44:
65:
297:
Kusy, Dominik; He, Jin‐Wu; Bybee, Seth M.; Motyka, Michal; Bi, Wen‐Xuan; Podsiadlowski, Lars; Li, Xue‐Yan; Bocak, Ladislav (2020-08-25). "Phylogenomic relationships of bioluminescent elateroids define the 'lampyroid' clade with clicking
Sinopyrophoridae as its earliest member".
335:"Sinopyrophorinae, a new subfamily of Elateridae (Coleoptera, Elateroidea) with the first record of a luminous click beetle in Asia and evidence for multiple origins of bioluminescence in Elateridae"
241:), phylogenomic analyses revealed they are phylogenetically outside the Elateridae proper, and therefore were elevated to their own independent family, the Sinopyrophoridae.
2368:
237:) known from Asia, however, despite their similar "clicking" mechanism & light organ structure when compared to the true bioluminescent click beetles (e.g.
43:
2443:
2293:
2355:
406:
2472:
2467:
2436:
64:
2477:
2429:
399:
2386:
2308:
255:
represents the earliest diverging member of the monophyletic clade of bioluminescent beetles known as the
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256:
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1545:
2346:
1764:
392:
59:
2373:
2033:
1450:
370:
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315:
2413:
2130:
1909:
1904:
1827:
1585:
601:
360:
342:
307:
264:
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247:
is only distantly related to the hard bodied true bioluminescent click beetles, such as the
1819:
1772:
1617:
1601:
1277:
1037:
1006:
921:
839:
730:
698:
223:, which was described in 2019 from the subtropical evergreen broadleaf forests of western
2222:
2173:
2028:
1835:
1503:
1487:
1426:
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1292:
1220:
1184:
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1029:
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642:
622:
518:
365:
334:
333:
Bi, Wen-Xuan; He, Jin-Wu; Chen, Chang-Chin; Kundrata, Robin; Li, Xue-Yan (2019-07-17).
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747:
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2267:
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2212:
2199:
2139:
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2065:
2049:
2041:
1977:
1935:
1927:
1896:
1811:
1790:
1756:
1748:
1661:
1593:
1577:
1569:
1532:
1519:
1495:
1413:
1405:
1355:
1347:
1305:
1269:
1240:
1212:
1207:
1123:
1118:
1110:
1060:
996:
991:
905:
870:
855:
847:
763:
755:
572:
268:
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126:
17:
2194:
2155:
2147:
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1867:
1859:
1679:
1627:
1511:
1442:
1313:
1297:
1253:
1194:
1158:
1138:
1102:
1011:
1001:
875:
609:
593:
347:
272:
234:
384:
356:
319:
2259:
2251:
2243:
2230:
2186:
1985:
1960:
1888:
1716:
1687:
1371:
1202:
1176:
1128:
1042:
944:
913:
885:
718:
693:
688:
669:
634:
523:
506:
498:
447:
435:
96:
76:
374:
233:
was previously thought to represent the only bioluminescent click beetles (
2325:
2163:
2113:
2057:
2016:
1965:
1622:
1553:
1434:
1146:
952:
926:
784:
617:
564:
543:
453:
2360:
2381:
1880:
1872:
1724:
1458:
1379:
1363:
311:
1471:
1331:
964:
441:
429:
416:
224:
116:
106:
86:
2302:
2306:
715:
666:
540:
472:
388:
213:, and is the sole member of the recently recognized family
51:
Male (labelled "2") and female (labelled "3") specimens of
787:(furniture beetles, death watch beetles, spider beetles)
2417:
2315:
2221:
2172:
2138:
2129:
2007:
1998:
1951:
1926:
1810:
1781:
1678:
1652:
1643:
1425:
1404:
1330:
1137:
1028:
884:
830:
821:
796:
738:
729:
554:
217:. The genus currently contains a single species,
1664:(jewel beetles, or metallic wood-boring beetles)
209:hard-bodied clicking beetles in the superfamily
1387:(true weevils, bark beetles, ambrosia beetles)
2437:
400:
8:
1630:(ironclad beetles, cylindrical bark beetles)
292:
290:
288:
2444:
2430:
2303:
2135:
2004:
1649:
827:
735:
726:
712:
663:
551:
537:
469:
407:
393:
385:
42:
31:
2294:List of subgroups of the order Coleoptera
1374:(straight snout weevils, New York weevil)
364:
346:
1801:(cicada beetle, cicada parasite beetles)
284:
1891:(false fireflies, long-lipped beetles)
7:
2405:
2403:
2387:6828CC35-47EA-4F3A-994E-4BE4C2C4AF8A
1917:(false metallic wood-boring beetles)
2416:. You can help Knowledge (XXG) by
341:(864). Pensoft Publishers: 79–97.
25:
1528:Perimylopidae, or Promecheilidae
1113:(minute brown scavenger beetles)
63:
1743:(variegated mud-loving beetles)
1:
1572:(narrow-waisted bark beetles)
1264:(parasitic flat bark beetles)
1215:(short-winged flower beetles)
1071:(lady beetles, or God's cows)
808:(tooth-necked fungus beetles)
2084:(sand-loving scarab beetles)
1474:(minute tree-fungus beetles)
1300:(silvanid flat bark beetles)
2233:(primitive carrion beetles)
2189:(minute mud-loving beetles)
2052:(bumble bee scarab beetles)
1822:(soft-bodied plant beetles)
1793:(soft bodied plant beetles)
596:(predaceous diving beetles)
2494:
2402:
2068:(scavenger scarab beetles)
2060:(enigmatic scarab beetles)
1233:Lamingtonium binnaberrense
947:(soft-wing flower beetles)
2291:
2207:(water scavenger beetles)
1940:Rhinorhipus tamborinensis
1719:(long-toed water beetles)
1612:(polypore fungus beetles)
1514:(palm and flower beetles)
1498:(tumbling flower beetles)
1445:(ant-like flower beetles)
1223:(lined flat bark beetles)
1197:(pleasing fungus beetles)
1092:(handsome fungus beetles)
725:
711:
676:
662:
637:(burrowing water beetles)
550:
536:
482:
468:
424:
348:10.3897/zookeys.864.26689
183:
176:
60:Scientific classification
58:
50:
41:
34:
2254:(feather-winged beetles)
1588:(false longhorn beetles)
1535:(jugular-horned beetles)
1482:(false darkling beetles)
1453:(cryptic fungus beetles)
1272:(shining flower beetles)
612:(crawling water beetles)
511:Sikhotealinia zhiltzovae
220:Sinopyrophorus schimmeli
187:Sinopyrophorus schimmeli
53:Sinopyrophorus schimmeli
2473:Monotypic beetle genera
1735:(forest stream beetles)
1596:(synchroa bark beetles)
1556:(dead log bark beetles)
1548:(fire-coloured beetles)
1437:(ant-like leaf beetles)
1171:(silken fungus beetles)
1105:(well polished beetles)
1079:(minute fungus beetles)
898:Acanthocnemus nigricans
2468:Bioluminescent insects
2412:-related article is a
2246:(round fungus beetles)
1580:(false flower beetles)
1564:(wedge-shaped beetles)
1522:(false blister beetle)
1506:(hairy fungus beetles)
1461:(conifer bark beetles)
1358:(leaf-rolling weevils)
1019:(bark-gnawing beetles)
779:(wounded-tree beetles)
645:(false ground beetles)
567:(trout-stream beetles)
27:Genus of click beetles
2158:(false clown beetles)
1980:(plate-thigh beetles)
1862:(false click beetles)
1767:(water-penny beetles)
1416:(ship-timber beetles)
1395:(pine flower weevils)
1243:(root-eating beetles)
1063:(minute bark beetles)
974:Phloiophilus edwardsi
501:(reticulated beetles)
300:Systematic Entomology
2036:(false stag beetles)
2021:Belohina inexpectata
1883:(net-winged beetles)
1846:(rare click beetles)
1759:(travertine beetles)
1751:(minute mud beetles)
1316:(dry-fungus beetles)
1151:Agapytho foveicollis
908:(false skin beetles)
771:(Jacobson's beetles)
491:Crowsoniella relicta
2034:Diphyllostomatidae
1366:(primitive weevils)
1179:(flat bark beetles)
929:(checkered beetles)
916:(fruitworm beetles)
604:(whirligig beetles)
1899:(glowworm beetles)
1604:(darkling beetles)
1451:Archeocrypticidae
1308:(palmetto beetles)
1055:(dry bark beetles)
842:(longhorn beetles)
312:10.1111/syen.12451
306:. Wiley: 111–123.
2478:Elateroidea stubs
2425:
2424:
2397:
2396:
2309:Taxon identifiers
2300:
2299:
2287:
2286:
2283:
2282:
2279:
2278:
2271:
2263:
2262:(carrion beetles)
2255:
2247:
2234:
2208:
2190:
2159:
2151:
2125:
2124:
2117:
2109:
2101:
2093:
2085:
2077:
2069:
2061:
2053:
2045:
2037:
2024:
1994:
1993:
1981:
1973:
1943:
1918:
1900:
1892:
1884:
1876:
1863:
1855:
1847:
1839:
1838:(soldier beetles)
1831:
1828:Brachypsectridae
1823:
1802:
1794:
1768:
1760:
1752:
1744:
1736:
1728:
1720:
1707:
1699:
1691:
1665:
1639:
1638:
1631:
1613:
1605:
1597:
1589:
1586:Stenotrachelidae
1581:
1573:
1565:
1557:
1549:
1536:
1523:
1515:
1507:
1499:
1491:
1490:(blister beetles)
1483:
1475:
1462:
1454:
1446:
1438:
1417:
1396:
1388:
1375:
1367:
1359:
1351:
1341:
1317:
1309:
1301:
1273:
1265:
1257:
1244:
1236:
1224:
1216:
1198:
1180:
1172:
1154:
1114:
1106:
1093:
1080:
1072:
1064:
1056:
1038:Akalyptoischiidae
1020:
977:
960:
948:
930:
917:
909:
901:
851:
843:
817:
816:
809:
788:
780:
772:
759:
751:
707:
706:
684:
658:
657:
654:
653:
646:
638:
630:
613:
605:
597:
589:
581:
568:
532:
531:
514:
502:
494:
259:, which includes
257:lampyroid beetles
198:
197:
193:Bi & Li, 2019
169:S. schimmeli
158:
157:Bi & Li, 2019
142:
141:Bi & Li, 2019
16:(Redirected from
2485:
2446:
2439:
2432:
2404:
2390:
2389:
2377:
2376:
2364:
2363:
2351:
2350:
2349:
2336:
2335:
2334:
2304:
2269:
2261:
2253:
2245:
2232:
2206:
2188:
2157:
2149:
2136:
2131:Staphyliniformia
2115:
2107:
2099:
2091:
2083:
2075:
2067:
2059:
2051:
2043:
2035:
2018:
2005:
1979:
1970:Declinia relicta
1967:
1937:
1916:
1910:Sinopyrophoridae
1905:Rhagophthalmidae
1898:
1890:
1882:
1874:
1861:
1853:
1845:
1837:
1829:
1821:
1820:Artematopodidae
1800:
1792:
1766:
1758:
1750:
1742:
1734:
1727:(riffle beetles)
1726:
1718:
1706:(turtle beetles)
1705:
1697:
1689:
1663:
1650:
1629:
1611:
1603:
1595:
1587:
1579:
1571:
1563:
1555:
1547:
1534:
1521:
1513:
1505:
1497:
1489:
1481:
1473:
1460:
1452:
1444:
1436:
1415:
1394:
1386:
1373:
1365:
1357:
1350:(fungus weevils)
1349:
1340:
1339:
1335:
1322:Tasmosalpingidae
1315:
1307:
1299:
1271:
1263:
1255:
1242:
1230:
1222:
1214:
1196:
1178:
1170:
1148:
1112:
1104:
1091:
1078:
1070:
1062:
1054:
1018:
971:
954:
946:
928:
915:
907:
895:
894:Acanthocnemidae
849:
841:
828:
807:
786:
778:
770:
757:
749:
736:
727:
713:
682:
664:
644:
636:
624:
611:
603:
595:
587:
580:(ground beetles)
579:
566:
552:
538:
508:
500:
488:
487:Crowsoniellidae
470:
409:
402:
395:
386:
379:
378:
368:
350:
330:
324:
323:
294:
265:Rhagophthalmidae
261:Sinopyrophoridae
215:Sinopyrophoridae
189:
156:
140:
137:Sinopyrophoridae
68:
67:
46:
32:
21:
18:Sinopyrophoridae
2493:
2492:
2488:
2487:
2486:
2484:
2483:
2482:
2453:
2452:
2451:
2450:
2400:
2398:
2393:
2385:
2380:
2372:
2367:
2359:
2354:
2345:
2344:
2339:
2330:
2329:
2324:
2311:
2301:
2296:
2275:
2217:
2168:
2150:(clown beetles)
2121:
2092:(betsy beetles)
2000:Scarabaeiformia
1990:
1947:
1922:
1854:(click beetles)
1830:(Texas beetles)
1806:
1777:
1773:Ptilodactylidae
1698:(cedar beetles)
1674:
1635:
1618:Trictenotomidae
1504:Mycetophagidae
1421:
1400:
1337:
1336:
1334:
1326:
1278:Phloeostichidae
1229:Lamingtoniidae
1221:Laemophloeidae
1169:Cryptophagidae
1133:
1024:
1007:Thanerocleridae
970:Phloiophilidae
957:Metaxina ornata
922:Chaetosomatidae
880:
813:
792:
750:(auger beetles)
731:Bostrichiformia
721:
703:
699:Torridincolidae
683:(skiff beetles)
681:Hydroscaphidae
672:
650:
643:Trachypachidae
588:(tiger beetles)
546:
528:
478:
464:
420:
413:
383:
382:
332:
331:
327:
296:
295:
286:
281:
194:
191:
185:
172:
155:
139:
62:
28:
23:
22:
15:
12:
11:
5:
2491:
2489:
2481:
2480:
2475:
2470:
2465:
2455:
2454:
2449:
2448:
2441:
2434:
2426:
2423:
2422:
2395:
2394:
2392:
2391:
2378:
2365:
2352:
2347:Sinopyrophorus
2337:
2321:
2319:
2317:Sinopyrophorus
2313:
2312:
2307:
2298:
2297:
2292:
2289:
2288:
2285:
2284:
2281:
2280:
2277:
2276:
2274:
2273:
2270:(rove beetles)
2268:Staphylinidae
2265:
2257:
2249:
2241:
2236:
2227:
2225:
2223:Staphylinoidea
2219:
2218:
2216:
2215:
2210:
2205:Hydrophilidae
2202:
2197:
2192:
2184:
2178:
2176:
2174:Hydrophiloidea
2170:
2169:
2167:
2166:
2161:
2153:
2144:
2142:
2133:
2127:
2126:
2123:
2122:
2120:
2119:
2116:(hide beetles)
2111:
2103:
2100:(rain beetles)
2095:
2087:
2079:
2076:(stag beetles)
2071:
2063:
2055:
2047:
2039:
2031:
2029:Bolboceratidae
2026:
2013:
2011:
2002:
1996:
1995:
1992:
1991:
1989:
1988:
1983:
1975:
1963:
1957:
1955:
1949:
1948:
1946:
1945:
1936:Rhinorhipidae
1932:
1930:
1928:Rhinorhipoidea
1924:
1923:
1921:
1920:
1912:
1907:
1902:
1894:
1886:
1878:
1870:
1865:
1857:
1849:
1841:
1833:
1825:
1816:
1814:
1808:
1807:
1805:
1804:
1796:
1787:
1785:
1779:
1778:
1776:
1775:
1770:
1762:
1754:
1746:
1741:Heteroceridae
1738:
1730:
1722:
1714:
1709:
1704:Chelonariidae
1701:
1696:Callirhipidae
1693:
1690:(pill beetles)
1684:
1682:
1676:
1675:
1673:
1672:
1667:
1658:
1656:
1647:
1641:
1640:
1637:
1636:
1634:
1633:
1625:
1620:
1615:
1607:
1602:Tenebrionidae
1599:
1591:
1583:
1575:
1567:
1559:
1551:
1543:
1538:
1530:
1525:
1517:
1509:
1501:
1493:
1485:
1477:
1469:
1464:
1456:
1448:
1440:
1431:
1429:
1427:Tenebrionoidea
1423:
1422:
1420:
1419:
1410:
1408:
1402:
1401:
1399:
1398:
1390:
1385:Curculionidae
1382:
1377:
1369:
1361:
1353:
1344:
1342:
1332:Curculionoidea
1328:
1327:
1325:
1324:
1319:
1311:
1303:
1295:
1293:Protocucujidae
1290:
1285:
1280:
1275:
1267:
1259:
1251:
1246:
1238:
1226:
1218:
1210:
1205:
1200:
1192:
1187:
1185:Cybocephalidae
1182:
1174:
1166:
1161:
1156:
1143:
1141:
1135:
1134:
1132:
1131:
1126:
1121:
1116:
1108:
1100:
1095:
1087:
1085:Discolomatidae
1082:
1074:
1069:Coccinellidae
1066:
1058:
1053:Bothrideridae
1050:
1045:
1040:
1034:
1032:
1030:Coccinelloidea
1026:
1025:
1023:
1022:
1017:Trogossitidae
1014:
1009:
1004:
999:
994:
989:
984:
979:
967:
962:
950:
942:
937:
935:Lophocateridae
932:
924:
919:
911:
903:
890:
888:
882:
881:
879:
878:
873:
868:
863:
858:
853:
850:(leaf beetles)
848:Chrysomelidae
845:
836:
834:
832:Chrysomeloidea
825:
819:
818:
815:
814:
812:
811:
802:
800:
794:
793:
791:
790:
782:
777:Nosodendridae
774:
769:Jacobsoniidae
766:
761:
758:(skin beetles)
753:
744:
742:
733:
723:
722:
716:
709:
708:
705:
704:
702:
701:
696:
691:
686:
677:
674:
673:
667:
660:
659:
656:
655:
652:
651:
649:
648:
640:
632:
627:Meru phyllisae
620:
615:
607:
599:
591:
583:
575:
570:
561:
559:
548:
547:
541:
534:
533:
530:
529:
527:
526:
521:
519:Micromalthidae
516:
504:
496:
483:
480:
479:
473:
466:
465:
463:
462:
456:
450:
444:
438:
432:
425:
422:
421:
414:
412:
411:
404:
397:
389:
381:
380:
325:
283:
282:
280:
277:
253:Sinopyrophorus
245:Sinopyrophorus
231:Sinopyrophorus
207:bioluminescent
205:is a genus of
202:Sinopyrophorus
196:
195:
192:
181:
180:
174:
173:
166:
164:
160:
159:
152:Sinopyrophorus
148:
144:
143:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
56:
55:
48:
47:
39:
38:
36:Sinopyrophorus
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2490:
2479:
2476:
2474:
2471:
2469:
2466:
2464:
2461:
2460:
2458:
2447:
2442:
2440:
2435:
2433:
2428:
2427:
2421:
2419:
2415:
2411:
2406:
2401:
2388:
2383:
2379:
2375:
2370:
2366:
2362:
2357:
2353:
2348:
2342:
2338:
2333:
2327:
2323:
2322:
2320:
2318:
2314:
2310:
2305:
2295:
2290:
2272:
2266:
2264:
2258:
2256:
2250:
2248:
2242:
2240:
2237:
2235:
2229:
2228:
2226:
2224:
2220:
2214:
2211:
2209:
2203:
2201:
2198:
2196:
2193:
2191:
2185:
2183:
2180:
2179:
2177:
2175:
2171:
2165:
2162:
2160:
2156:Sphaeritidae
2154:
2152:
2146:
2145:
2143:
2141:
2137:
2134:
2132:
2128:
2118:
2112:
2110:
2106:Scarabaeidae
2104:
2102:
2096:
2094:
2088:
2086:
2080:
2078:
2072:
2070:
2064:
2062:
2056:
2054:
2048:
2046:
2044:(dor beetles)
2040:
2038:
2032:
2030:
2027:
2025:
2022:
2015:
2014:
2012:
2010:
2009:Scarabaeoidea
2006:
2003:
2001:
1997:
1987:
1984:
1982:
1976:
1974:
1971:
1964:
1962:
1959:
1958:
1956:
1954:
1950:
1944:
1941:
1934:
1933:
1931:
1929:
1925:
1919:
1913:
1911:
1908:
1906:
1903:
1901:
1895:
1893:
1887:
1885:
1879:
1877:
1871:
1869:
1866:
1864:
1858:
1856:
1850:
1848:
1844:Cerophytidae
1842:
1840:
1834:
1832:
1826:
1824:
1818:
1817:
1815:
1813:
1809:
1803:
1799:Rhipiceridae
1797:
1795:
1789:
1788:
1786:
1784:
1780:
1774:
1771:
1769:
1763:
1761:
1755:
1753:
1747:
1745:
1739:
1737:
1733:Eulichadidae
1731:
1729:
1723:
1721:
1715:
1713:
1712:Cneoglossidae
1710:
1708:
1702:
1700:
1694:
1692:
1686:
1685:
1683:
1681:
1677:
1671:
1670:Schizopodidae
1668:
1666:
1660:
1659:
1657:
1655:
1651:
1648:
1646:
1645:Elateriformia
1642:
1632:
1626:
1624:
1621:
1619:
1616:
1614:
1610:Tetratomidae
1608:
1606:
1600:
1598:
1592:
1590:
1584:
1582:
1576:
1574:
1568:
1566:
1562:Ripiphoridae
1560:
1558:
1552:
1550:
1546:Pyrochroidae
1544:
1542:
1541:Pterogeniidae
1539:
1537:
1531:
1529:
1526:
1524:
1518:
1516:
1510:
1508:
1502:
1500:
1494:
1492:
1486:
1484:
1480:Melandryidae
1478:
1476:
1470:
1468:
1467:Chalcodryidae
1465:
1463:
1457:
1455:
1449:
1447:
1441:
1439:
1433:
1432:
1430:
1428:
1424:
1418:
1412:
1411:
1409:
1407:
1403:
1397:
1393:Nemonychidae
1391:
1389:
1383:
1381:
1378:
1376:
1370:
1368:
1362:
1360:
1354:
1352:
1346:
1345:
1343:
1333:
1329:
1323:
1320:
1318:
1312:
1310:
1304:
1302:
1296:
1294:
1291:
1289:
1288:Propalticidae
1286:
1284:
1283:Priasilphidae
1281:
1279:
1276:
1274:
1268:
1266:
1262:Passandridae
1260:
1258:
1256:(sap beetles)
1252:
1250:
1247:
1245:
1239:
1237:
1234:
1227:
1225:
1219:
1217:
1211:
1209:
1206:
1204:
1201:
1199:
1193:
1191:
1188:
1186:
1183:
1181:
1175:
1173:
1167:
1165:
1164:Cavognathidae
1162:
1160:
1157:
1155:
1152:
1145:
1144:
1142:
1140:
1136:
1130:
1127:
1125:
1122:
1120:
1117:
1115:
1109:
1107:
1101:
1099:
1098:Eupsilobiidae
1096:
1094:
1090:Endomychidae
1088:
1086:
1083:
1081:
1077:Corylophidae
1075:
1073:
1067:
1065:
1059:
1057:
1051:
1049:
1046:
1044:
1041:
1039:
1036:
1035:
1033:
1031:
1027:
1021:
1015:
1013:
1010:
1008:
1005:
1003:
1000:
998:
995:
993:
992:Protopeltidae
990:
988:
987:Prionoceridae
985:
983:
980:
978:
975:
968:
966:
963:
961:
958:
951:
949:
943:
941:
940:Mauroniscidae
938:
936:
933:
931:
925:
923:
920:
918:
912:
910:
904:
902:
899:
892:
891:
889:
887:
883:
877:
874:
872:
869:
867:
864:
862:
861:Megalopodidae
859:
857:
854:
852:
846:
844:
840:Cerambycidae
838:
837:
835:
833:
829:
826:
824:
820:
810:
806:Derodontidae
804:
803:
801:
799:
798:Derodontoidea
795:
789:
783:
781:
775:
773:
767:
765:
764:Endecatomidae
762:
760:
754:
752:
748:Bostrichidae
746:
745:
743:
741:
740:Bostrichoidea
737:
734:
732:
728:
724:
720:
714:
710:
700:
697:
695:
694:Sphaeriusidae
692:
690:
687:
685:
679:
678:
675:
671:
665:
661:
647:
641:
639:
633:
631:
628:
621:
619:
616:
614:
608:
606:
600:
598:
592:
590:
586:Cicindelidae
584:
582:
576:
574:
571:
569:
563:
562:
560:
557:
553:
549:
545:
539:
535:
525:
522:
520:
517:
515:
512:
505:
503:
497:
495:
492:
485:
484:
481:
477:
471:
467:
461:
457:
455:
451:
449:
445:
443:
439:
437:
433:
431:
427:
426:
423:
418:
410:
405:
403:
398:
396:
391:
390:
387:
376:
372:
367:
362:
358:
354:
349:
344:
340:
336:
329:
326:
321:
317:
313:
309:
305:
301:
293:
291:
289:
285:
278:
276:
274:
270:
266:
262:
258:
254:
250:
246:
242:
240:
236:
232:
228:
226:
222:
221:
216:
212:
208:
204:
203:
190:
188:
182:
179:
178:Binomial name
175:
171:
170:
165:
162:
161:
154:
153:
149:
146:
145:
138:
135:
132:
131:
128:
125:
123:Superfamily:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
66:
61:
57:
54:
49:
45:
40:
37:
33:
30:
19:
2418:expanding it
2407:
2399:
2316:
2195:Helophoridae
2187:Georissidae
2182:Epimetopidae
2098:Pleocomidae
2082:Ochodaeidae
2066:Hybosoridae
2050:Glaphyridae
2042:Geotrupidae
2020:
2017:Belohinidae
1978:Eucinetidae
1969:
1966:Decliniidae
1939:
1897:Phengodidae
1836:Cantharidae
1791:Dascillidae
1783:Dascilloidea
1765:Psephenidae
1757:Lutrochidae
1749:Limnichidae
1662:Buprestidae
1654:Buprestoidea
1594:Synchroidae
1578:Scraptiidae
1570:Salpingidae
1533:Prostomidae
1520:Oedemeridae
1496:Mordellidae
1414:Lymexylidae
1406:Lymexyloidea
1356:Attelabidae
1348:Anthribidae
1306:Smicripidae
1270:Phalacridae
1254:Nitidulidae
1249:Myraboliidae
1241:Monotomidae
1232:
1213:Kateretidae
1190:Cyclaxyridae
1150:
1147:Agapythidae
1111:Latridiidae
1061:Cerylonidae
1048:Anamorphidae
982:Phycosecidae
973:
956:
953:Metaxinidae
906:Biphyllidae
897:
866:Orsodacnidae
823:Cucujiformia
756:Dermestidae
626:
565:Amphizoidae
510:
490:
476:Archostemata
460:Holometabola
458:Superorder:
452:Infraclass:
338:
328:
303:
299:
252:
244:
243:
230:
229:
219:
218:
214:
201:
200:
199:
186:
184:
168:
167:
151:
150:
136:
52:
35:
29:
2463:Elateroidea
2410:Elateroidea
2341:Wikispecies
2239:Hydraenidae
2213:Spercheidae
2200:Hydrochidae
2164:Synteliidae
2148:Histeridae
2140:Histeroidea
2090:Passalidae
2058:Glaresidae
1915:Throscidae
1875:(fireflies)
1873:Lampyridae
1860:Eucnemidae
1852:Elateridae
1812:Elateroidea
1628:Zopheridae
1512:Mycteridae
1443:Anthicidae
1314:Sphindidae
1298:Silvanidae
1208:Hobartiidae
1195:Erotylidae
1124:Mycetaeidae
1119:Murmidiidae
1103:Euxestidae
997:Rentoniidae
871:Oxypeltidae
856:Disteniidae
689:Lepiceridae
618:Hygrobiidae
610:Haliplidae
594:Dytiscidae
573:Aspidytidae
269:Phengodidae
251:. Instead,
249:Pyrophorini
239:Pyrophorini
211:Elateroidea
127:Elateroidea
2457:Categories
2260:Silphidae
2252:Ptiliidae
2244:Leiodidae
2231:Agyrtidae
2074:Lucanidae
1953:Scirtoidea
1889:Omethidae
1868:Jurasaidae
1717:Dryopidae
1688:Byrrhidae
1680:Byrrhoidea
1372:Brentidae
1177:Cucujidae
1159:Boganiidae
1139:Cucujoidea
1012:Thymalidae
1002:Rhadalidae
945:Melyridae
914:Byturidae
876:Vesperidae
785:Ptiniidae
635:Noteridae
602:Gyrinidae
578:Carabidae
507:Jurodidae
499:Cupedidae
446:Subclass:
436:Arthropoda
417:Coleoptera
279:References
273:Lampyridae
235:Elateridae
117:Coleoptera
97:Arthropoda
2332:Q70078246
2114:Trogidae
2108:(scarabs)
1986:Scirtidae
1961:Clambidae
1554:Pythidae
1488:Meloidae
1435:Aderidae
1338:(weevils)
1203:Helotidae
1129:Teredidae
1043:Alexiidae
927:Cleridae
886:Cleroidea
719:Polyphaga
717:Suborder
670:Myxophaga
668:Suborder
623:Meruidae
542:Suborder
524:Ommatidae
474:Suborder
448:Pterygota
428:Kingdom:
357:1313-2970
320:0307-6970
227:, China.
163:Species:
83:Kingdom:
77:Eukaryota
2374:11927332
2361:10423792
2326:Wikidata
1881:Lycidae
1725:Elmidae
1623:Ulodidae
1459:Boridae
1364:Belidae
965:Peltidae
558:families
544:Adephaga
454:Neoptera
434:Phylum:
430:Animalia
419:families
375:31363346
133:Family:
93:Phylum:
87:Animalia
73:Domain:
2382:ZooBank
1472:Ciidae
1380:Caridae
442:Insecta
440:Class:
415:Extant
366:6656784
339:ZooKeys
147:Genus:
113:Order:
107:Insecta
103:Class:
556:Extant
373:
363:
355:
318:
271:, and
225:Yunnan
2408:This
2369:IRMNG
2414:stub
2356:GBIF
371:PMID
353:ISSN
316:ISSN
361:PMC
343:doi
308:doi
2459::
2384::
2371::
2358::
2343::
2328::
369:.
359:.
351:.
337:.
314:.
304:46
302:.
287:^
275:.
267:,
263:,
2445:e
2438:t
2431:v
2420:.
2023:)
2019:(
1972:)
1968:(
1942:)
1938:(
1235:)
1231:(
1153:)
1149:(
976:)
972:(
959:)
955:(
900:)
896:(
629:)
625:(
513:)
509:(
493:)
489:(
408:e
401:t
394:v
377:.
345::
322:.
310::
20:)
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