1538:
1543:
122:
1669:
1566:
794:
385:
The arms were short, with longer upper arms (humeri) than lower arms (ulna). The legs were long, and the upper leg (femur) was longer than the pelvic bones. The tail was relatively long and feather impressions were found near the tip. Additional feather impressions were identified along the neck.
375:
and six in the maxilla. The teeth were closely packed together and lacked serrations. The eyes were relatively large. The skull was generally triangular in shape but longer than other basal oviraptorosaurs like
394:
Ji et al., 2012 considered
Ningyuansaurus a basal member of Oviraptorosauria. However, some traits, such as the low iliofemoral ratio, low ischiopubic ratio and enlarged pedal ungual II are more similar to
599:
Numerous small oval-shaped structures were found in the body cavity of the type specimen, each 10 millimetres (0.39 in) or less in diameter. These may be the remains of seeds, indicating that
624:
Q. Ji, J.-c. Lü, X.-f. Wei and X.-r. Wang. 2012. A new oviraptorosaur from the Yixian
Formation of Jianching, western Liaoning Province, China. Geological Bulletin of China 31(12):2102-2107
321:
and a greater number of teeth in comparison to any other known oviraptorosaur. However, later research showed that it is a member of
Halszkaraptorinae. The generic name
665:"A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae"
1734:
644:
Ji Qiang; Lü Jun-Chang; Wei Xue-Fang; Wang Xu-Ri (2012). "A new oviraptorosaur from the Yixian
Formation of Jianchang, Western Liaoning Province, China".
827:
1618:
1710:
1739:
329:. The specific name honors Wang Qiuwu, the private owner of the specimen who donated it for scientific study. The specimen is now in the
1559:
612:
1754:
349:
is notable for having a large number of teeth compared to more advanced oviraptorosaurs, but the teeth in the back of the upper jaw (
1749:
1537:
820:
734:
Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019-07-10).
314:
121:
1744:
1703:
1542:
353:) are still reduced in number compared to most other non-avialan theropods. The reduced number of maxillary teeth in
813:
1729:
1569:
924:
736:"A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight"
1759:
330:
1696:
1586:
1644:
1209:
978:
676:
664:
230:
710:
1100:
367:. Though the skull is poorly preserved, the specimen preserved at least 14 teeth in the lower jaw (
951:
506:
432:
116:
1649:
1631:
1128:
1093:
775:
757:
692:
468:
422:
358:
1636:
399:. While a 2016 study recovered it as an Oviraptorosaur, the only non-Oviraptorosaur taxa are
317:. It was originally thought to be the basalmost species of oviraptorosaur, based on its long
1313:
1046:
1024:
990:
836:
799:
765:
747:
684:
298:
295:
30:
1238:
1151:
1135:
909:
267:
1668:
680:
1680:
1506:
1492:
1299:
1276:
1161:
1071:
770:
735:
363:
1723:
1478:
1456:
1397:
1381:
1328:
1202:
1188:
1112:
552:
413:
401:
279:
68:
1449:
1405:
1363:
1306:
1249:
1223:
893:
456:
443:
407:
238:
205:
688:
1499:
1432:
1370:
1292:
1285:
1121:
963:
378:
43:
1609:
1513:
1464:
1341:
1265:
1195:
1181:
1174:
875:
866:
789:
372:
179:
88:
53:
761:
696:
1471:
1420:
1413:
1348:
1216:
1078:
936:
884:
326:
306:
192:
133:
93:
37:
779:
1676:
1603:
1390:
1355:
1230:
1064:
860:
310:
282:
166:
153:
83:
78:
63:
58:
48:
1623:
1258:
752:
396:
368:
350:
98:
73:
854:
342:
302:
143:
1580:
805:
1485:
1441:
318:
285:
1584:
847:
809:
382:, with a straight lower jaw unlike most other oviraptorosaurs.
417:
1684:
663:
Funston, Gregory F.; Currie, Philip J. (2016-07-03).
245:
1593:
1431:
1380:
1327:
1275:
1248:
1160:
1149:
1111:
1044:
989:
962:
935:
908:
639:
637:
635:
633:
426:, which contains many Oviraptorosaurs recovers
325:is derived from Ningyuan, an ancient name for
1704:
821:
371:) and 10 teeth in the upper jaw, four in the
8:
1711:
1697:
1581:
1333:
1166:
1157:
1056:
1052:
1041:
968:
941:
914:
905:
844:
828:
814:
806:
20:
769:
751:
361:and other basal oviraptorosaurs such as
294:, known from a fossil specimen from the
629:
603:was at least partially a seed-eater.
7:
1665:
1663:
613:Timeline of oviraptorosaur research
531:
481:
474:
449:
439:
1735:Early Cretaceous dinosaurs of Asia
669:Journal of Vertebrate Paleontology
14:
288:. It contains the single species
1667:
1565:
1564:
1541:
1536:
792:
120:
1:
1740:Fossil taxa described in 2012
689:10.1080/02724634.2016.1160910
1683:. You can help Knowledge by
646:Geological Bulletin of China
1776:
1662:
436:within Halszkaraptorinae:
315:People's Republic of China
265:
16:Extinct genus of dinosaurs
1755:Monotypic dinosaur genera
1554:
1534:
1336:
1169:
1059:
1055:
1040:
1021:
971:
944:
917:
904:
843:
549:
536:
529:
503:
486:
479:
472:
454:
447:
244:
237:
117:Scientific classification
115:
23:
1750:Paleontology in Liaoning
420:The 2019 description of
305:stage, 124.6 Ma ago) of
266:Not to be confused with
430:as the sister taxon of
1679:-related article is a
1645:Paleobiology Database
331:Confuciusornis Museum
715:theropoddatabase.com
291:Ningyuansaurus wangi
250:Ningyuansaurus wangi
1745:Feathered dinosaurs
1101:Scansoriopterygidae
681:2016JVPal..36E0910F
357:is shared with the
753:10.7717/peerj.7247
359:scansoriopterygids
1692:
1691:
1660:
1659:
1632:Open Tree of Life
1587:Taxon identifiers
1578:
1577:
1550:
1549:
1532:
1531:
1528:
1527:
1524:
1523:
1323:
1322:
1145:
1144:
1129:Similicaudipteryx
1094:Protarchaeopteryx
1036:
1035:
1032:
1031:
1017:
1016:
1013:
1012:
1009:
1008:
591:
590:
582:
581:
573:
572:
564:
563:
518:
517:
469:Halszkaraptorinae
423:Hesperornithoides
264:
263:
233:
1767:
1713:
1706:
1699:
1671:
1664:
1653:
1652:
1640:
1639:
1627:
1626:
1614:
1613:
1612:
1582:
1568:
1567:
1545:
1540:
1334:
1314:Epichirostenotes
1167:
1158:
1057:
1053:
1047:Oviraptorosauria
1042:
1025:Oviraptorosauria
991:Oviraptorosauria
969:
942:
915:
906:
899:
898:
845:
837:Oviraptorosauria
830:
823:
816:
807:
802:
800:Dinosaurs portal
797:
796:
795:
784:
783:
773:
755:
731:
725:
724:
722:
721:
707:
701:
700:
660:
654:
653:
652:(12): 2102–2107.
641:
532:
482:
475:
450:
440:
299:Yixian Formation
296:Early Cretaceous
247:
224:
217:
125:
124:
110:
40:
31:Early Cretaceous
29:Temporal range:
21:
1775:
1774:
1770:
1769:
1768:
1766:
1765:
1764:
1730:Oviraptorosaurs
1720:
1719:
1718:
1717:
1661:
1656:
1648:
1643:
1635:
1630:
1622:
1617:
1608:
1607:
1602:
1589:
1579:
1574:
1546:
1520:
1427:
1376:
1319:
1271:
1244:
1239:Ojoraptorsaurus
1154:
1152:Caenagnathoidea
1141:
1136:Xingtianosaurus
1107:
1049:
1028:
1005:
985:
958:
931:
925:Avemetatarsalia
910:Avemetatarsalia
900:
850:
849:
839:
834:
798:
793:
791:
788:
787:
733:
732:
728:
719:
717:
709:
708:
704:
675:(4): e1160910.
662:
661:
657:
643:
642:
631:
621:
609:
597:
592:
583:
574:
565:
519:
392:
341:The only known
339:
271:
268:Lingyuanosaurus
260:
253:
223:
215:
119:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
1773:
1771:
1763:
1762:
1760:Theropod stubs
1757:
1752:
1747:
1742:
1737:
1732:
1722:
1721:
1716:
1715:
1708:
1701:
1693:
1690:
1689:
1672:
1658:
1657:
1655:
1654:
1641:
1628:
1615:
1599:
1597:
1595:Ningyuansaurus
1591:
1590:
1585:
1576:
1575:
1573:
1572:
1562:
1555:
1552:
1551:
1548:
1547:
1535:
1533:
1530:
1529:
1526:
1525:
1522:
1521:
1519:
1518:
1510:
1503:
1496:
1493:Machairasaurus
1489:
1482:
1475:
1468:
1461:
1453:
1446:
1437:
1435:
1429:
1428:
1426:
1425:
1417:
1410:
1402:
1394:
1386:
1384:
1378:
1377:
1375:
1374:
1367:
1360:
1352:
1345:
1337:
1331:
1325:
1324:
1321:
1320:
1318:
1317:
1310:
1303:
1300:Caenagnathasia
1296:
1289:
1281:
1279:
1277:Caenagnathinae
1273:
1272:
1270:
1269:
1262:
1254:
1252:
1246:
1245:
1243:
1242:
1235:
1227:
1220:
1213:
1206:
1199:
1192:
1185:
1178:
1170:
1164:
1162:Caenagnathidae
1155:
1150:
1147:
1146:
1143:
1142:
1140:
1139:
1132:
1125:
1117:
1115:
1109:
1108:
1106:
1105:
1097:
1090:
1087:Ningyuansaurus
1083:
1075:
1072:Incisivosaurus
1068:
1060:
1050:
1045:
1038:
1037:
1034:
1033:
1030:
1029:
1022:
1019:
1018:
1015:
1014:
1011:
1010:
1007:
1006:
1004:
1003:
1002:
1001:
995:
993:
987:
986:
984:
983:
982:
981:
972:
966:
960:
959:
957:
956:
955:
954:
945:
939:
933:
932:
930:
929:
928:
927:
918:
912:
902:
901:
897:
896:
887:
878:
869:
863:
857:
848:
841:
840:
835:
833:
832:
825:
818:
810:
804:
803:
786:
785:
726:
711:"Dromaeosaurs"
702:
655:
628:
627:
626:
625:
620:
617:
616:
615:
608:
605:
596:
593:
589:
588:
585:
584:
580:
579:
576:
575:
571:
570:
567:
566:
562:
561:
558:
557:
548:
545:
544:
541:
540:
535:
530:
528:
525:
524:
521:
520:
516:
515:
512:
511:
502:
499:
498:
495:
494:
490:Ningyuansaurus
485:
480:
478:
473:
471:
465:
464:
461:
460:
453:
448:
446:
438:
428:Ningyuansaurus
391:
388:
364:Incisivosaurus
355:Ningyuansaurus
338:
335:
333:in Xingcheng.
327:Xingcheng City
323:Ningyuansaurus
275:Ningyuansaurus
262:
261:
254:
242:
241:
235:
234:
220:Ningyuansaurus
213:
209:
208:
203:
196:
195:
190:
183:
182:
177:
170:
169:
164:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
25:Ningyuansaurus
15:
13:
10:
9:
6:
4:
3:
2:
1772:
1761:
1758:
1756:
1753:
1751:
1748:
1746:
1743:
1741:
1738:
1736:
1733:
1731:
1728:
1727:
1725:
1714:
1709:
1707:
1702:
1700:
1695:
1694:
1688:
1686:
1682:
1678:
1673:
1670:
1666:
1651:
1646:
1642:
1638:
1633:
1629:
1625:
1620:
1616:
1611:
1605:
1601:
1600:
1598:
1596:
1592:
1588:
1583:
1571:
1563:
1561:
1557:
1556:
1553:
1544:
1539:
1516:
1515:
1511:
1509:
1508:
1504:
1502:
1501:
1497:
1495:
1494:
1490:
1488:
1487:
1483:
1481:
1480:
1479:Jiangxisaurus
1476:
1474:
1473:
1469:
1467:
1466:
1462:
1459:
1458:
1457:Ganzhousaurus
1454:
1452:
1451:
1447:
1444:
1443:
1439:
1438:
1436:
1434:
1430:
1423:
1422:
1418:
1416:
1415:
1411:
1408:
1407:
1403:
1400:
1399:
1398:Corythoraptor
1395:
1393:
1392:
1388:
1387:
1385:
1383:
1382:Oviraptorinae
1379:
1373:
1372:
1368:
1366:
1365:
1361:
1358:
1357:
1353:
1351:
1350:
1346:
1344:
1343:
1339:
1338:
1335:
1332:
1330:
1329:Oviraptoridae
1326:
1316:
1315:
1311:
1309:
1308:
1304:
1302:
1301:
1297:
1295:
1294:
1290:
1288:
1287:
1283:
1282:
1280:
1278:
1274:
1268:
1267:
1263:
1261:
1260:
1256:
1255:
1253:
1251:
1247:
1241:
1240:
1236:
1233:
1232:
1228:
1226:
1225:
1221:
1219:
1218:
1214:
1212:
1211:
1210:Leptorhynchos
1207:
1205:
1204:
1203:Gigantoraptor
1200:
1198:
1197:
1193:
1191:
1190:
1189:Chirostenotes
1186:
1184:
1183:
1179:
1177:
1176:
1172:
1171:
1168:
1165:
1163:
1159:
1156:
1153:
1148:
1138:
1137:
1133:
1131:
1130:
1126:
1124:
1123:
1119:
1118:
1116:
1114:
1113:Caudipteridae
1110:
1103:
1102:
1098:
1096:
1095:
1091:
1089:
1088:
1084:
1081:
1080:
1076:
1074:
1073:
1069:
1067:
1066:
1062:
1061:
1058:
1054:
1051:
1048:
1043:
1039:
1027:
1026:
1020:
999:
998:
997:
996:
994:
992:
988:
980:
976:
975:
974:
973:
970:
967:
965:
961:
953:
949:
948:
947:
946:
943:
940:
938:
934:
926:
922:
921:
920:
919:
916:
913:
911:
907:
903:
895:
891:
888:
886:
882:
879:
877:
873:
870:
868:
864:
862:
858:
856:
852:
851:
846:
842:
838:
831:
826:
824:
819:
817:
812:
811:
808:
801:
790:
781:
777:
772:
767:
763:
759:
754:
749:
745:
741:
737:
730:
727:
716:
712:
706:
703:
698:
694:
690:
686:
682:
678:
674:
670:
666:
659:
656:
651:
647:
640:
638:
636:
634:
630:
623:
622:
618:
614:
611:
610:
606:
604:
602:
594:
587:
586:
578:
577:
569:
568:
560:
559:
556:
555:
554:
553:Halszkaraptor
547:
546:
543:
542:
539:
534:
533:
527:
526:
523:
522:
514:
513:
510:
509:
508:
501:
500:
497:
496:
493:
492:
491:
484:
483:
477:
476:
470:
467:
466:
463:
462:
459:
458:
452:
451:
445:
442:
441:
437:
435:
434:
429:
425:
424:
418:
416:
415:
414:Herrerasaurus
410:
409:
404:
403:
402:Archaeopteryx
398:
389:
387:
383:
381:
380:
374:
370:
366:
365:
360:
356:
352:
348:
344:
336:
334:
332:
328:
324:
320:
316:
312:
308:
304:
300:
297:
293:
292:
287:
284:
281:
280:pennaraptoran
277:
276:
269:
258:
252:
251:
243:
240:
236:
232:
228:
222:
221:
214:
211:
210:
207:
204:
201:
198:
197:
194:
191:
188:
185:
184:
181:
178:
175:
172:
171:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
1685:expanding it
1674:
1594:
1512:
1505:
1498:
1491:
1484:
1477:
1470:
1463:
1455:
1450:Conchoraptor
1448:
1440:
1419:
1412:
1406:Huanansaurus
1404:
1396:
1389:
1369:
1364:Tongtianlong
1362:
1354:
1347:
1340:
1312:
1307:Caenagnathus
1305:
1298:
1291:
1284:
1264:
1257:
1250:Elmisaurinae
1237:
1229:
1224:Microvenator
1222:
1215:
1208:
1201:
1194:
1187:
1180:
1173:
1134:
1127:
1120:
1099:
1092:
1086:
1085:
1077:
1070:
1063:
1023:
894:Pennaraptora
889:
880:
871:
743:
739:
729:
718:. Retrieved
714:
705:
672:
668:
658:
649:
645:
600:
598:
551:
550:
537:
505:
504:
489:
488:
487:
457:Unenlagiinae
455:
444:Unenlagiidae
431:
427:
421:
419:
412:
408:Velociraptor
406:
400:
393:
384:
377:
362:
354:
346:
345:specimen of
340:
322:
290:
289:
274:
273:
272:
256:
249:
248:
239:Type species
226:
219:
218:
206:Pennaraptora
199:
186:
173:
160:
24:
18:
1500:Nemegtomaia
1433:Heyuanninae
1293:Apatoraptor
1122:Caudipteryx
979:Maniraptora
964:Maniraptora
379:Caudipteryx
337:Description
36:124.6
1724:Categories
1558:See also:
1514:Shixinggia
1465:Gobiraptor
1342:Luoyanggia
1266:Elmisaurus
1196:Eoneophron
1182:Beibeilong
1175:Anomalipes
1000:see below↓
876:Dinosauria
867:Sauropsida
720:2024-08-24
619:References
538:ISMD-VP09
373:premaxilla
313:Province,
309:, western
180:Saurischia
167:Dinosauria
1472:Heyuannia
1421:Rinchenia
1414:Oviraptor
1349:Nankangia
1217:Hagryphus
952:Theropoda
937:Theropoda
885:Theropoda
853:Kingdom:
762:2167-8359
746:: e7247.
697:0272-4634
397:paravians
390:Phylogeny
307:Jianchang
193:Theropoda
140:Kingdom:
134:Eukaryota
1677:theropod
1610:Q6580233
1604:Wikidata
1570:Category
1560:Timeline
1391:Citipati
1356:Nomingia
1231:Nomingia
1065:Avimimus
861:Chordata
859:Phylum:
855:Animalia
780:31333906
607:See also
601:N. wangi
507:Mahakala
433:Mahakala
347:N. wangi
311:Liaoning
283:dinosaur
154:Chordata
150:Phylum:
144:Animalia
130:Domain:
1637:6149445
1624:8446236
1259:Citipes
865:Class:
771:6626525
677:Bibcode
369:dentary
351:maxilla
212:Genus:
105:↓
1650:261035
1507:Oksoko
1371:Yulong
778:
768:
760:
695:
343:fossil
303:Aptian
259:, 2012
257:et al.
227:et al.
1675:This
1486:Khaan
1442:Banji
890:Clade
881:Clade
872:Clade
740:PeerJ
319:skull
286:genus
278:is a
200:Clade
187:Clade
174:Clade
161:Clade
1681:stub
1619:GBIF
1286:Anzu
977:see
950:see
923:see
776:PMID
758:ISSN
693:ISSN
595:Diet
411:and
231:2012
44:PreꞒ
1079:Kol
766:PMC
748:doi
685:doi
255:Ji
225:Ji
1726::
1647::
1634::
1621::
1606::
892::
883::
874::
774:.
764:.
756:.
742:.
738:.
713:.
691:.
683:.
673:36
671:.
667:.
650:31
648:.
632:^
405:,
229:,
202::
189::
176::
163::
94:Pg
38:Ma
33:,
1712:e
1705:t
1698:v
1687:.
1517:?
1460:?
1445:?
1424:?
1409:?
1401:?
1359:?
1234:?
1104:?
1082:?
829:e
822:t
815:v
782:.
750::
744:7
723:.
699:.
687::
679::
301:(
270:.
246:†
216:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
