42:
68:
1063:
870:"convention On International Trade in Endangered Species of Wild Fauna and Flora: Sixteenth meeting of the Conference of the parties, Bangkok (Thailand), 3-14 March 2013. Consideration of Proposals for Amendment of Appendices I and II (CoP16 Prop. 32)"
749:. 2012. Turtles of the World, 2012 update: annotated checklist of taxonomy, synonymy, distribution, and conservation status. Chelonian Research Monographs No. 5, pp. 000.243–000.328, doi:10.3854/crm.5.000.checklist.v5.2012,
732:
Gray, John Edward. 1870. Supplement to the
Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). London: British Museum, 120 pp.
751:
4149:
382:
The intrafamilial taxonomy is not well established yet, due to the large number and diversity of species. The family is usually divided into two subfamilies and 19
4175:
361:
900:
710:
Theobald, William, Jr. 1868. Catalogue of
Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society, Extra Number, 88 pp.
695:. A joint China-US proposal for a March 2013 CITES participants' conference seeks to add 15 more Geoemydidae species to the convention's Appendix II.
4123:
784:"A ZZ/ZW microchromosome system in the spiny softshell turtle, Apalone spinifera, reveals an intriguing sex chromosome conservation in Trionychidae"
917:
719:
Gray, John Edward. 1869. Notes on the families and genera of tortoises (Testudinata), and on the characters afforded by the study of their skulls.
4275:
4162:
965:
1825:
1190:
1345:
1260:
1240:
869:
4237:
244:. The plastron is composed of 12 scutes and has no mesoplastron; the pectoral and abdominal scutes contact the marginal scutes.
224:
Geoemydidae are turtles of various sizes (from about 10 to 80 cm (4 to 30 in) in length) with often a high degree of
3858:
2941:
1180:
758:
4167:
3985:
Turtles of the World 2017 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
1794:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
746:
3835:
1485:
3983:
1793:
41:
4058:
3994:
2517:
1901:
1693:
1580:
1571:
1440:
1400:
1130:
934:
334:
324:
752:"Turtles of the World, 2012 update: annotated checklist of taxonomy, synonymy, distribution, and conservation status"
67:
4180:
4102:
2250:
1450:
1445:
1340:
4115:
1250:
1120:
958:
4270:
2896:
1818:
1761:
1710:
1195:
310:
54:
1291:
1299:
3752:
1703:
1683:
1535:
1390:
1255:
1155:
1125:
1110:
360:. In the 1980s, the subfamily was elevated to the family status and renamed to Geoemydidae according to the
4011:
3724:
2524:
2482:
2122:
1731:
1600:
1591:
1505:
1495:
1420:
1411:
1165:
1160:
849:
248:
4063:
1283:
4201:
2468:
1970:
1688:
1673:
1365:
1356:
1245:
1105:
924:
372:
4219:
4265:
4242:
4084:
3710:
2376:
2000:
1678:
1630:
1490:
1385:
1210:
951:
285:. Their habitats include freshwater ecosystems, coastal marine areas, and tropical forests. Most are
3695:
2054:
1811:
1698:
1625:
1620:
1555:
1530:
1500:
1480:
1175:
329:
3102:
3067:
2934:
2732:
2673:
2643:
1756:
1642:
1635:
1560:
1510:
1470:
1200:
1185:
1150:
819:
341:
318:
160:
62:
49:
4206:
2651:
2405:
423:
4188:
4071:
3645:
2628:
2580:
2571:
2454:
2031:
1668:
1663:
1320:
1311:
1235:
1230:
1205:
1170:
811:
803:
692:
252:
225:
4229:
4154:
4049:
4193:
3437:
2825:
2800:
2761:
2709:
2440:
1985:
1736:
1141:
1115:
795:
688:
387:
197:
876:
3885:
3533:
3262:
3230:
2867:
2776:
2552:
2138:
1883:
1747:
623:
276:
4110:
4097:
3780:
3770:
3667:
3517:
3357:
3059:
3004:
2746:
2681:
2221:
2212:
2181:
2089:
1978:
1892:
1475:
314:
839:(2005): On the hybridisation between two distantly related Asian turtles (Testudines:
4259:
3960:
3843:
3743:
3732:
3703:
3617:
3501:
3485:
3445:
3379:
3289:
3254:
3198:
3182:
3166:
3140:
3128:
3112:
3090:
3082:
2889:
2600:
2433:
2294:
2241:
2189:
1084:
1073:
1023:
782:
Badenhorst, Daleen; Stanyon, Roscoe; Engstrom, Tag; Valenzuela, Nicole (2013-04-01).
518:
412:
306:
229:
139:
857:
823:
309:
is common, produced several times a year. Some species have a temperature-dependent
301:, the males are usually much more active than females. A relatively small number of
4076:
3895:
3820:
3796:
3313:
3190:
3174:
3150:
3051:
2996:
2920:
2836:
2810:
2724:
2635:
2614:
2538:
2503:
2313:
2279:
2146:
2062:
2008:
1654:
1521:
1395:
667:
659:
640:
565:
376:
281:
272:
260:
256:
4136:
4043:
3873:
3828:
3674:
3659:
3581:
3573:
3565:
3541:
3421:
3405:
3389:
3305:
3297:
3246:
3222:
3214:
3206:
3074:
3026:
2949:
2874:
2739:
2695:
2621:
2607:
2559:
2334:
2272:
2265:
2258:
2173:
2115:
2107:
2024:
2016:
1963:
1953:
1781:
1772:
431:
357:
213:
3952:
3865:
3851:
3812:
3589:
3509:
3477:
3461:
3429:
3367:
3340:
3321:
3238:
3043:
2963:
2956:
2904:
2881:
2791:
2658:
2390:
2197:
2130:
2047:
2039:
1942:
1933:
1272:
1017:
1011:
799:
583:
201:
129:
4034:
807:
3944:
3804:
3788:
3717:
3681:
3631:
3624:
3549:
3525:
3469:
3413:
3120:
2912:
2860:
2818:
2531:
2489:
2397:
2383:
2369:
2362:
2355:
2348:
2320:
2165:
1993:
1611:
1431:
783:
601:
535:
479:
474:
294:
286:
79:
20:
916:
Phillip Q. Spinks; H. Bradley
Shaffer; John B. Iverson; William P. McCord.
815:
4028:
3759:
3688:
3608:
3597:
3493:
3158:
3035:
2927:
2769:
2754:
2716:
2665:
2591:
2496:
2475:
2461:
2447:
2303:
2230:
2099:
2078:
2070:
1852:
1461:
1376:
1331:
1221:
993:
544:
505:
496:
487:
465:
353:
290:
237:
233:
99:
3903:
3652:
3557:
3453:
3397:
2977:
2970:
2852:
2702:
2688:
2545:
2510:
2425:
2327:
1867:
1858:
1722:
1546:
1096:
1005:
999:
684:
As of the early 2013, six species of the family
Geoemydidae are on the
614:
592:
574:
439:
209:
205:
109:
4141:
3638:
3281:
1873:
1846:
1834:
987:
943:
368:
298:
119:
89:
4128:
4005:
938:
4214:
2845:
1912:
685:
452:
383:
241:
2341:
4009:
1807:
947:
271:
Geoemydidae live in tropics and subtropics of Asia, Europe and
302:
4089:
1803:
352:
Traditional systematics placed the geoemydids in the family
340:
About 70% of the extant species have been reported to be in
918:"Phylogenetic hypotheses for the turtle family Geoemydidae"
837:
Buskirk, James R.; Parham, James F. & Feldman, Chris R.
901:
China Backs
Tortoise in Race to Protect Endangered Species
398:
236:
flexibly. Their necks are drawn back vertically. Their
390:, which makes the systematics even more complicated.
419:
408:
375:
data support their close relationship to the family
4018:
3935:
3928:
3921:
3883:
3769:
3742:
3607:
3377:
3355:
3348:
3339:
3272:
3138:
3100:
3025:
3018:
2987:
2835:
2799:
2790:
2590:
2569:
2415:
2302:
2293:
2240:
2220:
2211:
2156:
2088:
1952:
1941:
1932:
1900:
1891:
1882:
1770:
1745:
1720:
1652:
1609:
1589:
1569:
1544:
1519:
1459:
1429:
1409:
1374:
1354:
1329:
1309:
1269:
1219:
1139:
1094:
1070:
1038:
721:Proceedings of the Zoological Society of London
3982:Phylogenetic arrangement of turtles based on
1819:
1792:Phylogenetic arrangement of turtles based on
959:
8:
255:, the lack of a hyomandibular branch of the
456:, Asian box turtle (10 species) (including
386:. Several species are known to give viable
228:. They usually have webbed toes, and their
4006:
3932:
3925:
3352:
3345:
3022:
2796:
2299:
2217:
1949:
1938:
1906:
1897:
1888:
1826:
1812:
1804:
1042:
966:
952:
944:
196:) are one of the largest and most diverse
40:
27:
34:Temporal range: Late Cretaceous - Present
703:
313:system, while others possess different
742:
740:
738:
247:Some other features include a single
7:
4220:726CA836-1A1F-D038-FF4B-FAE93239FA3E
4116:e5204723-403b-4f0f-93c0-4bf2c8c38e90
1191:Vietnamese three-striped box turtle
691:, and 30 more are on the treaty's
14:
1346:Ryukyu black-breasted leaf turtle
1261:Western black-bridged leaf turtle
1241:Eastern black-bridged leaf turtle
1061:
443:(six species, including part of
66:
4276:Taxa named by William Theobald
1181:Southern Vietnamese box turtle
55:Rhinoclemmys pulcherrima manni
1:
747:Turtle Taxonomy Working Group
1486:Chinese stripe-necked turtle
605:(four species) (formerly in
251:between the fifth and sixth
3995:List of Testudines families
1694:Mexican spotted wood turtle
1581:Malayan flat-shelled turtle
1441:Malayan snail-eating turtle
1401:Yellow-headed temple turtle
1131:Three-striped roofed turtle
644:, cane turtle (formerly in
627:(two species, one subgenus
320:Siebenrockiella crassicolis
212:pond and river turtles and
4292:
1451:Khorat snail-eating turtle
1446:Mekong snail-eating turtle
1341:Black-breasted leaf turtle
1059:
671:, Neotropical wood turtles
548:, pond turtles (including
208:. The family includes the
18:
3978:
1909:
1841:
1790:
1251:Myanmar brown leaf turtle
1121:Red-crowned roofed turtle
1045:
982:
800:10.1007/s10577-013-9343-2
348:Systematics and evolution
344:or vulnerable condition.
327:, while another species (
204:(turtles), with about 70
166:
159:
63:Scientific classification
61:
48:
39:
30:
1762:Philippine forest turtle
1711:Rhinoclemmys panamaensis
1292:Echmatemys stevensoniana
1196:Yellow-headed box turtle
19:Not to be confused with
1704:Spot-legged wood turtle
1684:Large-nosed wood turtle
1536:Tricarinate hill turtle
1391:Giant Asian pond turtle
1300:Echmatemys wyomingensis
1126:Southern river terrapin
1111:Northern river terrapin
335:ZZ/ZW sex determination
325:XX/XY sex determination
1601:Malaysian giant turtle
1506:Vietnamese pond turtle
1496:Red-necked pond turtle
1421:Sulawesi forest turtle
1161:Indochinese box turtle
1047:Species of the family
394:Subfamilies and genera
232:articulate with their
4202:Paleobiology Database
1689:Maracaibo wood turtle
1674:Colombian wood turtle
1366:Brahminy river turtle
1246:Enigmatic leaf turtle
1106:Burmese roofed turtle
317:; one known species (
279:and South America is
4111:Fauna Europaea (new)
1679:Furrowed wood turtle
1631:Indian roofed turtle
1491:Japanese pond turtle
1386:Arakan forest turtle
1256:Oldham's leaf turtle
1156:Bourret's box turtle
939:The Reptile Database
275:; the only genus in
1732:Beale's eyed turtle
1699:Painted wood turtle
1626:Brown roofed turtle
1621:Assam roofed turtle
1556:Burmese eyed turtle
1531:Indian black turtle
1501:Spanish pond turtle
1481:Chinese pond turtle
1284:Echmatemys septaria
1166:McCord's box turtle
1085:Banhxeochelys trani
788:Chromosome Research
263:bone in the skull.
192:(formerly known as
3103:Nanhsiungchelyidae
3068:Macrocephalochelys
1757:Black marsh turtle
1643:Pangshura tatrotia
1636:Indian tent turtle
1561:Indian eyed turtle
1511:Yellow pond turtle
1471:Balkan pond turtle
1201:Chinese box turtle
1186:Golden coin turtle
1151:Amboina box turtle
403:Family Geoemydidae
253:cervical vertebrae
50:Ornate wood turtle
4253:
4252:
4189:Open Tree of Life
4012:Taxon identifiers
4003:
4002:
3974:
3973:
3970:
3969:
3917:
3916:
3913:
3912:
3335:
3334:
3331:
3330:
3014:
3013:
2786:
2785:
2289:
2288:
2207:
2206:
1928:
1927:
1924:
1923:
1920:
1919:
1801:
1800:
1669:Brown wood turtle
1664:Black wood turtle
1321:Black pond turtle
1236:Assam leaf turtle
1231:Asian leaf turtle
1211:Zhou's box turtle
1206:Yunnan box turtle
1171:Keeled box turtle
1057:
1056:
937:(all species) at
631:, formerly under
596:(monotypic genus)
587:(monotypic genus)
500:(monotypic genus)
356:as the subfamily
330:Pangshura smithii
311:sex determination
240:have 24 marginal
226:sexual dimorphism
186:
185:
181:
176:
171:
155:
16:Family of turtles
4283:
4246:
4245:
4233:
4232:
4223:
4222:
4210:
4209:
4197:
4196:
4184:
4183:
4171:
4170:
4158:
4157:
4145:
4144:
4132:
4131:
4119:
4118:
4106:
4105:
4093:
4092:
4080:
4079:
4067:
4066:
4054:
4053:
4052:
4039:
4038:
4037:
4007:
3933:
3926:
3438:Chupacabrachelys
3353:
3346:
3023:
2801:Carettochelyidae
2797:
2300:
2218:
1950:
1939:
1907:
1898:
1889:
1828:
1821:
1814:
1805:
1737:Four-eyed turtle
1176:Pan's box turtle
1116:Painted terrapin
1065:
1064:
1043:
968:
961:
954:
945:
931:
929:
923:. Archived from
922:
904:
897:
891:
890:
888:
887:
881:
875:. Archived from
874:
866:
860:
838:
834:
828:
827:
779:
773:
772:
770:
769:
763:
757:. Archived from
756:
744:
733:
730:
724:
717:
711:
708:
660:Rhinoclemmydinae
618:, "eyed" turtles
421:
410:
179:
174:
169:
153:
71:
70:
44:
28:
4291:
4290:
4286:
4285:
4284:
4282:
4281:
4280:
4271:Turtle families
4256:
4255:
4254:
4249:
4241:
4236:
4228:
4226:
4218:
4213:
4205:
4200:
4192:
4187:
4179:
4174:
4166:
4161:
4153:
4148:
4140:
4135:
4127:
4122:
4114:
4109:
4101:
4096:
4088:
4083:
4075:
4070:
4062:
4057:
4048:
4047:
4042:
4033:
4032:
4027:
4014:
4004:
3999:
3966:
3909:
3886:Sahonachelyidae
3879:
3765:
3738:
3603:
3534:Phosphatochelys
3373:
3327:
3268:
3263:Terlinguachelys
3231:Pneumatoarthrus
3134:
3096:
3010:
2983:
2831:
2782:
2586:
2565:
2553:Siebenrockiella
2411:
2285:
2236:
2203:
2152:
2139:Mesodermochelys
2084:
1945:
1916:
1878:
1837:
1832:
1802:
1797:
1786:
1766:
1748:Siebenrockiella
1741:
1716:
1648:
1605:
1585:
1565:
1540:
1515:
1455:
1425:
1405:
1370:
1350:
1325:
1305:
1265:
1215:
1135:
1090:
1066:
1062:
1053:
1034:
978:
972:
927:
920:
915:
912:
907:
899:Dinny McMahon,
898:
894:
885:
883:
879:
872:
868:
867:
863:
836:
835:
831:
781:
780:
776:
767:
765:
761:
754:
750:
745:
736:
731:
727:
718:
714:
709:
705:
701:
682:
624:Siebenrockiella
539:(three species)
469:(seven species)
396:
350:
315:sex chromosomes
289:, but some are
269:
222:
220:Characteristics
177:
172:
170:Theobald 1868:9
152:
65:
52:
35:
32:
24:
17:
12:
11:
5:
4289:
4287:
4279:
4278:
4273:
4268:
4258:
4257:
4251:
4250:
4248:
4247:
4234:
4224:
4211:
4198:
4185:
4172:
4159:
4146:
4133:
4120:
4107:
4098:Fauna Europaea
4094:
4081:
4068:
4055:
4040:
4024:
4022:
4016:
4015:
4010:
4001:
4000:
3998:
3997:
3990:
3989:
3988:. † = extinct.
3979:
3976:
3975:
3972:
3971:
3968:
3967:
3965:
3964:
3956:
3948:
3939:
3937:
3930:
3923:
3919:
3918:
3915:
3914:
3911:
3910:
3908:
3907:
3899:
3890:
3888:
3881:
3880:
3878:
3877:
3869:
3862:
3855:
3847:
3839:
3832:
3824:
3816:
3808:
3800:
3792:
3784:
3781:Albertwoodemys
3775:
3773:
3771:Podocnemididae
3767:
3766:
3764:
3763:
3756:
3748:
3746:
3740:
3739:
3737:
3736:
3728:
3721:
3714:
3707:
3699:
3692:
3685:
3678:
3671:
3668:Lomalatachelys
3663:
3656:
3649:
3642:
3635:
3628:
3621:
3613:
3611:
3605:
3604:
3602:
3601:
3593:
3585:
3577:
3569:
3561:
3553:
3545:
3537:
3529:
3521:
3518:Labrostochelys
3513:
3505:
3497:
3489:
3481:
3473:
3465:
3457:
3449:
3441:
3433:
3425:
3417:
3409:
3401:
3393:
3384:
3382:
3375:
3374:
3372:
3371:
3362:
3360:
3358:Araripemydidae
3350:
3343:
3337:
3336:
3333:
3332:
3329:
3328:
3326:
3325:
3317:
3309:
3301:
3293:
3285:
3276:
3274:
3270:
3269:
3267:
3266:
3258:
3250:
3242:
3234:
3226:
3218:
3210:
3202:
3194:
3186:
3178:
3170:
3162:
3154:
3145:
3143:
3136:
3135:
3133:
3132:
3124:
3116:
3107:
3105:
3098:
3097:
3095:
3094:
3086:
3078:
3071:
3063:
3060:Emarginachelys
3055:
3047:
3039:
3031:
3029:
3020:
3016:
3015:
3012:
3011:
3009:
3008:
3005:Sinaspideretes
3000:
2991:
2989:
2985:
2984:
2982:
2981:
2974:
2967:
2960:
2953:
2945:
2938:
2931:
2924:
2916:
2908:
2900:
2893:
2885:
2878:
2871:
2864:
2856:
2849:
2841:
2839:
2833:
2832:
2830:
2829:
2822:
2814:
2805:
2803:
2794:
2788:
2787:
2784:
2783:
2781:
2780:
2773:
2765:
2758:
2750:
2743:
2736:
2728:
2720:
2713:
2706:
2699:
2692:
2685:
2682:Hesperotestudo
2677:
2669:
2662:
2655:
2647:
2639:
2632:
2625:
2618:
2611:
2604:
2596:
2594:
2588:
2587:
2585:
2584:
2576:
2574:
2572:Platysternidae
2567:
2566:
2564:
2563:
2556:
2549:
2542:
2535:
2528:
2521:
2514:
2507:
2500:
2493:
2486:
2479:
2472:
2465:
2458:
2451:
2444:
2437:
2429:
2421:
2419:
2413:
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2409:
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2394:
2387:
2380:
2373:
2366:
2359:
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2338:
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2324:
2317:
2308:
2306:
2297:
2291:
2290:
2287:
2286:
2284:
2283:
2276:
2269:
2262:
2254:
2246:
2244:
2238:
2237:
2235:
2234:
2226:
2224:
2222:Dermatemydidae
2215:
2213:Kinosternoidea
2209:
2208:
2205:
2204:
2202:
2201:
2193:
2185:
2182:Procolpochelys
2177:
2169:
2160:
2158:
2154:
2153:
2151:
2150:
2142:
2134:
2126:
2119:
2111:
2103:
2094:
2092:
2090:Dermochelyidae
2086:
2085:
2083:
2082:
2074:
2066:
2058:
2051:
2043:
2035:
2028:
2020:
2012:
2004:
1997:
1989:
1982:
1979:Carolinochelys
1974:
1967:
1958:
1956:
1947:
1936:
1930:
1929:
1926:
1925:
1922:
1921:
1918:
1917:
1910:
1904:
1895:
1886:
1880:
1879:
1877:
1876:
1870:
1861:
1855:
1849:
1842:
1839:
1838:
1833:
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1808:
1799:
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1527:
1525:
1517:
1516:
1514:
1513:
1508:
1503:
1498:
1493:
1488:
1483:
1478:
1476:Caspian turtle
1473:
1467:
1465:
1457:
1456:
1454:
1453:
1448:
1443:
1437:
1435:
1427:
1426:
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1026:
1020:
1014:
1008:
1002:
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983:
980:
979:
973:
971:
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956:
948:
942:
941:
932:
930:on 2006-09-05.
911:
910:External links
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906:
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829:
794:(2): 137–147.
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268:
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230:pelvic girdles
221:
218:
216:wood turtles.
184:
183:
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163:
157:
156:
154:Theobald, 1868
147:
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4017:
4013:
4008:
3996:
3992:
3991:
3987:
3986:
3981:
3980:
3977:
3963:
3962:
3961:Tacuarembemys
3957:
3955:
3954:
3949:
3947:
3946:
3941:
3940:
3938:
3934:
3931:
3927:
3924:
3920:
3906:
3905:
3900:
3898:
3897:
3896:Sahonachelys
3892:
3891:
3889:
3887:
3882:
3876:
3875:
3870:
3868:
3867:
3863:
3861:
3860:
3859:Peltocephalus
3856:
3854:
3853:
3848:
3846:
3845:
3844:Lapparentemys
3840:
3838:
3837:
3833:
3831:
3830:
3825:
3823:
3822:
3817:
3815:
3814:
3809:
3807:
3806:
3801:
3799:
3798:
3793:
3791:
3790:
3785:
3783:
3782:
3777:
3776:
3774:
3772:
3768:
3762:
3761:
3757:
3755:
3754:
3750:
3749:
3747:
3745:
3744:Pelomedusidae
3741:
3735:
3734:
3733:Yaminuechelys
3729:
3727:
3726:
3722:
3720:
3719:
3715:
3713:
3712:
3708:
3706:
3705:
3704:Prochelidella
3700:
3698:
3697:
3693:
3691:
3690:
3686:
3684:
3683:
3679:
3677:
3676:
3672:
3670:
3669:
3664:
3662:
3661:
3657:
3655:
3654:
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3634:
3633:
3629:
3627:
3626:
3622:
3620:
3619:
3618:Acanthochelys
3615:
3614:
3612:
3610:
3606:
3600:
3599:
3594:
3592:
3591:
3586:
3584:
3583:
3578:
3576:
3575:
3570:
3568:
3567:
3562:
3560:
3559:
3554:
3552:
3551:
3546:
3544:
3543:
3538:
3536:
3535:
3530:
3528:
3527:
3522:
3520:
3519:
3514:
3512:
3511:
3506:
3504:
3503:
3502:Kinkonychelys
3498:
3496:
3495:
3490:
3488:
3487:
3486:Itapecuruemys
3482:
3480:
3479:
3474:
3472:
3471:
3466:
3464:
3463:
3458:
3456:
3455:
3450:
3448:
3447:
3446:Eotaphrosphys
3442:
3440:
3439:
3434:
3432:
3431:
3426:
3424:
3423:
3418:
3416:
3415:
3410:
3408:
3407:
3402:
3400:
3399:
3394:
3392:
3391:
3386:
3385:
3383:
3381:
3380:Bothremydidae
3376:
3370:
3369:
3364:
3363:
3361:
3359:
3354:
3351:
3347:
3344:
3342:
3338:
3324:
3323:
3318:
3316:
3315:
3310:
3308:
3307:
3302:
3300:
3299:
3294:
3292:
3291:
3290:Argillochelys
3286:
3284:
3283:
3278:
3277:
3275:
3271:
3265:
3264:
3259:
3257:
3256:
3255:Santanachelys
3251:
3249:
3248:
3243:
3241:
3240:
3235:
3233:
3232:
3227:
3225:
3224:
3219:
3217:
3216:
3211:
3209:
3208:
3203:
3201:
3200:
3199:Desmatochelys
3195:
3193:
3192:
3187:
3185:
3184:
3183:Calcarichelys
3179:
3177:
3176:
3171:
3169:
3168:
3167:Atlantochelys
3163:
3161:
3160:
3155:
3153:
3152:
3147:
3146:
3144:
3142:
3141:Protostegidae
3137:
3131:
3130:
3129:Jiangxichelys
3125:
3123:
3122:
3117:
3115:
3114:
3113:Anomalochelys
3109:
3108:
3106:
3104:
3099:
3093:
3092:
3091:Protochelydra
3087:
3085:
3084:
3083:Planiplastron
3079:
3077:
3076:
3072:
3070:
3069:
3064:
3062:
3061:
3056:
3054:
3053:
3048:
3046:
3045:
3040:
3038:
3037:
3033:
3032:
3030:
3028:
3024:
3021:
3017:
3007:
3006:
3001:
2999:
2998:
2993:
2992:
2990:
2986:
2980:
2979:
2975:
2973:
2972:
2968:
2966:
2965:
2961:
2959:
2958:
2954:
2952:
2951:
2946:
2944:
2943:
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2923:
2922:
2917:
2915:
2914:
2909:
2907:
2906:
2901:
2899:
2898:
2894:
2892:
2891:
2890:Drazinderetes
2886:
2884:
2883:
2879:
2877:
2876:
2872:
2870:
2869:
2865:
2863:
2862:
2857:
2855:
2854:
2850:
2848:
2847:
2843:
2842:
2840:
2838:
2834:
2828:
2827:
2826:Carettochelys
2823:
2821:
2820:
2815:
2813:
2812:
2807:
2806:
2804:
2802:
2798:
2795:
2793:
2789:
2779:
2778:
2774:
2772:
2771:
2766:
2764:
2763:
2759:
2757:
2756:
2751:
2749:
2748:
2744:
2742:
2741:
2737:
2735:
2734:
2729:
2727:
2726:
2721:
2719:
2718:
2714:
2712:
2711:
2710:Malacochersus
2707:
2705:
2704:
2700:
2698:
2697:
2693:
2691:
2690:
2686:
2684:
2683:
2678:
2676:
2675:
2670:
2668:
2667:
2663:
2661:
2660:
2656:
2654:
2653:
2648:
2646:
2645:
2640:
2638:
2637:
2633:
2631:
2630:
2626:
2624:
2623:
2619:
2617:
2616:
2612:
2610:
2609:
2605:
2603:
2602:
2601:Aldabrachelys
2598:
2597:
2595:
2593:
2589:
2583:
2582:
2578:
2577:
2575:
2573:
2568:
2562:
2561:
2557:
2555:
2554:
2550:
2548:
2547:
2543:
2541:
2540:
2536:
2534:
2533:
2529:
2527:
2526:
2522:
2520:
2519:
2515:
2513:
2512:
2508:
2506:
2505:
2501:
2499:
2498:
2494:
2492:
2491:
2487:
2485:
2484:
2483:Leucocephalon
2480:
2478:
2477:
2473:
2471:
2470:
2466:
2464:
2463:
2459:
2457:
2456:
2452:
2450:
2449:
2445:
2443:
2442:
2438:
2436:
2435:
2434:Banhxeochelys
2430:
2428:
2427:
2423:
2422:
2420:
2418:
2414:
2408:
2407:
2402:
2400:
2399:
2395:
2393:
2392:
2388:
2386:
2385:
2381:
2379:
2378:
2374:
2372:
2371:
2367:
2365:
2364:
2360:
2358:
2357:
2353:
2351:
2350:
2346:
2344:
2343:
2339:
2337:
2336:
2332:
2330:
2329:
2325:
2323:
2322:
2318:
2316:
2315:
2310:
2309:
2307:
2305:
2301:
2298:
2296:
2295:Testudinoidea
2292:
2282:
2281:
2277:
2275:
2274:
2270:
2268:
2267:
2263:
2261:
2260:
2255:
2253:
2252:
2248:
2247:
2245:
2243:
2242:Kinosternidae
2239:
2233:
2232:
2228:
2227:
2225:
2223:
2219:
2216:
2214:
2210:
2200:
2199:
2194:
2192:
2191:
2190:Protosphargis
2186:
2184:
2183:
2178:
2176:
2175:
2170:
2168:
2167:
2162:
2161:
2159:
2155:
2149:
2148:
2143:
2141:
2140:
2135:
2133:
2132:
2127:
2125:
2124:
2120:
2118:
2117:
2112:
2110:
2109:
2104:
2102:
2101:
2096:
2095:
2093:
2091:
2087:
2081:
2080:
2075:
2073:
2072:
2067:
2065:
2064:
2059:
2057:
2056:
2052:
2050:
2049:
2044:
2042:
2041:
2036:
2034:
2033:
2029:
2027:
2026:
2021:
2019:
2018:
2013:
2011:
2010:
2005:
2003:
2002:
1998:
1996:
1995:
1990:
1988:
1987:
1983:
1981:
1980:
1975:
1973:
1972:
1968:
1966:
1965:
1960:
1959:
1957:
1955:
1951:
1948:
1946:(Sea turtles)
1944:
1940:
1937:
1935:
1931:
1915:
1914:
1908:
1905:
1903:
1899:
1896:
1894:
1890:
1887:
1885:
1881:
1875:
1871:
1869:
1865:
1862:
1860:
1856:
1854:
1850:
1848:
1844:
1843:
1840:
1836:
1829:
1824:
1822:
1817:
1815:
1810:
1809:
1806:
1795:
1789:
1783:
1780:
1779:
1777:
1775:
1774:
1769:
1763:
1760:
1758:
1755:
1754:
1752:
1750:
1749:
1744:
1738:
1735:
1733:
1730:
1729:
1727:
1725:
1724:
1719:
1713:
1712:
1707:
1705:
1702:
1700:
1697:
1695:
1692:
1690:
1687:
1685:
1682:
1680:
1677:
1675:
1672:
1670:
1667:
1665:
1662:
1661:
1659:
1657:
1656:
1651:
1645:
1644:
1639:
1637:
1634:
1632:
1629:
1627:
1624:
1622:
1619:
1618:
1616:
1614:
1613:
1608:
1602:
1599:
1598:
1596:
1594:
1593:
1588:
1582:
1579:
1578:
1576:
1574:
1573:
1568:
1562:
1559:
1557:
1554:
1553:
1551:
1549:
1548:
1543:
1537:
1534:
1532:
1529:
1528:
1526:
1524:
1523:
1518:
1512:
1509:
1507:
1504:
1502:
1499:
1497:
1494:
1492:
1489:
1487:
1484:
1482:
1479:
1477:
1474:
1472:
1469:
1468:
1466:
1464:
1463:
1458:
1452:
1449:
1447:
1444:
1442:
1439:
1438:
1436:
1434:
1433:
1428:
1422:
1419:
1418:
1416:
1414:
1413:
1412:Leucocephalon
1408:
1402:
1399:
1397:
1394:
1392:
1389:
1387:
1384:
1383:
1381:
1379:
1378:
1373:
1367:
1364:
1363:
1361:
1359:
1358:
1353:
1347:
1344:
1342:
1339:
1338:
1336:
1334:
1333:
1328:
1322:
1319:
1318:
1316:
1314:
1313:
1308:
1302:
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1296:
1294:
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1285:
1280:
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1277:
1275:
1274:
1268:
1262:
1259:
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1239:
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1234:
1232:
1229:
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1226:
1224:
1223:
1218:
1212:
1209:
1207:
1204:
1202:
1199:
1197:
1194:
1192:
1189:
1187:
1184:
1182:
1179:
1177:
1174:
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1169:
1167:
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1162:
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1144:
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1132:
1129:
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1119:
1117:
1114:
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1109:
1107:
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1101:
1099:
1098:
1093:
1087:
1086:
1081:
1080:
1078:
1076:
1075:
1074:Banhxeochelys
1069:
1052:
1051:
1044:
1041:
1037:
1031:
1027:
1025:
1024:Testudinoidea
1022:Superfamily:
1021:
1019:
1015:
1013:
1009:
1007:
1003:
1001:
997:
995:
991:
989:
985:
984:
981:
977:
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949:
946:
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936:
933:
926:
919:
914:
913:
909:
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896:
893:
882:on 2013-02-22
878:
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862:
859:
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821:
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797:
793:
789:
785:
778:
775:
764:on 2013-06-16
760:
753:
748:
743:
741:
739:
735:
729:
726:
723:1869:165–225.
722:
716:
713:
707:
704:
698:
696:
694:
690:
687:
679:
670:
669:
664:
663:
662:
661:
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651:
647:
643:
642:
637:
634:
630:
626:
625:
620:
617:
616:
611:
608:
604:
603:
598:
595:
594:
589:
586:
585:
580:
578:(two species)
577:
576:
571:
569:(two species)
568:
567:
562:
559:
555:
551:
547:
546:
541:
538:
537:
532:
529:
525:
522:(formerly in
521:
520:
519:Leucocephalon
515:
512:
509:(formerly in
508:
507:
502:
499:
498:
493:
491:(two species)
490:
489:
484:
481:
477:
476:
471:
468:
467:
462:
459:
455:
454:
449:
446:
442:
441:
436:
435:
434:
433:
428:
426:
425:
417:
415:
414:
413:Banhxeochelys
406:
405:
404:
401:
400:
399:
393:
391:
389:
385:
380:
378:
374:
370:
365:
363:
359:
355:
347:
345:
343:
338:
336:
332:
331:
326:
322:
321:
316:
312:
308:
304:
300:
296:
292:
288:
284:
283:
278:
274:
266:
264:
262:
258:
254:
250:
245:
243:
239:
235:
231:
227:
219:
217:
215:
211:
207:
203:
200:in the order
199:
195:
191:
182:
175:Gray 1869:185
165:
162:
158:
151:
148:
145:
144:
141:
140:Testudinoidea
138:
136:Superfamily:
135:
134:
131:
128:
125:
124:
121:
118:
115:
114:
111:
108:
105:
104:
101:
98:
95:
94:
91:
88:
85:
84:
81:
78:
75:
74:
69:
64:
60:
57:
56:
51:
47:
43:
38:
29:
26:
22:
4019:
3984:
3959:
3951:
3943:
3902:
3894:
3872:
3864:
3857:
3850:
3842:
3836:Erymnochelys
3834:
3827:
3821:Cerrejonemys
3819:
3811:
3803:
3797:Brontochelys
3795:
3787:
3779:
3758:
3751:
3731:
3723:
3716:
3711:Pseudemydura
3709:
3702:
3694:
3687:
3680:
3673:
3666:
3658:
3651:
3644:
3637:
3630:
3623:
3616:
3596:
3588:
3580:
3572:
3564:
3556:
3548:
3540:
3532:
3524:
3516:
3508:
3500:
3492:
3484:
3476:
3468:
3460:
3452:
3444:
3436:
3428:
3420:
3412:
3404:
3396:
3388:
3366:
3320:
3314:Prionochelys
3312:
3304:
3296:
3288:
3280:
3261:
3253:
3245:
3237:
3229:
3221:
3213:
3205:
3197:
3191:Cratochelone
3189:
3181:
3175:Bouliachelys
3173:
3165:
3157:
3151:Alienochelys
3149:
3127:
3119:
3111:
3089:
3081:
3073:
3066:
3058:
3052:Chelydropsis
3050:
3042:
3034:
3003:
2997:Basilochelys
2995:
2976:
2969:
2962:
2955:
2948:
2940:
2933:
2926:
2921:Khunnuchelys
2919:
2911:
2903:
2895:
2888:
2880:
2873:
2866:
2859:
2851:
2844:
2837:Trionychidae
2824:
2817:
2811:Allaeochelys
2809:
2775:
2768:
2762:Stigmochelys
2760:
2753:
2745:
2738:
2731:
2725:Megalochelys
2723:
2715:
2708:
2701:
2694:
2687:
2680:
2672:
2664:
2657:
2650:
2642:
2636:Cylindraspis
2634:
2627:
2620:
2615:Centrochelys
2613:
2606:
2599:
2592:Testudinidae
2581:Platysternon
2579:
2560:Vijayachelys
2558:
2551:
2544:
2539:Rhinoclemmys
2537:
2530:
2523:
2516:
2509:
2504:Melanochelys
2502:
2495:
2488:
2481:
2474:
2467:
2460:
2453:
2446:
2439:
2432:
2424:
2416:
2404:
2396:
2389:
2382:
2375:
2368:
2361:
2354:
2347:
2340:
2333:
2326:
2319:
2314:Acherontemys
2312:
2280:Sternotherus
2278:
2271:
2264:
2257:
2249:
2229:
2196:
2188:
2180:
2172:
2164:
2147:Psephophorus
2145:
2137:
2129:
2121:
2114:
2106:
2098:
2077:
2069:
2063:Pacifichelys
2061:
2053:
2046:
2038:
2032:Lepidochelys
2030:
2023:
2015:
2009:Gigantatypus
2007:
2001:Eretmochelys
1999:
1992:
1984:
1977:
1969:
1962:
1943:Chelonioidea
1911:
1863:
1796:. †=Extinct.
1773:Vijayachelys
1771:
1746:
1721:
1709:
1655:Rhinoclemmys
1653:
1641:
1610:
1590:
1570:
1545:
1522:Melanochelys
1520:
1460:
1430:
1410:
1396:Spiny turtle
1375:
1355:
1330:
1310:
1298:
1290:
1282:
1271:
1220:
1140:
1095:
1083:
1072:
1049:
1048:
1029:
975:
925:the original
903:, 2013-03-04
895:
884:. Retrieved
877:the original
864:
858:PDF fulltext
853:
848:
844:
840:
832:
791:
787:
777:
766:. Retrieved
759:the original
728:
720:
715:
706:
683:
680:Conservation
668:Rhinoclemmys
666:
657:
649:
645:
641:Vijayachelys
639:
632:
628:
622:
613:
606:
600:
591:
582:
573:
566:Melanochelys
564:
557:
553:
549:
543:
534:
527:
523:
517:
510:
504:
495:
486:
473:
464:
458:Cistoclemmys
457:
451:
444:
438:
429:
422:
411:
402:
397:
381:
377:Testudinidae
366:
351:
339:
328:
319:
297:species. In
282:Rhinoclemmys
280:
273:North Africa
270:
261:epipterygoid
257:facial nerve
249:articulation
246:
223:
193:
189:
187:
180:Gray 1870:17
178:Bataguridae
168:Geoemydidae
167:
149:
53:
25:
4266:Geoemydidae
4137:iNaturalist
4050:Geoemydidae
4044:Wikispecies
4020:Geoemydidae
3874:Stupendemys
3829:Cordichelys
3675:Mesoclemmys
3660:Hydromedusa
3582:Taphrosphys
3574:Sankuchemys
3566:Rhothonemys
3542:Polysternon
3422:Cearachelys
3406:Azabbaremys
3390:Araiochelys
3306:Ctenochelys
3298:Bashuchelys
3247:Rhinochelys
3223:Ocepechelon
3215:Notochelone
3207:Iserosaurus
3075:Macrochelys
3027:Chelydridae
2950:Palaeoamyda
2875:Cyclanorbis
2740:Psammobates
2733:Oligopherus
2696:Indotestudo
2644:Cymatholcus
2622:Chelonoidis
2608:Astrochelys
2417:Geoemydidae
2335:Deirochelys
2273:Staurotypus
2266:Kinosternon
2259:Hoplochelys
2174:Peritresius
2123:Dermochelys
2116:Cosmochelys
2108:Corsochelys
2025:Itilochelys
2017:Glarichelys
1964:Allopleuron
1954:Cheloniidae
1893:Superfamily
1782:Cane turtle
1050:Geoemydidae
1030:Geoemydidae
976:Geoemydidae
935:Geoemydidae
693:Appendix II
629:Panyaenemys
432:Geoemydinae
358:Batagurinae
333:) exhibits
323:) exhibits
295:carnivorous
287:herbivorous
214:Neotropical
194:Bataguridae
190:Geoemydidae
173:Batagurina
150:Geoemydidae
31:Geoemydidae
4260:Categories
3953:Caririemys
3866:Podocnemis
3852:Latentemys
3813:Carbonemys
3753:Pelomedusa
3590:Ummulisani
3510:Kurmademys
3478:Inaechelys
3462:Galianemys
3430:Chedighaii
3368:Araripemys
3341:Pleurodira
3322:Toxochelys
3239:Protostega
3044:Chelydrops
2964:Pelodiscus
2957:Pelochelys
2905:Gilmoremys
2882:Cycloderma
2792:Trionychia
2659:Geochelone
2652:Floridemys
2518:Notochelys
2406:Wilburemys
2377:Malaclemys
2231:Dermatemys
2198:Puppigerus
2131:Eosphargis
2048:Miocaretta
2040:Mexichelys
1934:Cryptodira
1874:Testudines
1835:Testudines
1572:Notochelys
1273:Echmatemys
1018:Cryptodira
1016:Suborder:
1012:Testudines
1004:Subclass:
886:2013-03-04
850:Salamandra
768:2014-04-19
699:References
689:Appendix I
658:Subfamily
584:Notochelys
554:Cathaiemys
430:Subfamily
424:Duboisemys
342:endangered
291:omnivorous
202:Testudines
130:Cryptodira
126:Suborder:
120:Testudines
3993:See also
3945:Caribemys
3805:Caninemys
3789:Bauruemys
3718:Rheodytes
3682:Myuchelys
3625:Chelodina
3550:Puentemys
3526:Nigeremys
3470:Ilatardia
3414:Bothremys
3121:Basilemys
2935:Nilssonia
2913:Hutchemys
2861:Axestemys
2819:Anosteira
2674:Hadrianus
2532:Pangshura
2490:Malayemys
2455:Geoclemys
2398:Trachemys
2391:Terrapene
2384:Pseudemys
2370:Graptemys
2363:Glyptemys
2356:Emydoidea
2349:Actinemys
2321:Chrysemys
2166:Euclastes
1994:Eochelone
1845:Kingdom:
1612:Pangshura
1432:Malayemys
1312:Geoclemys
986:Kingdom:
856:: 21-26.
808:1573-6849
602:Pangshura
550:Annamemys
536:Malayemys
480:monotypic
475:Geoclemys
373:molecular
259:, and an
238:carapaces
234:plastrons
86:Kingdom:
80:Eukaryota
21:Geomyidae
4029:Wikidata
3760:Pelusios
3725:Rhinemys
3696:Platemys
3689:Phrynops
3609:Chelidae
3598:Zolhafah
3494:Jainemys
3159:Archelon
3036:Chelydra
2928:Lissemys
2770:Stylemys
2755:Solitudo
2717:Manouria
2666:Gopherus
2629:Chersina
2497:Mauremys
2476:Heosemys
2469:Hardella
2462:Geoemyda
2448:Cyclemys
2304:Emydidae
2251:Claudius
2100:Arabemys
2079:Tasbacka
2071:Syllomus
1986:Chelonia
1884:Suborder
1868:Diapsida
1859:Reptilia
1853:Chordata
1851:Phylum:
1847:Animalia
1462:Mauremys
1377:Heosemys
1357:Hardella
1332:Geoemyda
1222:Cyclemys
1028:Family:
1006:Anapsida
1000:Reptilia
994:Chordata
992:Phylum:
988:Animalia
845:Mauremys
824:14434440
816:23512312
650:Heosemys
646:Geoemyda
633:Heosemys
545:Mauremys
528:Heosemys
524:Geoemyda
511:Geoemyda
506:Heosemys
497:Hardella
488:Geoemyda
466:Cyclemys
354:Emydidae
210:Eurasian
198:families
161:Synonyms
146:Family:
110:Reptilia
100:Chordata
96:Phylum:
90:Animalia
76:Domain:
4230:5457149
4035:Q446640
3904:Sokatra
3653:Emydura
3558:Rosasia
3454:Foxemys
3398:Arenila
2978:Trionyx
2971:Rafetus
2897:Dogania
2853:Apalone
2777:Testudo
2703:Kinixys
2689:Homopus
2546:Sacalia
2525:Orlitia
2511:Morenia
2426:Batagur
2328:Clemmys
2055:Natator
1971:Caretta
1872:Order:
1857:Class:
1723:Sacalia
1592:Orlitia
1547:Morenia
1097:Batagur
1010:Order:
998:Class:
974:Family
841:Sacalia
615:Sacalia
607:Kachuga
593:Orlitia
575:Morenia
558:Emmenia
445:Kachuga
440:Batagur
388:hybrids
364:rules.
277:Central
267:Ecology
206:species
116:Order:
106:Class:
4243:999897
4227:uBio:
4207:276087
4194:975162
4181:328320
4168:948953
4155:105314
4064:152912
3936:
3929:
3922:
3646:Elusor
3639:Elseya
3632:Chelus
3349:
3282:Adocus
3273:
3019:
2988:
2868:Chitra
2570:
2157:
1902:Family
1039:Genera
822:
814:
806:
665:Genus
638:Genus
621:Genus
612:Genus
599:Genus
590:Genus
581:Genus
572:Genus
563:Genus
542:Genus
533:Genus
516:Genus
503:Genus
494:Genus
485:Genus
482:genus)
472:Genus
463:Genus
450:Genus
437:Genus
418:Genus
407:Genus
384:genera
369:fossil
307:clutch
299:mating
242:scutes
4238:WoRMS
4215:Plazi
4150:IRMNG
4142:39886
4103:54068
4077:623VH
2942:Palea
2846:Amyda
2747:Pyxis
2441:Cuora
1913:Genus
1864:Clade
1142:Cuora
928:(PDF)
921:(PDF)
880:(PDF)
873:(PDF)
820:S2CID
762:(PDF)
755:(PDF)
686:CITES
453:Cuora
367:Most
4176:NCBI
4163:ITIS
4129:5463
4124:GBIF
4090:8128
4059:BOLD
2342:Emys
812:PMID
804:ISSN
648:and
556:and
526:and
371:and
362:ICZN
305:per
303:eggs
188:The
4085:EoL
4072:CoL
847:).
796:doi
293:or
4262::
4240::
4217::
4204::
4191::
4178::
4165::
4152::
4139::
4126::
4113::
4100::
4087::
4074::
4061::
4046::
4031::
1866::
854:41
843:×
818:.
810:.
802:.
792:21
790:.
786:.
737:^
552:,
379:.
337:.
3958:†
3950:†
3942:†
3901:†
3893:†
3884:†
3871:†
3849:†
3841:†
3826:†
3818:†
3810:†
3802:†
3794:†
3786:†
3778:†
3730:†
3701:†
3665:†
3595:†
3587:†
3579:†
3571:†
3563:†
3555:†
3547:†
3539:†
3531:†
3523:†
3515:†
3507:†
3499:†
3491:†
3483:†
3475:†
3467:†
3459:†
3451:†
3443:†
3435:†
3427:†
3419:†
3411:†
3403:†
3395:†
3387:†
3378:†
3365:†
3356:†
3319:†
3311:†
3303:†
3295:†
3287:†
3279:†
3260:†
3252:†
3244:†
3236:†
3228:†
3220:†
3212:†
3204:†
3196:†
3188:†
3180:†
3172:†
3164:†
3156:†
3148:†
3139:†
3126:†
3118:†
3110:†
3101:†
3088:†
3080:†
3065:†
3057:†
3049:†
3041:†
3002:†
2994:†
2947:†
2918:†
2910:†
2902:†
2887:†
2858:†
2816:†
2808:†
2767:†
2752:†
2730:†
2722:†
2679:†
2671:†
2649:†
2641:†
2431:†
2403:†
2311:†
2256:†
2195:†
2187:†
2179:†
2171:†
2163:†
2144:†
2136:†
2128:†
2113:†
2105:†
2097:†
2076:†
2068:†
2060:†
2045:†
2037:†
2022:†
2014:†
2006:†
1991:†
1976:†
1961:†
1827:e
1820:t
1813:v
1708:†
1640:†
1297:†
1289:†
1281:†
1270:†
1082:†
1071:†
967:e
960:t
953:v
889:.
826:.
798::
771:.
652:)
635:)
609:)
560:)
530:)
513:)
478:(
460:)
447:)
420:†
409:†
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
