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Cistecephalus Assemblage Zone

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Kammerer, C. (2008, September). A new therocephalian from the Cistecephalus Assemblage Zone of South Africa and new information on therocephalian systematics. In Journal of Vertebrate Paleontology (Vol. 28, No. 3, pp. 98A-99A). 60 REVERE DR, STE 500, NORTHBROOK, IL 60062 USA: SOC VERTEBRATE
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Kammerer, C. (2008, September). A new therocephalian from the Cistecephalus Assemblage Zone of South Africa and new information on therocephalian systematics. In Journal of Vertebrate Paleontology (Vol. 28, No. 3, pp. 98A-99A). 60 REVERE DR, STE 500, NORTHBROOK, IL 60062 USA: SOC VERTEBRATE
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have affected the uniformity in colour and texture of the biozone rocks in areas. This includes the colour of the fossilized bones, which range from a smooth, white appearance to being nearly black depending on their proximity to the
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Assemblage Zone is located in the lower portion of the Teekloof Formation west of 24°E, in the upper Middleton Formation and lower Balfour Formation east of 24°E. These formations all fall within the Adelaide Subgroup of the
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NICOLAS, MERRILL; RUBIDGE, BRUCE S. (March 2010). "Changes in Permo-Triassic terrestrial tetrapod ecological representation in the Beaufort Group (Karoo Supergroup) of South Africa".
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Smith, R.M.H. (1995-08-01). "Changing fluvial environments across the Permian-Triassic boundary in the Karoo Basin, South Africa and possible causes of tetrapod extinctions".
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Kammerer, Christian F.; Bandyopadhyay, Saswati; Ray, Sanghamitra (2016-10-03). "A new taxon of cistecephalid dicynodont from the upper Permian Kundaram Formation of India".
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species. It is arranged into lower and upper zones due to the appearance or frequency of certain fossils. The lower zones are characterized by the first appearance of the
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Sidor, Christian A.; Smith, Roger M. H. (2007-06-01). "A second burnetiamorph therapsid from the Permian Teekloof Formation of South Africa and its associated fauna".
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Nasterlack, Tobias; Canoville, Aurore; Chinsamy, Anusuya (2012-11-01). "New Insights into the Biology of the Permian Genus Cistecephalus (Therapsida, Dicynodontia)".
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Dias da Silva, Sérgio (2011-03-01). "Middle-Late Permian tetrapods from the Rio do Rasto Formation, Southern Brazil: a biostratigraphic reassessment".
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Hancox, P.J; Rubidge, B.S (2001-01-01). "Breakthroughs in the biodiversity, biogeography, biostratigraphy, and basin analysis of the Beaufort group".
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in the biozone are found in these interchannel mudrock layers as animal remains that came to rest on the floodplains were quickly buried by
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washed downstream. Articulated fossils are not commonly found. However, isolated skulls are fairly common and are usually found encased in
1684:"Dyke-sill relationships in Karoo dolerites as indicators of propagation and emplacement processes of mafic magmas in the shallow crust" 2017:"Pareiasaurids from the Rio do Rasto Formation, southern Brazil: Biostratigraphic implications for Permian faunas of the Paraná Basin" 2143: 1879:
Damiani, Ross J. (2004-01-01). "Temnospondyls from the Beaufort Group (Karoo Basin) of South Africa and Their Biostratigraphy".
1245:"Researches on the Structure, Organization, and Classification of the Fossil Reptilia. Part IX., Section 4. On the Gomphodontia" 2179: 177: 2021: 190: 1714: 1073: 1916:"A dicynodont trackway from the Cistecephalus Assemblage Zone in the Karoo, East of Graaff-Reinet, South Africa" 1830: 1817:
Sidor, C. A.; Welman, J. (2003-09-01). "A second specimen of lemurosaurus pricei (therapsida: burnetiamorpha)".
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Keyser, A. W. (1979). A review of the biostratigraphy of the Beaufort Group in the Karoo Basin of South Africa.
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divisions were approved by paleontologists of the time and were left largely unchanged for several decades. The
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Merrill, van der Walt; Michael, Day; Bruce, Rubidge; K., Cooper, Antony; Inge, Netterberg (December 2010).
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Watson, D. M. S. (September 1913). "II.—The Beaufort Beds of the Karroo System of South Africa".
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fossils are extremely rare apart from the uppermost portion. It was suggested that, due to the
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in 1856. However, it was not until 1892 that it was observed that the geological strata of the
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Vertebrate biozonation of the Beaufort Group with special reference to the western Karoo Basin
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Kitching, J. W. (1970). A short review of the Beaufort zoning in South Africa. In
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Kitching, J. W. (1984). A reassessment of the biozonation of the Beaufort Group.
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Rubidge, B. S. (1995). Biostratigraphy of the Beaufort Group(Karoo Supergroup).
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and its fossils being found throughout, the biozone should be renamed to the
314:, a majorly fossiliferous and geologically important geological group of the 878: 850: 830: 822: 807: 729: 721: 709: 395: 347: 95: 39: 2144:"Magnetostratigraphy – Background to stratigraphy across the PTB in Russia" 1261: 1244: 2035: 1730: 1089: 1325:"II.—The Zones of the Beaufort Beds of the Karroo System in South Africa" 896: 856: 763: 748: 717: 705: 303: 85: 80: 65: 60: 50: 1715:"The role of fossils in interpreting the development of the Karoo Basin" 1074:"The role of fossils in interpreting the development of the Karoo Basin" 935: 921: 779: 733: 713: 659: 645: 635: 618: 577: 554: 497: 455: 428: 399: 382: 367: 323: 307: 134: 100: 75: 2121: 1270: 991: 977: 800: 744: 672:
Assemblage Zone. However, due to the biozone first being named after
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Cisneros, Juan; Abdala, Fernando; Malabarba, Maria (2005-01-01).
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10.1671/0272-4634(2003)023[0631:ASSOLP]2.0.CO;2
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Philosophical Transactions of the Royal Society of London B
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were most abundant in a narrow band at the very top of the
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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10.1671/0272-4634(2007)27[420:ASBTFT]2.0.CO;2
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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List of fossiliferous stratigraphic units in South Africa
1364:"V.—On the Permian and Triassic Faunas of South Africa" 1286:"V.—On the Permian and Triassic Faunas of South Africa" 934:
fossils are rare until the uppermost portion of the
285: 275: 263: 258: 248: 230: 212: 207: 199: 186: 173: 151: 140: 130: 18: 1545:Palaeogeography, Palaeoclimatology, Palaeoecology 762:are often fragmented. In addition, Jurassic-aged 634:Band". Later, researchers sought to redefine the 567:in 1906, where Broom referred to the rock layers 1437:Second Gondwana Symposium Proceedings and Papers 1493:Geocongress, Geological Society of South Africa 576:were found in as the "Kistecephalus beds". The 580:was originally assigned to a far more broader 1478:Keyser, A. W., & Smith, R. M. H. (1978). 920:track way is also known in an outcrop of the 8: 1940:: CS1 maint: multiple names: authors list ( 1750:: CS1 maint: multiple names: authors list ( 1682:Coetzee, André; Kisters, Alex (2017-03-01). 1163:: CS1 maint: multiple names: authors list ( 1109:: CS1 maint: multiple names: authors list ( 848:, and numerous Gorgonopsian species such as 697:, sediments of which were formed in a large 458:, which he named (from oldest to youngest): 1194:Quarterly Journal of the Geological Society 799:biozone is well known for its diversity of 584:range between the uppermost occurrence of 117: 15: 2034: 2001:South African Committee of Stratigraphy. 1865:South African Committee of Stratigraphy. 1768:South African Committee of Stratigraphy. 1633:South African Committee of Stratigraphy. 1581:South African Committee of Stratigraphy. 1529:South African Committee of Stratigraphy. 1260: 962:zone to geological formations outside of 1713:J., Hancox, P.; S., Rubidge, B. (1997). 1072:J., Hancox, P.; S., Rubidge, B. (1997). 906:have been found in the upper zones. The 662:was then, for a brief time, renamed the 1064: 439:could be differentiated based on their 427:rocks that encompass the current eight 1933: 1743: 1156: 1102: 446:. The initial undertaking was done by 310:found in the Adelaide Subgroup of the 203:up to 984.252 ft (300.000 m) 476:Zone of "highly specialized group of 7: 1009:. The zone also correlates with the 563:Assemblage Zone was first coined by 2022:Revista Brasileira de Paleontologia 2005:. Pretoria, Council for Geoscience. 1869:. Pretoria, Council for Geoscience. 1772:. Pretoria, Council for Geoscience. 1637:. Pretoria, Council for Geoscience. 1585:. Pretoria, Council for Geoscience. 1533:. Pretoria, Council for Geoscience. 1005:of the Pranhita-Godavari Valley in 708:. The sedimentary rocks are mainly 648:containing the first appearance of 1957:Journal of Vertebrate Paleontology 1819:Journal of Vertebrate Paleontology 1784:Journal of Vertebrate Paleontology 1188:Bain, Andrew Geddes (1845-02-01). 402:, and the first appearance of the 14: 1649:Journal of African Earth Sciences 488:named were subdivided further by 2064:10.1111/j.1502-3931.2011.00263.x 1609:10.1111/j.1502-3931.2009.00171.x 1454:South African Journal of Science 1027: 366:Assemblage Zone is one of eight 235: 2085:Sidor, Christian (2015-04-09). 2175:South African assemblage zones 1206:10.1144/GSL.JGS.1845.001.01.72 826:. Other species found are the 1: 1914:J, de Klerk, William (2002). 1893:10.1016/S1342-937X(05)70315-4 1688:Journal of Structural Geology 1661:10.1016/S0899-5362(01)00081-1 1323:Watson, D. M. S. (May 1914). 1997:Rubidge, B. S. (ed.) 1995b. 1969:10.1080/02724634.2012.697410 1861:Rubidge, B. S. (ed.) 1995b. 1764:Rubidge, B. S. (ed.) 1995b. 1629:Rubidge, B. S. (ed.) 1995b. 1577:Rubidge, B. S. (ed.) 1995b. 1557:10.1016/0031-0182(94)00119-S 1525:Rubidge, B. S. (ed.) 1995b. 972:snout was discovered in the 740:that flanked these streams. 326:has outcrops located in the 986:fossils have been found in 500:(from oldest to youngest): 2201: 1362:Broom, R. (January 1906). 1284:Broom, R. (January 1906). 890:, and rare occurrences of 732:streams flowed, while the 374:, and is considered to be 2003:Biostratigraphic Series 1 1867:Biostratigraphic Series 1 1770:Biostratigraphic Series 1 1700:10.1016/j.jsg.2017.03.002 1635:Biostratigraphic Series 1 1583:Biostratigraphic Series 1 1531:Biostratigraphic Series 1 1415:10.1017/S0016756800127219 1380:10.1017/S001675680012271X 1341:10.1017/S001675680019675X 1302:10.1017/S001675680012271X 866:. In addition species of 758:. Fossils not encased in 621:. He named this area the 608:. Kitching observed that 124:Cistecephalus microrhinus 116: 26: 956:have been made with the 484:These proposed biozones 147:within Adelaide Supgroup 2110:Papers in Palaeontology 2091:Palaeontologia Africana 1920:Palaeontologia Africana 1719:Palaeontologia Africana 1143:Palaeontologia Africana 1125:Biostratigraphic series 1078:Palaeontologia Africana 1048:Geology of South Africa 496:proposed the following 492:between 1906 and 1909. 1439:(Vol. 1, pp. 309–312). 1262:10.1098/rstb.1895.0001 1243:Seeley, H. G. (1895). 1015:Vyazovskaya Formations 1001:was discovered in the 974:Rio do Rasto Formation 2036:10.4072/rbp.2005.1.02 658:Assemblage Zone. The 590:and the lowermost of 391:, a small, burrowing 2180:Permian South Africa 1450:"Inside front cover" 743:The majority of the 1403:Geological Magazine 1368:Geological Magazine 1329:Geological Magazine 1290:Geological Magazine 1214:2027/uc1.c034667778 1035:Paleontology portal 988:Madumabisa Mudstone 736:accumulated on the 450:who subdivided the 448:Harry Govier Seeley 431:were discovered by 423:to be found in the 306:assemblage zone or 163:Middleton Formation 28:Stratigraphic range 1003:Kundaram Formation 994:. Recently, a new 944:Band" is located. 749:alluvial sediments 716:layers containing 433:Andrew Geddes Bain 346:respectively from 344:Balfour Formations 328:Teekloof Formation 157:Teekloof Formation 2122:10.1002/spp2.1055 1881:Gondwana Research 1460:(2). 1979-02-01. 1053:Fremouw Formation 728:plains where low- 596:. Decades later, 293: 292: 167:Balfour Formation 38:257.5–254.5  2192: 2159: 2158: 2156: 2155: 2146:. Archived from 2140: 2134: 2133: 2105: 2099: 2098: 2082: 2076: 2075: 2047: 2041: 2040: 2038: 2012: 2006: 1995: 1989: 1988: 1963:(6): 1396–1410. 1952: 1946: 1945: 1939: 1931: 1911: 1905: 1904: 1876: 1870: 1859: 1853: 1849: 1843: 1842: 1814: 1808: 1807: 1779: 1773: 1762: 1756: 1755: 1749: 1741: 1739: 1738: 1729:. Archived from 1710: 1704: 1703: 1679: 1673: 1672: 1655:(3–4): 563–577. 1644: 1638: 1627: 1621: 1620: 1592: 1586: 1575: 1569: 1568: 1540: 1534: 1523: 1517: 1506: 1500: 1489: 1483: 1476: 1470: 1469: 1446: 1440: 1433: 1427: 1426: 1398: 1392: 1391: 1359: 1353: 1352: 1320: 1314: 1313: 1281: 1275: 1274: 1264: 1240: 1234: 1233: 1185: 1179: 1175: 1169: 1168: 1162: 1154: 1134: 1128: 1121: 1115: 1114: 1108: 1100: 1098: 1097: 1088:. Archived from 1069: 1037: 1032: 1031: 1030: 952:Correlations in 806:, especially of 704:in southwestern 602:biostratigraphic 381:The name of the 316:Karoo Supergroup 241: 239: 238: 165:& lowermost 121: 112: 42: 16: 2200: 2199: 2195: 2194: 2193: 2191: 2190: 2189: 2165: 2164: 2163: 2162: 2153: 2151: 2142: 2141: 2137: 2107: 2106: 2102: 2084: 2083: 2079: 2049: 2048: 2044: 2014: 2013: 2009: 1996: 1992: 1954: 1953: 1949: 1932: 1913: 1912: 1908: 1878: 1877: 1873: 1860: 1856: 1850: 1846: 1816: 1815: 1811: 1781: 1780: 1776: 1763: 1759: 1742: 1736: 1734: 1712: 1711: 1707: 1681: 1680: 1676: 1646: 1645: 1641: 1628: 1624: 1594: 1593: 1589: 1576: 1572: 1551:(1–2): 81–104. 1542: 1541: 1537: 1524: 1520: 1507: 1503: 1490: 1486: 1477: 1473: 1448: 1447: 1443: 1434: 1430: 1400: 1399: 1395: 1361: 1360: 1356: 1322: 1321: 1317: 1283: 1282: 1278: 1242: 1241: 1237: 1187: 1186: 1182: 1176: 1172: 1155: 1136: 1135: 1131: 1122: 1118: 1101: 1095: 1093: 1071: 1070: 1066: 1061: 1033: 1028: 1026: 1023: 954:biostratigraphy 950: 884:Therocephalians 842:species namely 791: 684: 628:acme zone" or " 417: 338:, in the upper 300:Assemblage Zone 236: 234: 194:Assemblage Zone 181:Assemblage Zone 160: 126: 111: 110: 109: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 37: 36: 34: 22:Assemblage Zone 12: 11: 5: 2198: 2196: 2188: 2187: 2182: 2177: 2167: 2166: 2161: 2160: 2135: 2116:(4): 569–584. 2100: 2077: 2058:(1): 109–120. 2042: 2007: 1990: 1947: 1906: 1887:(1): 165–173. 1871: 1854: 1844: 1825:(3): 631–642. 1809: 1790:(2): 420–430. 1774: 1757: 1705: 1674: 1639: 1622: 1587: 1570: 1535: 1518: 1501: 1484: 1471: 1441: 1428: 1409:(9): 388–393. 1393: 1354: 1335:(5): 203–208. 1315: 1276: 1235: 1200:(1): 317–318. 1180: 1170: 1129: 1116: 1063: 1062: 1060: 1057: 1056: 1055: 1050: 1045: 1039: 1038: 1022: 1019: 1011:Malokinelskaya 949: 946: 817:Aulacephalodon 790: 787: 775:Beaufort Group 702:foreland basin 695:Beaufort Group 683: 680: 665:Aulacephalodon 656:Aulacephalodon 651:Aulacephalodon 606:Beaufort Group 604:ranges of the 598:James Kitching 551: 550: 542: 534: 526: 518: 510: 482: 481: 474: 467: 464:Pareiasaurians 452:Beaufort Group 437:Beaufort Group 425:Beaufort Group 416: 413: 408:Aulacephalodon 372:Beaufort Group 330:north-west of 312:Beaufort Group 291: 290: 287: 283: 282: 277: 273: 272: 265: 261: 260: 256: 255: 250: 246: 245: 232: 228: 227: 214: 210: 209: 205: 204: 201: 197: 196: 188: 184: 183: 175: 171: 170: 153: 149: 148: 145:Beaufort Group 142: 138: 137: 132: 128: 127: 122: 114: 113: 106: 105: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 24: 23: 13: 10: 9: 6: 4: 3: 2: 2197: 2186: 2185:Wuchiapingian 2183: 2181: 2178: 2176: 2173: 2172: 2170: 2150:on 2019-02-16 2149: 2145: 2139: 2136: 2131: 2127: 2123: 2119: 2115: 2111: 2104: 2101: 2096: 2092: 2088: 2081: 2078: 2073: 2069: 2065: 2061: 2057: 2053: 2046: 2043: 2037: 2032: 2028: 2024: 2023: 2018: 2011: 2008: 2004: 2000: 1994: 1991: 1986: 1982: 1978: 1974: 1970: 1966: 1962: 1958: 1951: 1948: 1943: 1937: 1929: 1925: 1921: 1917: 1910: 1907: 1902: 1898: 1894: 1890: 1886: 1882: 1875: 1872: 1868: 1864: 1858: 1855: 1852:PALEONTOLOGY. 1848: 1845: 1840: 1836: 1832: 1828: 1824: 1820: 1813: 1810: 1805: 1801: 1797: 1793: 1789: 1785: 1778: 1775: 1771: 1767: 1761: 1758: 1753: 1747: 1733:on 2018-05-28 1732: 1728: 1724: 1720: 1716: 1709: 1706: 1701: 1697: 1693: 1689: 1685: 1678: 1675: 1670: 1666: 1662: 1658: 1654: 1650: 1643: 1640: 1636: 1632: 1626: 1623: 1618: 1614: 1610: 1606: 1602: 1598: 1591: 1588: 1584: 1580: 1574: 1571: 1566: 1562: 1558: 1554: 1550: 1546: 1539: 1536: 1532: 1528: 1522: 1519: 1515: 1511: 1505: 1502: 1498: 1494: 1488: 1485: 1481: 1475: 1472: 1467: 1463: 1459: 1455: 1451: 1445: 1442: 1438: 1432: 1429: 1424: 1420: 1416: 1412: 1408: 1404: 1397: 1394: 1389: 1385: 1381: 1377: 1373: 1369: 1365: 1358: 1355: 1350: 1346: 1342: 1338: 1334: 1330: 1326: 1319: 1316: 1311: 1307: 1303: 1299: 1295: 1291: 1287: 1280: 1277: 1272: 1268: 1263: 1258: 1254: 1250: 1246: 1239: 1236: 1231: 1227: 1223: 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675:Cistecephalus 671: 670: 669:Cistecephalus 666: 661: 657: 653: 652: 647: 643: 642: 641:Cistecephalus 637: 633: 632: 631:Cistecephalus 627: 626: 625:Cistecephalus 620: 616: 613: 612: 611:Cistecephalus 607: 603: 599: 595: 594: 589: 588: 583: 582:stratigraphic 579: 575: 572: 571: 570:Cistecephalus 566: 562: 561: 560:Cistecephalus 556: 548: 547: 543: 540: 539: 535: 532: 531: 527: 524: 523: 522:Kistecephalus 519: 516: 515: 511: 508: 507: 503: 502: 501: 499: 495: 491: 487: 479: 475: 472: 468: 465: 461: 460: 459: 457: 453: 449: 445: 442: 438: 434: 430: 426: 422: 414: 412: 410: 409: 405: 401: 397: 394: 390: 389: 388:Cistecephalus 384: 379: 377: 373: 370:found in the 369: 365: 364:Cistecephalus 361: 357: 356:Graaff-Reinet 353: 352:Northern Cape 349: 345: 341: 337: 333: 332:Beaufort West 329: 325: 321: 317: 313: 309: 305: 301: 299: 298:Cistecephalus 288: 284: 281: 278: 274: 271: 270: 269:Cistecephalus 266: 262: 257: 254: 251: 247: 244: 233: 229: 226: 222: 218: 215: 211: 206: 202: 198: 195: 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Index

Stratigraphic range
Late Permian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Biozone
Beaufort Group
Teekloof Formation
Middleton Formation
Balfour Formation
Daptocephalus Assemblage Zone
Tropidostoma Assemblage Zone
Eastern
Northern
Western Cape
South Africa
Karoo Basin
Cistecephalus
Robert Broom

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