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Phanaeini

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31: 204: 51: 632: 589: 574:"A Worldwide Checklist of the Tribes and Genera of Aegialiinae, Aphodiinae, Termitotroginae, Aulonocneminae, Scarabaeinae, and Coprinae (Coleoptera: Scarabaeoidea: Scarabaeidae)" 195:
includes some of the largest dung beetles), often with bright metallic colors, and often with horns on their heads (especially in the males, less frequently in the females).
808: 604: 731: 462:"A phylogenetic analysis of the New World tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae): Hypotheses on relationships and origins" 30: 531: 203: 801: 562: 736: 407: 590:"A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae)" 832: 794: 50: 648: 827: 744: 636: 352:
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
492: 269: 279: 45: 749: 330: 289: 718: 558: 527: 389: 371: 309: 588:
Vaz-de-Mello, Fernando Z.; Edmonds, W.D.; Ocampo, Federico C.; Schoolmeesters, Paul (2011).
548: 473: 379: 361: 155:. There are about 12 genera and 200 described species in Phanaeini. They are native to the 686: 778: 384: 172: 821: 461: 320: 229: 774: 723: 353: 239: 168: 164: 152: 122: 112: 542: 680: 148: 671: 572:
Mathison, Blaine A.; Hardy, Martin; Bezdek, Aleš; Schoolmeesters, Paul (2001).
477: 299: 249: 160: 375: 553: 524:
American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
522:
Arnett, R.H. Jr.; Thomas, M. C.; Skelley, P. E.; Frank, J. H., eds. (2002).
82: 62: 393: 366: 697: 631: 705: 665: 259: 219: 156: 710: 432: 102: 92: 72: 642: 202: 573: 646: 782: 547:. Smithsonian Miscellaneous Collections. Vol. 3. 655: 171:, but some of the least known genera appear to be 544:Classification of the Coleoptera of North America 460:Philips, T; Edmonds, W; Scholtz, Clarke (2004). 213:These 12 genera belong to the tribe Phanaeini: 493:"Scarabaeidae (Neotropical Region) – Coprinae" 802: 455: 453: 8: 354:"Family-group names in Coleoptera (Insecta)" 191:). They are medium-sized to large beetles ( 809: 795: 643: 29: 20: 552: 383: 365: 159:with the highest species richness in the 412:Integrated Taxonomic Information System 344: 497:Kaefer der Welt - Beetles of the World 7: 770: 768: 466:Insect Systematics & Evolution 14: 630: 603:. Magnolia Press. Archived from 49: 1: 433:"Phanaeini Tribe Information" 781:. You can help Knowledge by 849: 767: 478:10.1163/187631204788964664 304:Laporte de Castelnau, 1840 46:Scientific classification 44: 39:Coprophanaeus bonariensis 37: 28: 23: 554:10.5962/bhl.title.38459 541:LeConte, J.L. (1861). 367:10.3897/zookeys.88.807 210: 745:Paleobiology Database 206: 639:at Wikimedia Commons 183:, possibly also in 833:Scarabaeinae stubs 408:"Phanaeini Report" 234:D'Olsoufieff, 1924 211: 163:. They are mostly 790: 789: 760: 759: 649:Taxon identifiers 635:Media related to 326: 316:(rainbow scarabs) 315: 305: 295: 285: 275: 265: 255: 245: 235: 225: 142: 141: 138: 840: 811: 804: 797: 769: 753: 752: 740: 739: 727: 726: 714: 713: 701: 700: 691: 690: 689: 676: 675: 674: 644: 634: 619: 617: 615: 609: 594: 584: 582: 580: 568: 556: 537: 508: 507: 505: 503: 488: 482: 481: 457: 448: 447: 445: 443: 429: 423: 422: 420: 418: 404: 398: 397: 387: 369: 349: 325:Olsoufieff, 1924 324: 313: 303: 293: 284:Waterhouse, 1891 283: 273: 263: 253: 243: 233: 223: 208:Phanaeus tridens 136: 54: 53: 33: 21: 16:Tribe of beetles 848: 847: 843: 842: 841: 839: 838: 837: 818: 817: 816: 815: 765: 761: 756: 748: 743: 735: 730: 722: 717: 709: 704: 696: 694: 685: 684: 679: 670: 669: 664: 651: 627: 622: 613: 611: 610:on 14 July 2018 607: 592: 587: 578: 576: 571: 565: 540: 534: 521: 517: 515:Further reading 512: 511: 501: 499: 490: 489: 485: 459: 458: 451: 441: 439: 431: 430: 426: 416: 414: 406: 405: 401: 351: 350: 346: 341: 336: 201: 135: 48: 17: 12: 11: 5: 846: 844: 836: 835: 830: 820: 819: 814: 813: 806: 799: 791: 788: 787: 758: 757: 755: 754: 741: 728: 715: 702: 692: 677: 661: 659: 653: 652: 647: 641: 640: 626: 625:External links 623: 621: 620: 585: 569: 563: 538: 533:978-0849309540 532: 518: 516: 513: 510: 509: 483: 449: 424: 399: 343: 342: 340: 337: 335: 334: 327: 317: 306: 296: 286: 276: 274:Janssens, 1932 266: 256: 246: 236: 226: 215: 200: 197: 173:myrmecophilous 151:in the family 147:is a tribe of 140: 139: 130: 126: 125: 120: 116: 115: 110: 106: 105: 100: 96: 95: 90: 86: 85: 80: 76: 75: 70: 66: 65: 60: 56: 55: 42: 41: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 845: 834: 831: 829: 826: 825: 823: 812: 807: 805: 800: 798: 793: 792: 786: 784: 780: 777:article is a 776: 771: 766: 763: 751: 746: 742: 738: 733: 729: 725: 720: 716: 712: 707: 703: 699: 693: 688: 682: 678: 673: 667: 663: 662: 660: 658: 654: 650: 645: 638: 633: 629: 628: 624: 606: 602: 598: 591: 586: 575: 570: 566: 560: 555: 550: 546: 545: 539: 535: 529: 526:. CRC Press. 525: 520: 519: 514: 498: 494: 487: 484: 479: 475: 471: 467: 463: 456: 454: 450: 438: 434: 428: 425: 413: 409: 403: 400: 395: 391: 386: 381: 377: 373: 368: 363: 360:(88): 1–972. 359: 355: 348: 345: 338: 333: 332: 328: 323: 322: 321:Sulcophanaeus 318: 314:MacLeay, 1819 312: 311: 307: 302: 301: 297: 292: 291: 287: 282: 281: 277: 272: 271: 267: 262: 261: 257: 252: 251: 247: 242: 241: 237: 232: 231: 230:Coprophanaeus 227: 222: 221: 217: 216: 214: 209: 205: 198: 196: 194: 193:Coprophanaeus 190: 186: 182: 178: 174: 170: 166: 162: 158: 154: 150: 146: 134: 131: 128: 127: 124: 121: 118: 117: 114: 111: 108: 107: 104: 101: 98: 97: 94: 91: 88: 87: 84: 81: 78: 77: 74: 71: 68: 67: 64: 61: 58: 57: 52: 47: 43: 40: 36: 32: 27: 22: 19: 828:Scarabaeinae 783:expanding it 775:Scarabaeinae 772: 764: 762: 656: 612:. Retrieved 605:the original 600: 596: 577:. Retrieved 543: 523: 500:. Retrieved 496: 491:Schmidt, U. 486: 472:(1): 43–63. 469: 465: 440:. Retrieved 437:BugGuide.net 436: 427: 415:. Retrieved 411: 402: 357: 347: 329: 319: 308: 298: 288: 278: 270:Homalotarsus 268: 264:Brullé, 1839 258: 254:Gistel, 1857 248: 240:Dendropaemon 238: 228: 224:Harold, 1868 218: 212: 207: 192: 188: 185:Homalotarsus 184: 180: 177:Dendropaemon 176: 169:necrophagous 165:coprophagous 153:Scarabaeidae 149:dung beetles 144: 143: 132: 123:Scarabaeinae 113:Scarabaeidae 38: 18: 681:Wikispecies 294:Bates, 1870 280:Megatharsis 244:Perty, 1830 189:Megatharsis 181:Tetramereia 175:(likely in 119:Subfamily: 822:Categories 564:0665100558 339:References 331:Tetrameira 300:Oxysternon 250:Diabroctis 161:Neotropics 137:Hope, 1838 103:Coleoptera 83:Arthropoda 24:Phanaeini 687:Phanaeini 657:Phanaeini 637:Phanaeini 614:8 January 579:7 January 442:7 January 417:7 January 376:1313-2989 290:Oruscatus 145:Phanaeini 133:Phanaeini 69:Kingdom: 63:Eukaryota 706:BugGuide 695:BioLib: 672:Q4046812 666:Wikidata 394:21594053 310:Phanaeus 260:Gromphas 220:Bolbites 157:Americas 109:Family: 79:Phylum: 73:Animalia 59:Domain: 597:Zootaxa 502:16 July 385:3088472 358:ZooKeys 129:Tribe: 99:Order: 93:Insecta 89:Class: 750:284528 737:926267 711:871349 698:465629 561:  530:  392:  382:  374:  199:Genera 773:This 724:8RYXN 608:(PDF) 593:(PDF) 779:stub 732:ITIS 616:2019 601:2854 581:2019 559:ISBN 528:ISBN 504:2023 444:2019 419:2019 390:PMID 372:ISSN 187:and 179:and 719:CoL 549:doi 474:doi 380:PMC 362:doi 167:or 824:: 747:: 734:: 721:: 708:: 683:: 668:: 599:. 595:. 557:. 495:. 470:35 468:. 464:. 452:^ 435:. 410:. 388:. 378:. 370:. 356:. 810:e 803:t 796:v 785:. 618:. 583:. 567:. 551:: 536:. 506:. 480:. 476:: 446:. 421:. 396:. 364::

Index


Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Scarabaeinae
Phanaeini
dung beetles
Scarabaeidae
Americas
Neotropics
coprophagous
necrophagous
myrmecophilous

Bolbites
Coprophanaeus
Dendropaemon
Diabroctis
Gromphas
Homalotarsus
Megatharsis
Oruscatus
Oxysternon
Phanaeus
Sulcophanaeus

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