders 4561:Pliocene taxonomic orders 4127: 4078:(kingfishers and rollers) 3932:(albatrosses and petrels) 3816:(swifts and hummingbirds) 3798:(nightjars and relatives) 3648: 3622:(pheasants and relatives) 3180: 3149: 3088: 3084: 3073: 3069: 2605: 2556:10.1017/S1477201906001957 2093:but almost the size of a 1842:10.1101/2021.07.15.452585 1724:10.1134/s0031030114130024 1658:10.1134/s0031030113110026 1126: 1110: 1103: 1087: 1080: 1055: 1039: 1032: 1025: 1009: 1002: 961:– seedsnipes (4 species) 956: 940: 933: 908: 892: 885: 878: 862: 855: 847: 786: 770: 763: 747: 740: 724: 717: 701: 694: 660: 644: 637: 621: 614: 607: 599: 455:– sheathbills (2 species) 244: 239: 147:Scientific classification 145: 140: 116: 23: 4556:Miocene taxonomic orders 3977:(seriemas and relatives) 3956:(pelicans and relatives) 2142:10.5479/SI.00810266.89.1 1333:boundary, roughly 35-30 526:– seedsnipes (4 species) 396:Sibley-Ahlquist taxonomy 330:. It includes about 390 4546:Eocene taxonomic orders 4070:(hornbills and hoopoes) 3985:(falcons and relatives) 1930:Cracraft, Joel (2013). 1855:Fain & Houde (2004) 1703:Paleontological Journal 1638:Paleontological Journal 729:– plovers (69 species) 4054:(trogons and quetzals) 3866:(cranes and relatives) 3136:(emus and cassowaries) 3026:Glossary of bird terms 2838:Confuciusornithiformes 2505:Supplementary Material 2491:10.1186/1471-2148-4-28 2456:Mol. Phylogenet. Evol. 2385:Mol. Phylogenet. Evol. 2356:Mol. Phylogenet. Evol. 2291:10.1186/1471-2148-3-16 1981:10.1098/rsbl.2006.0606 1876:(2004a,b), van Tuinen 1803:10.1093/molbev/msaa191 913:– jacanas (8 species) 469:– plovers (69 species) 373:and their allies (or " 134: 4467:Paleobiology Database 3895:(kagu and sunbittern) 3858:(gulls and relatives) 2863:Songlingornithiformes 2828:Omnivoropterygiformes 1756:Mayr, Gerald (2016). 1582:(Paleocene of France) 1414:"Totanus" teruelensis 512:– jacanas (8 species) 412:DNA-DNA hybridization 133: 4343:Fauna Europaea (new) 3717:Phoenicopteriformes 2543:J. Syst. Palaeontol. 2081:S42681, S42736) and 2062:double-banded plover 1900:, eds. (July 2021). 1503:- phoenicopteriform? 1295:convergent evolution 1247:improve this article 1219:Evolutionary history 1131:– auks (25 species) 1115:– skuas (7 species) 649:– Magellanic plover 3688:Mesitornithiformes 3682:(doves and pigeons) 3016:Families and orders 2878:Ichthyornithiformes 2853:Patagopterygiformes 1715:2014PalJ...48.1441B 1650:2013PalJ...47.1282B 1424:") are a generally 1390:"Larus" desnoyersii 583:– auks (25 species) 576:– skuas (7 species) 448:– Magellanic plover 434:Suborder Charadrii 4030:(eagles and hawks) 3930:Procellariiformes 3835:Opisthocomiformes 3804:Steatornithiformes 3031:List by population 2868:Hongshanornithidae 2793:Evolution of birds 1896:; Donsker, David; 1587:Ouled Abdoun Basin 1460:(Late Cretaceous?) 945:– plains-wanderer 706:– Egyptian plover 135: 4508: 4507: 4454:Open Tree of Life 4192:Taxon identifiers 4183: 4182: 4123: 4122: 4119: 4118: 4115: 4114: 4111: 4110: 4107: 4106: 4103: 4102: 4099: 4098: 4095: 4094: 3916:(loons or divers) 3885:Phaethontiformes 3877:Phaethontimorphae 3827:Opisthocomiformes 3796:Caprimulgiformes 3736: 3735: 3725:Podicipediformes 3644: 3643: 3640: 3639: 3636: 3635: 3362: 3361: 3145: 3144: 3102:Struthioniformes 2970:Waterfowl hunting 2903:Gastornithiformes 2898:Aepyornithiformes 2888:Lithornithiformes 2558: 2538:Worthy, Trevor H. 2531: 2500: 2471: 2407: 2378: 2324: 2308:(11): 2558–2573. 2293: 2182:): Bourdon (2005) 2091:black-billed gull 1949:978-0-9568611-0-8 1898:Rasmussen, Pamela 1767:978-1-119-02076-9 1566:(Late Cretaceous) 1511:- presbyornithid? 1396:"Larus" pristinus 1297:brought about by 1279: 1278: 1271: 1216: 1215: 1194: 1193: 1185: 1184: 1176: 1175: 1167: 1166: 1158: 1157: 1149: 1148: 1140: 1139: 1069: 1068: 988: 987: 979: 978: 970: 969: 922: 921: 836: 835: 827: 826: 818: 817: 809: 808: 800: 799: 683: 682: 674: 673: 519:– plains-wanderer 462:– Egyptian plover 251: 250: 235: 111: 4578: 4501: 4500: 4488: 4487: 4475: 4474: 4462: 4461: 4449: 4448: 4439: 4438: 4426: 4425: 4423:NBNSYS0000159655 4413: 4412: 4400: 4399: 4387: 4386: 4374: 4373: 4361: 4360: 4351: 4350: 4338: 4337: 4325: 4324: 4312: 4311: 4299: 4298: 4286: 4285: 4273: 4272: 4270:7FC8B7ED29E2F6B4 4260: 4259: 4247: 4246: 4234: 4233: 4232: 4219: 4218: 4217: 4187: 4171: 4170: 4159: 4147: 4146: 4135: 4134: 4087: 4079: 4071: 4063: 4060:Leptosomiformes 4055: 4047: 4039: 4031: 4028:Accipitriformes 4023: 4002: 4001:(perching birds) 3994: 3986: 3978: 3957: 3949: 3941: 3933: 3925: 3922:Sphenisciformes 3917: 3896: 3888: 3867: 3859: 3856:Charadriiformes 3838: 3817: 3799: 3778: 3770: 3767:Musophagiformes 3762: 3747: 3728: 3720: 3699: 3691: 3683: 3668: 3659: 3650: 3623: 3610: 3388: 3384: 3377: 3290: 3283: 3247: 3234: 3230: 3221: 3217: 3182: 3178: 3151: 3137: 3129: 3121: 3113: 3105: 3090: 3086: 3075: 3071: 2938:Bird collections 2893:Dinornithiformes 2798:Darwin's finches 2788:Origin of flight 2728:Seabird breeding 2718:Sexual selection 2587: 2580: 2573: 2564: 2550: 2523: 2485: 2463: 2446: 2392: 2363: 2309: 2285: 2252: 2249: 2243: 2240: 2234: 2230: 2224: 2223: 2189: 2183: 2172: 2166: 2165: 2163: 2161: 2121: 2115: 2108: 2102: 2075: 2069: 2058: 2052: 2045:red-necked stint 2026: 2020: 2017: 2011: 2010: 2000: 1960: 1954: 1953: 1936:Remsen, J.V. Jr. 1927: 1918: 1917: 1915: 1913: 1890: 1881: 1862: 1856: 1853: 1847: 1846: 1844: 1832: 1826: 1825: 1815: 1805: 1781: 1772: 1771: 1753: 1747: 1746: 1736: 1726: 1694: 1688: 1687: 1669: 1633: 1274: 1267: 1263: 1260: 1254: 1231: 1223: 1206:Pamela Rasmussen 1106: 1083: 1035: 1028: 1005: 936: 888: 881: 858: 850: 766: 750:Recurvirostridae 743: 720: 697: 640: 617: 610: 602: 597:Charadriiformes 594: 589: 588: 474:Recurvirostridae 311: 310: 307: 306: 303: 300: 297: 294: 291: 288: 285: 282: 279: 276: 273: 270: 267: 264: 230: 155: 154: 121: 106: 105: 42: 27:Temporal range: 21: 4586: 4585: 4581: 4580: 4579: 4577: 4576: 4575: 4521:Charadriiformes 4511: 4510: 4509: 4504: 4496: 4491: 4483: 4478: 4470: 4465: 4457: 4452: 4444: 4442: 4434: 4429: 4421: 4416: 4408: 4403: 4395: 4390: 4382: 4377: 4369: 4364: 4358:charadriiformes 4356: 4354: 4346: 4341: 4333: 4328: 4320: 4315: 4307: 4302: 4294: 4289: 4281: 4276: 4268: 4263: 4257:Charadriiformes 4255: 4250: 4244:Charadriiformes 4242: 4237: 4230:Charadriiformes 4228: 4227: 4222: 4213: 4212: 4207: 4200:Charadriiformes 4194: 4184: 4179: 4091: 4085: 4077: 4069: 4068:Bucerotiformes 4062:(cuckoo-roller) 4061: 4053: 4045: 4037: 4029: 4021: 4020:Cathartiformes 4006: 4000: 3992: 3991:Psittaciformes 3984: 3976: 3961: 3955: 3954:Pelecaniformes 3947: 3939: 3931: 3923: 3915: 3900: 3894: 3893:Eurypygiformes 3886: 3871: 3865: 3857: 3842: 3836: 3821: 3815: 3797: 3782: 3776: 3768: 3760: 3732: 3726: 3718: 3703: 3697: 3696:Pterocliformes 3689: 3681: 3632: 3621: 3608: 3593: 3517: 3476: 3414: 3379: 3378: 3371: 3370: 3358: 3338: 3311: 3288: 3281: 3241: 3240: 3228: 3227: 3211: 3210: 3207: 3205: 3203: 3201: 3199: 3197: 3195: 3193: 3191: 3189: 3187: 3172: 3170: 3168: 3166: 3164: 3162: 3160: 3158: 3156: 3141: 3135: 3134:Casuariiformes 3127: 3126:Apterygiformes 3119: 3111: 3103: 3080: 3065: 3036:Lists by region 3004: 2914: 2907: 2848:Chaoyangiformes 2833:Jeholornithidae 2807: 2771:Origin of birds 2757: 2738:Brood parasites 2684: 2614: 2601: 2591: 2559:(HTML abstract) 2478:BMC Evol. Biol. 2472:(HTML abstract) 2435: 2414: 2408:(HTML abstract) 2379:(HTML abstract) 2278:BMC Evol. Biol. 2261: 2256: 2255: 2250: 2246: 2241: 2237: 2231: 2227: 2212: 2191: 2190: 2186: 2180:Ciconia ciconia 2173: 2169: 2159: 2157: 2123: 2122: 2118: 2109: 2105: 2076: 2072: 2059: 2055: 2027: 2023: 2018: 2014: 1969:Biology Letters 1962: 1961: 1957: 1950: 1932:Dickinson, E.C. 1929: 1928: 1921: 1911: 1909: 1892: 1891: 1884: 1872:(2003), Thomas 1863: 1859: 1854: 1850: 1834: 1833: 1829: 1783: 1782: 1775: 1768: 1755: 1754: 1750: 1696: 1695: 1691: 1635: 1634: 1630: 1625: 1613: 1604: 1307:Late Cretaceous 1275: 1264: 1258: 1255: 1244: 1232: 1221: 1195: 1186: 1177: 1168: 1159: 1150: 1141: 1070: 989: 980: 971: 923: 837: 828: 819: 810: 801: 773:Ibidorhynchidae 684: 675: 481:Ibidorhynchidae 424: 352: 261: 257: 254:Charadriiformes 229: 227:Charadriiformes 149: 136: 132: 119: 112: 109:molecular clock 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 37: 36: 25: 17: 12: 11: 5: 4584: 4582: 4574: 4573: 4568: 4563: 4558: 4553: 4548: 4543: 4538: 4533: 4528: 4523: 4513: 4512: 4506: 4505: 4503: 4502: 4489: 4476: 4463: 4450: 4440: 4427: 4414: 4401: 4388: 4375: 4362: 4352: 4339: 4330:Fauna Europaea 4326: 4313: 4300: 4287: 4274: 4261: 4248: 4235: 4220: 4204: 4202: 4196: 4195: 4190: 4181: 4180: 4178: 4177: 4165: 4153: 4141: 4128: 4125: 4124: 4121: 4120: 4117: 4116: 4113: 4112: 4109: 4108: 4105: 4104: 4101: 4100: 4097: 4096: 4093: 4092: 4090: 4089: 4081: 4076:Coraciiformes 4073: 4065: 4057: 4052:Trogoniformes 4049: 4041: 4033: 4025: 4016: 4014: 4008: 4007: 4005: 4004: 3999:Passeriformes 3996: 3988: 3983:Falconiformes 3980: 3975:Cariamiformes 3971: 3969: 3963: 3962: 3960: 3959: 3951: 3943: 3938:Ciconiiformes 3935: 3927: 3919: 3910: 3908: 3902: 3901: 3899: 3898: 3890: 3881: 3879: 3873: 3872: 3870: 3869: 3861: 3852: 3850: 3848:Cursorimorphae 3844: 3843: 3841: 3840: 3831: 3829: 3823: 3822: 3820: 3819: 3811: 3806: 3801: 3792: 3790: 3784: 3783: 3781: 3780: 3772: 3764: 3755: 3753: 3744: 3738: 3737: 3734: 3733: 3731: 3730: 3722: 3713: 3711: 3709:Mirandornithes 3705: 3704: 3702: 3701: 3693: 3685: 3680:Columbiformes 3676: 3674: 3672:Columbimorphae 3665: 3656: 3646: 3645: 3642: 3641: 3638: 3637: 3634: 3633: 3631: 3630: 3625: 3617: 3612: 3607:Meleagridinae 3603: 3601: 3595: 3594: 3592: 3591: 3584: 3577: 3570: 3563: 3556: 3549: 3542: 3535: 3527: 3525: 3523:Odontophoridae 3519: 3518: 3516: 3515: 3508: 3501: 3494: 3486: 3484: 3478: 3477: 3475: 3474: 3467: 3460: 3453: 3446: 3439: 3432: 3424: 3422: 3416: 3415: 3413: 3412: 3407: 3402: 3396: 3394: 3385: 3364: 3363: 3360: 3359: 3357: 3356: 3348: 3346: 3340: 3339: 3337: 3336: 3329: 3321: 3319: 3313: 3312: 3310: 3309: 3304: 3302:Stictonettinae 3299: 3297:Dendrocygninae 3294: 3293: 3292: 3285: 3273: 3272: 3271: 3266: 3261: 3250: 3248: 3231: 3218: 3175: 3147: 3146: 3143: 3142: 3140: 3139: 3131: 3123: 3115: 3107: 3098: 3096: 3082: 3081: 3078: 3067: 3066: 3064: 3063: 3062: 3061: 3056: 3050:Notable birds 3048: 3043: 3038: 3033: 3028: 3023: 3018: 3012: 3010: 3006: 3005: 3003: 3002: 3000:Egg collecting 2997: 2992: 2987: 2982: 2977: 2972: 2967: 2962: 2957: 2952: 2951: 2950: 2940: 2935: 2930: 2925: 2919: 2917: 2909: 2908: 2906: 2905: 2900: 2895: 2890: 2885: 2883:Hesperornithes 2880: 2875: 2870: 2865: 2860: 2858:Ambiortiformes 2855: 2850: 2845: 2843:Enantiornithes 2840: 2835: 2830: 2825: 2817: 2815: 2809: 2808: 2806: 2805: 2800: 2795: 2790: 2785: 2784: 2783: 2778: 2767: 2765: 2759: 2758: 2756: 2755: 2750: 2745: 2740: 2735: 2730: 2725: 2720: 2715: 2710: 2705: 2700: 2694: 2692: 2686: 2685: 2683: 2682: 2677: 2672: 2667: 2662: 2657: 2652: 2647: 2646: 2645: 2635: 2630: 2624: 2622: 2616: 2615: 2613: 2612: 2606: 2603: 2602: 2592: 2590: 2589: 2582: 2575: 2567: 2561: 2560: 2535: 2522:(3): 191–194. 2516:J. Avian Biol. 2511: 2507: 2473: 2462:(3): 516–526. 2447: 2433: 2412: 2409: 2380: 2362:(3): 657–667. 2351: 2347: 2344: 2341: 2328: 2315:10.1554/04-235 2297: 2273: 2260: 2257: 2254: 2253: 2244: 2235: 2225: 2210: 2184: 2167: 2116: 2103: 2070: 2053: 2041:carpometacarpi 2021: 2012: 1975:(2): 205–210. 1955: 1948: 1919: 1882: 1868:(2003), Paton 1857: 1848: 1827: 1773: 1766: 1748: 1689: 1627: 1626: 1624: 1621: 1620: 1619: 1612: 1609: 1603: 1600: 1599: 1598: 1590: 1583: 1575: 1567: 1559: 1551: 1541: 1529:incertae sedis 1525: 1524: 1523: 1522: 1519: 1512: 1504: 1496: 1488: 1480: 1463: 1462: 1461: 1418: 1417: 1411: 1408: 1405: 1402: 1399: 1393: 1387: 1379: 1371: 1363: 1355: 1277: 1276: 1235: 1233: 1226: 1220: 1217: 1214: 1213: 1197: 1196: 1192: 1191: 1188: 1187: 1183: 1182: 1179: 1178: 1174: 1173: 1170: 1169: 1165: 1164: 1161: 1160: 1156: 1155: 1152: 1151: 1147: 1146: 1143: 1142: 1138: 1137: 1134: 1133: 1125: 1122: 1121: 1118: 1117: 1113:Stercorariidae 1109: 1104: 1102: 1099: 1098: 1095: 1094: 1086: 1081: 1079: 1076: 1075: 1072: 1071: 1067: 1066: 1063: 1062: 1054: 1051: 1050: 1047: 1046: 1044:– crab-plover 1038: 1033: 1031: 1026: 1024: 1021: 1020: 1017: 1016: 1008: 1003: 1001: 995: 994: 991: 990: 986: 985: 982: 981: 977: 976: 973: 972: 968: 967: 964: 963: 955: 952: 951: 948: 947: 939: 934: 932: 929: 928: 925: 924: 920: 919: 916: 915: 907: 904: 903: 900: 899: 891: 886: 884: 879: 877: 874: 873: 870: 869: 861: 856: 854: 848: 846: 843: 842: 839: 838: 834: 833: 830: 829: 825: 824: 821: 820: 816: 815: 812: 811: 807: 806: 803: 802: 798: 797: 794: 793: 789:Haematopodidae 785: 782: 781: 778: 777: 769: 764: 762: 759: 758: 755: 754: 746: 741: 739: 736: 735: 732: 731: 723: 718: 716: 713: 712: 709: 708: 700: 695: 693: 690: 689: 686: 685: 681: 680: 677: 676: 672: 671: 668: 667: 659: 656: 655: 652: 651: 647:Pluvianellidae 643: 638: 636: 633: 632: 629: 628: 620: 615: 613: 608: 606: 600: 598: 592: 587: 586: 585: 584: 577: 574:Stercorariidae 570: 563: 556: 549: 536: 535: 534: 527: 520: 513: 506: 493: 492: 491: 488:Haematopodidae 484: 477: 470: 463: 456: 449: 446:Pluvianellidae 442: 423: 420: 392: 391: 382: 365: 351: 348: 249: 248: 242: 241: 237: 236: 224: 220: 219: 214: 207: 206: 201: 197: 196: 191: 187: 186: 181: 177: 176: 171: 167: 166: 161: 157: 156: 143: 142: 138: 137: 117: 114: 113: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 26: 16:Order of birds 15: 13: 10: 9: 6: 4: 3: 2: 4583: 4572: 4569: 4567: 4564: 4562: 4559: 4557: 4554: 4552: 4549: 4547: 4544: 4542: 4539: 4537: 4534: 4532: 4529: 4527: 4524: 4522: 4519: 4518: 4516: 4499: 4494: 4490: 4486: 4481: 4477: 4473: 4468: 4464: 4460: 4455: 4451: 4447: 4441: 4437: 4432: 4428: 4424: 4419: 4415: 4411: 4406: 4402: 4398: 4393: 4389: 4385: 4380: 4376: 4372: 4367: 4363: 4359: 4355:Featherbase: 4353: 4349: 4344: 4340: 4336: 4331: 4327: 4323: 4318: 4314: 4310: 4305: 4301: 4297: 4292: 4288: 4284: 4279: 4275: 4271: 4266: 4262: 4258: 4253: 4249: 4245: 4240: 4236: 4231: 4225: 4221: 4216: 4210: 4206: 4205: 4203: 4201: 4197: 4193: 4188: 4176: 4175: 4166: 4164: 4163: 4158: 4154: 4152: 4151: 4142: 4140: 4139: 4130: 4129: 4126: 4088: 4082: 4080: 4074: 4072: 4066: 4064: 4058: 4056: 4050: 4048: 4042: 4040: 4036:Strigiformes 4034: 4032: 4026: 4024: 4018: 4017: 4015: 4013: 4009: 4003: 3997: 3995: 3989: 3987: 3981: 3979: 3973: 3972: 3970: 3968: 3964: 3958: 3952: 3950: 3944: 3942: 3936: 3934: 3928: 3926: 3920: 3918: 3912: 3911: 3909: 3907: 3903: 3897: 3891: 3889: 3887:(tropicbirds) 3883: 3882: 3880: 3878: 3874: 3868: 3862: 3860: 3854: 3853: 3851: 3849: 3845: 3839: 3833: 3832: 3830: 3828: 3824: 3818: 3812: 3810: 3809:Podargiformes 3807: 3805: 3802: 3800: 3794: 3793: 3791: 3789: 3785: 3779: 3773: 3771: 3765: 3763: 3759:Cuculiformes 3757: 3756: 3754: 3752: 3748: 3745: 3743: 3739: 3729: 3723: 3721: 3715: 3714: 3712: 3710: 3706: 3700: 3694: 3692: 3686: 3684: 3678: 3677: 3675: 3673: 3669: 3666: 3664: 3660: 3657: 3655: 3651: 3647: 3629: 3626: 3624: 3618: 3616: 3613: 3611: 3605: 3604: 3602: 3600: 3596: 3590: 3589: 3585: 3583: 3582: 3578: 3576: 3575: 3571: 3569: 3568: 3564: 3562: 3561: 3557: 3555: 3554: 3550: 3548: 3547: 3543: 3541: 3540: 3536: 3534: 3533: 3529: 3528: 3526: 3524: 3520: 3514: 3513: 3509: 3507: 3506: 3502: 3500: 3499: 3495: 3493: 3492: 3488: 3487: 3485: 3483: 3479: 3473: 3472: 3468: 3466: 3465: 3461: 3459: 3458: 3457:Macrocephalon 3454: 3452: 3451: 3447: 3445: 3444: 3440: 3438: 3437: 3433: 3431: 3430: 3426: 3425: 3423: 3421: 3417: 3411: 3408: 3406: 3403: 3401: 3398: 3397: 3395: 3393: 3389: 3386: 3382: 3375: 3369: 3365: 3355: 3354: 3350: 3349: 3347: 3345: 3344:Anseranatidae 3341: 3335: 3334: 3330: 3328: 3327: 3323: 3322: 3320: 3318: 3314: 3308: 3305: 3303: 3300: 3298: 3295: 3291: 3286: 3284: 3279: 3278: 3277: 3274: 3270: 3267: 3265: 3262: 3260: 3257: 3256: 3255: 3252: 3251: 3249: 3245: 3239: 3235: 3232: 3226: 3222: 3219: 3215: 3209: 3183: 3179: 3176: 3174: 3152: 3148: 3138: 3132: 3130: 3124: 3122: 3118:Tinamiformes 3116: 3114: 3108: 3106: 3100: 3099: 3097: 3095: 3094:Palaeognathae 3091: 3087: 3083: 3076: 3072: 3068: 3060: 3057: 3055: 3052: 3051: 3049: 3047: 3044: 3042: 3039: 3037: 3034: 3032: 3029: 3027: 3024: 3022: 3019: 3017: 3014: 3013: 3011: 3007: 3001: 2998: 2996: 2993: 2991: 2988: 2986: 2983: 2981: 2980:Pigeon racing 2978: 2976: 2973: 2971: 2968: 2966: 2963: 2961: 2958: 2956: 2953: 2949: 2946: 2945: 2944: 2941: 2939: 2936: 2934: 2931: 2929: 2926: 2924: 2921: 2920: 2918: 2916: 2910: 2904: 2901: 2899: 2896: 2894: 2891: 2889: 2886: 2884: 2881: 2879: 2876: 2874: 2871: 2869: 2866: 2864: 2861: 2859: 2856: 2854: 2851: 2849: 2846: 2844: 2841: 2839: 2836: 2834: 2831: 2829: 2826: 2824: 2823: 2822:Archaeopteryx 2819: 2818: 2816: 2814: 2810: 2804: 2801: 2799: 2796: 2794: 2791: 2789: 2786: 2782: 2779: 2777: 2774: 2773: 2772: 2769: 2768: 2766: 2764: 2760: 2754: 2751: 2749: 2746: 2744: 2741: 2739: 2736: 2734: 2731: 2729: 2726: 2724: 2721: 2719: 2716: 2714: 2711: 2709: 2706: 2704: 2701: 2699: 2696: 2695: 2693: 2691: 2687: 2681: 2678: 2676: 2673: 2671: 2668: 2666: 2663: 2661: 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2077:Premaxillae ( 2074: 2071: 2067: 2063: 2057: 2054: 2050: 2046: 2042: 2038: 2034: 2030: 2025: 2022: 2016: 2013: 2008: 2004: 1999: 1994: 1990: 1986: 1982: 1978: 1974: 1970: 1966: 1959: 1956: 1951: 1945: 1941: 1937: 1933: 1926: 1924: 1920: 1907: 1903: 1899: 1895: 1889: 1887: 1883: 1879: 1875: 1871: 1867: 1861: 1858: 1852: 1849: 1843: 1838: 1831: 1828: 1823: 1819: 1814: 1809: 1804: 1799: 1795: 1791: 1787: 1780: 1778: 1774: 1769: 1763: 1759: 1752: 1749: 1744: 1740: 1735: 1730: 1725: 1720: 1716: 1712: 1708: 1704: 1700: 1693: 1690: 1685: 1681: 1677: 1673: 1668: 1663: 1659: 1655: 1651: 1647: 1643: 1639: 1632: 1629: 1622: 1618: 1615: 1614: 1610: 1608: 1601: 1596: 1595: 1594:Fluviatitavis 1591: 1588: 1584: 1581: 1580: 1576: 1573: 1572: 1568: 1565: 1564: 1560: 1557: 1556: 1552: 1549: 1547: 1542: 1539: 1538: 1534: 1533: 1532: 1530: 1520: 1518: 1517: 1513: 1510: 1509: 1505: 1502: 1501: 1497: 1494: 1493: 1489: 1486: 1485: 1481: 1478: 1477: 1473: 1472: 1471: 1469: 1464: 1459: 1458: 1454: 1453: 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882: 876: 875: 872: 871: 868: 866: 860: 859: 852: 851: 845: 844: 841: 840: 832: 831: 823: 822: 814: 813: 805: 804: 796: 795: 792: 790: 784: 783: 780: 779: 776: 774: 768: 767: 761: 760: 757: 756: 753: 751: 745: 744: 738: 737: 734: 733: 730: 728: 722: 721: 715: 714: 711: 710: 707: 705: 699: 698: 692: 691: 688: 687: 679: 678: 670: 669: 666: 664: 658: 657: 654: 653: 650: 648: 642: 641: 635: 634: 631: 630: 627: 625: 619: 618: 612: 611: 604: 603: 596: 595: 591: 590: 582: 578: 575: 571: 568: 564: 561: 557: 555:– crab-plover 554: 550: 547: 543: 542: 541: 537: 532: 528: 525: 521: 518: 514: 511: 507: 504: 503:Rostratulidae 500: 499: 498: 494: 489: 485: 482: 478: 475: 471: 468: 464: 461: 457: 454: 450: 447: 443: 440: 436: 435: 433: 432: 431: 429: 421: 419: 416: 413: 409: 408:Ciconiiformes 405: 404:birds of prey 401: 397: 389: 388: 383: 380: 376: 372: 371: 366: 363: 362: 357: 356: 355: 349: 347: 345: 341: 337: 336:invertebrates 333: 329: 325: 321: 317: 316: 309: 255: 247: 243: 238: 233: 228: 225: 222: 221: 218: 215: 212: 209: 208: 205: 202: 199: 198: 195: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 153: 148: 144: 139: 115: 110: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 34: 30: 22: 19: 4531:Wading birds 4199: 4172: 4160: 4148: 4136: 4046:(mousebirds) 4044:Coliiformes 3914:Gaviiformes 3906:Aequornithes 3855: 3814:Apodiformes 3775:Otidiformes 3751:Otidimorphae 3698:(sandgrouse) 3620:Phasianinae 3586: 3579: 3572: 3567:Odontophorus 3565: 3558: 3551: 3544: 3537: 3530: 3510: 3503: 3496: 3489: 3469: 3462: 3455: 3448: 3441: 3434: 3427: 3405:Oreophasinae 3351: 3331: 3324: 3229:(waterfowls) 3225:Anseriformes 2975:Cockfighting 2960:Conservation 2955:Bird feeding 2943:Birdwatching 2933:Ornithomancy 2873:Gansuiformes 2820: 2813:Fossil birds 2703:Intelligence 2546: 2541: 2533:PDF fulltext 2519: 2514: 2502:PDF fulltext 2481: 2476: 2459: 2454: 2450: 2416: 2388: 2383: 2359: 2354: 2339:PDF fulltext 2334: 2331: 2326:PDF fulltext 2305: 2300: 2295:PDF 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Retrieved 2133: 2129: 2119: 2111: 2106: 2098: 2073: 2065: 2056: 2048: 2024: 2015: 1972: 1968: 1958: 1939: 1910:. 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3553:Dactylortyx 3420:Megapodidae 3410:Penelopinae 3368:Galliformes 3110:Rheiformes 3104:(ostriches) 3054:individuals 2928:Ornithology 2915:interaction 2670:Preen gland 2549:(1): 1-39. 2391:: 268–278. 2176:white stork 2160:23 February 2136:: 261–266. 1912:19 December 1894:Gill, Frank 1796:: 108–127. 1734:11336/12701 1579:Eupterornis 1555:Palintropus 1492:Telmatornis 1367:Boutersemia 1359:Jiliniornis 1351:nomen nudum 1311:Vega Island 1299:semiaquatic 1058:Glareolidae 775:– ibisbill 704:Pluvianidae 560:Glareolidae 460:Pluvianidae 4515:Categories 3924:(penguins) 3777:(bustards) 3615:Perdicinae 3560:Dendrortyx 3532:Callipepla 3464:Megapodius 3429:Aepypodius 3307:Tadorninae 3289:true geese 3120:(tinamous) 3079:Neornithes 2990:Pheasantry 2965:Aviculture 2733:Incubation 2723:Lek mating 2337:: 33–78. 2259:References 1667:11336/7192 1589:, Morocco) 1537:Ceramornis 1516:Dakotornis 1500:Scaniornis 1476:Graculavus 1429:form taxon 1344:Morsoravis 1319:thick-knee 1287:Neornithes 1202:Frank Gill 1042:Dromadidae 1012:Turnicidae 853:Scolopaci 624:Burhinidae 605:Charadrii 553:Dromadidae 546:Turnicidae 483:– ibisbill 439:Burhinidae 344:shorebirds 322:of family 320:type genus 315:Charadrius 246:See text. 204:Neognathae 38:55–0  24:Shorebirds 3993:(parrots) 3837:(hoatzin) 3788:Strisores 3769:(turacos) 3761:(cuckoos) 3690:(mesites) 3609:(turkeys) 3581:Philortyx 3498:Agelastes 3491:Acryllium 3482:Numididae 3471:Talegalla 3381:gamebirds 3374:landfowls 3353:Anseranas 3317:Anhimidae 3276:Anserinae 3059:fictional 2781:dinosaurs 2776:Theropoda 2763:Evolution 2708:Migration 2690:Behaviour 2443:0065-3454 2302:Evolution 2220:0065-3454 2150:140700031 2114:(1998-99) 2097:: Worthy 2095:kelp gull 2064:: Worthy 2047:: Worthy 1989:1744-9561 1743:0031-0301 1676:0031-0301 1623:Footnotes 1437:flamingos 1433:waterfowl 1331:Oligocene 1323:Paleogene 1259:June 2021 911:Jacanidae 663:Chionidae 538:Suborder 510:Jacanidae 497:Scolopaci 495:Suborder 453:Chionidae 428:suborders 240:Families 170:Kingdom: 164:Eukaryota 4526:Seabirds 4209:Wikidata 4138:Category 4012:Afroaves 3940:(storks) 3742:Passerea 3727:(grebes) 3663:Columbea 3574:Oreortyx 3546:Cyrtonyx 3436:Alectura 3400:Cracinae 3392:Cracidae 3269:Oxyurini 3259:Aythyini 3254:Anatinae 3238:Anatidae 2985:Falconry 2948:big year 2803:Seabirds 2713:Foraging 2660:Feathers 2510:251–257. 2498:15329156 2405:13678682 2376:16531074 2350:283–293. 2322:15612298 2233:251–257. 2154:Archived 2083:proximal 2029:Proximal 2007:17284401 1938:(eds.). 1906:Archived 1864:Ericson 1822:32781465 1684:85141394 1611:See also 1571:Volgavis 1508:Zhylgaia 1426:Mesozoic 1375:Turnipax 422:Families 400:seabirds 184:Chordata 180:Phylum: 174:Animalia 160:Domain: 33:Holocene 4371:7192402 4309:2917972 4265:Avibase 4150:Commons 3654:Neoaves 3539:Colinus 3505:Guttera 3443:Eulipoa 3264:Mergini 3128:(kiwis) 3112:(rheas) 2923:Ringing 2748:Hybrids 2743:Nesting 2698:Singing 2675:Plumage 2650:Dactyly 2620:Anatomy 2610:Outline 2600:: Aves) 2284:: 16. 2087:scapula 2033:humerus 1998:2375939 1837:bioRxiv 1813:7783168 1711:Bibcode 1646:Bibcode 1563:Torotix 1457:Laornis 1383:Elorius 1129:Alcidae 1090:Laridae 581:Alcidae 579:Family 572:Family 567:Laridae 565:Family 558:Family 551:Family 544:Family 529:Family 522:Family 515:Family 508:Family 501:Family 486:Family 479:Family 472:Family 465:Family 458:Family 451:Family 444:Family 437:Family 340:pelagic 332:species 312:, from 223:Order: 217:Neoaves 190:Class: 4459:313124 4443:NZOR: 4410:176445 4322:1CHDRO 4215:Q25978 4162:Portal 4038:(owls) 3512:Numida 3450:Leipoa 3333:Chauna 3326:Anhima 3021:Genera 2995:Imping 2913:Human 2753:Colony 2680:Vision 2665:Flight 2496:  2484:: 28. 2441:  2431:  2403:  2374:  2320:  2218:  2208:  2148:  2112:et al. 2101:(2007) 2099:et al. 2085:right 2068:(2007) 2066:et al. 2051:(2007) 2049:et al. 2031:right 2005:  1995:  1987:  1946:  1878:et al. 1874:et al. 1870:et al. 1866:et al. 1839:  1820:  1810:  1764:  1741:  1682:  1674:  1327:Eocene 361:waders 318:, the 234:, 1867 232:Huxley 29:Eocene 4493:WoRMS 4480:Plazi 4472:36640 4397:10265 4392:IRMNG 4384:67561 4335:10769 3282:swans 3244:ducks 3214:fowls 3009:Lists 2633:Brain 2598:class 2594:Birds 2146:S2CID 1680:S2CID 1315:basal 394:The 379:skuas 370:gulls 328:birds 211:Clade 4498:2682 4436:8906 4431:NCBI 4405:ITIS 4366:GBIF 4317:EPPO 4278:BOLD 2655:Eggs 2643:milk 2638:Crop 2628:Beak 2494:PMID 2439:ISSN 2429:ISBN 2401:PMID 2372:PMID 2318:PMID 2216:ISSN 2206:ISBN 2162:2024 2110:GĂĄl 2003:PMID 1985:ISSN 1944:ISBN 1914:2021 1818:PMID 1762:ISBN 1739:ISSN 1672:ISSN 1435:and 999:Lari 540:Lari 402:and 387:auks 384:The 375:Lari 367:The 358:The 194:Aves 46:PreꞒ 4418:NBN 4304:EoL 4291:CoL 4283:300 4252:AFD 4239:ADW 2552:doi 2525:doi 2487:doi 2465:doi 2421:doi 2394:doi 2365:doi 2311:doi 2287:doi 2198:doi 2138:doi 2079:MNZ 2037:MNZ 1993:PMC 1977:doi 1808:PMC 1798:doi 1729:hdl 1719:doi 1662:hdl 1654:doi 1441:mya 1335:mya 1309:of 1249:by 296:ɔːr 4517:: 4495:: 4482:: 4469:: 4456:: 4433:: 4420:: 4407:: 4394:: 4381:: 4368:: 4345:: 4332:: 4319:: 4306:: 4296:X3 4293:: 4280:: 4267:: 4254:: 4241:: 4226:: 4211:: 2520:35 2460:30 2437:. 2427:. 2389:29 2360:39 2335:23 2306:58 2269:. 2214:. 2204:. 2152:. 2144:. 2134:89 2132:. 2128:. 2001:. 1991:. 1983:. 1971:. 1967:. 1934:; 1922:^ 1885:^ 1816:. 1806:. 1794:38 1792:. 1788:. 1776:^ 1737:. 1727:. 1717:. 1707:48 1705:. 1701:. 1678:. 1670:. 1660:. 1652:. 1642:47 1640:. 1305:, 1204:, 346:. 302:iː 213:: 96:Pg 40:Ma 35:, 3383:) 3376:- 3372:( 3246:) 3242:( 3216:) 3212:( 3208:e 3206:a 3204:r 3202:e 3200:s 3198:n 3196:a 3194:o 3192:l 3190:l 3188:a 3186:G 3173:e 3171:a 3169:h 3167:t 3165:a 3163:n 3161:g 3159:o 3157:e 3155:N 2596:( 2586:e 2579:t 2572:v 2554:: 2547:5 2527:: 2489:: 2482:4 2467:: 2451:b 2445:. 2423:: 2396:: 2367:: 2313:: 2289:: 2282:3 2222:. 2200:: 2178:( 2164:. 2140:: 2035:( 2009:. 1979:: 1973:3 1952:. 1916:. 1845:. 1824:. 1800:: 1770:. 1745:. 1731:: 1721:: 1713:: 1686:. 1664:: 1656:: 1648:: 1548:" 1544:" 1470:" 1466:" 1354:? 1346:" 1342:" 1329:- 1272:) 1266:( 1261:) 1257:( 1243:. 308:/ 305:z 299:m 293:f 290:ÉȘ 287:. 284:i 281:r 278:d 275:ĂŠ 272:r 269:ˈ 266:ə 263:k 260:/ 256:( 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ 31:-

Index

Eocene
Holocene
Ma
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N
molecular clock
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Neognathae
Neoaves
Charadriiformes
Huxley
/kəˈrĂŠdri.ÉȘfɔːrmiːz/
Charadrius
type genus
Charadriidae

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