729:
oval-shaped parapophyses that are oriented posteriorly, and each one begins a pronounced ridge that reaches the centrum's posterior edge. The dorsoventral and mediolateral sides of the centra are concave, forming distinct but shallow lateral fossae that make contact with the neural arch dorsally. At the base of the neural arch, the diapophyses are positioned centrally. The best-preserved vertebra reveals that they are oval in cross-section, bent ventrolaterally, and somewhat elongate. In the back, the neural canal is rounded. The matching centrum face is shorter than the complete neural arch. The zygapophyses are well formed, long, and inclined at a 45-degree angle.
1923:
126:
422:
814:. In contrast to the equilateral triangles observed in the trunk series, the form of that protuberance is more like a right triangle in the cervical paramedian osteoderms. The osteoderms are slightly arched when seen from the back. As is characteristic for aetosaurians, the median margins are thick dorsoventrally and sigmoidal in medial view. The "tongue-and-groove" lateral articular surfaces found in other aetosaurias are absent in
797:
1941:
382:. The Camp Butte, Lot's Wife, Jasper Forest, Jim Camp Wash, and Martha's Butte beds may be found in the PEFO area of the Sonsela Member. Lot's Wife, Jasper Forest, and Martha's Butte beds form cliffs, whereas the Lot's Wife and Marthas Butte beds are slope-forming units with a larger proportion of mudrocks than sandstones. These locations all show neighboring floodplain facies connected to a braided river system.
108:
780:
ventrolateral hook, a thin bone flange that starts on the dorsal side of the tuberculum continues laterally and confluences with the rib body. The posterodorsal surface of the rib is lined by a deep, elongate groove created by the flange. The rib is flattened dorsally and develops a slender anterior blade. The transverse processes of the dorsal vertebrae are fused with the single-headed, posteriormost ribs.
447:, which played a significant role in a variety of Late Triassic terrestrial faunas with a widespread range and high species diversity. Measuring between one and three meters in length, with strongly armored bodies and tails, it had a crucial ecological function of being the earliest and one of only two clades of big herbivorous archosaurs from the Late Triassic. Aetosauria is divided into two main clades:
679:
anterolateral edges are dorsoventrally thick, rugose, anteromedially sloping regions that are posteriorly bordered by a narrow, curving ridge. For articulation with the nasals, the front borders of the frontals are thick and rugose. Almost transverse is the frontal/nasal suture. The distinct, elevated midline ridge that is found in other aertosaurs is however absent from the frontal of
742:
arch of the previous vertebra when combined with the flat prezygapophyses. Between the ventromedial borders of the prezygapophyses, a horizontal ventral bar covers the neural canal's entrance. The centroprezygapophyseal fossa (cprf), which is ventrolateral to the prezygapophysis, is a deep depression that is posteriorly bounded by the major strut of the transverse process.
822:
curvature of the anterior and posterior plate edges is minor. These osteoderms are dorsoventrally thinner and less arched in posterior view than the osteoderms that are located more anterior. Although there might be an individual difference, the dorsal eminence is significantly offset medially and slightly more developed, becoming elevated and more pyramidal in form.
838:
The related lateral osteoderm has a minor overlap with the sigmoidal lateral border in the anterior region, which is just posterior to the anterior bar. The osteoderm is hardly curved in the posterior view. The osteoderm is more sharply arched in those that are thought to be located more posteriorly.
762:
differ noticeably from each other, and this coincides with the loss of distinct parapophyses and diapophyses along the series. The centra also become dorsoventrally taller than they are anteroposteriorly. Regarding the existence of the different vertebral laminae and related fossae, the properties of
749:
In comparison to the centrum height, the neural spine is short (32.3 mm). The spine is mediolaterally enlarged at the distal end and is anteroposteriorly elongate, measuring the same length as the neural arch's proximal section. Spinoprezygapophyseal and spinopostzygapophyseal laminae are paired
674:
The frontal's exceptional dorsoventral thickness is obvious, with dorsoventral thickness 0.35 times the element's midline length. The posterior frontal edge is inclined posterolaterally in the dorsal view so that the lateral frontal margin is longer than the medial frontal margin, creating a separate
788:
An extended, thin "tube" that bends anteroventrally and widens medially into two broad sheets of bone that meet in a median symphysis makes up the pubis' body. The front and posterior contours of the pubic are convex, rarely extending over the puboischiadic plate's ventral edge dorsoventrally due to
834:
The anterior bar is consistently wide over the osteoderm's lateral section, but it dramatically narrows medially before swelling once again at the anteromedial projection, making it profoundly concave. This is known as scalloping. This "scalloping" of the anterior bar is a synapomorphy found in all
855:
The dorsal eminence is reduced to a raised, anteroposteriorly elongate keel with a posterior projection that reaches beyond the posterior boundary of the osteoderm, and the posterior dorsal caudal paramedians become longer than broad. They presumably continue until they develop into long strips of
809:
The anterior bars of the cervical osteoderms, which are well-developed and have pronounced anteromedial projections, are thick dorsoventrally. The lateral edges are extremely sigmoidal and devoid of anterolateral projections. The ornamentation on the dorsal surface is minimally developed, with few
741:
The prezygapophyses are inclined at around 45 degrees from the horizontal. There is a large, sub-triangular spinoprezygapophyseal fossa between the prezygapophyses and ventral to the base of the neural spine that is well-developed. This produces a wide shelf for the posterior section of the neural
728:
In both the anterior and posterior views, the centrum faces are subcircular, somewhat concave, and have slightly flared rims. Each centrum has two concave, ventromedially inclined, rectangular surfaces that are separated by a sharp, deep mid-line keel. The anteroventral corners of the centrum have
408:
One of the most often discovered vertebrate fossils in the
Petrified Forest National Park (PEFO), Arizona's Upper Triassic Chinle Formation are aetosaurians. Four fragmentary skeletons that were believed to be a new taxon were discovered during paleontological research in the park between 2001 and
830:
While it can be challenging to distinguish the osteoderm transition between the cervical and trunk series, anterior dorsal trunk osteoderms are considered to have greater width/length ratios and be dorsoventrally thinner than the cervical paramedian osteoderms. The triangular protuberance is also
821:
While comparable and have a wider width/length ratio of 2.05 than their more anterior counterparts, the cervical paramedian osteoderms lack a pronounced sigmoidal lateral border. Even though the margin has a considerable anterolateral projection, it remains sigmoidal. Moreover, the anterolateral
779:
The anterior and mid-trunk ribs are double-headed. They extend laterally and then sharply turn ventrolaterally and then gradually turn more ventrally. The rib body is oval in cross-section, flattening more distally; it is broadest at the point of the sharp ventrolateral turn. Just lateral to the
737:
The centra's articular faces are rounded, somewhat concave, and have wide rims that flare outward. The centrum has a ventral surface that is thin and smooth, profoundly concave lateral sides, and is longer (45.78 mm) than it is tall (41.81 mm). The neural canal is broad, and the neural
665:
The element's main body is thick in the dorsoventral region, and the dorsomedial portion of the element is slightly twisted such that the dorsal surface is visibly concave. A thin layer of hematite covers any surface ornamentation. The sutural surface for the ascending process of the maxilla is
847:
For
Scutarx deltatylus, the trunk-caudal transition exhibits a noticeable height increase from the mid-trunk area to the anterior dorsal caudal region. The dorsal prominence is a large pyramid with a vertical keel at the back. The anterior caudal osteoderms are identical to those in the trunk.
745:
The postzygapophyses are likewise situated around 45 degrees above the horizontal in posterior view. On the posterior aspect, they have a triangular shape and a lateral postzygadiapophyseal lamina that is well developed. In dorsal view, that lamina creates a wide dorsal ledge of the transverse
746:
process and continues laterally to the diapophysis. Along the transverse process, the shelf is larger proximally and narrower distally. There are two shallow postzygapophyseal spinodiapophyseal fossae located along the shelf's dorsal surface, between the postzygapophyses and the neural spine.
678:
The frontals' dorsal surfaces are rugose and decorated with deep indentations, some of which are connected to longer grooves. There are broader, anteroposteriorly directed grooves anteriorly and laterally over the rounded orbits that indicate the higher center of the frontals. The frontals'
691:
The frontal/parietal suture is clearly apparent at the posterior boundary of the frontals, making it clear that these parts were not fused. The posterolateral section defines the dorsal boundary of the supratemporal fenestra. The top part of the skull's back is made up of posterior flanges
763:
the neural spine are the same as those of the mid-trunk vertebrae. Fossil records show that the prezygodiapophyseal laminae of the mid and posterior trunk vertebrae have significantly more pronounced development and lateral extension when compared to neural spines.
810:
features. The posterior plate edge is touched by the low, wide, and mounded dorsal eminence. Also medially offset and closer to the midline margin is the eminence. The osteoderm's posteromedial corner contains the distinctive triangle protuberance used to identify
1337:
692:
posteroventrally sloping contours that make connection with the opisthotics' paroccipital processes ventrally. The posterior process of the squamosal is in touch with the parietal flanges laterally and the supraoccipital in medial position.
436:. The Late Triassic (Carnian/Norian) aetosaurs were armored archosaurian reptiles. They are commonly found in India, North Africa, North America, and South America. These rare, highly derived archosaurs are easily distinguished from other
789:
its short length. As opposed to the robust, knob-like projections, the distal extremities of the pubes are shaped like elongate commata, which are thin and curve towards the symphysis. A unique character of the pubis of
851:
The breadth and length of dorsal mid-caudal paramedians are almost comparable. The prominent dorsal eminence and the anteromedial and anterolateral projections of the anterior bar are still present in those osteoderms.
675:
posterolateral process. Its anterior part connects posteriorly to the parietal and laterally to the postfrontal. The frontal creates the dorsal edge of the orbit just in front of the posterolateral process.
1264:"Anatomical notes and discussion of the first described aetosaur Stagonolepis robertsoni (Archosauria: Suchia) from the Upper Triassic of Europe, and the use of plesiomorphies in aetosaur biochronology"
373:
of Texas. Four partial skeletons including much of the carapace, vertebral column as well as the shoulder and pelvic girdles have been collected from PEFO between 2002 and 2009. The locations where the
333:
The presence of a large, triangular boss in the posteromedial corner of the dorsal surface of the dorsal paramedian osteoderms is the primary morphological characteristic that distinguishes
683:. The frontals' ventral surfaces are smooth and ventrally concave. The laterosphenoid joins the orbital fossa at a prominent, somewhat curved ridge that runs medially to the orbital fossa.
1338:"Cranial Anatomy and Dentition of the Aetosaur Typothorax coccinarum (Archosauria: Pseudosuchia) from the Upper Triassic (Revueltian–Mid Norian) Chinle Formation of Arizona"
771:
A robust, wide sacral rib that laterally extends anterodorsally to touch the posterodorsal border of the left ilium is a recognizable feature of this element.
1993:
1128:"Revised phylogenetic analysis of the Aetosauria (Archosauria: Pseudosuchia); assessing the effects of incongruent morphological character sets"
2069:
2034:
839:
There is a little embayment on the underside of the dorsal eminence and a smooth ventral surface to the dorsal trunk paramedian osteoderms.
2044:
1393:
2049:
1322:
1922:
666:
markedly posteroventrally concave. The dorsal edge of the external naris is formed by the anterior nasal narrowing mediolaterally.
2039:
960:"Taxonomy and phylogeny of the Aetosauria (Archosauria: Pseudosuchia) including a new species from the Upper Triassic of Arizona"
405:
from the early
Revueltian of Arizona, this significantly increases the record of non-desmatosuchin aetosaurs in the Revueltian.
306:
Greek for “triangular protuberance” -delta meaning triangle and -tylos meaning “knob, knot, swelling, callous, protuberance”).
366:
1198:
2064:
399:
is notable because it is from the lowermost
Revueltian teilzone. With the sole other record being a reported specimen of
385:
In Texas, the Cooper Canyon
Formation's uppermost and the majority of its middle part are Revueltian in age. While the
1184:"New insights into Late Triassic dinosauromorph-bearing assemblages from Texas using apomorphy-based identifications"
1218:
Nesbitt, Sterling; Stocker, Michelle; Parker, William; Wood, Thomas; Sidor, Christian; Angielczyk, Kenneth (2017).
570:
125:
1386:
2054:
370:
2059:
1961:
1220:"The braincase and endocast of Parringtonia gracilis, a Middle Triassic suchian (Archosaur: Pseudosuchia)"
831:
more equally sized. These osteoderms are difficult to distinguish from those in the anterior caudal area.
440:
archosaurs by the presence of an elongated, occasionally spiky, premaxilla and substantial dermal armor.
2006:
1481:
585:
357:
is prevalent in the fossil record because it may be an indicator taxon for the late
Adamanian biozone.
1219:
1379:
1944:
1357:
1303:
1239:
1067:
120:
2011:
1089:
1805:
1679:
1295:
1159:
1059:
793:
is the presence of two large pubic foremen, a characteristic seen in only one other aetosaur.
738:
arch's canal-lateral borders are medium-thin and have strong anterior edges in anterior view.
510:
1829:
1752:
1567:
1349:
1285:
1275:
1231:
1149:
1139:
1049:
1039:
870:
523:
421:
391:
379:
235:
1780:
1710:
1450:
535:
475:
448:
433:
291:
209:
1028:"Osteology of the Late Triassic aetosaur Scutarx deltatylus (Archosauria: Pseudosuchia)"
959:
758:
The neural spines and transverse processes of the mid- and posterior trunk vertebrae of
1894:
1849:
1762:
1703:
1617:
1290:
1263:
1154:
1127:
1090:"Cranial anatomy of Desmatosuchus haplocerus (Reptilia: Archosauria: Stagonolepididae)"
1054:
1027:
548:
2028:
1872:
1865:
1821:
1813:
1797:
1743:
1724:
1661:
1646:
1638:
1597:
1555:
1361:
1183:
875:
708:
470:
464:
400:
349:
343:
319:
65:
30:
1307:
1243:
720:
also possess deep, round parabasisphenoid recess, a trait apomorphic to this taxon.
107:
1769:
1717:
1654:
1631:
1577:
1466:
1441:
1071:
796:
702:
498:
452:
444:
261:
193:
1353:
1235:
302:
Latin for “shield fortress” -scutum meaning “shield” and -arx meaning “fortress”;
1984:
389:
fossils were found at around the same level as the lowest recorded occurrence of
1880:
1696:
1323:"Pelvic girdle morphology in Stagonolepis, with remarks on aetosaur systematics"
40:
1887:
1688:
1668:
1590:
1432:
485:
85:
50:
1605:
865:
180:
137:
90:
1299:
1163:
1063:
1978:
1541:
1519:
1493:
1426:
1402:
881:
462:
has been identified through phylogenetic analysis as the sister taxon of
437:
327:
222:
157:
80:
75:
60:
55:
45:
409:
2009. This genoholotype was later named in 2016 by
William G. Parker as
1998:
1280:
1144:
1044:
750:
vertically along the anterior and posterior edges of the neural spine.
378:
fossils were gathered are all found in the lower
Sonsela Member of the
365:
Fossils of
Scutarx have been found in the Triassic Chinle Formation of
311:
167:
95:
70:
1420:
315:
147:
1955:
395:, which indicates the local base of the Revueltian, this finding of
420:
337:
from other aetosaurs. Both morphologically and stratigraphically,
326:
are “medium sized” paramedian osteoderms belonging to the clade
1959:
1413:
1375:
330:, a heavily armored and more herbivorous cousin of crocodiles.
1371:
1199:"Archosaur Phylogeny and the Relationships of the Crocodylia"
864:
It was originally believed that aetosaurs were primarily
880:, demonstrated that at least some aetosaur species were
712:, the basal tubercle and the basipterygoid processes of
432:
1336:
Reyes, William; Parker, William; Marsh, Adam (2021).
341:
can be distinguished from the closely related taxas
268:
1968:
1848:
1779:
1751:
1742:
1678:
1616:
1576:
1565:
1539:
1492:
1465:
1321:Desmet, Hilde; Antczak, Mateusz; Bodzioch, Adam.
888:, like more aetosaurs however, are herbivores.
1182:Lessner, Emily; Parker, William; Marsh, Adam.
310:lived around 230 million years ago during the
1387:
868:. Recent aetosaur skull finds, such those of
294:, most commonly regarded by its species name
8:
716:have a broad contact and are not elongate.
1956:
1789:
1748:
1623:
1582:
1573:
1547:
1536:
1471:
1462:
1410:
1394:
1380:
1372:
443:Aetosauria is an early-diverging clade of
106:
20:
1289:
1279:
1153:
1143:
1053:
1043:
795:
700:Unlike those of other aetosaurians like
897:
1121:
1119:
1021:
1019:
1017:
1015:
1013:
1011:
1009:
1007:
1005:
1003:
1001:
999:
997:
995:
993:
991:
989:
953:
951:
949:
947:
945:
943:
941:
939:
937:
935:
933:
931:
929:
927:
925:
923:
921:
7:
1257:
1255:
1253:
1213:
1211:
1177:
1175:
1173:
1117:
1115:
1113:
1111:
1109:
1107:
1105:
1103:
1101:
1099:
1083:
1081:
987:
985:
983:
981:
979:
977:
975:
973:
971:
969:
919:
917:
915:
913:
911:
909:
907:
905:
903:
901:
563:
541:
516:
491:
481:
1342:Journal of Vertebrate Paleontology
1224:Journal of Vertebrate Paleontology
115:Life restoration and size diagram
14:
1940:
1939:
1921:
124:
367:Petrified Forest National Park
1:
2070:Fossil taxa described in 2016
1354:10.1080/02724634.2020.1876080
1236:10.1080/02724634.2017.1393431
474:, all forming a clade within
2035:Late Triassic pseudosuchians
425:Desmatosuchia skeletal mount
2045:Late Triassic North America
2086:
2050:Aetosaurs of North America
571:Adamanasuchus eisenhardtae
465:Adamanasuchus eisenhardtae
350:Adamanasuchus eisenhardtae
1935:
1918:
1792:
1626:
1585:
1550:
1535:
1516:
1474:
1461:
1409:
598:
583:
568:
561:
546:
539:
521:
514:
496:
489:
267:
260:
121:Scientific classification
119:
114:
105:
23:
16:Extinct genus of reptiles
1262:Parker, William (2018).
1197:Benton, Micheal (1988).
1126:Parker, William (2016).
1026:Parker, William (2016).
958:Parker, William (2014).
871:Neoaetosauroides engaeus
754:Posterior trunk vertebra
2040:Paleontology in Arizona
703:Stagonolepis robertsoni
371:Cooper Canyon Formation
290:is an extinct genus of
877:Aetosauroides scagliai
801:
709:Aetosauroides scagliai
426:
2007:Paleobiology Database
1088:Small, Bryan (2001).
799:
724:Post cranial skeleton
586:Calyptosuchus wellesi
424:
344:Calyptosuchus wellesi
2065:Triassic extinctions
471:Calytosuchus wellesi
402:Desmatosuchus smalli
361:History of discovery
800:Aetosaur osteoderms
369:of Arizona and the
353:. Because of this,
1281:10.7717/peerj.5455
1145:10.7717/peerj.1583
1045:10.7717/peerj.1583
886:Scutarx deltatylus
816:Scutarx deltatylus
812:Scutarx deltatylus
802:
791:Scutarx deltatylus
760:Scutarx deltatylus
733:Mid trunk vertebra
718:Scutarx deltatylus
714:Scutarx deltatylus
601:Scutarx deltatylus
427:
397:Scutarx deltatylus
296:Scutarx deltatylus
273:Scutarx deltatylus
2022:
2021:
1962:Taxon identifiers
1953:
1952:
1931:
1930:
1916:
1915:
1912:
1911:
1908:
1907:
1904:
1903:
1853:(Desmatosuchinae
1806:Aetobarbakinoides
1738:
1737:
1734:
1733:
1680:Paratypothoracini
1531:
1530:
1527:
1526:
1512:
1511:
647:
646:
638:
637:
629:
628:
620:
619:
611:
610:
511:Stagonolepidoidea
283:
282:
256:
2077:
2015:
2014:
2002:
2001:
1989:
1988:
1987:
1957:
1943:
1942:
1926:
1925:
1830:Neoaetosauroides
1790:
1753:Stagonolepidinae
1749:
1624:
1583:
1574:
1568:Stagonolepididae
1548:
1537:
1472:
1463:
1456:
1455:
1411:
1396:
1389:
1382:
1373:
1366:
1365:
1333:
1327:
1326:
1318:
1312:
1311:
1293:
1283:
1259:
1248:
1247:
1215:
1206:
1205:
1203:
1194:
1188:
1187:
1179:
1168:
1167:
1157:
1147:
1123:
1094:
1093:
1085:
1076:
1075:
1057:
1047:
1023:
964:
963:
955:
696:Parabasisphenoid
564:
542:
524:Stagonolepidinae
517:
492:
482:
392:Machaeroprosopus
380:Chinle Formation
270:
254:
247:
236:Stagonolepididae
234:
221:
208:
129:
128:
110:
100:
37:
29:Temporal range:
21:
2085:
2084:
2080:
2079:
2078:
2076:
2075:
2074:
2025:
2024:
2023:
2018:
2010:
2005:
1997:
1992:
1983:
1982:
1977:
1964:
1954:
1949:
1927:
1920:
1900:
1854:
1852:
1844:
1783:
1781:Desmatosuchinae
1775:
1730:
1711:Rioarribasuchus
1674:
1612:
1570:
1561:
1544:
1523:
1508:
1488:
1457:
1451:Aetosauriformes
1416:
1415:
1405:
1400:
1370:
1369:
1348:(6): e1876080.
1335:
1334:
1330:
1320:
1319:
1315:
1261:
1260:
1251:
1217:
1216:
1209:
1201:
1196:
1195:
1191:
1181:
1180:
1171:
1125:
1124:
1097:
1087:
1086:
1079:
1025:
1024:
967:
957:
956:
899:
894:
862:
845:
828:
807:
786:
777:
769:
767:Sacral vertebra
756:
735:
726:
698:
689:
672:
663:
658:
653:
648:
639:
630:
621:
612:
536:Desmatosuchinae
476:Desmatosuchinae
434:Aetosauriformes
419:
363:
292:Aetosauriformes
279:
276:
253:
245:
232:
219:
210:Aetosauriformes
206:
123:
101:
99:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
35:
34:
27:
17:
12:
11:
5:
2083:
2081:
2073:
2072:
2067:
2062:
2057:
2055:Carnian genera
2052:
2047:
2042:
2037:
2027:
2026:
2020:
2019:
2017:
2016:
2003:
1990:
1974:
1972:
1966:
1965:
1960:
1951:
1950:
1948:
1947:
1936:
1933:
1932:
1929:
1928:
1919:
1917:
1914:
1913:
1910:
1909:
1906:
1905:
1902:
1901:
1899:
1898:
1895:Sierritasuchus
1891:
1884:
1877:
1869:
1861:
1859:
1850:Desmatosuchini
1846:
1845:
1843:
1842:
1834:
1826:
1818:
1810:
1802:
1793:
1787:
1777:
1776:
1774:
1773:
1766:
1763:Polesinesuchus
1758:
1756:
1746:
1740:
1739:
1736:
1735:
1732:
1731:
1729:
1728:
1721:
1714:
1707:
1704:Paratypothorax
1700:
1693:
1684:
1682:
1676:
1675:
1673:
1672:
1665:
1658:
1651:
1643:
1635:
1627:
1621:
1618:Typothoracinae
1614:
1613:
1611:
1610:
1602:
1594:
1586:
1580:
1571:
1566:
1563:
1562:
1560:
1559:
1551:
1545:
1540:
1533:
1532:
1529:
1528:
1525:
1524:
1517:
1514:
1513:
1510:
1509:
1507:
1506:
1505:
1504:
1498:
1496:
1490:
1489:
1487:
1486:
1485:
1484:
1475:
1469:
1459:
1458:
1454:
1453:
1444:
1435:
1429:
1423:
1414:
1407:
1406:
1401:
1399:
1398:
1391:
1384:
1376:
1368:
1367:
1328:
1313:
1249:
1207:
1189:
1169:
1095:
1077:
965:
896:
895:
893:
890:
861:
858:
844:
841:
827:
824:
806:
803:
785:
782:
776:
773:
768:
765:
755:
752:
734:
731:
725:
722:
697:
694:
688:
685:
671:
668:
662:
659:
657:
654:
652:
649:
645:
644:
641:
640:
636:
635:
632:
631:
627:
626:
623:
622:
618:
617:
614:
613:
609:
608:
605:
604:
597:
594:
593:
590:
589:
582:
579:
578:
575:
574:
567:
562:
560:
557:
556:
553:
552:
549:Desmatosuchini
545:
540:
538:
532:
531:
528:
527:
520:
515:
513:
507:
506:
503:
502:
495:
490:
488:
480:
445:Pseudosuchians
418:
417:Classification
415:
362:
359:
281:
280:
277:
265:
264:
258:
257:
243:
239:
238:
230:
226:
225:
217:
213:
212:
204:
197:
196:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
135:
131:
130:
117:
116:
112:
111:
103:
102:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
38:
36:Carnian–Norian
28:
15:
13:
10:
9:
6:
4:
3:
2:
2082:
2071:
2068:
2066:
2063:
2061:
2060:Norian genera
2058:
2056:
2053:
2051:
2048:
2046:
2043:
2041:
2038:
2036:
2033:
2032:
2030:
2013:
2008:
2004:
2000:
1995:
1991:
1986:
1980:
1976:
1975:
1973:
1971:
1967:
1963:
1958:
1946:
1938:
1937:
1934:
1924:
1897:
1896:
1892:
1890:
1889:
1885:
1883:
1882:
1878:
1875:
1874:
1873:Gorgetosuchus
1870:
1868:
1867:
1866:Desmatosuchus
1863:
1862:
1860:
1857:
1856:sensu stricto
1851:
1847:
1840:
1839:
1835:
1832:
1831:
1827:
1824:
1823:
1822:Chilenosuchus
1819:
1816:
1815:
1814:Calyptosuchus
1811:
1808:
1807:
1803:
1800:
1799:
1798:Adamanasuchus
1795:
1794:
1791:
1788:
1786:
1782:
1778:
1772:
1771:
1767:
1765:
1764:
1760:
1759:
1757:
1754:
1750:
1747:
1745:
1744:Desmatosuchia
1741:
1727:
1726:
1725:Venkatasuchus
1722:
1720:
1719:
1715:
1713:
1712:
1708:
1706:
1705:
1701:
1699:
1698:
1694:
1691:
1690:
1686:
1685:
1683:
1681:
1677:
1671:
1670:
1666:
1664:
1663:
1662:Redondasuchus
1659:
1657:
1656:
1652:
1649:
1648:
1647:Gorgetosuchus
1644:
1641:
1640:
1639:Chilenosuchus
1636:
1634:
1633:
1629:
1628:
1625:
1622:
1619:
1615:
1608:
1607:
1603:
1600:
1599:
1598:Coahomasuchus
1595:
1593:
1592:
1588:
1587:
1584:
1581:
1579:
1575:
1572:
1569:
1564:
1558:
1557:
1556:Aetosauroides
1553:
1552:
1549:
1546:
1543:
1538:
1534:
1522:
1521:
1515:
1502:
1501:
1500:
1499:
1497:
1495:
1491:
1483:
1479:
1478:
1477:
1476:
1473:
1470:
1468:
1464:
1460:
1452:
1448:
1445:
1443:
1439:
1436:
1434:
1430:
1428:
1424:
1422:
1418:
1417:
1412:
1408:
1404:
1397:
1392:
1390:
1385:
1383:
1378:
1377:
1374:
1363:
1359:
1355:
1351:
1347:
1343:
1339:
1332:
1329:
1324:
1317:
1314:
1309:
1305:
1301:
1297:
1292:
1287:
1282:
1277:
1273:
1269:
1265:
1258:
1256:
1254:
1250:
1245:
1241:
1237:
1233:
1229:
1225:
1221:
1214:
1212:
1208:
1200:
1193:
1190:
1185:
1178:
1176:
1174:
1170:
1165:
1161:
1156:
1151:
1146:
1141:
1137:
1133:
1129:
1122:
1120:
1118:
1116:
1114:
1112:
1110:
1108:
1106:
1104:
1102:
1100:
1096:
1091:
1084:
1082:
1078:
1073:
1069:
1065:
1061:
1056:
1051:
1046:
1041:
1037:
1033:
1029:
1022:
1020:
1018:
1016:
1014:
1012:
1010:
1008:
1006:
1004:
1002:
1000:
998:
996:
994:
992:
990:
988:
986:
984:
982:
980:
978:
976:
974:
972:
970:
966:
961:
954:
952:
950:
948:
946:
944:
942:
940:
938:
936:
934:
932:
930:
928:
926:
924:
922:
920:
918:
916:
914:
912:
910:
908:
906:
904:
902:
898:
891:
889:
887:
883:
879:
878:
873:
872:
867:
859:
857:
853:
849:
842:
840:
836:
832:
825:
823:
819:
817:
813:
804:
798:
794:
792:
783:
781:
774:
772:
766:
764:
761:
753:
751:
747:
743:
739:
732:
730:
723:
721:
719:
715:
711:
710:
705:
704:
695:
693:
686:
684:
682:
676:
669:
667:
660:
655:
650:
643:
642:
634:
633:
625:
624:
616:
615:
607:
606:
603:
602:
596:
595:
592:
591:
588:
587:
581:
580:
577:
576:
573:
572:
566:
565:
559:
558:
555:
554:
551:
550:
544:
543:
537:
534:
533:
530:
529:
526:
525:
519:
518:
512:
509:
508:
505:
504:
501:
500:
494:
493:
487:
484:
483:
479:
477:
473:
472:
467:
466:
461:
458:
454:
450:
449:Desmatosuchia
446:
441:
439:
435:
431:
423:
416:
414:
412:
406:
404:
403:
398:
394:
393:
388:
383:
381:
377:
372:
368:
360:
358:
356:
352:
351:
346:
345:
340:
336:
331:
329:
325:
321:
320:Late Triassic
318:stage of the
317:
313:
309:
305:
301:
297:
293:
289:
288:
275:
274:
266:
263:
259:
252:
251:
244:
241:
240:
237:
231:
228:
227:
224:
218:
215:
214:
211:
205:
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
127:
122:
118:
113:
109:
104:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
32:
31:Late Triassic
26:
22:
19:
1969:
1893:
1886:
1879:
1871:
1864:
1855:
1837:
1836:
1828:
1820:
1812:
1804:
1796:
1784:
1770:Stagonolepis
1768:
1761:
1723:
1718:Tecovasuchus
1716:
1709:
1702:
1695:
1687:
1667:
1660:
1655:Kryphioparma
1653:
1645:
1637:
1632:Apachesuchus
1630:
1604:
1596:
1589:
1578:Aetosaurinae
1554:
1518:
1482:Pseudosuchia
1467:Pseudosuchia
1446:
1442:Pseudosuchia
1437:
1345:
1341:
1331:
1316:
1271:
1267:
1227:
1223:
1192:
1135:
1131:
1035:
1031:
885:
876:
869:
863:
854:
850:
846:
837:
833:
829:
820:
815:
811:
808:
790:
787:
778:
770:
759:
757:
748:
744:
740:
736:
727:
717:
713:
707:
701:
699:
690:
680:
677:
673:
664:
600:
599:
584:
569:
547:
522:
499:Aetosaurinae
497:
469:
463:
459:
456:
453:Aetosaurinae
442:
429:
428:
410:
407:
401:
396:
390:
386:
384:
375:
364:
354:
348:
342:
338:
334:
332:
323:
307:
303:
299:
295:
286:
285:
284:
278:Parker, 2016
272:
271:
262:Type species
255:Parker, 2016
249:
248:
200:
194:Pseudosuchia
187:
174:
24:
18:
1881:Longosuchus
1697:Kocurypelta
1230:: 122–141.
835:aetosaurs.
651:Description
181:Archosauria
2029:Categories
1888:Lucasuchus
1785:sensu lato
1689:Garzapelta
1669:Typothorax
1591:Aetosaurus
1542:Aetosauria
1520:Aetosauria
1503:see below↓
1494:Aetosauria
1433:Sauropsida
1403:Aetosauria
892:References
882:omnivorous
866:herbivores
486:Aetosauria
460:deltatylus
328:Aetosauria
304:deltatylus
223:Aetosauria
1985:Q22911737
1606:Stenomyti
1419:Kingdom:
1362:233616969
1274:: e5455.
1138:: e1583.
1038:: e1583.
805:Osteoderm
144:Kingdom:
138:Eukaryota
1979:Wikidata
1945:Category
1427:Chordata
1425:Phylum:
1421:Animalia
1308:52158424
1300:30186682
1244:89657063
1164:26819845
1064:26819845
687:Parietal
438:Triassic
229:Family:
168:Reptilia
158:Chordata
154:Phylum:
148:Animalia
134:Domain:
1999:8824841
1970:Scutarx
1838:Scutarx
1431:Class:
1291:6118205
1155:4727975
1072:5317139
1055:4727975
681:Scutarx
670:Frontal
457:Scutarx
430:Scutarx
411:Scutarx
387:Scutarx
376:Scutarx
355:Scutarx
339:Scutarx
335:Scutarx
324:Scutarx
312:Carnian
308:Scutarx
300:Scutarx
287:Scutarx
250:Scutarx
242:Genus:
216:Order:
164:Class:
25:Scutarx
2012:347478
1360:
1306:
1298:
1288:
1242:
1162:
1152:
1070:
1062:
1052:
856:bone.
843:Caudal
316:Norian
1447:Clade
1438:Clade
1358:S2CID
1304:S2CID
1268:PeerJ
1240:S2CID
1202:(PDF)
1132:PeerJ
1068:S2CID
1032:PeerJ
826:Trunk
784:Pubis
661:Nasal
656:Skull
201:Clade
188:Clade
175:Clade
1994:GBIF
1480:see
1296:PMID
1160:PMID
1060:PMID
874:and
860:Diet
706:and
468:and
451:and
347:and
314:and
41:Preęž’
1350:doi
1286:PMC
1276:doi
1232:doi
1150:PMC
1140:doi
1050:PMC
1040:doi
775:Rib
2031::
2009::
1996::
1981::
1449::
1440::
1356:.
1346:40
1344:.
1340:.
1302:.
1294:.
1284:.
1270:.
1266:.
1252:^
1238:.
1228:37
1226:.
1222:.
1210:^
1172:^
1158:.
1148:.
1134:.
1130:.
1098:^
1080:^
1066:.
1058:.
1048:.
1034:.
1030:.
968:^
900:^
884:.
818:.
478:.
455:.
413:.
322:.
203::
190::
177::
91:Pg
33:,
1876:?
1858:)
1841:?
1833:?
1825:?
1817:?
1809:?
1801:?
1755:?
1692:?
1650:?
1642:?
1620:?
1609:?
1601:?
1395:e
1388:t
1381:v
1364:.
1352::
1325:.
1310:.
1278::
1272:6
1246:.
1234::
1204:.
1186:.
1166:.
1142::
1136:4
1092:.
1074:.
1042::
1036:4
962:.
298:(
269:â€
246:â€
233:â€
220:â€
207:â€
96:N
86:K
81:J
76:T
71:P
66:C
61:D
56:S
51:O
46:ęž’
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