268:
33:
55:
255:
789:
Cupello, M.; Vaz-de-Mello, F.Z. (2014). "Revalidation of the
Brazilian Atlantic Forest dung beetle species Coprophanaeus (Metallophanaeus) machadoi (Pereira & d'Andretta, 1955) (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) based on morphological and distributional evidence".
823:
Ururahy-Rodrigues, A.; Rafael, J.A.; Wanderley, R.F.; Marques, H.; Pujol-Luz, J.R. (2008). "Coprophanaeus lancifer (Linnaeus, 1767) (Coleoptera, Scarabaeidae) activity moves a man-size pig carcass: Relevant data for forensic taphonomy".
242:. They are paracoprids, meaning that adults dig tunnels into the soil under the food source and move parts of the food source to a nest chamber where the eggs are laid. Because of their feeding on carrion, they are of interest to
734:
Maldaner, M.E.; Vaz-de-Mello, F.Z.; Takiya, D.M.; Ferreira, D.C. (2008). "Molecular
Phylogeny of Coprophanaeus (Megaphanaeus) d'Olsoufieff, 1924 (Coleoptera: Scarabaeidae: Scarabaeinae) and the position of C. bellicosus".
851:
Cantil, L.F.; Sánche, M.V.; Sarzetti, L.; Molina, A.; Genise, J.F. (2015). "Nest and brood balls of
Coprophanaeus (Coprophanaeus) cyanescens (Olsoufiefff, 1924) (Coleoptera: Scarabaeidae: Scarabaeinae)".
215:) and the largest in the Americas. They often have a horn on the head, and are typically a bright metallic color, most often blue or green (other colors also occur, for example red), or black. These
762:
Maldaner, M.E.; Vaz-de-Mello, F.Z.; Takiya, D.M.; Ferreira, D.C. (2019). "Genetic and chromatic variation of
Coprophanaeus (Megaphanaeus) ensifer (Germar, 1821) (Coleoptera: Scarabaeidae)".
1037:
1063:
683:
Edmonds, W.D.; Zidek, J. (2010). "A taxonomic review of the
Neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)".
267:
1011:
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32:
658:
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199:
sometimes exceeding 5 cm (2 in) in length, making these two some of the largest
187:
in the United States. They are medium-sized to large beetles, with the South
American
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170:
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277:, a common and variably colored species where only the male has a distinct horn
803:
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1016:
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1029:
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263:, a large species where both sexes have a large horn on the head
900:
1003:
296:, but it doubtful that the first and last of these are
818:
816:
814:
909:
226:Although part of the true dung beetle subfamily
223:beetles are excellent diggers and good fliers.
173:(scarab beetles). The genus is almost entirely
729:
727:
678:
676:
674:
672:
670:
8:
897:
303:The genus includes the following species:
31:
20:
873:
626:
7:
203:in the world (together with certain
737:Insect Systematics & Evolution
14:
284:is divided into three subgenera:
53:
838:10.1016/j.forsciint.2008.09.009
826:Forensic Science International
645:Nomen.at - animals and plants
1:
381:Coprophanaeus christophorowi
238:, but many species may also
183:, ranging into southernmost
1121:
866:10.1649/0010-065X-69.1.153
854:The Coleopterists Bulletin
448:Kohlmann & Solís, 2012
389:Coprophanaeus conocephalus
373:Coprophanaeus chiriquensis
804:10.11646/zootaxa.3869.4.7
776:10.1016/j.jcz.2019.09.001
749:10.1163/1876312X-00002198
613:Coprophanaeus thalassinus
469:Coprophanaeus ignecinctus
333:Coprophanaeus bonariensis
177:, with a single species,
50:Scientific classification
48:
39:
30:
23:
573:Coprophanaeus saphirinus
536:Howden & Young, 1981
421:Coprophanaeus degallieri
405:Coprophanaeus cyanescens
349:Coprophanaeus callegarii
325:Coprophanaeus bellicosus
661:March 27, 2015, at the
565:Coprophanaeus punctatus
525:Coprophanaeus parvulus
485:Coprophanaeus lancifer
413:Coprophanaeus dardanus
397:Coprophanaeus corythus
365:Coprophanaeus cerberus
357:Coprophanaeus caroliae
341:Coprophanaeus boucardi
317:Coprophanaeus acrisius
278:
264:
42:Coprophanaeus lancifer
764:Zoologischer Anzeiger
605:Coprophanaeus telamon
597:Coprophanaeus suredai
549:Coprophanaeus pessoai
509:Coprophanaeus morenoi
445:Coprophanaeus gephyra
429:Coprophanaeus ensifer
270:
257:
707:species Information"
589:Coprophanaeus spitzi
581:Coprophanaeus solisi
541:Coprophanaeus pertyi
517:Coprophanaeus ohausi
493:Coprophanaeus magnoi
477:Coprophanaeus jasius
437:Coprophanaeus gamezi
1105:Scarabaeidae genera
705:Coprophanaeus pluto
557:Coprophanaeus pluto
533:Coprophanaeus pecki
501:Coprophanaeus milon
461:Coprophanaeus horus
453:Coprophanaeus gilli
244:forensic entomology
464:(Waterhouse, 1891)
309:Coprophanaeus abas
279:
265:
230:, adult and young
153:D'Olsoufieff, 1924
1092:
1091:
1077:Open Tree of Life
903:Taxon identifiers
568:(Olsufieff, 1924)
544:(Olsufieff, 1924)
528:(Olsufieff, 1924)
504:(Blanchard, 1846)
408:(Olsufieff, 1924)
392:(Olsufieff, 1924)
384:(Olsufieff, 1924)
376:(Olsufieff, 1924)
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832:(1–3): e19–e22.
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608:(Erichson, 1847)
488:(Linnaeus, 1767)
344:(Nevinson, 1891)
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16:Genus of beetles
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663:Wayback Machine
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552:(Pereira, 1949)
520:(Felsche, 1911)
480:(Olivier, 1789)
472:(Felsche, 1909)
416:(MacLeay, 1819)
328:(Olivier, 1789)
320:(MacLeay, 1819)
312:(MacLeay, 1819)
294:Metallophanaeus
252:
236:feed on carrion
151:
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860:(1): 153–158.
843:
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798:(4): 435.451.
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743:(5): 241–255.
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169:in the family
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941:Coprophanaeus
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286:Coprophanaeus
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282:Coprophanaeus
276:
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274:C. saphirinus
269:
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241:
240:feed on feces
237:
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232:Coprophanaeus
229:
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162:Coprophanaeus
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25:Coprophanaeus
22:
19:
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736:
714:. Retrieved
711:BugGuide.net
710:
704:
697:
688:
684:
651:
640:
629:
611:
603:
600:Arnaud, 1996
595:
587:
584:Arnaud, 1997
579:
571:
563:
555:
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531:
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512:Arnaud, 1982
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496:Arnaud, 2002
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456:Arnaud, 1997
451:
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419:
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371:
363:
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352:Arnaud, 2002
347:
339:
336:(Gory, 1844)
331:
323:
315:
307:
302:
298:monophyletic
293:
290:Megaphanaeus
289:
285:
281:
280:
272:
258:
231:
228:Scarabaeinae
225:
210:
204:
201:dung beetles
194:
188:
178:
171:Scarabaeidae
161:
160:
159:
147:
146:
127:Scarabaeinae
117:Scarabaeidae
40:
24:
18:
1025:iNaturalist
935:Wikispecies
875:11336/50474
770:: 150–160.
221:crepuscular
212:Heliocopris
196:C. lancifer
175:Neotropical
123:Subfamily:
621:References
260:C. ensifer
234:primarily
206:Catharsius
190:C. ensifer
107:Coleoptera
87:Arthropoda
137:Phanaeini
73:Kingdom:
67:Eukaryota
1099:Category
973:BugGuide
949:BioLib:
926:Q5168943
920:Wikidata
691:: 1–111.
659:Archived
180:C. pluto
113:Family:
83:Phylum:
77:Animalia
63:Domain:
1082:1008443
1043:1112536
1017:1086344
792:Zootaxa
716:29 June
271:Female
250:Species
217:diurnal
143:Genus:
133:Tribe:
103:Order:
97:Insecta
93:Class:
1069:206694
1056:926428
1030:210587
978:344893
965:168036
952:465631
889:Biolib
1038:IRMNG
1004:17093
185:Texas
167:genus
165:is a
1064:NCBI
1051:ITIS
1012:GBIF
991:3TRJ
960:BOLD
796:3869
718:2023
689:0129
656:GBIF
634:ITIS
292:and
209:and
193:and
999:EoL
986:CoL
870:hdl
862:doi
834:doi
830:182
800:doi
772:doi
768:283
745:doi
219:or
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858:69
856:.
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813:^
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246:.
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836::
808:|
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703:"
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