389:(CSHL) for 18 months before joining the MRC Human Genetics Unit as a junior group leader. While at CSHL he decided to switch his focus to investigating chromosomal elements in the genetically tractable fission yeast. At the MRC HGU, Edinburgh (1990 - 2002), and subsequently at the Wellome Trust Centre for Cell Biology, University of Edinburgh (2002–present), he discovered that genes are silenced when placed within fission yeast centromeres and telomeres, and then utilised this gene silencing to gain fundamental insights into the processes of chromosome segregation, and heterochromatin and kinetochore CENP-A chromatin establishment and maintenance. He is particularly interested in the epigenetic mechanisms that allow the persistence of specialised chromatin domains through multiple cell divisions and meiosis. He has investigated how
393:(RNAi) mediates heterochromatin formation and shown that splicing factors contribute to heterochromatin integrity via siRNA generation and RNAi. He has provided insight into how transcription and resulting non-coding RNA might influence the assembly of specialised CENP-A chromatin and demonstrated that some acts of lncRNA transcription are responsive to environmental stimuli and regulate neighbouring genes by transcriptional interference. Recently using fission yeast his team discovered an epigenetic mechanism that allows fungi to develop resistance to antifungal drugs without alterations to their DNA. This finding is important for understanding how pathogenic fungi become resistant to the limited number of available antifungal agents in both clinical and agricultural arenas.
449:
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where he discovered that mammalian telomeres are composed of simple repetitive sequences similar to those of unicellular eukaryotes and that telomere length in blood cells shorten with age and are further eroded in cancerous cells. This work resulted from following the fate of fission yeast
626:
Allshire, Robin C; Gosden, John R; Cross, Sally H; Cranston, Gwen; Rout, Derek; Sugawara, Neal; Szostak, Jack W; Fantes, Peter A; Hastie, Nicholas D (1988). "Telomeric repeat from T. Thermophila cross hybridizes with human telomeres".
1189:
Bannister, Andrew J; Zegerman, Philip; Partridge, Janet F; Miska, Eric A; Thomas, Jean O; Allshire, Robin C; Kouzarides, Tony (2001). "Selective recognition of methylated lysine 9 on histone H3 by the HP1 chromo domain".
1431:
Pidoux, Alison L; Choi, Eun Shik; Abbott, Johanna K.R; Liu, Xingkun; Kagansky, Alexander; Castillo, Araceli G; Hamilton, Georgina L; Richardson, William; Rappsilber, Juri; He, Xiangwei; Allshire, Robin C (2009).
1996:
Buscaino, Alessia; White, Sharon A; Houston, Douglas R; Lejeune, Erwan; Simmer, Femke; De Lima Alves, Flavia; Diyora, Piyush T; Urano, Takeshi; Bayne, Elizabeth H; Rappsilber, Juri; Allshire, Robin C (2012).
1946:
Bayne, Elizabeth H; White, Sharon A; Kagansky, Alexander; Bijos, Dominika A; Sanchez-Pulido, Luis; Hoe, Kwang-Lae; Kim, Dong-Uk; Park, Han-Oh; Ponting, Chris P; Rappsilber, Juri; Allshire, Robin C (2010).
678:
Hastie, Nicholas D; Dempster, Maureen; Dunlop, Malcolm G; Thompson, Alastair M; Green, Daryll K; Allshire, Robin C (1990). "Telomere reduction in human colorectal carcinoma and with ageing".
1015:
Pidoux, A. L; Uzawa, S; Perry, P. E; Cande, W. Z; Allshire, R. C (2000). "Live analysis of lagging chromosomes during anaphase and their effect on spindle elongation rate in fission yeast".
342:, in 1981 where he was motivated by the inspirational teaching of David McConnell and colleagues at the Dept of Genetics to undertake post-graduate studies. He subsequently joined the
385:) telomeres after introdroducing fission yeast chromosomes into mouse cell in collaboration with Peter Fantes. In 1989 he took a position as an independent visiting scientist at
2655:
1382:
Dunleavy, Elaine M; Pidoux, Alison L; Monet, Marie; Bonilla, Carolina; Richardson, William; Hamilton, Georgina L; Ekwall, Karl; McLaughlin, Paul J; Allshire, Robin C (2007).
3006:
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Centre for Cell
Biology focuses on the epigenetic mechanisms governing the assembly of specialised domains of chromatin and their transmission through cell division.
1097:
Ekwall, K; Javerzat, J. P; Lorentz, A; Schmidt, H; Cranston, G; Allshire, R (1995). "The chromodomain protein Swi6: A key component at fission yeast centromeres".
729:
Allshire, R. C; Cranston, G; Gosden, J. R; Maule, J. C; Hastie, N. D; Fantes, P. A (1987). "A fission yeast chromosome can replicate autonomously in mouse cells".
3021:
1897:
Simmer, Femke; Buscaino, Alessia; Kos-Braun, Isabelle C; Kagansky, Alexander; Boukaba, Abdelhalim; Urano, Takeshi; Kerr, Alastair R W; Allshire, Robin C (2010).
330:, Co Dublin 1960–1978. His parents were Arthur Gordon Allshire (1925-2012) who was a Pharmacist and Freda Margaret (née Schmutz; 1933–2014). He was awarded his
964:
Nimmo, Elaine R; Pidoux, Alison L; Perry, Paul E; Allshire, Robin C (1998). "Defective meiosis in telomere-silencing mutants of
Schizosaccharomyces pombe".
1046:
Bernard, P; Maure, J. F; Partridge, J. F; Genier, S; Javerzat, J. P; Allshire, R. C (2001). "Requirement of
Heterochromatin for Cohesion at Centromeres".
2207:
Choi, Eun Shik; StrĂĄlfors, Annelie; Catania, Sandra; Castillo, Araceli G; Svensson, J. Peter; Pidoux, Alison L; Ekwall, Karl; Allshire, Robin C (2012).
44:
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1243:"Cis-acting DNA from fission yeast centromeres mediates histone H3 methylation and recruitment of silencing factors and cohesin to an ectopic site"
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2209:"Factors That Promote H3 Chromatin Integrity during Transcription Prevent Promiscuous Deposition of CENP-ACnp1 in Fission Yeast"
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Allshire, R. C; Javerzat, J. P; Redhead, N. J; Cranston, G (1994). "Position effect variegation at fission yeast centromeres".
526:
102:
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Bayne, E. H; Portoso, M; Kagansky, A; Kos-Braun, I. C; Urano, T; Ekwall, K; Alves, F; Rappsilber, J; Allshire, R. C (2008).
1636:
Kagansky, A; Folco, H. D; Almeida, R; Pidoux, A. L; Boukaba, A; Simmer, F; Urano, T; Hamilton, G. L; Allshire, R. C (2009).
2105:
Bayne, Elizabeth H; Bijos, Dominika A; White, Sharon A; Alves, Flavia de Lima; Rappsilber, Juri; Allshire, Robin C (2014).
74:
386:
235:
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Choi, Eun Shik; Strålfors, Annelie; Castillo, Araceli G; Durand-Dubief, Mickaël; Ekwall, Karl; Allshire, Robin C (2011).
1150:"Transient inhibition of histone deacetylation alters the structural and functional imprint at fission yeast centromeres"
374:'s research group at the MRC Human Genetics Unit, Edinburgh (formerly MRC Clinical and Population Cytogentics Unit) as a
2107:"A systematic genetic screen identifies new factors influencing centromeric heterochromatin integrity in fission yeast"
2647:
2360:"Long non-coding RNA-mediated transcriptional interference of a permease gene confers drug tolerance in fission yeast"
1999:"Raf1 is a DCAF for the Rik1 DDB1-Like Protein and Has Separable Roles in siRNA Generation and Chromatin Modification"
406:
142:
81:
774:"Introduction of large linear minichromosomes into Schizosaccharomyces pombe by an improved transformation procedure"
1791:
Buscaino, Alessia; Lejeune, Erwan; Audergon, Pauline; Hamilton, Georgina; Pidoux, Alison; Allshire, Robin C (2013).
1384:"A NASP (N1/N2)-Related Protein, Sim3, Binds CENP-A and is Required for Its Deposition at Fission Yeast Centromeres"
1331:
Castillo, A. G; Mellone, B. G; Partridge, J. F; Richardson, W; Hamilton, G. L; Allshire, R. C; Pidoux, A. L (2007).
2920:
40:
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2260:"Sequence Features and Transcriptional Stalling within Centromere DNA Promote Establishment of CENP-A Chromatin"
88:
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917:"Telomere-associated chromosome breakage in fission yeast results in variegated expression of adjacent genes"
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Audergon, P. N. C. B; Catania, S; Kagansky, A; Tong, P; Shukla, M; Pidoux, A. L; Allshire, R. C (2015).
594:
359:
1899:"Hairpin RNA induces secondary small interfering RNA synthesis and silencing in trans in fission yeast"
1793:"Distinct roles for Sir2 and RNAi in centromeric heterochromatin nucleation, spreading and maintenance"
1949:"Stc1: A Critical Link between RNAi and Chromatin Modification Required for Heterochromatin Integrity"
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1638:"Synthetic Heterochromatin Bypasses RNAi and Centromeric Repeats to Establish Functional Centromeres"
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2417:"Transcription-coupled changes to chromatin underpin gene silencing by transcriptional interference"
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2158:"Identification of Noncoding Transcripts from within CENP-A Chromatin at Fission Yeast Centromeres"
876:"Mutations derepressing silent centromeric domains in fission yeast disrupt chromosome segregation"
2689:
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One or more of the preceding sentences incorporates text from the royalsociety.org website where:
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1744:"The JmjC domain protein Epe1 prevents unregulated assembly and disassembly of heterochromatin"
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1333:"Plasticity of fission yeast CENP-A chromatin driven by relative levels of histone H3 and H4"
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Trewick, Sarah C; Minc, Elsa; Antonelli, Richard; Urano, Takeshi; Allshire, Robin C (2007).
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Partridge, J. F; Scott, K. S; Bannister, A. J; Kouzarides, T; Allshire, R. C (2002).
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1695:"Distinct protein interaction domains and protein spreading in a complex centromere"
1581:"Heterochromatin and RNAi Are Required to Establish CENP-A Chromatin at Centromeres"
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Sanchez-Pulido, Luis; Pidoux, Alison L; Ponting, Chris P; Allshire, Robin C (2009).
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54:. Contentious material about living persons that is unsourced or poorly sourced
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Proceedings of the
National Academy of Sciences of the United States of America
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1532:"Eic1 links Mis18 with the CCAN/Mis6/Ctf19 complex to promote CENP-A assembly"
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Allshire, R. C; Nimmo, E. R; Ekwall, K; Javerzat, J. P; Cranston, G (1995).
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595:"ALLSHIRE, Freda Margaret : Death notice - Irish Times Family Notices"
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Ekwall, K; Olsson, T; Turner, B. M; Cranston, G; Allshire, R. C (1997).
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2466:"Epigenetic gene silencing by heterochromatin primes fungal resistance"
2383:
2050:"Splicing Factors Facilitate RNAi-Directed Silencing in Fission Yeast"
475:
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Subramanian, L; Toda, N. R. T; Rappsilber, J; Allshire, R. C (2014).
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549:"Wellcome Centre for Cell Biology - Wellcome Centre for Cell Biology"
242:
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Construction and analysis of vectors based on bovine papilloma virus
249:
Construction and analysis of vectors based on bovine papilloma virus
612:
577:"ALLSHIRE, Arthur : Death notice - Irish Times Family Notices"
248:
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2581:"Professor Robin Allshire | the Academy of Medical Sciences"
1282:
Pidoux, Alison L; Richardson, William; Allshire, Robin C (2003).
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as a chassis for mammalian artificial chromosome construction.
351:
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Catania, Sandra; Pidoux, Alison L; Allshire, Robin C (2015).
306:(born 19 May 1960) is a British academic who is Professor of
1579:
Folco, H. D; Pidoux, A. L; Urano, T; Allshire, R. C (2008).
548:
1483:"Common Ancestry of the CENP-A Chaperones Scm3 and HJURP"
292:
1842:"Restricted epigenetic inheritance of H3K9 methylation"
2558:
Creative
Commons Attribution 4.0 International License
1693:
Partridge, J. F; Borgstrøm, B; Allshire, R. C (2000).
527:"Principal Research Fellowships: people we've funded"
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915:Nimmo, E. R; Cranston, G; Allshire, R. C (1994).
2358:Ard, Ryan; Tong, Pin; Allshire, Robin C (2014).
354:in 1985 under the guidance of Chris Bostock and
2554:
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8:
2562:Royal Society Terms, conditions and policies
326:Allshire grew up in the fishing village of
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2309:Shukla M, Manu; Allshire, Robin C (2018).
447:
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3007:Fellows of the Royal Society of Edinburgh
3002:Wellcome Trust Principal Research Fellows
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411:Fellow of the Academy of Medical Sciences
310:Biology at University of Edinburgh and a
122:Learn how and when to remove this message
461:
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403:Fellow of the Royal Society of Edinburgh
312:Wellcome Trust Principal Research Fellow
3012:Fellows of the Royal Society of Biology
613:"EPIGENETICS AND SPECIALIZED CHROMATIN"
470:(PhD thesis). University of Edinburgh.
430:
3022:Alumni of the University of Edinburgh
2415:Ard, Ryan; Allshire, Robin C (2016).
7:
436:
434:
346:(MRC) Mammalian Genome Unit at the
933:10.1002/j.1460-2075.1994.tb06691.x
14:
466:Allshire, Robin Campbell (1985).
3027:Alumni of Trinity College Dublin
2965: This article incorporates
2960:
497:
25:
2992:21st-century British biologists
2987:20th-century British biologists
2162:Journal of Biological Chemistry
2545:"Professor Robin Allshire FRS"
1:
2599:"Genetics Society Medal 2013"
1260:10.1016/S0960-9822(02)01177-6
1167:10.1016/S0092-8674(00)80492-4
387:Cold Spring Harbor Laboratory
236:Cold Spring Harbor Laboratory
52:secondary or tertiary sources
2464:Torres-Garcia, Sito (2020).
2277:10.1371/journal.pgen.1004986
2226:10.1371/journal.pgen.1002985
2016:10.1371/journal.pgen.1002499
1450:10.1016/j.molcel.2009.01.019
1400:10.1016/j.molcel.2007.10.010
1350:10.1371/journal.pgen.0030121
845:10.1016/0092-8674(94)90180-5
743:10.1016/0092-8674(87)90493-4
314:. His research group at the
274:Other academic advisors
58:, especially if potentially
36:biography of a living person
2585:Academy of Medical Sciences
1288:The Journal of Cell Biology
1019:. 113 Pt 23 (23): 4177–91.
407:Fellow of the Royal Society
174:1960 (age 63–64)
56:must be removed immediately
3043:
2568: (archived 2016-11-11)
1965:10.1016/j.cell.2010.01.038
1499:10.1016/j.cell.2009.06.010
551:. University of Edinburgh.
2490:10.1038/s41586-020-2706-x
2327:10.1016/j.cub.2018.10.049
2124:10.1186/s13059-014-0481-4
382:Schizosaccharomyces pombe
358:investigating the use of
283:
194:
154:
1760:10.1038/sj.emboj.7601892
443:publications indexed by
370:In 1985 Allshire joined
344:Medical Research Council
322:Early life and education
2175:10.1074/jbc.M111.228510
2074:10.1126/science.1164029
1866:10.1126/science.1260638
1699:Genes & Development
1662:10.1126/science.1172026
1605:10.1126/science.1150944
1119:10.1126/science.7660126
1068:10.1126/science.1064027
1025:10.1242/jcs.113.23.4177
1017:Journal of Cell Science
880:Genes & Development
799:10.1073/pnas.87.11.4043
772:Allshire, R. C (1990).
401:Allshire was elected a
376:postdoctoral researcher
348:University of Edinburgh
304:Robin Campbell Allshire
232:University of Edinburgh
188:University of Edinburgh
171:Robin Campbell Allshire
2570:
2421:Nucleic Acids Research
1915:10.1038/embor.2009.273
513:"Allshire lab members"
419:Genetics Society Medal
350:where he obtained his
340:Trinity College Dublin
183:Trinity College Dublin
161:Robin Allshire in 2019
50:Please help by adding
2364:Nature Communications
1809:10.1038/emboj.2013.72
1300:10.1083/jcb.200212110
409:(FRS) in 2011 and a
360:bovine papillomavirus
2969:available under the
2841:Konstantin Novoselov
1711:10.1101/gad.14.7.783
563:"Wellcome Trust lab"
2931:Phillip Allen Sharp
2482:2020Natur.585..453T
2427:(22): 10619–10630.
2376:2014NatCo...5.5576A
2066:2008Sci...322..602B
1858:2015Sci...348..132A
1654:2009Sci...324.1716K
1597:2008Sci...319...94F
1548:10.1098/rsob.140043
1204:2001Natur.410..120B
1111:1995Sci...269.1429E
1060:2001Sci...294.2539B
978:1998Natur.392..825N
893:10.1101/gad.9.2.218
790:1990PNAS...87.4043A
692:1990Natur.346..866H
641:1988Natur.332..656A
413:(FMedSci) in 2020.
366:Career and research
39:relies too much on
2787:David Manolopoulos
2601:. Genetics Society
2433:10.1093/nar/gkw801
2384:10.1038/ncomms6576
565:. 11 January 2024.
492:uk.bl.ethos.355979
397:Awards and honours
2956:
2955:
2770:Steffen Lauritzen
2476:(7825): 453–458.
2321:(24): 3924–3936.
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301:
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196:Scientific career
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2941:Thomas A. Steitz
2792:Gerhard Materlik
2765:Robert Kennicutt
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1262:
1238:
1232:
1231:
1212:10.1038/35065138
1186:
1180:
1179:
1169:
1145:
1139:
1138:
1094:
1088:
1087:
1043:
1037:
1036:
1012:
1006:
1005:
961:
955:
954:
944:
921:The EMBO Journal
912:
906:
905:
895:
871:
865:
864:
828:
822:
821:
811:
801:
769:
763:
762:
726:
720:
719:
700:10.1038/346866a0
675:
669:
668:
649:10.1038/332656a0
623:
617:
616:
609:
603:
602:
591:
585:
584:
573:
567:
566:
559:
553:
552:
545:
539:
538:
523:
517:
516:
509:
503:
502:
501:
495:
463:
452:
451:
438:
391:RNA interference
332:Bachelor of Arts
297:
294:
261:Doctoral advisor
255:
159:
149:
134:
127:
120:
116:
113:
107:
105:
71:"Robin Allshire"
64:
29:
21:
16:British academic
3042:
3041:
3037:
3036:
3035:
3033:
3032:
3031:
2977:
2976:
2957:
2952:
2899:
2755:Colin Humphreys
2745:Janet Hemingway
2715:Nicholas Franks
2695:Doreen Cantrell
2680:Jeremy Baumberg
2675:Andrew Balmford
2658:
2645:
2615:
2614:
2604:
2602:
2597:
2596:
2592:
2579:
2578:
2574:
2566:Wayback Machine
2542:
2541:
2537:
2527:
2525:
2520:
2519:
2515:
2463:
2462:
2458:
2414:
2413:
2409:
2357:
2356:
2352:
2315:Current Biology
2308:
2307:
2303:
2270:(3): e1004986.
2257:
2256:
2252:
2219:(9): e1002985.
2206:
2205:
2201:
2168:(26): 23600–7.
2155:
2154:
2150:
2104:
2103:
2099:
2060:(5901): 602–6.
2047:
2046:
2042:
2009:(2): e1002499.
1995:
1994:
1990:
1945:
1944:
1940:
1896:
1895:
1891:
1852:(6230): 132–5.
1839:
1838:
1834:
1790:
1789:
1785:
1754:(22): 4670–82.
1741:
1740:
1736:
1692:
1691:
1687:
1635:
1634:
1630:
1578:
1577:
1573:
1529:
1528:
1524:
1480:
1479:
1475:
1430:
1429:
1425:
1381:
1380:
1376:
1330:
1329:
1325:
1281:
1280:
1276:
1253:(19): 1652–60.
1247:Current Biology
1240:
1239:
1235:
1198:(6824): 120–4.
1188:
1187:
1183:
1147:
1146:
1142:
1096:
1095:
1091:
1045:
1044:
1040:
1014:
1013:
1009:
972:(6678): 825–8.
963:
962:
958:
927:(16): 3801–11.
914:
913:
909:
873:
872:
868:
830:
829:
825:
771:
770:
766:
728:
727:
723:
686:(6287): 866–8.
677:
676:
672:
635:(6165): 656–9.
625:
624:
620:
611:
610:
606:
599:The Irish Times
593:
592:
588:
581:The Irish Times
575:
574:
570:
561:
560:
556:
547:
546:
542:
525:
524:
520:
511:
510:
506:
496:
465:
464:
455:
439:
432:
427:
399:
372:Nicholas Hastie
368:
324:
291:
278:Nicholas Hastie
266:
253:
234:
220:
216:
212:
210:Heterochromatin
208:
186:
179:Alma mater
173:
172:
162:
150:
141:
139:
128:
117:
111:
108:
65:
63:
49:
45:primary sources
30:
17:
12:
11:
5:
3040:
3038:
3030:
3029:
3024:
3019:
3014:
3009:
3004:
2999:
2994:
2989:
2979:
2978:
2954:
2953:
2951:
2950:
2947:Edward Stolper
2943:
2938:
2936:Carla J. Shatz
2933:
2928:
2923:
2921:Mikhail Gromov
2918:
2913:
2911:David Chandler
2907:
2905:
2901:
2900:
2898:
2897:
2892:
2887:
2885:Angela Vincent
2882:
2877:
2872:
2867:
2860:
2855:
2848:
2843:
2838:
2833:
2828:
2823:
2816:
2809:
2807:David McMurtry
2804:
2799:
2797:James McKernan
2794:
2789:
2784:
2777:
2772:
2767:
2762:
2757:
2752:
2747:
2742:
2737:
2732:
2727:
2725:Steven Gamblin
2722:
2717:
2712:
2707:
2702:
2697:
2692:
2687:
2682:
2677:
2672:
2670:Robin Allshire
2666:
2664:
2660:
2659:
2646:
2644:
2643:
2636:
2629:
2621:
2613:
2612:
2590:
2572:
2535:
2513:
2456:
2407:
2350:
2301:
2250:
2199:
2148:
2111:Genome Biology
2097:
2040:
1988:
1938:
1889:
1832:
1803:(9): 1250–64.
1783:
1734:
1685:
1628:
1591:(5859): 94–7.
1571:
1522:
1473:
1444:(3): 299–311.
1438:Molecular Cell
1423:
1394:(6): 1029–44.
1388:Molecular Cell
1374:
1323:
1294:(2): 295–307.
1274:
1233:
1181:
1160:(7): 1021–32.
1140:
1089:
1038:
1007:
956:
907:
866:
823:
784:(11): 4043–7.
764:
737:(3): 391–403.
721:
670:
618:
604:
586:
568:
554:
540:
535:Wellcome Trust
531:wellcome.ac.uk
518:
504:
453:
445:Google Scholar
441:Robin Allshire
429:
428:
426:
423:
422:
421:
398:
395:
367:
364:
356:Edwin Southern
323:
320:
316:Wellcome Trust
299:
298:
289:
285:
284:
281:
280:
275:
271:
270:
268:Edwin Southern
263:
257:
256:
245:
239:
238:
229:
225:
224:
203:
199:
198:
192:
191:
180:
176:
175:
170:
168:
164:
163:
160:
152:
151:
140:
138:Robin Allshire
137:
130:
129:
33:
31:
24:
15:
13:
10:
9:
6:
4:
3:
2:
3039:
3028:
3025:
3023:
3020:
3018:
3017:Living people
3015:
3013:
3010:
3008:
3005:
3003:
3000:
2998:
2995:
2993:
2990:
2988:
2985:
2984:
2982:
2975:
2974:
2972:
2968:
2963:
2949:
2948:
2944:
2942:
2939:
2937:
2934:
2932:
2929:
2927:
2926:Thomas Hughes
2924:
2922:
2919:
2917:
2914:
2912:
2909:
2908:
2906:
2902:
2896:
2895:Robert Watson
2893:
2891:
2888:
2886:
2883:
2881:
2878:
2876:
2873:
2871:
2868:
2866:
2865:
2864:Mark Randolph
2861:
2859:
2856:
2854:
2853:
2849:
2847:
2844:
2842:
2839:
2837:
2834:
2832:
2829:
2827:
2824:
2822:
2821:
2820:Arthur Milner
2817:
2815:
2814:
2810:
2808:
2805:
2803:
2800:
2798:
2795:
2793:
2790:
2788:
2785:
2783:
2782:
2778:
2776:
2773:
2771:
2768:
2766:
2763:
2761:
2760:Alex Kacelnik
2758:
2756:
2753:
2751:
2748:
2746:
2743:
2741:
2738:
2736:
2733:
2731:
2728:
2726:
2723:
2721:
2718:
2716:
2713:
2711:
2708:
2706:
2703:
2701:
2698:
2696:
2693:
2691:
2690:Béla Bollobás
2688:
2686:
2683:
2681:
2678:
2676:
2673:
2671:
2668:
2667:
2665:
2661:
2657:
2653:
2652:Royal Society
2649:
2642:
2637:
2635:
2630:
2628:
2623:
2622:
2619:
2600:
2594:
2591:
2586:
2582:
2576:
2573:
2569:
2567:
2563:
2559:
2550:
2546:
2543:Anon (2011).
2539:
2536:
2523:
2517:
2514:
2509:
2505:
2500:
2495:
2491:
2487:
2483:
2479:
2475:
2471:
2467:
2460:
2457:
2452:
2448:
2443:
2438:
2434:
2430:
2426:
2422:
2418:
2411:
2408:
2403:
2399:
2394:
2389:
2385:
2381:
2377:
2373:
2369:
2365:
2361:
2354:
2351:
2346:
2342:
2337:
2332:
2328:
2324:
2320:
2316:
2312:
2305:
2302:
2297:
2293:
2288:
2283:
2278:
2273:
2269:
2265:
2264:PLOS Genetics
2261:
2254:
2251:
2246:
2242:
2237:
2232:
2227:
2222:
2218:
2214:
2213:PLOS Genetics
2210:
2203:
2200:
2195:
2191:
2186:
2181:
2176:
2171:
2167:
2163:
2159:
2152:
2149:
2144:
2140:
2135:
2130:
2125:
2120:
2116:
2112:
2108:
2101:
2098:
2093:
2089:
2084:
2079:
2075:
2071:
2067:
2063:
2059:
2055:
2051:
2044:
2041:
2036:
2032:
2027:
2022:
2017:
2012:
2008:
2004:
2003:PLOS Genetics
2000:
1992:
1989:
1984:
1980:
1975:
1970:
1966:
1962:
1959:(5): 666–77.
1958:
1954:
1950:
1942:
1939:
1934:
1930:
1925:
1920:
1916:
1912:
1908:
1904:
1900:
1893:
1890:
1885:
1881:
1876:
1871:
1867:
1863:
1859:
1855:
1851:
1847:
1843:
1836:
1833:
1828:
1824:
1819:
1814:
1810:
1806:
1802:
1798:
1794:
1787:
1784:
1779:
1775:
1770:
1765:
1761:
1757:
1753:
1749:
1745:
1738:
1735:
1730:
1726:
1721:
1716:
1712:
1708:
1705:(7): 783–91.
1704:
1700:
1696:
1689:
1686:
1681:
1677:
1672:
1667:
1663:
1659:
1655:
1651:
1647:
1643:
1639:
1632:
1629:
1624:
1620:
1615:
1610:
1606:
1602:
1598:
1594:
1590:
1586:
1582:
1575:
1572:
1567:
1563:
1558:
1553:
1549:
1545:
1542:(4): 140043.
1541:
1537:
1533:
1526:
1523:
1518:
1514:
1509:
1504:
1500:
1496:
1493:(7): 1173–4.
1492:
1488:
1484:
1477:
1474:
1469:
1465:
1460:
1455:
1451:
1447:
1443:
1439:
1435:
1427:
1424:
1419:
1415:
1410:
1405:
1401:
1397:
1393:
1389:
1385:
1378:
1375:
1370:
1366:
1361:
1356:
1351:
1346:
1342:
1338:
1337:PLOS Genetics
1334:
1327:
1324:
1319:
1315:
1310:
1305:
1301:
1297:
1293:
1289:
1285:
1278:
1275:
1270:
1266:
1261:
1256:
1252:
1248:
1244:
1237:
1234:
1229:
1225:
1221:
1217:
1213:
1209:
1205:
1201:
1197:
1193:
1185:
1182:
1177:
1173:
1168:
1163:
1159:
1155:
1151:
1144:
1141:
1136:
1132:
1128:
1124:
1120:
1116:
1112:
1108:
1104:
1100:
1093:
1090:
1085:
1081:
1077:
1073:
1069:
1065:
1061:
1057:
1053:
1049:
1042:
1039:
1034:
1030:
1026:
1022:
1018:
1011:
1008:
1003:
999:
995:
991:
987:
986:10.1038/33941
983:
979:
975:
971:
967:
960:
957:
952:
948:
943:
938:
934:
930:
926:
922:
918:
911:
908:
903:
899:
894:
889:
886:(2): 218–33.
885:
881:
877:
870:
867:
862:
858:
854:
850:
846:
842:
839:(1): 157–69.
838:
834:
827:
824:
819:
815:
810:
805:
800:
795:
791:
787:
783:
779:
775:
768:
765:
760:
756:
752:
748:
744:
740:
736:
732:
725:
722:
717:
713:
709:
705:
701:
697:
693:
689:
685:
681:
674:
671:
666:
662:
658:
654:
650:
646:
642:
638:
634:
630:
622:
619:
614:
608:
605:
600:
596:
590:
587:
582:
578:
572:
569:
564:
558:
555:
550:
544:
541:
536:
532:
528:
522:
519:
514:
508:
505:
500:
493:
489:
485:
481:
477:
473:
469:
462:
460:
458:
454:
450:
446:
442:
437:
435:
431:
424:
420:
416:
415:
414:
412:
408:
404:
396:
394:
392:
388:
384:
383:
377:
373:
365:
363:
361:
357:
353:
349:
345:
341:
337:
333:
329:
321:
319:
317:
313:
309:
305:
296:
290:
286:
282:
279:
276:
272:
269:
265:Chris Bostock
264:
262:
258:
251:
250:
246:
244:
240:
237:
233:
230:
226:
223:
219:
215:
211:
207:
204:
200:
197:
193:
189:
184:
181:
177:
169:
165:
158:
153:
148:
144:
135:
126:
123:
115:
104:
101:
97:
94:
90:
87:
83:
80:
76:
73: –
72:
68:
67:Find sources:
61:
57:
53:
47:
46:
42:
37:
32:
28:
23:
22:
19:
2959:
2958:
2945:
2916:Joanne Chory
2890:Mark Walport
2880:Simon Tavaré
2870:Len Stephens
2862:
2858:Fiona Powrie
2850:
2818:
2813:Mervyn Miles
2811:
2779:
2750:Ian Horrocks
2685:Hagan Bayley
2669:
2603:. Retrieved
2593:
2584:
2575:
2555:
2548:
2538:
2526:. Retrieved
2516:
2473:
2469:
2459:
2424:
2420:
2410:
2367:
2363:
2353:
2318:
2314:
2304:
2267:
2263:
2253:
2216:
2212:
2202:
2165:
2161:
2151:
2114:
2110:
2100:
2057:
2053:
2043:
2006:
2002:
1991:
1956:
1952:
1941:
1909:(2): 112–8.
1906:
1903:EMBO Reports
1902:
1892:
1849:
1845:
1835:
1800:
1796:
1786:
1751:
1747:
1737:
1702:
1698:
1688:
1645:
1641:
1631:
1588:
1584:
1574:
1539:
1536:Open Biology
1535:
1525:
1490:
1486:
1476:
1441:
1437:
1426:
1391:
1387:
1377:
1340:
1336:
1326:
1291:
1287:
1277:
1250:
1246:
1236:
1195:
1191:
1184:
1157:
1153:
1143:
1102:
1098:
1092:
1051:
1047:
1041:
1016:
1010:
969:
965:
959:
924:
920:
910:
883:
879:
869:
836:
832:
826:
781:
777:
767:
734:
730:
724:
683:
679:
673:
632:
628:
621:
607:
598:
589:
580:
571:
557:
543:
530:
521:
507:
467:
400:
380:
369:
325:
303:
302:
247:
228:Institutions
195:
118:
109:
99:
92:
85:
78:
66:
55:
38:
18:
2997:1960 births
2875:Patrick Tam
2852:John Parkes
2836:Werner Nahm
2826:John Morton
2802:Tom McLeish
2781:Ian Manners
2735:Alan Grafen
2730:John Goodby
2710:Alun Davies
2705:Alan Cowman
2700:Stan Cowley
2522:"biography"
2117:(10): 481.
1343:(7): e121.
405:in 2005, a
293:allshirelab
222:Kinetochore
206:Epigenetics
62:or harmful.
2981:Categories
2846:Mark Pagel
2831:Sean Munro
2740:Clare Grey
2720:Ian Frazer
2605:4 December
2528:4 December
476:1842/11176
425:References
334:degree in
308:Chromosome
218:Centromere
82:newspapers
41:references
2971:CC BY 4.0
2775:Dennis Lo
484:606010479
214:Chromatin
112:July 2018
2973:license.
2654:elected
2508:32908306
2451:27613421
2402:25428589
2370:: 5576.
2345:30503616
2296:25738810
2245:23028377
2194:21531710
2143:25274039
2092:18948543
2035:22319459
1983:20211136
1933:20062003
1884:25838386
1827:23572080
1778:17948055
1729:10766735
1680:19556509
1623:18174443
1566:24789708
1517:19563746
1468:19217404
1418:18158900
1369:17677001
1318:12719471
1269:12361567
1220:11242054
1135:38678389
1084:31166180
1076:11598266
1033:11069763
861:42369473
336:Genetics
60:libelous
2904:Foreign
2663:Fellows
2656:in 2011
2650:of the
2648:Fellows
2564:at the
2499:7116710
2478:Bibcode
2442:5159543
2393:4255232
2372:Bibcode
2336:6303189
2287:4349457
2236:3447972
2185:3123123
2134:4210515
2083:2585287
2062:Bibcode
2054:Science
2026:3271066
1974:2875855
1924:2828748
1875:4397586
1854:Bibcode
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