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Kembawacela

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The skull has various features associated with burrowing, being broad and wedge-shaped. The large, forward facing eyes may be related to seeing in low-light environments, with binocular vision providing greater light sensitivity, although this is not typical of modern fossorial mammals which have
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was sexually dimorphic, as two of the seven known specimens appear to have lacked tusks. Notably, one of these specimens is the largest known individual, implying that the lack of tusks was not due to it not being fully grown yet. Sexual dimorphism is known from other cistecephalids, such as the
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Angielczyk, K.D.; Steyer, J.‐S.; Sidor, C.A.; Smith, R.M.H.; Whatley, R.L.; Tolan, S. (2014). "Permian and Triassic dicynodont (Therapsida: Anomodontia) faunas of the Luangwa Basin, Zambia: taxonomic update and implications for dicynodont biogeography and biostratigraphy". In Kammerer, C.F.;
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In 2016, a cistecephalid skull (DMMM-PK-16-1) was found encased in a rock nodule during exploration of the Chiweta Beds in the Mount Waller Area of Northern Malawi. Due its small size and delicate nature, the examination of the specimen was performed using
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lacks those traits the presence or absence of tusks is suggested to be an indicator of sexual dimorphism more broadly in dicynodonts. Unfortunately, the relatively small sample size means this possibility cannot be statistically analysed.
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by Freeman in 1993. Further examinations and the discovery of an additional four specimens proposed that the Luangwa cistecephalid was an entirely new genus. In 2019, the taxon was formally described in detail and named
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broadly resembled other cistecephalids in size and shape. It was a small dicynodont (skull length roughly 5 centimetres (2.0 in) long along the base) and had a highly specialised body plan for digging.
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with paired extensions that reach up from the back of the skull to flank the pineal foramen. There are also various other slight differences of the skull and skeleton between it and other cistecephalids.
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Angielczyk, K.D.; Benoit, J.; Rubidge, B.S. (2019). "A new tusked cistecephalid dicynodont (Therapsida, Anomodontia) from the upper Permian upper Madumabisa Mudstone Formation, Luangwa Basin, Zambia".
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The specimens were collected from various different localities in the Luangwa Basin, but were all from the upper Madumabisa Mudstone Formation. This formation is believed to overlap in time with the
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in this species. These tusks face slightly out to the sides, but do not sit out on a prominent caniniform process projecting from the jaw margin like in some other dicynodonts. Aside from the tusks,
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are also notable as one specimen preserves unerupted tusks within the jaw bones. There is no evidence that these tusks are replacements of a previous pair (as has been suggested to occur in
656:(NHCC LB18)—have been found in, as well referring to the proposed digging lifestyle of cistecephalids. The species was named to honour James Kitching, who collected the first specimens of 1663:
Huttenlocker, A. K.; Sidor, C. A. (2016). "The first karenitid (Therapsida, Therocephalia) from the upper Permian of Gondwana and the biogeography of Permo-Triassic therocephalians".
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and other cistecephalids is a likely factor in its endemism, as its unique ecology would have limited its ability to disperse and so restricted its range to within the Luangwa Basin.
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shares various features with other cistecephalids associated with a fossorial (burrowing) lifestyle. The ulna, the only known bone from the forelimb, has the typically large
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may not have erupted until quite late in the animal's life, close to their mature body size. It's possible that this relatively late eruption of the tusks in
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by Kenneth Angielczyk, Julien Benoit and Bruce Rubidge. The genus name is from the phrase "kemba wacela", translated to "iron digger" in the locally spoken
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AZ in the Karoo, South Africa, suggesting the two regions shared similar biomes and were broadly connected. However, the presence of endemic species like
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beak at the tips of its jaws, as is typical of dicynodonts. The beak was relatively broad and blunt, and the tip of the upper jaw was arched upwards.
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was taken even further in other cistecephalids, where their development was suppressed and prevented entirely from erupting in the tuskless species.
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Kammerer, C.F.; Bandyopadhyay, S.; Ray, S. (2016). "A new taxon of cistecephalid dicynodont from the upper Permian Kundaram Formation of India".
739:. The skull is mostly complete, missing only the tip of the snout, its left zygomatic arch and postorbital bar behind the eye, and parts of the 3712: 3697: 697:. The specimens are housed at both the Evolutionary Studies Institute in South Africa and at the National Heritage Conservation Commission in 1880: 1567: 1098:
is primarily distinguished from other cistecephalids by its tusks, but can also be uniquely diagnosed by a trough on the underside of the
1620: 1466: 1616:"A new eutherocephalian (Therapsida, Therocephalia) from the upper Permian Madumabisa Mudstone Formation (Luangwa Basin) of Zambia" 1361:
are indicative of fine scale variations between the two localities, such as environmental differences. The fossorial lifestyle of
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coexisted with a wide variety and abundance of other dicynodonts in the upper Madumabisa Mudstone Formation, including
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with a very short, tapered snout. It had large, strongly forward facing eyes like some other cistecephalids (including
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are large, associated with hearing low-frequency sounds underground, although its inner-ear is not as specialised as
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Sidor, C. (2015). "The first biarmosuchian from the upper Madumabisa Mudstone Formation (Luangwa Basin) of Zambia".
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in the Luangwa Basin of Zambia. They reported discovering at least 13 specimens that they preliminarily assigned to
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it is hidden on the internal side. Further, the anterior process of the jugal beneath the eye is notably taller in
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project out almost laterally behind the eyes and curve back almost 90 degrees to the back of the skull. The
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The environment of the upper Madumabisa Mudstone Formation was a wet, vegetated floodplain within a wide
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with well developed forward and backward pointing processes, and a large, robust ulna in the arm. Unlike
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n. sp., a new cistecephalid species (Dicynodontia: Emydopoidea) from the Upper Permian of Malawi"
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and various vertebrae. Although broadly similar in superficial appearance, the two species
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surrounding the eye, and they are remarkably smaller than is suggested by the size of the
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is known from skulls, lower jaws and various pieces of postcrania, including parts of the
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is most obviously distinguished by the prominent tusks in its upper jaw. The majority of
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Much of the fauna of the Madumabisa Mudstone Formation is shared with that of the
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AraĂșjo, R.; Macungo, Z.; Fernandez, V.; Chindebvu, E. G.; Jacobs, L. L. (2022).
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Huttenlocker, Adam K.; Sidor, Christian A.; Angielczyk, Kenneth D. (2015).
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having a much narrower ~46° angle between them compared to the ~61° of
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is the only known cistecephalid known to preserve pieces of the bony
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Its skull is typical for cistecephalids, with a broad head and large
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in front) on the roof of the mouth also differs between them, with
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rimming the eye socket is exposed on the back of the bar, while in
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Kenneth D. Angielczyk (2019). "First occurrence of the dicynodont
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The specimens were later suggested to belong to a new species of
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associated with scratch-digging found in other cistecephalids.
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specimens have these tusks, and it is possible that they were
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Cistecephalidae in the phylogenetic analysis of Angielczyk
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can be distinguished by details of the skull architecture.
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various specimens from the Luangwa Basin—including the
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were first discovered and collected in the 1960s by
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"The primitive cynodont 1451: 1449: 1447: 556:resembles that of other cistecephalids like 1556:Early evolutionary history of the Synapsida 762:(2019), where it was found as a relatively 3604: 3418: 3407: 3253: 3070: 3059: 3046: 2841: 2646: 2635: 2622: 2397: 2388: 2254: 2243: 2230: 2137: 2030: 2019: 2006: 1952: 1943: 1897: 1881: 1867: 1859: 1409: 493:, the ascending process that joins to the 20: 1760: 1548: 1546: 1544: 1542: 1487: 1407: 1405: 1403: 1401: 1399: 1397: 1395: 1393: 1391: 1389: 470:was otherwise toothless, and possessed a 345:described in 2019, and a second species, 1819: 1554:Angielczyk, K.D.; Fröbisch, J. (eds.). 1385: 521:it emerges and runs laterally from the 517:and its blood vessels in the snout. In 262:Angielczyk, Benoit & Rubidge, 2019 3688:Wuchiapingian genus first appearances 754:is well supported as a member of the 7: 983: 959: 935: 911: 884: 860: 810: 803: 779: 769: 727:combines the surname Juwayeyi with 1665:Journal of Vertebrate Paleontology 1621:Journal of Vertebrate Paleontology 1337:. Reptiles are represented by the 1120:prominent ridges over the eyes in 533:(sheets of bone connecting to the 236:Angielczyk, Benoit & Rubidge, 14: 1790:from the Upper Permian of Zambia" 1467:Journal of African Earth Sciences 1312:, as well as the more widespread 641:. It was named for the iron-rich 3393: 3032: 3027: 2608: 2603: 2216: 1846: 1834: 1822: 1489:10.1016/j.jafrearsci.2022.104726 375:species of cistecephalid in the 115: 3693:Changhsingian genus extinctions 685:, which have been dated to the 606:University of the Witwatersrand 324:. The genus contains two known 1324:, along with the semi-aquatic 709:to image the bones inside the 602:Evolutionary Studies Institute 1: 3713:Fossil taxa described in 2019 3698:Lopingian synapsids of Africa 1685:10.1080/02724634.2016.1111897 339:Madumabisa Mudstone Formation 1642:10.1080/02724634.2015.969400 16:Extinct genus of dicynodonts 1192:The footplates of the bony 3729: 3417: 3406: 3391: 3256: 3073: 3058: 3045: 3025: 2844: 2649: 2634: 2621: 2601: 2400: 2257: 2242: 2229: 2214: 2140: 2033: 2018: 2005: 1955: 1942: 1896: 1321:Ictidosuchoides longiceps 1005: 988: 981: 964: 957: 940: 933: 916: 909: 889: 882: 865: 858: 832: 815: 808: 801: 784: 777: 594:Cistecephalus microrhinus 272: 267: 251: 244: 112:Scientific classification 110: 23: 1788:Pareiasuchus nasicornis 1749:Palaeontologia Africana 1589:Palaeontologia Africana 1513:Papers in Palaeontology 1417:Papers in Palaeontology 1372:with plentiful sinuous 1247:, a species similar to 596:and the now synonymous 1734:10.1098/rstb.1979.0001 1562:. pp. 93–138 In. 1460:Kembawacela yajuwayeyi 363:was specialised for a 302:("iron digger") is an 3655:Paleobiology Database 1333:and an indeterminate 1315:Theriognathus microps 1256:Odontocyclops whaitsi 766:member of the clade: 707:x-ray microtomography 634:Kembawacela kitchingi 334:Kembawacela kitchingi 257:Kembawacela kitchingi 3210:Shaanbeikannemeyeria 1115:It is possible that 576:History of discovery 3571:Paraphyletic groups 1726:1979RSPTB.285...73K 1677:2016JVPal..36E1897H 1634:2015JVPal..35E9400H 1560:Springer, Dordrecht 1480:2022JAfES.19604726A 3065:Kannemeyeriiformes 3052:Kannemeyeriiformes 2997:Kannemeyeriiformes 1474:: Article 104726. 1232:Diictodon feliceps 1177:). Interestingly, 1169:small eyes (as in 1104:interparietal bone 623:in an unpublished 464:sexually dimorphic 3708:Fossils of Malawi 3703:Fossils of Zambia 3670: 3669: 3610:Taxon identifiers 3601: 3600: 3597: 3596: 3593: 3592: 3402: 3401: 3389: 3388: 3385: 3384: 3381: 3380: 3224:Uralokannemeyeria 3041: 3040: 3023: 3022: 3019: 3018: 3015: 3014: 2810:Rhachiocephalidae 2617: 2616: 2599: 2598: 2595: 2594: 2591: 2590: 2567: 2566: 2225: 2224: 2212: 2211: 2208: 2207: 2204: 2203: 2001: 2000: 1997: 1996: 1993: 1992: 1569:978-94-007-6841-3 1525:10.1002/spp2.1055 1430:10.1002/spp2.1285 1374:meandering rivers 1293:and a species of 1163:olecranon process 1092: 1091: 1083: 1082: 1074: 1073: 1065: 1064: 1056: 1055: 1047: 1046: 1038: 1037: 1029: 1028: 1020: 1019: 847: 846: 737:Chichewa language 733:possessive prefix 489:of the cheek. In 440:. Similarly, the 420:temporal fenestra 295: 294: 289: 240: 3720: 3663: 3662: 3650: 3649: 3637: 3636: 3635: 3605: 3419: 3408: 3397: 3254: 3238:Xiyukannemeyeria 3217:Sinokannemeyeria 3182:Parakannemeyeria 3071: 3060: 3047: 3036: 3031: 2842: 2819:Kitchinganomodon 2785:Australobarbarus 2717:Interpresosaurus 2647: 2636: 2623: 2612: 2607: 2530:Cistecephaloides 2398: 2389: 2255: 2244: 2231: 2220: 2174:Anomocephaloidea 2138: 2058:Anomocephaloidea 2031: 2020: 2007: 1953: 1944: 1937: 1936: 1898: 1883: 1876: 1869: 1860: 1851: 1850: 1849: 1839: 1838: 1837: 1827: 1826: 1825: 1818: 1807: 1806: 1794: 1781: 1775: 1774: 1764: 1744: 1738: 1737: 1720:(1005): 73–122. 1703: 1697: 1696: 1660: 1654: 1653: 1611: 1605: 1604: 1580: 1574: 1573: 1550: 1537: 1536: 1508: 1502: 1501: 1491: 1453: 1442: 1441: 1411: 1346:, a herbivorous 1268:Kitchinganomodon 1262:Oudenodon bainii 1229:, the burrowing 1171:Cistecephaloides 1009:Cistecephaloides 984: 960: 936: 912: 885: 861: 811: 804: 780: 770: 562:sacral vertebrae 515:trigeminal nerve 495:postorbital bone 442:zygomatic arches 431:Cistecephaloides 281: 277: 254: 235: 228: 215: 202: 186: 120: 119: 105: 42: 29:Temporal range: 21: 3728: 3727: 3723: 3722: 3721: 3719: 3718: 3717: 3673: 3672: 3671: 3666: 3658: 3653: 3645: 3640: 3631: 3630: 3625: 3612: 3602: 3589: 3565: 3546:Platypodosaurus 3525:Opisthoctenodon 3413: 3398: 3377: 3337:Eubrachiosaurus 3328:Stahleckeriinae 3322: 3267: 3249:Stahleckeriidae 3243: 3152:Kannemeyeriidae 3146: 3111:Shansiodontidae 3105: 3093:Elephantosaurus 3086:Dinodontosaurus 3067: 3054: 3037: 3011: 2991: 2955:Euptychognathus 2932: 2837:Dicynodontoidea 2831: 2826:Rhachiocephalus 2804: 2770: 2729: 2695: 2643: 2630: 2613: 2587: 2563: 2521:Cistecephalidae 2515: 2496:Dicynodontoides 2481: 2454: 2427: 2378: 2344: 2309:Pylaecephalidae 2303: 2251: 2238: 2221: 2200: 2180: 2127: 2079: 2052: 2027: 2014: 1989: 1969: 1938: 1903: 1902: 1892: 1887: 1857: 1847: 1845: 1835: 1833: 1823: 1821: 1813: 1811: 1810: 1792: 1783: 1782: 1778: 1746: 1745: 1741: 1705: 1704: 1700: 1671:(4): e1111897. 1662: 1661: 1657: 1613: 1612: 1608: 1582: 1581: 1577: 1570: 1552: 1551: 1540: 1510: 1509: 1505: 1455: 1454: 1445: 1413: 1412: 1387: 1382: 1299:therocephalians 1244:Dicynodontoides 1206: 1156: 1124:, and although 1113: 1093: 1084: 1075: 1066: 1057: 1048: 1039: 1030: 1021: 906:Cistecephalidae 848: 749: 668:Assemblage Zone 578: 560:. It had three 531:pterygoid bones 523:maxillary sinus 499:postorbital bar 385: 263: 260: 234: 226: 217:Cistecephalidae 213: 200: 184: 114: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 34: 33: 27: 17: 12: 11: 5: 3726: 3724: 3716: 3715: 3710: 3705: 3700: 3695: 3690: 3685: 3675: 3674: 3668: 3667: 3665: 3664: 3651: 3638: 3622: 3620: 3614: 3613: 3608: 3599: 3598: 3595: 3594: 3591: 3590: 3588: 3587: 3582: 3575: 3573: 3567: 3566: 3564: 3563: 3556: 3549: 3542: 3539:Planitorostris 3535: 3528: 3521: 3514: 3507: 3500: 3493: 3486: 3479: 3472: 3465: 3458: 3455:Chelyoposaurus 3451: 3444: 3437: 3429: 3427: 3415: 3414: 3412:Other articles 3411: 3404: 3403: 3400: 3399: 3392: 3390: 3387: 3386: 3383: 3382: 3379: 3378: 3376: 3375: 3368: 3361: 3354: 3347: 3344:Ischigualastia 3340: 3332: 3330: 3324: 3323: 3321: 3320: 3313: 3306: 3299: 3292: 3285: 3277: 3275: 3269: 3268: 3266: 3265: 3257: 3251: 3245: 3244: 3242: 3241: 3234: 3227: 3220: 3213: 3206: 3199: 3192: 3185: 3178: 3171: 3164: 3156: 3154: 3148: 3147: 3145: 3144: 3141:Rhinodicynodon 3137: 3130: 3123: 3115: 3113: 3107: 3106: 3104: 3103: 3096: 3089: 3082: 3074: 3068: 3063: 3056: 3055: 3050: 3043: 3042: 3039: 3038: 3026: 3024: 3021: 3020: 3017: 3016: 3013: 3012: 3010: 3009: 3008: 3007: 3001: 2999: 2993: 2992: 2990: 2989: 2981: 2974: 2967: 2959: 2951: 2942: 2940: 2938:Lystrosauridae 2934: 2933: 2931: 2930: 2923: 2916: 2909: 2906:Repelinosaurus 2902: 2895: 2888: 2881: 2874: 2867: 2860: 2853: 2845: 2839: 2833: 2832: 2830: 2829: 2822: 2814: 2812: 2806: 2805: 2803: 2802: 2795: 2788: 2780: 2778: 2776:Oudenodontidae 2772: 2771: 2769: 2768: 2761: 2754: 2747: 2744:Aulacephalodon 2739: 2737: 2731: 2730: 2728: 2727: 2720: 2713: 2705: 2703: 2697: 2696: 2694: 2693: 2686: 2679: 2672: 2665: 2658: 2655:Daqingshanodon 2650: 2644: 2639: 2632: 2631: 2626: 2619: 2618: 2615: 2614: 2602: 2600: 2597: 2596: 2593: 2592: 2589: 2588: 2586: 2585: 2584: 2583: 2577: 2575: 2569: 2568: 2565: 2564: 2562: 2561: 2554: 2547: 2540: 2533: 2525: 2523: 2517: 2516: 2514: 2513: 2506: 2499: 2491: 2489: 2483: 2482: 2480: 2479: 2472: 2464: 2462: 2456: 2455: 2453: 2452: 2445: 2437: 2435: 2429: 2428: 2426: 2425: 2417: 2409: 2401: 2395: 2386: 2380: 2379: 2377: 2376: 2369: 2362: 2354: 2352: 2350:Endothiodontia 2346: 2345: 2343: 2342: 2335: 2328: 2321: 2313: 2311: 2305: 2304: 2302: 2301: 2294: 2287: 2284:Lanthanostegus 2280: 2273: 2266: 2263:Brachyprosopus 2258: 2252: 2247: 2240: 2239: 2234: 2227: 2226: 2223: 2222: 2215: 2213: 2210: 2209: 2206: 2205: 2202: 2201: 2199: 2198: 2197: 2196: 2190: 2188: 2182: 2181: 2179: 2178: 2170: 2163: 2156: 2149: 2141: 2135: 2129: 2128: 2126: 2125: 2118: 2111: 2104: 2097: 2089: 2087: 2085:Venyukovioidea 2081: 2080: 2078: 2077: 2070: 2062: 2060: 2054: 2053: 2051: 2050: 2042: 2034: 2028: 2023: 2016: 2015: 2010: 2003: 2002: 1999: 1998: 1995: 1994: 1991: 1990: 1988: 1987: 1986: 1985: 1979: 1977: 1971: 1970: 1968: 1967: 1966: 1965: 1956: 1950: 1940: 1939: 1935: 1934: 1925: 1916: 1910: 1901: 1894: 1893: 1888: 1886: 1885: 1878: 1871: 1863: 1856: 1855: 1843: 1831: 1809: 1808: 1776: 1739: 1698: 1655: 1628:(5): e969400. 1606: 1575: 1568: 1538: 1519:(4): 569–584. 1503: 1443: 1384: 1383: 1381: 1378: 1205: 1202: 1183:sclerotic ring 1155: 1152: 1112: 1109: 1090: 1089: 1086: 1085: 1081: 1080: 1077: 1076: 1072: 1071: 1068: 1067: 1063: 1062: 1059: 1058: 1054: 1053: 1050: 1049: 1045: 1044: 1041: 1040: 1036: 1035: 1032: 1031: 1027: 1026: 1023: 1022: 1018: 1017: 1014: 1013: 1004: 1001: 1000: 997: 996: 987: 982: 980: 977: 976: 973: 972: 963: 958: 956: 953: 952: 949: 948: 939: 934: 932: 929: 928: 925: 924: 915: 910: 908: 902: 901: 898: 897: 888: 883: 881: 878: 877: 874: 873: 864: 859: 857: 854: 853: 850: 849: 845: 844: 841: 840: 831: 828: 827: 824: 823: 814: 809: 807: 802: 800: 797: 796: 793: 792: 783: 778: 776: 768: 748: 747:Classification 745: 639:Bemba language 590:James Kitching 577: 574: 535:palatine bones 446:pineal foramen 384: 381: 371:was a locally 293: 292: 291: 290: 270: 269: 268:Other species 265: 264: 261: 249: 248: 242: 241: 224: 220: 219: 211: 207: 206: 198: 191: 190: 182: 178: 177: 172: 165: 164: 159: 152: 151: 146: 142: 141: 136: 132: 131: 126: 122: 121: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 43: 28: 15: 13: 10: 9: 6: 4: 3: 2: 3725: 3714: 3711: 3709: 3706: 3704: 3701: 3699: 3696: 3694: 3691: 3689: 3686: 3684: 3681: 3680: 3678: 3661: 3656: 3652: 3648: 3643: 3639: 3634: 3628: 3624: 3623: 3621: 3619: 3615: 3611: 3606: 3586: 3583: 3580: 3577: 3576: 3574: 3572: 3568: 3562: 3561: 3557: 3555: 3554: 3553:Putillosaurus 3550: 3548: 3547: 3543: 3541: 3540: 3536: 3534: 3533: 3532:Parvobestiola 3529: 3527: 3526: 3522: 3520: 3519: 3515: 3513: 3512: 3508: 3506: 3505: 3501: 3499: 3498: 3494: 3492: 3491: 3487: 3485: 3484: 3480: 3478: 3477: 3476:Dicranozygoma 3473: 3471: 3470: 3466: 3464: 3463: 3459: 3457: 3456: 3452: 3450: 3449: 3445: 3443: 3442: 3438: 3436: 3435: 3431: 3430: 3428: 3426: 3425: 3420: 3416: 3409: 3405: 3396: 3374: 3373: 3369: 3367: 3366: 3362: 3360: 3359: 3355: 3353: 3352: 3348: 3346: 3345: 3341: 3339: 3338: 3334: 3333: 3331: 3329: 3325: 3319: 3318: 3314: 3312: 3311: 3307: 3305: 3304: 3300: 3298: 3297: 3293: 3291: 3290: 3286: 3284: 3283: 3282:Argodicynodon 3279: 3278: 3276: 3274: 3270: 3264: 3263: 3259: 3258: 3255: 3252: 3250: 3246: 3240: 3239: 3235: 3233: 3232: 3228: 3226: 3225: 3221: 3219: 3218: 3214: 3212: 3211: 3207: 3205: 3204: 3200: 3198: 3197: 3193: 3191: 3190: 3186: 3184: 3183: 3179: 3177: 3176: 3172: 3170: 3169: 3165: 3163: 3162: 3158: 3157: 3155: 3153: 3149: 3143: 3142: 3138: 3136: 3135: 3131: 3129: 3128: 3124: 3122: 3121: 3117: 3116: 3114: 3112: 3108: 3102: 3101: 3097: 3095: 3094: 3090: 3088: 3087: 3083: 3081: 3080: 3076: 3075: 3072: 3069: 3066: 3061: 3057: 3053: 3048: 3044: 3035: 3030: 3005: 3004: 3003: 3002: 3000: 2998: 2994: 2987: 2986: 2985:Sintocephalus 2982: 2980: 2979: 2975: 2973: 2972: 2971:Kwazulusaurus 2968: 2965: 2964: 2960: 2957: 2956: 2952: 2949: 2948: 2944: 2943: 2941: 2939: 2935: 2929: 2928: 2924: 2922: 2921: 2917: 2915: 2914: 2910: 2908: 2907: 2903: 2901: 2900: 2896: 2894: 2893: 2889: 2887: 2886: 2882: 2880: 2879: 2875: 2873: 2872: 2868: 2866: 2865: 2864:Delectosaurus 2861: 2859: 2858: 2857:Daptocephalus 2854: 2852: 2851: 2847: 2846: 2843: 2840: 2838: 2834: 2828: 2827: 2823: 2821: 2820: 2816: 2815: 2813: 2811: 2807: 2801: 2800: 2796: 2794: 2793: 2789: 2787: 2786: 2782: 2781: 2779: 2777: 2773: 2767: 2766: 2762: 2760: 2759: 2755: 2753: 2752: 2748: 2746: 2745: 2741: 2740: 2738: 2736: 2732: 2726: 2725: 2721: 2719: 2718: 2714: 2712: 2711: 2707: 2706: 2704: 2702: 2698: 2692: 2691: 2687: 2685: 2684: 2680: 2678: 2677: 2676:Odontocyclops 2673: 2671: 2670: 2666: 2664: 2663: 2659: 2657: 2656: 2652: 2651: 2648: 2645: 2642: 2637: 2633: 2629: 2624: 2620: 2611: 2606: 2581: 2580: 2579: 2578: 2576: 2574: 2570: 2560: 2559: 2555: 2553: 2552: 2548: 2546: 2545: 2544:Kawingasaurus 2541: 2539: 2538: 2537:Cistecephalus 2534: 2532: 2531: 2527: 2526: 2524: 2522: 2518: 2512: 2511: 2510:Thliptosaurus 2507: 2505: 2504: 2500: 2498: 2497: 2493: 2492: 2490: 2488: 2484: 2478: 2477: 2473: 2471: 2470: 2466: 2465: 2463: 2461: 2457: 2451: 2450: 2446: 2444: 2443: 2439: 2438: 2436: 2434: 2430: 2423: 2422: 2418: 2415: 2414: 2410: 2408: 2407: 2403: 2402: 2399: 2396: 2394: 2390: 2387: 2385: 2384:Therochelonia 2381: 2375: 2374: 2370: 2368: 2367: 2363: 2361: 2360: 2356: 2355: 2353: 2351: 2347: 2341: 2340: 2336: 2334: 2333: 2329: 2327: 2326: 2322: 2320: 2319: 2315: 2314: 2312: 2310: 2306: 2300: 2299: 2295: 2293: 2292: 2288: 2286: 2285: 2281: 2279: 2278: 2274: 2272: 2271: 2267: 2265: 2264: 2260: 2259: 2256: 2253: 2250: 2245: 2241: 2237: 2232: 2228: 2219: 2194: 2193: 2192: 2191: 2189: 2187: 2183: 2176: 2175: 2171: 2169: 2168: 2164: 2162: 2161: 2157: 2155: 2154: 2150: 2148: 2147: 2143: 2142: 2139: 2136: 2134: 2130: 2124: 2123: 2119: 2117: 2116: 2112: 2110: 2109: 2105: 2103: 2102: 2098: 2096: 2095: 2091: 2090: 2088: 2086: 2082: 2076: 2075: 2071: 2069: 2068: 2067:Anomocephalus 2064: 2063: 2061: 2059: 2055: 2048: 2047: 2043: 2041: 2040: 2036: 2035: 2032: 2029: 2026: 2021: 2017: 2013: 2008: 2004: 1983: 1982: 1981: 1980: 1978: 1976: 1972: 1964: 1960: 1959: 1958: 1957: 1954: 1951: 1949: 1945: 1941: 1933: 1929: 1926: 1924: 1920: 1917: 1915: 1911: 1909: 1905: 1904: 1899: 1895: 1891: 1884: 1879: 1877: 1872: 1870: 1865: 1864: 1861: 1854: 1844: 1842: 1832: 1830: 1820: 1816: 1805:(2): 307–335. 1804: 1800: 1799: 1798:Palaeontology 1791: 1789: 1780: 1777: 1772: 1768: 1763: 1758: 1754: 1750: 1743: 1740: 1735: 1731: 1727: 1723: 1719: 1715: 1714: 1709: 1708:Procynosuchus 1702: 1699: 1694: 1690: 1686: 1682: 1678: 1674: 1670: 1666: 1659: 1656: 1651: 1647: 1643: 1639: 1635: 1631: 1627: 1623: 1622: 1617: 1610: 1607: 1602: 1598: 1594: 1590: 1586: 1579: 1576: 1571: 1565: 1561: 1557: 1549: 1547: 1545: 1543: 1539: 1534: 1530: 1526: 1522: 1518: 1514: 1507: 1504: 1499: 1495: 1490: 1485: 1481: 1477: 1473: 1469: 1468: 1463: 1461: 1452: 1450: 1448: 1444: 1439: 1435: 1431: 1427: 1423: 1419: 1418: 1410: 1408: 1406: 1404: 1402: 1400: 1398: 1396: 1394: 1392: 1390: 1386: 1379: 1377: 1375: 1371: 1366: 1364: 1360: 1356: 1355:Cistecephalus 1351: 1349: 1345: 1344: 1340: 1336: 1335:biarmosuchian 1332: 1331: 1330:Procynosuchus 1327: 1323: 1322: 1317: 1316: 1311: 1310: 1305: 1304: 1300: 1296: 1292: 1288: 1287: 1282: 1281: 1276: 1275: 1274:Daptocephalus 1270: 1269: 1264: 1263: 1258: 1257: 1252: 1251: 1246: 1245: 1240: 1239: 1234: 1233: 1228: 1227: 1222: 1221: 1216: 1215: 1210: 1204:Palaeoecology 1203: 1201: 1199: 1198:Kawingasaurus 1195: 1190: 1188: 1184: 1180: 1176: 1175:Kawingasaurus 1172: 1166: 1164: 1160: 1153: 1151: 1149: 1145: 1141: 1140: 1135: 1132:The tusks of 1130: 1127: 1123: 1122:Cistecephalus 1118: 1111:Palaeobiology 1110: 1108: 1105: 1101: 1097: 1088: 1087: 1079: 1078: 1070: 1069: 1061: 1060: 1052: 1051: 1043: 1042: 1034: 1033: 1025: 1024: 1016: 1015: 1012: 1011: 1010: 1003: 1002: 999: 998: 995: 994: 993: 992:Kawingasaurus 986: 985: 979: 978: 975: 974: 971: 970: 969: 962: 961: 955: 954: 951: 950: 947: 946: 945: 938: 937: 931: 930: 927: 926: 923: 922: 921: 920:Cistecephalus 914: 913: 907: 904: 903: 900: 899: 896: 895: 894: 887: 886: 880: 879: 876: 875: 872: 871: 870: 863: 862: 856: 855: 852: 851: 843: 842: 839: 838: 837: 830: 829: 826: 825: 822: 821: 820: 813: 812: 806: 805: 799: 798: 795: 794: 791: 790: 789: 782: 781: 775: 774:Kistecephalia 772: 771: 767: 765: 761: 757: 753: 746: 744: 742: 738: 734: 730: 726: 722: 721: 720:K. yajuwayeyi 716: 712: 708: 702: 700: 696: 692: 691:Changhsingian 688: 687:Wuchiapingian 684: 680: 676: 674: 673:Daptocephalus 669: 667: 666:Cistecephalus 661: 659: 655: 651: 650:type specimen 647: 644: 640: 636: 635: 629: 626: 622: 617: 615: 611: 607: 603: 599: 595: 591: 587: 586:Alan Drysdall 583: 580:Specimens of 575: 573: 571: 570:Cistecephalus 567: 563: 559: 558:Cistecephalus 555: 551: 546: 544: 540: 539:K. yajuwayeyi 536: 532: 528: 527:K. yajuwayeyi 524: 520: 516: 512: 508: 507:K. yajuwayeyi 504: 503:K. yajuwayeyi 500: 496: 492: 488: 484: 483:K. yajuwayeyi 480: 475: 473: 469: 465: 461: 457: 453: 452: 447: 443: 439: 438: 437:Kawingasaurus 433: 432: 427: 426: 425:Cistecephalus 421: 416: 414: 413:K. yajuwayeyi 410: 406: 402: 398: 394: 389: 382: 380: 378: 377:Luangwa Basin 374: 370: 366: 362: 358: 354: 350: 349: 348:K. yajuwayeyi 344: 340: 336: 335: 331: 327: 323: 319: 315: 312: 311:cistecephalid 308: 305: 301: 300: 288: 285: 280: 279:K. yajuwayeyi 274: 273: 271: 266: 259: 258: 250: 247: 243: 239: 233: 232: 225: 222: 221: 218: 212: 209: 208: 205: 199: 196: 193: 192: 189: 183: 180: 179: 176: 173: 170: 167: 166: 163: 160: 157: 154: 153: 150: 147: 144: 143: 140: 137: 134: 133: 130: 127: 124: 123: 118: 113: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 40:Changhsingian 37: 36:Wuchiapingian 32: 26: 22: 19: 3617: 3579:Cryptodontia 3558: 3551: 3544: 3537: 3530: 3523: 3518:Nasoplanites 3516: 3509: 3504:Keirognathus 3502: 3497:Haughtoniana 3495: 3488: 3481: 3474: 3467: 3460: 3453: 3446: 3439: 3434:Azarifeneria 3432: 3424:Nomina dubia 3422: 3372:Ufudocyclops 3370: 3365:Stahleckeria 3363: 3356: 3349: 3342: 3335: 3317:Zambiasaurus 3315: 3308: 3301: 3294: 3287: 3280: 3260: 3236: 3229: 3222: 3215: 3208: 3203:Rhadiodromus 3201: 3196:Rechnisaurus 3194: 3189:Rabidosaurus 3187: 3180: 3175:Kannemeyeria 3173: 3168:Dolichuranus 3166: 3161:Acratophorus 3159: 3139: 3132: 3125: 3118: 3098: 3091: 3084: 3079:Angonisaurus 3077: 2983: 2978:Lystrosaurus 2976: 2969: 2961: 2953: 2945: 2927:Vivaxosaurus 2925: 2918: 2911: 2904: 2897: 2890: 2883: 2876: 2869: 2862: 2855: 2848: 2824: 2817: 2799:Tropidostoma 2797: 2790: 2783: 2763: 2756: 2749: 2742: 2722: 2715: 2708: 2688: 2681: 2674: 2667: 2660: 2653: 2558:Sauroscaptor 2556: 2550: 2549: 2542: 2535: 2528: 2508: 2501: 2494: 2474: 2469:Myosauroides 2467: 2447: 2440: 2419: 2411: 2404: 2371: 2364: 2357: 2337: 2330: 2323: 2316: 2296: 2289: 2282: 2275: 2270:Colobodectes 2268: 2261: 2249:Dicynodontia 2236:Dicynodontia 2186:Dicynodontia 2172: 2167:Patranomodon 2165: 2158: 2151: 2144: 2133:Chainosauria 2120: 2113: 2106: 2099: 2092: 2072: 2065: 2044: 2037: 1927: 1918: 1829:Paleontology 1802: 1796: 1787: 1779: 1752: 1748: 1742: 1717: 1711: 1707: 1701: 1668: 1664: 1658: 1625: 1619: 1609: 1592: 1588: 1584: 1578: 1555: 1516: 1512: 1506: 1471: 1465: 1459: 1421: 1415: 1367: 1362: 1358: 1354: 1352: 1343:Pareiasuchus 1341: 1328: 1319: 1313: 1307: 1301: 1295:lystrosaurid 1290: 1284: 1278: 1272: 1266: 1260: 1254: 1248: 1242: 1236: 1230: 1224: 1218: 1212: 1208: 1207: 1197: 1191: 1178: 1174: 1170: 1167: 1158: 1157: 1147: 1143: 1137: 1133: 1131: 1125: 1121: 1116: 1114: 1095: 1094: 1007: 1006: 990: 989: 968:Sauroscaptor 966: 965: 943: 942: 941: 918: 917: 891: 890: 867: 866: 834: 833: 817: 816: 786: 785: 759: 751: 750: 728: 725:species name 719: 718: 714: 703: 695:Late Permian 683:South Africa 672: 665: 662: 657: 653: 633: 632: 621:Cistcephalus 620: 618: 614:South Africa 610:Johannesburg 598:C. planiceps 597: 593: 581: 579: 569: 557: 554:K. kitchingi 553: 549: 548:The body of 547: 543:K. kitchingi 542: 538: 526: 519:K. kitchingi 518: 511:K. kitchingi 510: 506: 502: 497:to form the 491:K. kitchingi 490: 482: 479:K. kitchingi 478: 476: 467: 459: 455: 451:Sauroscaptor 449: 435: 429: 423: 417: 412: 409:K. kitchingi 408: 392: 387: 386: 368: 360: 353:Chiweta Beds 347: 346: 333: 332: 330:type species 318:Late Permian 298: 297: 296: 283: 278: 256: 255: 246:Type species 230: 229: 204:Dicynodontia 194: 168: 155: 31:Late Permian 24: 18: 3618:Kembawacela 3585:Dromasauria 3483:Elatosaurus 3469:Diaelurodon 3462:Cristonasus 3448:Calleonasus 3358:Sangusaurus 3303:Pentasaurus 3296:Moghreberia 3273:Placeriinae 3231:Wadiasaurus 3127:Tetragonias 2878:Dinanomodon 2850:Counillonia 2765:Pelanomodon 2751:Bulbasaurus 2662:Idelesaurus 2551:Kembawacela 2487:Kingoriidae 2460:Myosauridae 2413:Emydorhinus 2393:Emydopoidea 2366:Endothiodon 2332:Prosictodon 2298:Pristerodon 2277:Eodicynodon 2101:Parasuminia 2074:Tiarajudens 2046:Novocynodon 2025:Anomodontia 2012:Anomodontia 1975:Anomodontia 1890:Anomodontia 1762:10539/17373 1601:10539/26832 1595:: 219–225. 1424:: 405–446. 1370:rift valley 1363:Kembawacela 1359:Kembawacela 1348:parareptile 1303:Ichibengops 1226:Pristerodon 1214:Endothiodon 1209:Kembawacela 1179:Kembawacela 1159:Kembawacela 1148:Kembawacela 1144:Kembawacela 1134:Kembawacela 1126:Kembawacela 1117:Kembawacela 1096:Kembawacela 944:Kembawacela 788:Kingoriidae 752:Kembawacela 715:Kembawacela 711:rock matrix 679:Karoo Basin 658:Kembawacela 654:Kembawacela 625:BSc honours 582:Kembawacela 550:Kembawacela 468:Kembawacela 460:Kembawacela 456:Kembawacela 393:Kembawacela 388:Kembawacela 383:Description 379:of Zambia. 369:Kembawacela 361:Kembawacela 351:, from the 322:East Africa 299:Kembawacela 231:Kembawacela 188:Anomodontia 25:Kembawacela 3683:Emydopoids 3677:Categories 3560:Taognathus 3490:Eurycarpus 3351:Jachaleria 3120:Shansiodon 3100:Woznikella 3006:see below↓ 2920:Turfanodon 2641:Bidentalia 2628:Bidentalia 2582:see below↓ 2573:Bidentalia 2433:Emydopidae 2421:Palemydops 2195:see below↓ 2146:Galechirus 2122:Venyukovia 2039:Biseridens 1984:see below↓ 1932:Therapsida 1380:References 1339:pareiasaur 701:, Zambia. 487:jugal bone 472:keratinous 314:dicynodont 181:Suborder: 175:Therapsida 3647:159712724 3633:Q71708346 3310:Placerias 3289:Lisowicia 3262:Sungeodon 2963:Jimusaria 2947:Basilodon 2913:Taoheodon 2899:Peramodon 2871:Dicynodon 2792:Oudenodon 2735:Geikiidae 2503:Kombuisia 2476:Myosaurus 2442:Compsodon 2406:Digalodon 2373:Niassodon 2318:Diictodon 2291:Nyaphulia 1963:Synapsida 1948:Synapsida 1923:Synapsida 1906:Kingdom: 1771:2410-4418 1693:130994874 1585:Digalodon 1498:252393501 1438:210304700 1286:Digalodon 1220:Compsodon 1154:Burrowing 1139:Diictodon 893:Myosaurus 869:Digalodon 819:Compsodon 689:to early 670:(AZ) and 365:fossorial 337:from the 316:from the 162:Synapsida 135:Kingdom: 129:Eukaryota 3627:Wikidata 3441:Baiopsis 3134:Vinceria 2892:Kunpania 2885:Gordonia 2724:Katumbia 2701:Elphidae 2683:Rastodon 2669:Keyseria 2359:Abajudon 2339:Robertia 2325:Eosimops 2094:Otsheria 1914:Chordata 1912:Phylum: 1908:Animalia 1650:83554630 1533:88833541 1326:cynodont 1291:rubidgei 1250:Katumbia 643:hematite 509:than in 477:Between 210:Family: 149:Chordata 145:Phylum: 139:Animalia 125:Domain: 2449:Emydops 2160:Galepus 2153:Galeops 2115:Ulemica 2108:Suminia 1815:Portals 1755:: 1–7. 1722:Bibcode 1673:Bibcode 1630:Bibcode 1476:Bibcode 1309:Mupashi 1238:Emydops 1102:and an 836:Emydops 741:occiput 735:in the 693:in the 677:of the 646:nodules 604:at the 564:and an 373:endemic 326:species 304:extinct 282:AraĂșjo 223:Genus: 3660:398004 3511:Koupia 2758:Geikia 1853:Malawi 1841:Zambia 1769:  1691:  1648:  1566:  1531:  1496:  1436:  1194:stapes 1187:orbits 760:et al. 756:family 731:, the 699:Lusaka 628:thesis 397:pelvis 357:Malawi 343:Zambia 328:, the 284:et al. 2690:Syops 1928:Clade 1919:Clade 1793:(PDF) 1689:S2CID 1646:S2CID 1529:S2CID 1494:S2CID 1434:S2CID 1280:Syops 1100:vomer 764:basal 566:ilium 401:femur 307:genus 195:Clade 169:Clade 156:Clade 3642:GBIF 2710:Elph 1961:see 1767:ISSN 1564:ISBN 1318:and 1306:and 1289:cf. 1173:and 588:and 481:and 434:and 411:and 405:ulna 287:2022 238:2019 46:PreꞒ 1757:hdl 1730:doi 1718:285 1681:doi 1638:doi 1597:hdl 1521:doi 1484:doi 1472:196 1426:doi 729:Ya- 681:in 652:of 608:in 454:). 355:of 341:of 320:of 309:of 3679:: 3657:: 3644:: 3629:: 1930:: 1921:: 1803:40 1801:. 1795:. 1765:. 1753:49 1751:. 1728:. 1716:. 1687:. 1679:. 1669:36 1667:. 1644:. 1636:. 1626:35 1624:. 1618:. 1593:53 1591:. 1558:. 1541:^ 1527:. 1515:. 1492:. 1482:. 1470:. 1464:. 1446:^ 1432:. 1420:. 1388:^ 1350:. 1283:, 1277:, 1271:, 1265:, 1259:, 1253:, 1241:, 1235:, 1223:, 1217:, 1200:. 743:. 717:, 675:AZ 660:. 612:, 545:. 403:, 399:, 197:: 171:: 158:: 96:Pg 3581:? 2988:? 2966:? 2958:? 2950:? 2424:? 2416:? 2177:? 2049:? 1882:e 1875:t 1868:v 1817:: 1773:. 1759:: 1736:. 1732:: 1724:: 1695:. 1683:: 1675:: 1652:. 1640:: 1632:: 1603:. 1599:: 1572:. 1535:. 1523:: 1517:2 1500:. 1486:: 1478:: 1458:" 1440:. 1428:: 1422:7 276:† 253:† 227:† 214:† 201:† 185:† 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ 38:–

Index

Late Permian
Wuchiapingian
Changhsingian
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Synapsida
Therapsida
Anomodontia
Dicynodontia
Cistecephalidae
Kembawacela
2019
Type species
2022
extinct

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