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Eurycnema goliath

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from the dark brown to a mottled greyish brown colour. This colouration will consist of less brown and more grey in successive moults. Following the third moult, along with changes in size and colour, sexual dimorphism between nymphs becomes visible; this involves both the development of male genitalia as well differences in body dimensions, with larger bodied females, and longer antennae in males. After the fourth moult, there is a further increase in size and spine development on the legs increases. In the fifth and final nymphal moult there is no increase in the body size of the nymph, yet wings will grow considerably during this period. The final moult to become an adult
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11–24 months. The body of female goliath stick insects becomes very large during the egg-laying period. When eggs are laid, the female will ‘flick’ the egg outwards to the ground using her abdomen, this aids in egg dispersal. Egg production may vary based on seasonal conditions and food abundance, with females recorded laying 200–800 eggs.
740:, the goliath stick insect has evolved to resemble the leaf, or branch of a tree with its coloration and shape. This is accompanied by a swaying body movement from side to side that is thought to mimic the swaying of leaves and branches. This adaptation assists the goliath stick insect in avoiding detection by predators. 535:
are large and run down the majority of the abdomen, two distinct white bands run continuously down the forewing and hindwings. Forewings are the same apple-green as the body, with a vibrant red ventral surface in females. Hindwings are transparent and green veined. Despite being fully winged, females
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become sexually mature following their final moult, where they can consistently reproduce throughout the rest of their adult lives. Reproduction can occur throughout the entire year in optimal conditions. In the wild, females lay eggs from summer through to autumn, with eggs hatching in a period of
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is dark brown with a very slender body and legs, cerci are prominent and antennae are very short. Following the first moult, the insect will increase in size but no other physical changes are apparent. After the second moult, along with a further increase in size, the nymph will change in colour
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in plant seeds, that serves the purpose of attracting ants to collect and disperse eggs. The fatty acid composition of this capitulum is key to attracting ant species to locate and carry the egg, with a reduction in dispersal rates seen when the capitula is removed.  Ants will often bring the
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due to the visual similarity between the insects. Key distinguishing features between the two species include differences in spines on the hind legs, forewing length and antennal segments. As a result of this confusion in identification and classification of Australian stick insects, various
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is bred in captivity on a global scale, with nymphs and eggs available to buy at online retailers to be shipped globally. Adults raised in captivity are said to live 6–7 months. The insect is also often used in educational displays at museums and zoos. Due to their size, sightings of wild
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are long and slender, with a mottled yellow and green colouration. All legs are lined with serrate spines, with the largest spines occurring on the inside of the hind tibia. In males, these spines are more pronounced, with enlarged spines on both the mid and hind-tibia.
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is thick and flexible, with the final three segments much shorter than the previous six segments. The operculum structure of females is distinct, extending well beyond the abdomen and tapering to a tip; this forms the 'chute' used for flicking both eggs and
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The phasmid has a large head coloured yellow with three longitudinal green bands running from the eyes to the back of the head. Three glossy ocelli are on the top of the head in a triangular formation, sitting between two brown compound eyes.
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are extremely frequent. Bushfires in this region causes considerable habitat loss, killing individual insects and potentially leading to ecosystem conversion. The increasing frequency and intensity of bushfires due to
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in its operculum, and using a whole body flicking motion, it will throw them away to the ground. This is said to be an adaptation to avoid predators detecting the location of the insect through seeing nearby frass.
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occurring between each stage. Nymphs usually hatch during the night and can be active immediately, where they will instinctually climb nearby trees just after hatching. The first instar of the
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species. Adult insects are said to eat approximately 10 medium-sized leaves per day. In the wild the species primarily feeds at night. Nymphs will tend to eat younger and more tender leaves.
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populations. Despite the magnitude of this threat, due to the wide distribution range of the E. goliath, the likelihood of bushfires threatening the overall survival of the species is low.
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further increases the direct impact of these events on the insect, and increases the period needed for habitat recovery. These factors bushfire event are the largest threats to
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As they are easy to care for and effectively harmless to humans, the goliath stick insect is commonly kept as an exotic pet and it is particularly popular among children.
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egg underground into nests, which protects the egg from the environment and predation. Although numerous species of ants are expected to carry and disperse the eggs, only
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on the metacoxae will be revealed. A "swooshing" sound, made by rubbing the wings together, also accompanies the display.  This display is an act of startle or
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are incapable of flight, this is likely due to their size and weight. Males have larger wings and are significantly smaller than females, enabling strong flight.
2064: 528:, like many phasmids, has adhesive pad structures on its legs which allow it to stick to surfaces, enabling ease of movement throughout the trees it inhabits. 2115: 903: 1706:
O’Hanlon, James C.; Jones, Braxton R.; Bulbert, Matthew W. (2020-07-31). "The dynamic eggs of the Phasmatodea and their apparent convergence with plants".
2012: 555:, where females can lay viable eggs without the requirement for male fertilisation, despite this, fertilised reproduction will still occur.  764:
will expose the bright red ventral surface of its wings and spreads out its hind legs, then rapidly strikes them together. Additionally, false
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takes over an hour, with the development of their distinct bright apple-green colouration and wings unfolding to reach their full size.
1108:"Studies on the Stick-Insect Genus Eurycnema Audinet-Serville (Phasmida; Phasmatidae) with Particular Reference to Australian Species" 1034: 1847: 563:
Eggs are large, dark brown and oval shaped, with a thick bark-like ridged exterior. Eggs resemble plant seeds such as those from
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CLARK, J. T. (April 1976). "The eggs of stick insects (Phasmida): a review with descriptions of the eggs of eleven species".
1817:"Breeding biology and behaviour of a pair of Pacific Bazas Aviceda subcristata in central-coastal Queensland over 10 years" 775:
The striking of the heavily spined hind legs also serves as an attack strategy to injure both predators and rival females.
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and spines are present on both dorsal and ventral sides of the body, usually accompanied by bluish green colouration. The
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species they inhabit. In captivity, the insect has been recorded eating a wider range of foods, such as leaves from
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are short (32 mm) with 29 segments in females, while being much longer in males (57 mm), with 27 segments.
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in 1834, using two female specimens, one of which came from the collection belonging to British entomologist
1571:"Egg Dispersal in the Phasmatodea: Convergence in Chemical Signaling Strategies Between Plants and Animals?" 1426:
Büscher, Thies H.; Buckley, Thomas R.; Grohmann, Constanze; Gorb, Stanislav N.; Bradler, Sven (2018-05-24).
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travels very little, and will spend a large portion, if not all of their life inhabiting a singular tree.
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The number of mature individuals present in the wild, and the population trend is unknown. Despite this,
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and considered one of the largest species of stick insects in the country. The species has the
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The egg also contains a large yellow capitulum, an edible fleshy structure similar to that of
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British Museum (Natural History).; History), British Museum (Natural; Kirby, W. F. (1904).
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on a Eucalyptus tree, a key habitat for the species and one of its primary food sources.
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With an adult body length of 105–146 mm in males, and 155–204 mm in females,
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birds, with the insect serving as a key food source for nestlings of species such as
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is smooth and glossy in appearance, it is also three to four times shorter than the
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Captive insects: the male abdomen top left shows the characteristically large
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The goliath stick insect is widespread throughout Australia's east coast in
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is considered common across its distribution range, and was classified as ‘
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Little is known about the predation of the insect in Australia, however,
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Stanton, Anthony O.; Dias, Daniel A.; O’Hanlon, James C. (August 2015).
772:, performed in an attempt to scare off, startle or distract a predator. 508:. Females have simple ovaries with approximately 200 ovarioles in total 2017: 1135: 830: 737: 624:. Small populations have also been reported in northern Queensland and 565: 492: 378: 2030: 1107: 819: 651: 592: 501: 460: 390: 124: 104: 1935: 1245:"The Development of the Phasmid Species List: Part One PSG No. 1-50" 1119: 796:
There is no recorded instances of competition between the males of
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at the Museum of Victoria. The species is named after the biblical
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distribution range, especially in southeastern Australia, large
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Phasmida Species File (Version 5.0/5.0; retrieved 23 May 2021)
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Complete Field Guide to Stick and Leaf Insects of Australia
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from the Australian Museum in 1900 by English entomologist
394:. In 1835, Gray then transferred the insect to the genus 817:, and in the wild primarily feeds on the leaves of the 1778:, Dordrecht: Springer Netherlands, pp. 205–219, 1387:"Studies on Female Genitalia of Queensland Phasmida" 1948: 385:, and refers to the impressive size of the insect. 381:from the popular story of David vs Goliath in the 957:10.2305/IUCN.UK.2017-3.RLTS.T79003367A79003460.en 646:The insect also has a preference for inhabiting 479:, with yellow banding across the upper body. The 1848:"Pheasant Coucal takes parenting very seriously" 388:Gray originally placed the species in the genus 783:When defecating, the E. goliath will catch its 1480:Bedford, G.O.; Chinnick, L.J. (October 1966). 894:are also often mentioned in local newspapers. 588:has been recorded undertaking the behaviour. 584:an ant common within the distribution of the 432:the species has often been confused with the 414:The insect was placed into its present genus 8: 1906:"Keeping live mini-beasts as classroom pets" 1281:. London: Printed by order of the Trustees . 1017:Brock, Paul D.; Hasenpusch, Jack W. (2009). 904:List of Australian stick insects and mantids 760:In response to being attacked or disturbed, 1542:"Husbandry Manual for Goliath Stick Insect" 1186:"Species Information - Phasmid Study Group" 1936: 56: 29: 20: 1770:Honan, Patrick (2008), New, T. R. (ed.), 1443: 1402: 955: 475:in Australia. General body coloration is 1626:"Biology and Ecology of the Phasmatodea" 595:stages prior to becoming an adult, with 443:have appeared in literature relating to 343: 1356:Die Königin, Cameron (September 2021). 1351: 1349: 1347: 1345: 1343: 1341: 1339: 1337: 1335: 1333: 915: 857:has been recorded being predated on by 673:, following a 2016 species assessment. 654:tree species, its primary food plants. 403:7 years later in 1842, Dutch zoologist 1931:Phasmid Study Group: Eurycnema goliath 1810: 1808: 1765: 1763: 1761: 1535: 1533: 1531: 1529: 1527: 1525: 1523: 1331: 1329: 1327: 1325: 1323: 1321: 1319: 1317: 1315: 1313: 1865:Gondwanaland Trading Company (2021). 1564: 1562: 1560: 1558: 1556: 1554: 1475: 1473: 1471: 1101: 1099: 1097: 1095: 1093: 1091: 1089: 1087: 1085: 1083: 1081: 1079: 1077: 1012: 1010: 1008: 1006: 1004: 1002: 1000: 998: 996: 994: 407:transferred the goliath to the genus 7: 1270: 1268: 1238: 1236: 1215: 1213: 1211: 1209: 1207: 1205: 1180: 1178: 1153: 1151: 1149: 1147: 1145: 1075: 1073: 1071: 1069: 1067: 1065: 1063: 1061: 1059: 1057: 992: 990: 988: 986: 984: 982: 980: 978: 976: 974: 2116:IUCN Red List least concern species 1642:10.1146/annurev.en.23.010178.001013 1278:A synonymic catalogue of Orthoptera 943:IUCN Red List of Threatened Species 1677:10.1111/j.1365-3113.1976.tb00342.x 1432:Frontiers in Ecology and Evolution 1404:10.1111/j.1440-6055.1965.tb00643.x 1295:Elementary text-book of entomology 428:Within literature surrounding the 14: 471:is one of the largest species of 1391:Australian Journal of Entomology 1221:"Goliath Stick Insect – Bugs Ed" 1106:Brock, Paul D. (December 1998). 81: 1846:Pearce, Clare (25 April 2019). 1776:Insect Conservation and Islands 1243:Harman, A. J. E. (April 2011). 363:was first described by English 1365:Phasmid Study Group Newsletter 1252:Phasmid Study Group Newsletter 1112:Journal of Orthoptera Research 932:Rudolf, E.; Brock, P. (2017). 1: 2090:Phasmida Species File (old): 1624:Bedford, G O (January 1978). 1498:10.1016/s0003-3472(66)80053-2 685:are primarily in the form of 1821:Australian Field Ornithology 1784:10.1007/978-1-4020-8782-0_17 813:The goliath stick insect is 1630:Annual Review of Entomology 1575:Journal of Chemical Ecology 1298:. London: S. Sonnenschein. 2147: 2131:Endemic fauna of Australia 1728:10.1007/s00114-020-01690-1 544:Goliath stick insects are 1904:Love, J. (January 2005). 1587:10.1007/s10886-015-0604-8 723:Anti-predator adaptations 641:Atlas of Living Australia 225: 218: 203: 196: 78:Scientific classification 76: 54: 45: 37: 28: 23: 950:: e.T79003367A79003460. 635:Reported occurrences of 612:Distribution and habitat 312:, is a large species of 304:, commonly known as the 1881:"Care of Stick Insects" 1445:10.3389/fevo.2018.00069 1385:Heather, N. W. (1965). 1163:www.brisbaneinsects.com 881:Interaction with humans 580:Rhytidoponera metallica 16:Species of stick insect 1910:Science Education News 1867:"Goliath Stick Insect" 1815:Briggs, Allan (2018). 1540:Bearman, Tara (2007). 1304:10.5962/bhl.title.8139 757: 643: 464: 357: 354:William Joseph Rainbow 273:(Kirby); Günther, 1929 259:Cyphocrania herculeana 229:Phasma (Diura) acheron 1885:The Australian Museum 1708:The Science of Nature 1665:Systematic Entomology 1292:Kirby, W. F. (1892). 1190:phasmidstudygroup.org 1027:10.1071/9780643097087 874:Centropus phasianinus 751: 634: 458: 423:William Forsell Kirby 347: 289:Eurycnema viridissima 255:(Gray); de Haan, 1842 24:Goliath stick insect 2126:Insects of Australia 1833:10.20938/afo35095101 306:goliath stick insect 249:(Gray); Brullé, 1835 1720:2020SciNa.107...34O 868:Aviceda subcristata 330:Phasmid Study Group 283:Eurycnema magnifica 279:(Gray); Kirby, 1904 253:Cyphocrania goliath 48:Conservation status 804:Diet and predation 758: 734:passive camouflage 644: 465: 430:Eurycnema goliath, 368:George Robert Gray 358: 348:Drawing of a male 310:regal stick insect 247:Cyphocrana goliath 241:Clemacantha regale 235:Acrophylla goliath 2103: 2102: 2078:Open Tree of Life 1979:Eurycnema_goliath 1950:Eurycnema goliath 1942:Taxon identifiers 1793:978-1-4020-8782-0 936:Eurycnema goliath 855:Eurycnema goliath 770:deimatic behavior 754:Eurycnema goliath 712:Eurycnema goliath 683:Eurycnema goliath 663:Eurycnema goliath 650:and broad-leafed 639:according to the 637:Eurycnema goliath 469:Eurycnema goliath 445:Eurycnema goliath 350:Eurycnema goliath 301:Eurycnema goliath 297: 296: 277:Eurycnema goliath 261:Charpentier, 1841 207:Eurycnema goliath 71: 2138: 2096: 2095: 2086: 2085: 2073: 2072: 2060: 2059: 2047: 2046: 2034: 2033: 2021: 2020: 2008: 2007: 1995: 1994: 1982: 1981: 1969: 1968: 1967: 1937: 1918: 1917: 1901: 1895: 1894: 1892: 1891: 1877: 1871: 1870: 1862: 1856: 1855: 1843: 1837: 1836: 1812: 1803: 1802: 1801: 1800: 1767: 1756: 1755: 1703: 1697: 1696: 1660: 1654: 1653: 1621: 1615: 1614: 1566: 1549: 1548: 1546: 1537: 1518: 1517: 1486:Animal Behaviour 1477: 1466: 1465: 1447: 1423: 1417: 1416: 1406: 1382: 1376: 1375: 1373: 1371: 1362: 1353: 1308: 1307: 1289: 1283: 1282: 1272: 1263: 1262: 1260: 1258: 1249: 1240: 1231: 1230: 1228: 1227: 1217: 1200: 1199: 1197: 1196: 1182: 1173: 1172: 1170: 1169: 1155: 1140: 1139: 1103: 1052: 1047: 1041: 1040: 1014: 969: 968: 966: 964: 959: 929: 435:Eurycnema osiris 209: 86: 85: 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591:There are five 553:parthenogenetic 551:The species is 542: 453: 405:Wilhelm de Haan 342: 271:Diura magnifica 214: 211: 205: 192: 189:E. goliath 80: 72: 61: 57: 50: 17: 12: 11: 5: 2144: 2142: 2134: 2133: 2128: 2123: 2118: 2108: 2107: 2101: 2100: 2098: 2097: 2087: 2074: 2061: 2048: 2035: 2022: 2009: 1996: 1983: 1970: 1954: 1952: 1946: 1945: 1940: 1934: 1933: 1926: 1925:External links 1923: 1920: 1919: 1896: 1872: 1857: 1838: 1804: 1792: 1757: 1698: 1655: 1636:(1): 125–149. 1616: 1581:(8): 689–695. 1550: 1519: 1492:(4): 518–521. 1467: 1418: 1377: 1309: 1284: 1264: 1232: 1201: 1174: 1141: 1053: 1042: 1035: 970: 914: 913: 911: 908: 907: 906: 899: 896: 882: 879: 850: 847: 810: 807: 805: 802: 793: 790: 780: 779:Faeces removal 777: 745: 742: 729: 726: 724: 721: 719: 716: 708:climate change 694: 691: 678: 675: 613: 610: 546:hemimetabolous 541: 538: 452: 449: 383:Book of Samuel 341: 338: 316:in the family 295: 294: 293: 292: 286: 280: 274: 268: 262: 256: 250: 244: 238: 232: 223: 222: 216: 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Index


Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Phasmatodea
Phasmatidae
Phasmatinae
Phasmatini
Eurycnema
Binomial name
Synonyms
stick insect
Phasmatidae
endemic
Australia
Phasmid Study Group

William Joseph Rainbow
zoologist
George Robert Gray
John Curtis
giant
Goliath
Book of Samuel

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