3413:
61:
2971:
2954:
38:
2815:
1747:
298:
3202:
incomplete hemocyanin as trace component. The combined data show that B. glabrata hemoglobin evolved from pulmonate myoglobin, possibly to replace a less-efficient hemocyanin, and reveals a surprisingly simple evolutionary mechanism to create a high molecular mass respiratory protein from 78 similar globin domains.
2908:
in their shell coiling, as is proved by their internal anatomy (the respiratory and the genital orifice are situated on the left side), however the animals carry their shells with what would normally be the ventral (i.e. umbilical) surface uppermost, and because of this, the shells appear to be
3201:
We present here the complete cDNA and predicted amino acid sequence of two hemoglobin polypeptides from the planorbid
Biomphalaria glabrata...Moreover, we identified a previously undescribed rosette-like hemolymph protein that has been mistaken for hemoglobin. We also detected expression of an
2991:
are hard to understand in terms of their coiling and orientation. Many of the shells of species in this family are almost planispiral in coiling such that one side of the shell often looks rather like the other side, but it is important to bear in mind that nonetheless there is an
238:. The foot and head of planorbids are rather small, while their thread-like tentacles are relatively long. Many of the species in this family have coiled shells that are planispiral, in other words, the shells are more or less coiled flat, rather than having an elevated
1735:. The following list of genera is organized according to the 2005 taxonomy, because Albrecht's 2007 taxonomy is not available for all genera of Planorbidae. Genera in the family Planorbidae include (subgenera listed according to Glöer (2002):
3155:
2919:
Most species of coiled planorbids have a rather thin and moderately smooth shell, although more distinct sculpture such as a keel occurs in, and is diagnostic of, certain species. In the flat, keeled species, the whorls tend to overlap.
3000:
side of the shell. In addition these are in fact sinistral shells, despite the fact that the snail carries its shell as if it were a normal dextral shell. To make sense of the shell coiling, the following facts are useful:
3034:
on the left and facing the observer, then the sunken spire side of the shell is uppermost. This is a convenience for understanding the shell, but is the opposite of the way the shell is actually carried in life.
3078:
Species in this family occur worldwide. In
Northwest Europe about 20 species are known (including non-indigenous species). In this region, various extinct taxa are known to have occurred, starting in the
2916:
shells. Although it is now understood that these species are sinistral in shell coiling, disk-like
Planorbid shells are often still shown in illustrations oriented as if they were dextral.
3310:
306:
282:
3905:
3485:
Manuel de
Conchyliologie et de Paléontologie conchyliologique ou histoire naturelle des Mollusques vivants et fossiles suivi d'un appendice sur les Brachiopodes par D. Oehlert.
2927:
has a sharp outer lip. A peristome can be present, but often the lip is not thickened nor reflected. Those planorbid species which have a high-spired shell may have a narrow
3046:
of the shell is angled to face downwards also, so the aperture faces a little towards the spire, not away from it, as is usually the case in other shelled gastropods.
2897:
of most species in this family are disk-like or button-like, being coiled in one plane, although several groups have shells that are more higher-spired, and some are
3779:
3818:
3054:
Most species of planorbids live only in fresh water, such as ponds, lakes, and slow moving rivers. A minority of species are able to survive in brackish water.
2876:
3222:
Albrecht C., Kuhn K. & Streit B. (2007). "A molecular phylogeny of
Planorboidea (Gastropoda, Pulmonata): insights from enhanced taxon sampling".
3042:
in the living animal, contrary to what is the case in almost all other shelled gastropods. Because the shell is carried "upside down" like this, the
3753:
2938:
In height most species vary between 6 mm and 6 cm, however, disk-like shells are usually less than about 2 cm in maximum dimension.
3792:
3026:
In some species the umbilicus is not as deeply "dished" as the sunken spire is, so superficially it can be hard to tell one from the other.
541:
3434:
3512:
3456:
3328:
3260:
2862:
250:, the shell of coiled planorbids is in fact sinistral in coiling, but is carried upside down, which makes it appear to be dextral.
3867:
217:
more efficiently than other molluscs. The presence of hemoglobin gives the body a reddish colour. This is especially apparent in
3548:
3400:. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. <www.iucnredlist.org>. Downloaded on 08 March 2011.
2840:
3797:
3564:
2912:
Indeed, formerly planorbids were thought to have dextral shells, and so species of this family were figured as if they had
3805:
3473:
Gittenberger, E., Janssen, A.W., Kuijper, W.J., Kuiper, J.G.J., Meijer, T., Velde, G. van der & Vries, J.N. de (1998)
2777:
Wenz, 1923 is no longer considered to be a planorbid. It was recognised by Curry (1965, p. 360) as a euthecosomatous
137:
2836:
3900:
3675:
3652:
3477:
Nederlandse Fauna 2. Nationaal
Natuurhistorisch Museum Naturalis, KNNV Uitgeverij & EIS-Nederland, Leiden, 288 pp.
2296:
3427:
3421:
2825:
60:
3732:
2844:
2829:
3823:
3745:
3438:
2885:
2792:
2518:
2046:
1102:
694:
1732:
3556:
2788:
2042:
1515:
1342:
1220:
1179:
3592:
3370:
2089:
625:
281:. Alternatively, sometimes each one of them is raised to the level of a family. However, according to the
112:
1835:
1203:
3854:
2942:
2010:
1633:
1498:
954:
917:
47:
3680:
2484:
1591:
537:
2558:
3895:
3872:
3833:
3701:
3639:
3577:
3164:
2975:
2958:
2578:
1756:
1406:
1284:
3030:
However, once it is understood that the planorbid shell is sinistral, if the shell is held with the
2773:
2741:
2404:
2384:
2201:
1881:
1381:
1276:
911:
705:
505:
353:
3010:
2993:
2980:
2928:
2548:
2291:
2263:
2177:
1650:
1423:
1008:
494:
2875:
2668:
2344:
2227:
2163:
1966:
1944:
791:
518:
383:
374:
3302:
3043:
3031:
2924:
2731:
2698:
2638:
2444:
2123:
1085:
1047:
674:
461:
55:
3116:& Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater".
2761:
2751:
2708:
2658:
2648:
2628:
2588:
2568:
2538:
2464:
2434:
1913:
1780:
723:
323:
3859:
2618:
2598:
2364:
2334:
2314:
2143:
1990:
984:
941:
536:(2007) analyzed a limited number of genera of Planorbidae, based on sequences of mitochondrial
394:
3841:
3688:
3508:
3334:
3324:
3256:
3224:
3192:
2970:
2258:
2101:
2069:
2055:
1824:
1574:
1474:
567:
424:
403:
3784:
3630:
2110:
1813:
301:
Albino planorbid clearly shows the reddish-colored body tissues due to the pigment hemoglobin
3846:
3378:
3298:
3232:
3182:
3172:
3132:
3123:
1325:
948:
259:
258:
For several taxa, no consensus exists as to whether the taxa should even be assigned to the
189:
185:
886:
that shows the phylogenic relationships within the
Planorbidae according to Albrecht 2007:
19:
For "ramshorn snails" in the aquarium trade, some but not all of which are planorbids, see
3020:
of the shell is quite sunken in many species. In addition, it is carried facing downwards.
3017:
2997:
2988:
2963:
2953:
2932:
2894:
2880:
1751:
1061:
247:
243:
239:
3395:
3475:
De
Nederlandse zoetwatermollusken. Recente en fossiele weekdieren uit zoet en brak water
3168:
3740:
3727:
3306:
3187:
3150:
3113:
2510:
289:
within the family
Planorbidae, and that is the taxonomic system that is followed here.
193:
20:
544:(COI) genes, and on the basis of the results, they rearranged the taxonomy like this:
37:
3889:
3351:
Starobogatov, Y. I. (1967). On the systematization of freshwater pulmonate molluscs.
3236:
3118:
2424:
2414:
2271:
2001:
1792:
770:
740:
584:
3644:
1746:
3693:
3657:
3535:
In: H. Schindewolf (ed.), Handbuch der Paläozoologie, 6(2): pp. I-XII + 1–834.
2797:
2721:
2212:
2017:
1922:
1901:
1890:
1628:
1624:
837:
712:
645:
226:
160:
3810:
3766:
3624:
3315:
3151:"Red blood with blue-blood ancestry: Intriguing structure of a snail hemoglobin"
2814:
2713:
2454:
2374:
2324:
2186:
2027:
1297:
1154:
857:
664:
314:
177:
122:
3615:
3156:
Proceedings of the
National Academy of Sciences of the United States of America
3136:
2688:
2494:
2354:
1975:
1953:
1725:
867:
798:
278:
210:
206:
102:
3338:
3714:
3497:
Jena, (1), 1–376, 1929; (2), 377–778, 1931; 779–1022, 1934; 1023–1134, 1935.
3177:
2905:
2678:
2608:
2528:
2504:
2394:
2304:
2281:
2238:
2153:
2132:
1932:
1847:
1025:
937:
883:
818:
808:
780:
684:
615:
267:
199:
196:
72:
3382:
3196:
205:. Unlike most molluscs, the blood of ram's horn snails contains iron-based
2253:
Subfamily = ? (other genera that are not yet sorted are listed here)
297:
3609:
3080:
3067:
3063:
2778:
2474:
1859:
1802:
1375:
994:
905:
760:
750:
635:
595:
556:
340:
286:
202:
92:
3667:
3038:
The side of the shell which is in fact the spire (a sunken spire) faces
2913:
2078:
2037:
1868:
847:
605:
574:
273:
263:
3128:
246:. Although they carry their shell in a way that makes it appear to be
3771:
3572:
2898:
1769:
730:
218:
214:
82:
3758:
3586:
3719:
3101:
Analyse de la Nature ou tableau de l'univers et des corps organisés
2969:
2952:
2941:
Like all pulmonate aquatic snails, ramshorn shells do not have an
2874:
1745:
296:
277:. Both of these genera have sometimes been assigned to the family
3706:
3062:
Ancestors of ramshorn snails are known with certainty since the
235:
231:
3590:
3406:
3255:. Die Tierwelt Deutschlands, ConchBooks, Hackenheim, 326 pp.,
2808:
3009:
compared to the normal gastropod shell orientation, with the
262:
Planorbidae. This is certainly the case with the freshwater
3557:
Systema naturae 2000 (classification) - Taxon: Planorbidae
307:
taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
283:
taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
309:, this family consists of the following subfamilies:
3311:
Classification and nomenclator of gastropod families
3023:
The umbilicus of the shell is very wide and shallow.
2781:(Heterobranchia) (note by Arie W. Janssen, 092507).
3599:
285:, these genera are currently placed in the tribe
3582:, The University of Illinois press, Urbana, 1945
3253:Die Süßwassergastropoden Nord- und Mitteleuropas
3323:(1–2). Hackenheim, Germany: ConchBooks: 1–397.
3293:
3291:
3289:
3218:
3216:
3214:
3212:
3210:
3287:
3285:
3283:
3281:
3279:
3277:
3275:
3273:
3271:
3269:
213:. As a result, planorbids are able to breathe
3309:; Valdés, Ángel & Warén, Anders (2005). "
3149:Lieb, Bernhard; et al. (8 August 2006).
3005:In life, these pond snails hold their shells
8:
3507:American Malacologists Inc., Melbourne USA,
2879:Apical, apertural and umbilical view of the
3906:Taxa named by Constantine Samuel Rafinesque
3495:Handbuch der Systematischen Weichtierkunde.
3364:Liu L et al. (2010) "The phylogeography of
2843:. Unsourced material may be challenged and
3587:
3525:Fossilium Catalogus I. (4 vols.), 3387 pp.
3353:Trudy Zoologicheskogo Instituta, Leningrad
36:
27:
3457:Learn how and when to remove this message
3186:
3176:
2863:Learn how and when to remove this message
1555:new subfamily? "C‑clade"
1142:new subfamily? "B‑clade"
3420:This article includes a list of general
2219:- type genus of subfamily Neoplanorbinae
3092:
2245:- type genus of subfamily Rhodacmeinae
900:new family? "A‑clade"
3066:period. Modern taxa developed in the
7:
3834:0ac30e78-5508-4a68-b9bd-33490593b97e
3746:c60285d4-f517-482a-86a6-07f757f288a8
3505:A classification of living mollusca.
3368:(Gastropoda: Planorbidae) in Asia".
2931:, but frequently this is covered by
2841:adding citations to reliable sources
1897:- type genus of tribe Biomphalariini
528:2007 taxonomy for part of the family
2193:- type genus of tribe Plesiophysini
2085:- type genus of subfamily Bulininae
1616:
1566:
1559:
1490:
1466:
1398:
1367:
1317:
1310:
1303:
1272:
1195:
1171:
1146:
1077:
1053:
1000:
990:
980:
973:
933:
896:
889:
3573:http://mkohl1.net/Planorbidae.html
3549:NCBI Taxonomy Browser: Planorbidae
3426:it lacks sufficient corresponding
3305:; Frýda, Jiri; Hausdorf, Bernard;
1982:- type genus of tribe Segmentinini
1960:- type genus of tribe Planorbulini
452:Starobogatov & Prozorova, 1990
14:
3523:Gastropoda extramarina tertiaria.
3394:Mollusc Specialist Group (1996).
2904:All coiled shell Planorbidae are
2139:- type genus of tribe Miratestini
3580:The Molluscan Family Planorbidae
3411:
3237:10.1111/j.1463-6409.2006.00258.x
2813:
59:
2966:(held facing downwards in life)
2419:Brown & Mandahl-Barth, 1973
2286:Brown & Mandahl-Barth, 1973
765:Brown & Mandahl-Barth, 1973
224:Being air breathers like other
2117:- type genus of tribe Coretini
1875:- type genus of tribe Ancylini
1:
3103:. Palermo, 223 pp., page 143.
2945:to close the shell aperture.
2276:Özdikmen & Darilmaz, 2007
2006:Annandale & Prashad, 1920
167:About 250 freshwater species
3515:& 0-915826-22-6, 195 pp.
3112:Strong E. E., Gargominy O.,
465:Sararsin & Sarasin, 1897
329:- synonyms: Choanomphalinae
2703:Sarasin & Sarasin, 1897
2369:Laubrière & Carez, 1880
2329:Harry & Hubendick, 1964
2181:Bequaert & Clench, 1939
2137:Sarasin & Sarasin, 1897
2127:Sarasin & Sarasin, 1897
498:Bequaert & Clench, 1939
369:Harry & Hubendick, 1964
184:or ram's horn snails, is a
3922:
3483:Fischer, P.H. (1880-1887)
2073:Fischer & Crosse, 1880
1994:Fischer & Crosse, 1880
1906:Crosse & Fischer, 1880
430:- synonyms: Pompholicinae
407:Fischer & Crosse, 1880
398:Fischer & Crosse, 1880
346:- synonym: Pseudancylinae
18:
3137:10.1007/s10750-007-9012-6
3074:Geographical distribution
1647:
1621:
1614:
1588:
1571:
1564:
1557:
1512:
1495:
1488:
1471:
1464:
1420:
1403:
1396:
1372:
1365:
1339:
1322:
1315:
1308:
1301:
1281:
1270:
1217:
1200:
1193:
1176:
1169:
1151:
1144:
1099:
1082:
1075:
1058:
1051:
1022:
1005:
998:
988:
978:
971:
945:
931:
902:
894:
828:"C-Clade" sensu Albrecht
689:Walker & Ponder, 2001
655:"B-clade" sensu Albrecht
599:Rafinesque-Schmaltz, 1815
547:"A-clade" sensu Albrecht
467:- synonyms: Ferrissiinae
409:- synonyms: Laevapicinae
331:Fisher & Crosse, 1880
254:General taxonomic context
230:, planorbids do not have
166:
159:
150:
145:
56:Scientific classification
54:
44:
35:
30:
2983:(held uppermost in life)
2886:Biomphalaria tenagophila
2795:, 1855 is a synonym for
2736:Gutiérrez Gregoric, 2014
1103:Hebetancylus excentricus
209:instead of copper-based
3493:Thiele, J. (1929-1935)
3441:more precise citations.
3371:Parasites & Vectors
3178:10.1073/pnas.0601861103
3099:Rafinesque C. S. 1815.
2603:Tapparone-Canefri, 1883
2008:- contains one species
1516:Polypylis hemisphaerula
1343:Bathyomphalus contortus
852:Adams & Adams, 1855
511:- synonym: Payettiinae
242:as is the case in most
3531:Zilch, A. (1959-1960)
3383:10.1186/1756-3305-3-57
2987:Flat-coiled planorbid
2984:
2967:
2890:
1760:
1739:subfamily Planorbinae
1221:Leichhardtia sisurnius
542:cytochrome-c oxidase I
359:- synonyms: Acrorbini
302:
234:, but instead, have a
3855:Paleobiology Database
3578:Frank Collins Baker,
3521:Wenz, W. (1923-1930)
3366:Indoplanorbis exustus
2973:
2962:. View of the sunken
2956:
2878:
2011:Indoplanorbis exustus
1749:
1634:Biomphalaria glabrata
1499:Hippeutis complanatus
1180:Protancylus adhaerens
955:Indoplanorbis exustus
918:Burnupia stenochorias
487:; Patelloplanorbidae
300:
48:Indoplanorbis exustus
45:A live individual of
3741:Fauna Europaea (new)
3503:Vaught, K.C. (1989)
2976:Planorbarius corneus
2959:Planorbarius corneus
2889:. Scale bar is 3 mm.
2837:improve this section
1757:Anisus septemgyratus
1407:Choanomphalus maacki
1285:Planorbarius corneus
363:; Drepanothrematini
16:Family of gastropods
3303:Rocroi, Jean-Pierre
3169:2006PNAS..10312011L
3163:(32): 12011–12016.
2726:Mandahl-Barth, 1954
2533:Mandahl-Barth, 1954
1750:View of the sunken
1651:Planorbula armigera
1424:Planorbis planorbis
1204:Kessneria papillosa
1009:Ancylus fluviatilis
882:The following is a
442:; Pompholycodeinae
438:; Megasystrophinae
348:Walker, 1923 (inv.)
3901:Gastropod families
3397:Culmenella rezvoji
3058:Geological history
2985:
2968:
2891:
2766:Meek in Dall, 1870
1927:Starobogatov, 1967
1829:Starobogatov, 1967
1761:
1728:of this family is
1086:Gundlachia radiata
481:Starobogatov, 1958
456:Starobogatov, 1990
440:Tryon, 1871 (inv.)
434:; Camptoceratinae
419:Starobogatov, 1967
361:Starobogatov, 1958
303:
3883:
3882:
3842:Open Tree of Life
3593:Taxon identifiers
3565:ITIS: Planorbidae
3487:– XXIV + 1369 pp.
3467:
3466:
3459:
3299:Bouchet, Philippe
3225:Zoologica Scripta
2873:
2872:
2865:
2805:Shell description
2784:The generic name
2767:
2757:
2747:
2737:
2727:
2717:
2704:
2694:
2684:
2674:
2664:
2654:
2644:
2634:
2624:
2614:
2604:
2594:
2584:
2574:
2564:
2554:
2544:
2543:(Meek) Dall, 1870
2534:
2524:
2514:
2500:
2490:
2480:
2470:
2460:
2450:
2440:
2430:
2420:
2410:
2400:
2390:
2380:
2370:
2360:
2350:
2340:
2330:
2320:
2310:
2300:
2287:
2277:
2267:
2244:
2232:
2218:
2206:
2192:
2182:
2169:
2159:
2149:
2138:
2128:
2116:
2106:
2095:
2084:
2074:
2061:
2050:
2033:
2023:
2007:
1995:
1981:
1971:
1959:
1949:
1938:
1928:
1918:
1907:
1896:
1886:
1874:
1864:
1853:
1852:Charpentier, 1837
1841:
1830:
1819:
1808:
1807:Charpentier, 1837
1798:
1797:Charpentier, 1837
1786:
1775:
1742:
1716:
1715:
1707:
1706:
1698:
1697:
1689:
1688:
1680:
1679:
1671:
1670:
1662:
1661:
1603:
1602:
1592:Planorbella tenue
1575:Menetus dilatatus
1545:
1544:
1536:
1535:
1527:
1526:
1475:Segmentina nitida
1453:
1452:
1444:
1443:
1435:
1434:
1354:
1353:
1259:
1258:
1250:
1249:
1241:
1240:
1232:
1231:
1132:
1131:
1123:
1122:
1114:
1113:
1037:
1036:
873:
863:
853:
843:
824:
814:
813:Charpentier, 1837
804:
786:
776:
766:
756:
755:Charpentier, 1837
746:
745:Charpentier, 1837
736:
718:
700:
690:
680:
670:
651:
641:
631:
621:
611:
600:
590:
580:
562:
538:18S ribosomal DNA
523:
514:
510:
499:
490:
486:
482:
478:
474:
470:
466:
457:
453:
449:
445:
441:
437:
433:
432:Dall, 1866 (inv.)
429:
420:
416:
412:
408:
399:
388:
379:
370:
366:
362:
358:
349:
345:
336:
333:; Orygoceratidae
332:
328:
319:
305:According to the
190:freshwater snails
188:of air-breathing
171:
170:
141:
3913:
3876:
3875:
3863:
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3836:
3827:
3826:
3814:
3813:
3811:NHMSYS0001702169
3801:
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3788:
3787:
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3442:
3437:this article by
3428:inline citations
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2989:gastropod shells
2949:Sinistral shells
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2559:Patelloplanorbis
2552:
2542:
2532:
2522:
2508:
2498:
2488:
2478:
2468:
2458:
2448:
2438:
2429:Gerstfeldt, 1859
2428:
2418:
2408:
2398:
2388:
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2368:
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2328:
2319:Sandberger, 1875
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1957:
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1926:
1917:Rafinesque, 1815
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1872:
1863:Rafinesque, 1815
1862:
1851:
1839:
1828:
1817:
1806:
1796:
1784:
1773:
1754:of the shell of
1741:Rafinesque, 1815
1740:
1617:
1567:
1560:
1491:
1467:
1399:
1368:
1326:Anisus spirorbis
1318:
1311:
1304:
1273:
1196:
1172:
1147:
1078:
1054:
1001:
991:
981:
974:
949:Bulinus tropicus
934:
897:
890:
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861:
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841:
822:
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802:
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775:Gerstfeldt, 1859
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497:
488:
484:
480:
476:
475:; Protancylinae
472:
471:; Ancylastrinae
468:
464:
455:
454:; Planorbariini
451:
447:
446:; Helisomatinae
443:
439:
435:
431:
427:
418:
417:; Gundlachiinae
414:
410:
406:
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386:
377:
368:
364:
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356:
347:
344:Rafinesque, 1815
343:
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327:Rafinesque, 1815
326:
318:Rafinesque, 1815
317:
244:gastropod shells
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3446:
3443:
3433:Please help to
3432:
3416:
3412:
3405:
3404:
3393:
3389:
3363:
3359:
3350:
3346:
3331:
3307:Ponder, Winston
3297:
3296:
3267:
3251:Glöer P. 2002.
3248:
3247:
3243:
3221:
3208:
3148:
3147:
3143:
3127:
3111:
3107:
3098:
3094:
3089:
3076:
3060:
3052:
3013:facing upwards.
2951:
2869:
2858:
2852:
2849:
2834:
2818:
2807:
2579:Pentagoniostoma
2563:Hubendick, 1957
2489:Annandale, 1922
2278:
1774:S. Studer, 1820
1722:
1717:
1708:
1699:
1690:
1681:
1672:
1663:
1604:
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1537:
1528:
1454:
1445:
1436:
1355:
1260:
1251:
1242:
1233:
1133:
1124:
1115:
1062:Laevapex fuscus
1038:
880:
589:Annandale, 1921
530:
479:; Physastrinae
450:; Bayardellini
415:Annandale, 1922
295:
256:
182:ramshorn snails
135:
58:
24:
17:
12:
11:
5:
3919:
3917:
3909:
3908:
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3898:
3888:
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3864:
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3802:
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3750:
3737:
3728:Fauna Europaea
3724:
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3672:
3662:
3649:
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3540:External links
3538:
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3355:, 42, 280–304.
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3206:
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3122:595: 149-166.
3105:
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3014:
2979:. View of the
2950:
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2870:
2821:
2819:
2812:
2806:
2803:
2774:Camptoceratops
2769:
2768:
2758:
2748:
2742:Syrioplanorbis
2738:
2728:
2718:
2705:
2695:
2685:
2675:
2665:
2655:
2653:Haldeman, 1842
2645:
2635:
2625:
2615:
2605:
2595:
2593:Hannibal, 1912
2585:
2575:
2565:
2555:
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2525:
2515:
2501:
2491:
2481:
2479:Swainson, 1840
2471:
2461:
2451:
2441:
2431:
2421:
2411:
2409:Polinski, 1929
2405:Carinogyraulis
2401:
2391:
2385:Camptoceratops
2381:
2371:
2361:
2351:
2341:
2331:
2321:
2311:
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2248:
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2246:
2224:
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2220:
2205:Hannibal, 1912
2202:Neoplanorbinae
2198:
2197:
2196:
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2194:
2174:
2173:
2172:
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2170:
2150:
2140:
2120:
2119:
2118:
2098:
2097:
2096:
2094:Pfeiffer, 1849
2086:
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2065:
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2062:
2034:
2032:Haldeman, 1842
2024:
2014:
1987:
1986:
1985:
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1983:
1963:
1962:
1961:
1958:Haldeman, 1840
1941:
1940:
1939:
1929:
1910:
1909:
1908:
1898:
1882:Biomphalariini
1878:
1877:
1876:
1856:
1855:
1854:
1844:
1843:
1842:
1840:Hartmann, 1843
1831:
1820:
1799:
1789:
1788:
1787:
1785:Boettger, 1944
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1419:
1416:
1415:
1412:
1411:
1402:
1397:
1395:
1392:
1391:
1388:
1387:
1382:Caronogyraulus
1371:
1366:
1364:
1361:
1360:
1357:
1356:
1352:
1351:
1348:
1347:
1338:
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1321:
1316:
1314:
1309:
1307:
1302:
1300:
1294:
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1289:
1280:
1277:Camptoceratini
1271:
1269:
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1265:
1262:
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1257:
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1111:
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964:
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959:
944:
932:
930:
927:
926:
923:
922:
912:Burnupia cafra
901:
895:
893:
888:
879:
876:
875:
874:
872:Haldeman, 1843
864:
862:Haldeman, 1843
854:
844:
826:
825:
815:
805:
788:
787:
777:
767:
757:
747:
737:
720:
719:
706:Camptoceratini
702:
701:
691:
681:
671:
653:
652:
642:
632:
630:Pfeiffer, 1849
622:
612:
592:
591:
581:
564:
563:
529:
526:
525:
524:
515:
509:Hannibal, 1912
506:Neoplanorbinae
502:
501:
500:
491:
483:; Ameriannini
458:
444:Lindholm, 1927
421:
411:Hannibal, 1912
391:
390:
389:
380:
371:
354:Biomphalariini
350:
337:
294:
291:
255:
252:
169:
168:
164:
163:
157:
156:
148:
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85:
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76:
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65:
52:
51:
42:
41:
33:
32:
21:Ramshorn snail
15:
13:
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9:
6:
4:
3:
2:
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3907:
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3641:
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3611:
3607:
3606:
3604:
3602:
3598:
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3558:
3552:
3550:
3544:
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3514:
3513:0-915826-22-4
3510:
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3408:
3399:
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3330:3-925919-72-4
3326:
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3290:
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3272:
3270:
3266:
3263:, page 19-20.
3262:
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3184:
3179:
3174:
3170:
3166:
3162:
3158:
3157:
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3145:
3142:
3138:
3134:
3130:
3125:
3121:
3120:
3119:Hydrobiologia
3115:
3109:
3106:
3102:
3096:
3093:
3086:
3084:
3082:
3073:
3071:
3069:
3065:
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3055:
3049:
3047:
3045:
3041:
3036:
3033:
3025:
3022:
3019:
3015:
3012:
3008:
3004:
3003:
3002:
2999:
2995:
2990:
2982:
2978:
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2910:
2907:
2902:
2900:
2896:
2888:
2887:
2882:
2877:
2867:
2864:
2856:
2846:
2842:
2838:
2832:
2831:
2827:
2822:This section
2820:
2816:
2811:
2810:
2804:
2802:
2800:
2799:
2794:
2790:
2787:
2782:
2780:
2776:
2775:
2764:
2763:
2759:
2754:
2753:
2749:
2744:
2743:
2739:
2734:
2733:
2729:
2724:
2723:
2719:
2715:
2711:
2710:
2706:
2701:
2700:
2696:
2691:
2690:
2686:
2683:Pilsbry, 1906
2681:
2680:
2676:
2673:Pilsbry, 1924
2671:
2670:
2666:
2663:Fischer, 1883
2661:
2660:
2656:
2651:
2650:
2646:
2643:Pilsbry, 1935
2641:
2640:
2636:
2633:Edwards, 1860
2631:
2630:
2626:
2621:
2620:
2616:
2611:
2610:
2606:
2601:
2600:
2596:
2591:
2590:
2586:
2583:Branson, 1935
2581:
2580:
2576:
2571:
2570:
2566:
2561:
2560:
2556:
2551:
2550:
2549:Paraplanorbis
2546:
2541:
2540:
2536:
2531:
2530:
2526:
2521:
2520:
2516:
2512:
2509:Walker &
2507:
2506:
2502:
2497:
2496:
2492:
2487:
2486:
2482:
2477:
2476:
2472:
2467:
2466:
2462:
2457:
2456:
2452:
2449:Pilsbry, 1934
2447:
2446:
2442:
2437:
2436:
2432:
2427:
2426:
2425:Choanomphalus
2422:
2417:
2416:
2415:Ceratophallus
2412:
2407:
2406:
2402:
2397:
2396:
2392:
2387:
2386:
2382:
2377:
2376:
2372:
2367:
2366:
2362:
2357:
2356:
2352:
2349:Pilsbry, 1934
2347:
2346:
2342:
2339:Clessin, 1884
2337:
2336:
2332:
2327:
2326:
2322:
2317:
2316:
2312:
2309:Pilsbry, 1906
2307:
2306:
2302:
2298:
2294:
2293:
2292:Afroplanorbis
2289:
2284:
2283:
2274:
2273:
2272:Africanogyrus
2269:
2265:
2261:
2260:
2256:
2255:
2254:
2241:
2240:
2236:
2235:
2234:
2233:
2229:
2225:
2217:Pilsbry, 1906
2215:
2214:
2210:
2209:
2208:
2207:
2203:
2199:
2191:Fischer, 1883
2189:
2188:
2184:
2183:
2179:
2178:Plesiophysini
2175:
2168:Iredale, 1943
2166:
2165:
2161:
2160:
2156:
2155:
2151:
2146:
2145:
2141:
2135:
2134:
2130:
2129:
2125:
2121:
2113:
2112:
2108:
2107:
2103:
2099:
2092:
2091:
2087:
2081:
2080:
2076:
2075:
2071:
2067:
2060:Clessin, 1885
2058:
2057:
2052:
2051:
2048:
2044:
2040:
2039:
2035:
2030:
2029:
2025:
2022:Duméril, 1806
2020:
2019:
2015:
2013:
2012:
2004:
2003:
2002:Indoplanorbis
1999:
1998:
1997:
1996:
1992:
1988:
1980:Fleming, 1818
1978:
1977:
1973:
1972:
1968:
1964:
1956:
1955:
1951:
1950:
1948:Pilsbry, 1934
1946:
1942:
1935:
1934:
1930:
1925:
1924:
1920:
1919:
1915:
1911:
1904:
1903:
1899:
1895:Preston, 1910
1893:
1892:
1888:
1887:
1883:
1879:
1871:
1870:
1866:
1865:
1861:
1857:
1850:
1849:
1845:
1838:
1837:
1832:
1827:
1826:
1821:
1816:
1815:
1810:
1809:
1805:
1804:
1800:
1795:
1794:
1793:Bathyomphalus
1790:
1783:
1782:
1777:
1776:
1772:
1771:
1767:
1766:
1765:
1764:
1763:
1762:
1759:
1758:
1753:
1748:
1738:
1737:
1736:
1734:
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1727:
1719:
1712:
1711:
1703:
1702:
1694:
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1684:
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1667:
1666:
1658:
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1654:
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1599:
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1569:
1568:
1562:
1561:
1554:
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1550:
1549:
1541:
1540:
1532:
1531:
1523:
1522:
1519:
1518:
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1510:
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1506:
1505:
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1501:
1500:
1493:
1492:
1486:
1485:
1482:
1481:
1478:
1477:
1476:
1469:
1468:
1462:
1461:
1458:
1457:
1449:
1448:
1440:
1439:
1431:
1430:
1427:
1426:
1425:
1418:
1417:
1414:
1413:
1410:
1409:
1408:
1401:
1400:
1394:
1393:
1390:
1389:
1386:
1384:
1383:
1378:
1377:
1370:
1369:
1363:
1362:
1359:
1358:
1350:
1349:
1346:
1345:
1344:
1337:
1336:
1333:
1332:
1329:
1328:
1327:
1320:
1319:
1313:
1312:
1306:
1305:
1299:
1296:
1295:
1292:
1291:
1288:
1287:
1286:
1278:
1275:
1274:
1268:
1267:
1264:
1263:
1255:
1254:
1246:
1245:
1237:
1236:
1228:
1227:
1224:
1223:
1222:
1215:
1214:
1211:
1210:
1207:
1206:
1205:
1198:
1197:
1191:
1190:
1187:
1186:
1183:
1182:
1181:
1174:
1173:
1167:
1166:
1163:
1162:
1159:
1157:
1156:
1149:
1148:
1141:
1140:
1137:
1136:
1128:
1127:
1119:
1118:
1110:
1109:
1106:
1105:
1104:
1097:
1096:
1093:
1092:
1089:
1088:
1087:
1080:
1079:
1073:
1072:
1069:
1068:
1065:
1064:
1063:
1056:
1055:
1049:
1046:
1045:
1042:
1041:
1033:
1032:
1029:
1028:
1027:
1020:
1019:
1016:
1015:
1012:
1011:
1010:
1003:
1002:
996:
993:
992:
986:
983:
982:
976:
975:
969:
966:
965:
962:
961:
958:
957:
956:
951:
950:
943:
940: /
939:
936:
935:
929:
928:
925:
924:
921:
920:
919:
914:
913:
908:
907:
899:
898:
892:
891:
887:
885:
877:
870:
869:
865:
860:
859:
855:
850:
849:
845:
842:Preston, 1910
840:
839:
835:
834:
833:
831:
823:Pilsbry, 1906
821:
820:
816:
811:
810:
806:
803:Fleming, 1818
801:
800:
796:
795:
794:
793:
783:
782:
778:
773:
772:
771:Choanomphalus
768:
763:
762:
758:
753:
752:
748:
743:
742:
741:Bathyomphalus
738:
733:
732:
728:
727:
726:
725:
717:Duméril, 1806
715:
714:
710:
709:
708:
707:
697:
696:
692:
687:
686:
682:
679:Sarasin, 1897
677:
676:
672:
667:
666:
662:
661:
660:
658:
650:Pilsbry, 1914
648:
647:
643:
638:
637:
633:
628:
627:
623:
618:
617:
613:
608:
607:
603:
602:
601:
597:
587:
586:
585:Indoplanorbis
582:
577:
576:
572:
571:
570:
569:
559:
558:
554:
553:
552:
550:
545:
543:
539:
535:
527:
520:
516:
507:
503:
496:
495:Plesiophysini
492:
463:
459:
426:
422:
413:; Isidorinae
405:
401:
400:
396:
392:
385:
381:
378:Pilsbry, 1934
376:
372:
355:
351:
342:
338:
335:Brusina, 1882
325:
321:
320:
316:
312:
311:
310:
308:
299:
293:2005 taxonomy
292:
290:
288:
284:
280:
276:
275:
270:
269:
265:
261:
253:
251:
249:
245:
241:
237:
233:
229:
228:
222:
220:
216:
212:
208:
204:
201:
198:
195:
191:
187:
183:
179:
175:
165:
162:
158:
155:
154:
149:
144:
139:
134:
131:
128:
127:
124:
121:
119:Superfamily:
118:
117:
114:
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
62:
57:
53:
50:
49:
43:
39:
34:
29:
26:
22:
3600:
3579:
3570:(in English)
3562:(in English)
3554:(in English)
3546:(in English)
3532:
3528:
3522:
3518:
3504:
3501:(in English)
3494:
3490:
3484:
3474:
3453:
3447:January 2009
3444:
3425:
3396:
3390:
3374:
3369:
3365:
3360:
3352:
3347:
3320:
3314:
3252:
3244:
3228:
3223:
3200:
3160:
3154:
3144:
3117:
3114:Ponder W. F.
3108:
3100:
3095:
3077:
3061:
3053:
3039:
3037:
3029:
3006:
2986:
2974:
2957:
2940:
2937:
2922:
2918:
2911:
2903:
2892:
2884:
2859:
2853:January 2016
2850:
2835:Please help
2823:
2798:Biomphalaria
2796:
2785:
2783:
2772:
2770:
2760:
2756:Benson, 1855
2750:
2740:
2730:
2722:Segmentorbis
2720:
2707:
2697:
2687:
2677:
2669:Platytaphius
2667:
2657:
2647:
2637:
2627:
2617:
2613:Krauss, 1848
2607:
2597:
2587:
2577:
2573:Taylor, 1985
2567:
2557:
2547:
2537:
2527:
2523:Walker, 1988
2519:Leichhardtia
2517:
2503:
2493:
2483:
2473:
2469:Benson, 1855
2463:
2459:Crosse, 1872
2453:
2443:
2439:Clench, 1927
2433:
2423:
2413:
2403:
2399:Binney, 1865
2393:
2383:
2379:Benson, 1843
2373:
2363:
2353:
2345:Australorbis
2343:
2333:
2323:
2313:
2303:
2290:
2280:
2270:
2257:
2252:
2243:Walker, 1917
2237:
2231:Walker, 1917
2228:Rhodacmeinae
2213:Neoplanorbis
2211:
2185:
2164:Pettancyclus
2162:
2158:Walker, 1903
2152:
2148:Strand, 1928
2142:
2131:
2109:
2088:
2083:Müller, 1781
2077:
2054:
2036:
2026:
2018:Planorbarius
2016:
2009:
2000:
1974:
1967:Segmentinini
1952:
1945:Planorbulini
1937:Müller, 1773
1931:
1923:Afrogyrorbis
1921:
1902:Drepanotrema
1900:
1891:Biomphalaria
1889:
1885:Watson, 1954
1873:Müller, 1773
1867:
1846:
1834:
1823:
1812:
1801:
1791:
1779:
1768:
1755:
1729:
1723:
1649:
1648:
1632:
1625:Biomphalaria
1623:
1622:
1590:
1589:
1573:
1572:
1514:
1513:
1497:
1496:
1473:
1472:
1422:
1421:
1405:
1404:
1380:
1374:
1373:
1341:
1340:
1324:
1323:
1283:
1282:
1219:
1218:
1202:
1201:
1178:
1177:
1153:
1152:
1101:
1100:
1084:
1083:
1060:
1059:
1024:
1023:
1007:
1006:
967:
953:
947:
946:
916:
910:
904:
903:
881:
866:
856:
846:
838:Biomphalaria
836:
829:
827:
817:
807:
797:
792:Segmentinini
789:
785:Müller, 1774
779:
769:
759:
749:
739:
735:Studer, 1820
729:
721:
713:Planorbarius
711:
703:
699:Walker, 1988
695:Leichhardtia
693:
683:
673:
669:Crosse, 1872
663:
656:
654:
646:Hebetancylus
644:
640:Walker, 1903
634:
624:
620:Walker, 1903
614:
610:Müller, 1774
604:
593:
583:
579:Müller, 1781
573:
565:
561:Walker, 1912
555:
548:
546:
533:
531:
522:Walker, 1917
519:Rhodacmeinae
477:Walker, 1923
473:Walker, 1923
469:Walker, 1917
384:Segmentinini
375:Planorbulini
367:; Taphiinae
357:Watson, 1954
304:
272:
266:
257:
227:Panpulmonata
225:
223:
181:
173:
172:
152:
151:
132:
109:Superorder:
46:
31:Planorbidae
25:
3896:Planorbidae
3767:iNaturalist
3658:Planorbidae
3645:Planorbidae
3631:Planorbidae
3625:Wikispecies
3601:Planorbidae
3533:Euthyneura.
3481:(in French)
3439:introducing
3316:Malacologia
3249:(in German)
3007:upside down
2996:side and a
2746:Baker, 1945
2732:Sineancylus
2699:Protancylus
2693:Baker, 1935
2639:Planorbifex
2623:Baker, 1945
2553:Hanna, 1922
2455:Glyptophysa
2445:Fossulorbis
2375:Camptoceras
2359:Burch, 1977
2325:Antillorbis
2279:- synonym:
2187:Plesiophysa
2124:Miratestini
2028:Planorbella
1970:Baker, 1945
1298:Planorbinae
1155:Glyptophysa
1048:Laevapecini
968:Planorbidae
858:Planorbella
675:Protancylus
665:Glyptophysa
489:Franc, 1968
485:Zilch, 1959
462:Miratestini
448:Baker, 1928
387:Baker, 1945
365:Zilch, 1959
315:Planorbinae
178:common name
174:Planorbidae
133:Planorbidae
123:Lymnaeoidea
3890:Categories
3471:(in Dutch)
3422:references
3129:10088/7390
3087:References
2771:The genus
2762:Vorticifex
2752:Trochorbis
2709:Pygmanisus
2689:Promenetus
2659:Platyphysa
2649:Planorbina
2629:Pitharella
2589:Perrinilla
2569:Pecosorbis
2539:Macrophysa
2499:Tate, 1896
2495:Isidorella
2465:Helicorbis
2435:Culmenella
2389:Wenz, 1923
2355:Bayardella
2226:subfamily
2200:subfamily
2115:Gray, 1847
2105:Gray, 1847
2090:Gundlachia
1989:subfamily
1976:Segmentina
1954:Planorbula
1914:Planorbini
1818:Dall, 1905
1781:Disculifer
1726:type genus
868:Planorbula
799:Segmentina
724:Planorbini
626:Gundlachia
517:subfamily
513:Dall, 1924
504:subfamily
436:Dall, 1870
428:Gray, 1847
393:subfamily
324:Planorbini
313:subfamily
279:Lymnaeidae
211:hemocyanin
207:hemoglobin
138:Rafinesque
113:Hygrophila
103:Gastropoda
3339:0076-2997
3011:umbilicus
2994:umbilical
2981:umbilicus
2943:operculum
2929:umbilicus
2909:dextral.
2906:sinistral
2824:does not
2679:Polypylis
2619:Pingiella
2609:Physopsis
2599:Physastra
2529:Lentorbis
2505:Kessneria
2395:Carinifex
2365:Berellaia
2335:Armigerus
2315:Anisopsis
2305:Amphigyra
2282:Afrogyrus
2239:Rhodacmea
2154:Ferrissia
2144:Amerianna
2133:Miratesta
2053:subgenus
1991:Bulininae
1933:Planorbis
1848:Hippeutis
1833:subgenus
1822:subgenus
1811:subgenus
1778:subgenus
1730:Planorbis
1026:Ferrissia
985:Ancylinae
942:Bulininae
938:Bulinidae
884:cladogram
878:Cladogram
832:, (2007)
819:Polypylis
809:Hippeutis
781:Planorbis
685:Kessneria
659:, (2007)
616:Ferrissia
532:Albrecht
395:Bulininae
268:Ferrissia
221:animals.
200:gastropod
197:pulmonate
161:Diversity
79:Kingdom:
73:Eukaryota
3707:46447618
3665:BioLib:
3616:Q1475666
3610:Wikidata
3529:(German)
3519:(German)
3491:(German)
3231:: 27-39.
3197:16877545
3083:period.
3081:Jurassic
3068:Jurassic
3064:Jurassic
3044:aperture
3032:aperture
2925:aperture
2779:pteropod
2475:Helisoma
2259:Acrorbis
2102:Coretini
2070:Bulinini
2056:Dilatata
1860:Ancylini
1825:Lamorbis
1803:Gyraulus
1376:Gyraulus
995:Ancylini
906:Burnupia
761:Hovorbis
751:Gyraulus
636:Laevapex
596:Ancylini
568:Bulinini
557:Burnupia
425:Coretini
404:Bulinini
341:Ancylini
287:Ancylini
203:molluscs
153:See text
129:Family:
93:Mollusca
89:Phylum:
83:Animalia
69:Domain:
3435:improve
3188:1567689
3165:Bibcode
3050:Habitat
2914:dextral
2901:-like.
2845:removed
2830:sources
2786:Taphius
2714:Iredale
2111:Coretus
2079:Bulinus
2038:Menetus
1869:Ancylus
1836:Armiger
1814:Torquis
848:Menetus
790:Tribus
722:Tribus
704:Tribus
606:Ancylus
594:Tribus
575:Bulinus
566:Tribus
551:(2007)
274:Ancylus
264:limpets
248:dextral
194:aquatic
146:Genera
99:Class:
3873:153908
3847:693697
3831:NZOR:
3785:114548
3720:1PLANF
3511:
3424:, but
3377:: 57.
3337:
3327:
3259:
3195:
3185:
2933:callus
2899:limpet
2895:shells
2791:&
2716:, 1943
2513:, 2001
2511:Ponder
2299:, 1931
2297:Thiele
2266:, 1937
2264:Odhner
2176:tribe
2122:tribe
2100:tribe
2068:tribe
2049:, 1855
2045:&
1965:tribe
1943:tribe
1912:tribe
1880:tribe
1858:tribe
1770:Anisus
1733:Müller
1720:Genera
1385:) sp.
830:et al.
731:Anisus
657:et al.
549:et al.
534:et al.
493:tribe
460:tribe
423:tribe
402:tribe
382:tribe
373:tribe
352:tribe
339:tribe
322:tribe
271:, and
260:family
219:albino
215:oxygen
186:family
140:, 1815
3868:WoRMS
3860:61804
3798:76591
3780:IRMNG
3772:85990
3733:11410
3694:7NKHX
3018:spire
2998:spire
2964:spire
2881:shell
2793:Adams
2789:Adams
2485:Intha
2047:Adams
2043:Adams
1752:spire
240:spire
232:gills
3824:6524
3819:NCBI
3793:ITIS
3759:2851
3754:GBIF
3715:EPPO
3681:1476
3676:BOLD
3668:2575
3509:ISBN
3335:ISSN
3325:ISBN
3257:ISBN
3193:PMID
3040:down
3016:The
2923:The
2893:The
2828:any
2826:cite
1724:The
1158:sp.
952:and
915:and
540:and
236:lung
180:the
3806:NBN
3702:EoL
3689:CoL
3653:AFD
3640:ADW
3379:doi
3313:".
3233:doi
3183:PMC
3173:doi
3161:103
3133:doi
3124:hdl
2883:of
2839:by
3892::
3870::
3857::
3844::
3821::
3808::
3795::
3782::
3769::
3756::
3743::
3730::
3717::
3704::
3691::
3678::
3655::
3642::
3627::
3612::
3333:.
3321:47
3319:.
3301:;
3268:^
3229:36
3209:^
3199:.
3191:.
3181:.
3171:.
3159:.
3153:.
3070:.
2935:.
2801:.
1631:-
1279:?
909:;
192:,
176:,
3460:)
3454:(
3449:)
3445:(
3431:.
3385:.
3381::
3375:3
3341:.
3239:.
3235::
3175::
3167::
3139:.
3135::
3126::
2866:)
2860:(
2855:)
2851:(
2847:.
2833:.
1629:►
1379:(
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
